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<document id="512A6F4CC0AAC269A54D6567D0D39241" ID-DOI="10.11646/zootaxa.3716.3.4" ID-GBIF-Dataset="a997e15d-6b78-4fcd-b241-95216cd7d3ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="222975" ID-ZooBank="798449F3-C14D-4D9D-B2E1-505EE2BC6115" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460682175734" checkinUser="plazi" docAuthor="Nunes, Valéria Juliete Da Silva Benedito Mendes &amp; Fernandes, José Antônio Marin" docDate="2013" docId="03E387ABFF80BE1545895F48FB18F84E" docLanguage="en" docName="zt03716p416.pdf" docOrigin="Zootaxa 3716 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraedessa ecuadoriensis Silva &amp; Fernandes, sp. nov." docType="treatment" docVersion="8" lastPageNumber="405" masterDocId="FFDAFFD3FF89BE1F451E5C2FFFBBFFBB" masterDocTitle="Paraedessa, a new genus of Edessinae (Hemiptera: Heteroptera: Pentatomidae)" masterLastPageNumber="416" masterPageNumber="395" pageNumber="404" updateTime="1698350568411" updateUser="plazi">
<mods:mods id="571AACFC89ED1AFBB955E302116C47ED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6D54FC53E8136475D38972B60B81C754">
<mods:title id="4AFB7F9C1BE1E305C866E30EBEBB156E">Paraedessa, a new genus of Edessinae (Hemiptera: Heteroptera: Pentatomidae)</mods:title>
</mods:titleInfo>
<mods:name id="210BE260DC6EE1D6B1153DD90F775CF0" type="personal">
<mods:role id="986F941C2D0A051626D768C38AB1D3E8">
<mods:roleTerm id="AFAA0EC1F5C35BB48AF479CBF5304D5B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A2AC41BC352AEC63644062840C321986">Nunes, Valéria Juliete Da Silva Benedito Mendes</mods:namePart>
</mods:name>
<mods:name id="0E8CD0951B2199B816BA4F3CAB0ABBB8" type="personal">
<mods:role id="3247F258BDE121709CD1D967BE7E28FF">
<mods:roleTerm id="84BA65516383AB357559F7010BC3B70A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D76818DE8BA3D374AC3A62B3D6659EFE">Fernandes, José Antônio Marin</mods:namePart>
</mods:name>
<mods:typeOfResource id="E4DDE55AFE0E5F0FBF6012EADD93FC34">text</mods:typeOfResource>
<mods:relatedItem id="78ABE85D2477CD0824823848FE817B00" type="host">
<mods:titleInfo id="5657E1D7DEA0CB6D5FB025FA06DC8006">
<mods:title id="A3B93CC8D1932D19EDD878EFE7926CC2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E2984498EBB05E28C1A0CD038561CA42">
<mods:date id="7495664C78C87D19B44F5183D9E58F30">2013</mods:date>
<mods:detail id="3BA24324AA852F07179B98ABF5246AB3" type="volume">
<mods:number id="289DC857EAC5FD7B889A1007B4D2F9D0">3716</mods:number>
</mods:detail>
<mods:detail id="70C775F5A6C48CB33F2FA9CDB3C51138" type="issue">
<mods:number id="B23C7D3A8327416C58AD619E094FD1BF">3</mods:number>
</mods:detail>
<mods:extent id="869464C5124BFD7B078D7144ED8311E4" unit="page">
<mods:start id="E2556F7A376772D725633C613C94C326">395</mods:start>
<mods:end id="26F219165A672CD35F87341820CB5E12">416</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FC5E99B57192D7A91A0A3626DFD60C3A">journal article</mods:classification>
<mods:identifier id="DE8387C8AF587E9D2FF5350DE55B3B8C" type="DOI">10.11646/zootaxa.3716.3.4</mods:identifier>
<mods:identifier id="CA2F5E3C53DF7156EAE8491DAA1682BB" type="GBIF-Dataset">a997e15d-6b78-4fcd-b241-95216cd7d3ca</mods:identifier>
<mods:identifier id="9C26E2E2BB2F4131E796A49D03F942A7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="84882744420FCFAA1FB84759C1003CFB" type="Zenodo-Dep">222975</mods:identifier>
<mods:identifier id="67B6247B04F9B72EFD68B84B2BC16C71" type="ZooBank">798449F3-C14D-4D9D-B2E1-505EE2BC6115</mods:identifier>
</mods:mods>
<treatment id="03E387ABFF80BE1545895F48FB18F84E" ID-DOI="http://doi.org/10.5281/zenodo.6158044" ID-GBIF-Taxon="119580386" ID-Zenodo-Dep="6158044" LSID="urn:lsid:plazi:treatment:03E387ABFF80BE1545895F48FB18F84E" httpUri="http://treatment.plazi.org/id/03E387ABFF80BE1545895F48FB18F84E" lastPageId="10" lastPageNumber="405" pageId="9" pageNumber="404">
<subSubSection id="C3506536FF80BE1645895F48FE1DFC18" pageId="9" pageNumber="404" type="nomenclature">
<paragraph id="8BF536BDFF80BE1645895F48FC99FC39" blockId="9.[151,802,871,931]" box="[151,802,871,898]" pageId="9" pageNumber="404">
<heading id="D0BD81D1FF80BE1645895F48FC99FC39" bold="true" box="[151,802,871,898]" fontSize="11" level="1" pageId="9" pageNumber="404" reason="1">
<emphasis id="B93EEAAFFF80BE1645895F48FC99FC39" bold="true" box="[151,802,871,898]" pageId="9" pageNumber="404">
<taxonomicName id="4C4A4D3EFF80BE1645895F48FD06FC39" authority="Silva &amp; Fernandes" authorityName="Silva &amp; Fernandes" box="[151,701,871,898]" class="Insecta" family="Pentatomidae" genus="Paraedessa" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="404" phylum="Arthropoda" rank="species" species="ecuadoriensis" status="sp. nov.">
<emphasis id="B93EEAAFFF80BE1645895F48FE76FC3A" bold="true" box="[151,461,871,897]" italics="true" pageId="9" pageNumber="404">Paraedessa ecuadoriensis</emphasis>
Silva &amp; Fernandes
</taxonomicName>
<taxonomicNameLabel id="A20D57D4FF80BE1647DA5F47FC99FC39" box="[708,802,872,898]" pageId="9" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF536BDFF80BE1645895FA5FE1DFC18" blockId="9.[151,802,871,931]" box="[151,422,906,931]" pageId="9" pageNumber="404">(Figures: 21–25, 53, 59)</paragraph>
</subSubSection>
<subSubSection id="C3506536FF80BE1645895FFDFAF5FC50" box="[151,1358,978,1003]" pageId="9" pageNumber="404" type="etymology">
<paragraph id="8BF536BDFF80BE1645895FFDFAF5FC50" blockId="9.[151,1437,978,2011]" box="[151,1358,978,1003]" pageId="9" pageNumber="404">
<emphasis id="B93EEAAFFF80BE1645895FFDFEA5FC50" bold="true" box="[151,286,978,1003]" pageId="9" pageNumber="404">Etymology.</emphasis>
The specific name makes reference to the country where most of the specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="C3506536FF80BE1645D95FD9FE7DF990" pageId="9" pageNumber="404" type="materials_examined">
<paragraph id="8BF536BDFF80BE1645D95FD9FE00FB88" blockId="9.[151,1437,978,2011]" pageId="9" pageNumber="404">
<emphasis id="B93EEAAFFF80BE1645D95FD9FE08FBB4" bold="true" box="[199,435,1014,1039]" pageId="9" pageNumber="404">Material examined.</emphasis>
<typeStatus id="54F1881FFF80BE1644A25FD9FD9FFBB4" box="[444,548,1014,1039]" pageId="9" pageNumber="404" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F35D762DFF80BE16476B5FD9FCB9FBB4" box="[629,770,1014,1039]" name="Ecuador" pageId="9" pageNumber="404">
<emphasis id="B93EEAAFFF80BE16476B5FD9FCB9FBB4" bold="true" box="[629,770,1014,1039]" pageId="9" pageNumber="404">ECUADOR</emphasis>
</collectingCountry>
.
<emphasis id="B93EEAAFFF80BE16460D5FD9FCE6FBB4" bold="true" box="[787,861,1014,1039]" pageId="9" pageNumber="404">Napo:</emphasis>
Tena,
<date id="FFF4107DFF80BE1646B35FD8FB8DFBB4" box="[941,1078,1015,1039]" pageId="9" pageNumber="404" value="1982-03-07">7-Mar-1982</date>
, G. Couturier &amp; G. Onoré coll. Bord du Fleuve (MNHN).
</paragraph>
<paragraph id="8BF536BDFF80BE1645D95810FE7DF990" blockId="9.[151,1437,978,2011]" pageId="9" pageNumber="404">
<typeStatus id="54F1881FFF80BE1645D95810FE83FBEC" box="[199,312,1087,1111]" pageId="9" pageNumber="404" type="paratype">Paratypes</typeStatus>
.
<emphasis id="B93EEAAFFF80BE16445A5811FDE3FBEC" bold="true" box="[324,600,1086,1111]" pageId="9" pageNumber="404">
<collectingCountry id="F35D762DFF80BE16445A5811FE5EFBEC" box="[324,485,1086,1111]" name="Colombia" pageId="9" pageNumber="404">COLOMBIA</collectingCountry>
. Meta: ♂
</emphasis>
Villavicencio, Bosque de Bavaria, nr. Rio Guatiquia
<date id="FFF4107DFF80BE1641A35811FAE0FBEC" box="[1213,1371,1086,1111]" pageId="9" pageNumber="404" value="2013-07-03" valueMax="2013-07-05" valueMin="2013-07-03">3–5-VII-2013</date>
, J. E. Eger &amp; A. A. Calixo, coll.,
<geoCoordinate id="EE7E507AFF80BE1644D0584CFDD2FBC0" box="[462,617,1122,1147]" direction="north" orientation="latitude" pageId="9" pageNumber="404" precision="1" value="4.1776166">N 04°10.657’</geoCoordinate>
,
<geoCoordinate id="EE7E507AFF80BE16476B584CFC9EFBC0" box="[629,805,1122,1147]" direction="west" orientation="longitude" pageId="9" pageNumber="404" precision="1" value="-73.64753">W 073°38.852’</geoCoordinate>
,
<quantity id="4CB29B58FF80BE16462F584DFC3EFBC0" box="[817,901,1122,1147]" metricMagnitude="2" metricUnit="m" metricValue="5.132832" pageId="9" pageNumber="404" unit="ft" value="1684.0">1684 ft</quantity>
. elev. (JEE); ♂ 2♀ Vic. GunavicheEstadero, nr. Rio Guatiquia,
<date id="FFF4107DFF80BE16445258A9FE51FB24" box="[332,490,1158,1183]" pageId="9" pageNumber="404" value="2013-07-03" valueMax="2013-07-05" valueMin="2013-07-03">3–5-VII-2013</date>
, J. E. Eger &amp; A. A. Calixto, coll.,
<geoCoordinate id="EE7E507AFF80BE16468C58A8FB89FB24" box="[914,1074,1158,1183]" direction="north" orientation="latitude" pageId="9" pageNumber="404" precision="1" value="4.1751">N 04°10.506’</geoCoordinate>
,
<geoCoordinate id="EE7E507AFF80BE16415C58A8FB4DFB24" box="[1090,1270,1159,1183]" direction="west" orientation="longitude" pageId="9" pageNumber="404" precision="1" value="-73.637215">W 073°38.233’</geoCoordinate>
,
<quantity id="4CB29B58FF80BE16401858A9FAE5FB24" box="[1286,1374,1158,1183]" metricMagnitude="2" metricUnit="m" metricValue="4.46532" pageId="9" pageNumber="404" unit="ft" value="1465.0">1465 ft</quantity>
. elev (JEE).
<collectingCountry id="F35D762DFF80BE1645C35885FED1FB78" box="[221,362,1194,1219]" name="Ecuador" pageId="9" pageNumber="404">
<emphasis id="B93EEAAFFF80BE1645C35885FED1FB78" bold="true" box="[221,362,1194,1219]" pageId="9" pageNumber="404">ECUADOR</emphasis>
</collectingCountry>
.
<emphasis id="B93EEAAFFF80BE1644655885FDBCFB78" bold="true" box="[379,519,1194,1219]" pageId="9" pageNumber="404">Sucumbios:</emphasis>
♂ Shushufinde, F 2. 3 c10. d
<date id="FFF4107DFF80BE16467A5884FC76FB78" box="[868,973,1195,1219]" pageId="9" pageNumber="404" value="1982-11-11">11-XI-82</date>
, R. Desmier de Chenon (UFRG); ♀ San Pablo de Kantesiya,
<date id="FFF4107DFF80BE16446258E1FDABFB5C" box="[380,528,1230,1255]" pageId="9" pageNumber="404" value="1985-04-17">17-Abr-1985</date>
, G. Couturier rec. Fauchage chauy de
<taxonomicName id="4C4A4D3EFF80BE1646A858E0FB9FFB5D" authority="Mays" authorityName="Mays" box="[950,1060,1231,1254]" class="Liliopsida" family="Poaceae" genus="Zea" kingdom="Plantae" order="Poales" pageId="9" pageNumber="404" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAAFFF80BE1646A858E0FB9FFB5D" box="[950,1060,1231,1254]" italics="true" pageId="9" pageNumber="404">Zea Mays</emphasis>
</taxonomicName>
(UFRG);
<emphasis id="B93EEAAFFF80BE16418B58E1FB65FB5C" bold="true" box="[1173,1246,1230,1255]" pageId="9" pageNumber="404">Napo:</emphasis>
♀ Santa Cecilia,
<date id="FFF4107DFF80BE16458958DDFEF0FAB0" box="[151,331,1266,1291]" pageId="9" pageNumber="404" value="1966-07-20" valueMax="1966-07-28" valueMin="1966-07-20">20–28-Jul-1966</date>
, C. R. Patrick (JEE); ♂ Coca, Juillet, 1982, G. Onoré Coll. Sur Les Peuilles de
<taxonomicName id="4C4A4D3EFF80BE1641C958DDFA27FAB0" box="[1239,1436,1266,1291]" class="Liliopsida" family="Commelinaceae" genus="Elasis" kingdom="Plantae" order="Commelinales" pageId="9" pageNumber="404" phylum="Tracheophyta" rank="species" species="guinaensis">
<emphasis id="B93EEAAFFF80BE1641C958DDFA27FAB0" box="[1239,1436,1266,1291]" italics="true" pageId="9" pageNumber="404">Elasis guinaensis</emphasis>
</taxonomicName>
(MNHN); ♂
<date id="FFF4107DFF80BE1644325939FE0CFA94" box="[300,439,1302,1327]" pageId="9" pageNumber="404" value="1982-03-06">6-Mar-1982</date>
. G. Couturier &amp; G. Onoré Coll,
<quantity id="4CB29B58FF80BE16463F5938FCD2FA94" box="[801,873,1302,1327]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="404" unit="km" value="32.0">32 km</quantity>
de Puyo chamo de cannes (MNHN);
<specimenCount id="9D4CFD34FF80BE1640065938FA80FA94" box="[1304,1339,1303,1327]" pageId="9" pageNumber="404" type="generic">3♂</specimenCount>
3♀ Vic. Puerto Misahuallí,
<quantity id="4CB29B58FF80BE16446E5915FDB0FAE8" box="[368,523,1338,1363]" metricMagnitude="2" metricUnit="m" metricValue="5.4102" metricValueMax="5.7912" metricValueMin="5.0291999999999994" pageId="9" pageNumber="404" unit="ft" value="1775.0" valueMax="1900.0" valueMin="1650.0">1650–1900 ft</quantity>
,
<date id="FFF4107DFF80BE1647065915FD00FAE8" box="[536,699,1338,1363]" pageId="9" pageNumber="404" value="1998-09-06" valueMax="1998-09-19" valueMin="1998-09-06">6–19-IX-1998</date>
, J. E. Eger, coll.;
<geoCoordinate id="EE7E507AFF80BE1646975914FBABFAE8" box="[905,1040,1339,1363]" direction="south" orientation="latitude" pageId="9" pageNumber="404" precision="1" value="-1.0345">1°02’4.2” S</geoCoordinate>
lat.,
<geoCoordinate id="EE7E507AFF80BE1641555914FB43FAE9" box="[1099,1272,1339,1363]" direction="west" orientation="longitude" pageId="9" pageNumber="404" precision="1" value="-77.663666">77°39’49.2” W</geoCoordinate>
lon., Mercury vapor and Ultraviolet Lights (JEE); ♀ Estación Biológia Jatun Sacha,
<date id="FFF4107DFF80BE1646B05971FBFDFACC" box="[942,1094,1374,1399]" pageId="9" pageNumber="404" value="1989-07-31">July-31-1989</date>
, Paul. H. Freytag, Tom Myers (JEE);
<emphasis id="B93EEAAFFF80BE1645F759ADFEE0FA20" bold="true" box="[233,347,1410,1435]" pageId="9" pageNumber="404">Orellana:</emphasis>
♂ Estación Cientifica Yasuní.
<date id="FFF4107DFF80BE1647D859ADFC85FA20" box="[710,830,1410,1435]" pageId="9" pageNumber="404" value="1999-10-05">5-10-1999</date>
.
<geoCoordinate id="EE7E507AFF80BE16465259ACFC4AFA20" box="[844,1009,1411,1435]" direction="south" orientation="latitude" pageId="9" pageNumber="404" precision="15" value="-0.67444444">00° 40’ 28” S</geoCoordinate>
,
<geoCoordinate id="EE7E507AFF80BE16411E59ADFB1EFA21" box="[1024,1189,1410,1435]" direction="west" orientation="longitude" pageId="9" pageNumber="404" precision="15" value="-76.64722">76° 38’50” W</geoCoordinate>
.UV light. Coll. E. G. Riley,
<quantity id="4CB29B58FF80BE1645FE5989FE9BFA04" box="[224,288,1446,1471]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="9" pageNumber="404" unit="m" value="215.0">215m</quantity>
(TAMU); ♂ 4 ♀ Yasuni National Park, Yasuni Research Station,
<geoCoordinate id="EE7E507AFF80BE16411B5989FBD7FA04" box="[1029,1132,1446,1471]" direction="west" orientation="longitude" pageId="9" pageNumber="404" precision="925" value="-76.6">76°36’W</geoCoordinate>
<geoCoordinate id="EE7E507AFF80BE16416D5988FB70FA04" box="[1139,1227,1447,1471]" direction="south" orientation="latitude" pageId="9" pageNumber="404" precision="925" value="-0.6333333">0° 38’S</geoCoordinate>
,
<date id="FFF4107DFF80BE1641C95988FAC1FA04" box="[1239,1402,1447,1471]" pageId="9" pageNumber="404" value="1998-11-03" valueMax="1998-11-20" valueMin="1998-11-03">3–20-XI-1998</date>
, T. Pape &amp; B. Vicklund (NHRS); ♂ Yasuni Research Satation,
<date id="FFF4107DFF80BE16462559E4FC4CFA58" box="[827,1015,1483,1507]" pageId="9" pageNumber="404" value="1998-10-19" valueMax="1998-10-30" valueMin="1998-10-19">19–30-Oct-1998</date>
, W. J. Hanson,
<quantity id="4CB29B58FF80BE1641B659E5FB4BFA59" box="[1192,1264,1482,1507]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="404" unit="m" value="250.0">250 m</quantity>
(JEE); ♂ Lake Limoncocha, Sacha Lodge Station 900’ El.
<date id="FFF4107DFF80BE1647B159C0FCCFF9BC" box="[687,884,1519,1543]" pageId="9" pageNumber="404" value="1980-06-24" valueMax="1980-06-27" valueMin="1980-06-24">24–27-June-1980</date>
col. Dan Bogar (JEE); ♂
<date id="FFF4107DFF80BE1641DB59C0FA8FF9BC" box="[1221,1332,1519,1543]" pageId="9" pageNumber="404" value="1974-02-09">
9-II-
<quantity id="4CB29B58FF80BE1641E659C0FA8FF9BC" box="[1272,1332,1519,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.9743" pageId="9" pageNumber="404" unit="m" value="1974.3">1974</quantity>
</date>
.
<specimenCount id="9D4CFD34FF80BE16405859C0FA27F9BD" box="[1350,1436,1519,1543]" pageId="9" pageNumber="404" type="generic">300 m.</specimenCount>
Drummond, B. coll. (JEE).
</paragraph>
</subSubSection>
<subSubSection id="C3506536FF80BE1545D95A19FCC1F942" lastPageId="10" lastPageNumber="405" pageId="9" pageNumber="404" type="description">
<paragraph id="8BF536BDFF80BE1645D95A19FE11F92C" blockId="9.[151,1437,978,2011]" pageId="9" pageNumber="404">
<emphasis id="B93EEAAFFF80BE1645D95A19FEC6F9F4" bold="true" box="[199,381,1590,1615]" pageId="9" pageNumber="404">Measurements.</emphasis>
Total length: 11.2–12.5; head length: 1.2–1.4; head width: 2.5–2.6; pronotal length: 2.2–2.5; pronotal width: 6.6–7.3; abdominal width: 6.1–6.8; length of antennal segments (I: 0.6–0.8; II: 1.1–1.2; III: 1.5– 1.6; IV: 2.5–2.7; V: 2.7).
</paragraph>
<paragraph id="8BF536BDFF80BE1645D95A8DFC4AF860" blockId="9.[151,1437,978,2011]" pageId="9" pageNumber="404">
Male: Pygophore with dorsal rim slightly concave (
<figureCitation id="13712A38FF80BE1646235A8DFC2CF900" box="[829,919,1698,1723]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="9" pageNumber="404">Fig. 21</figureCitation>
), furrowed and posterolateral angle poorly developed (
<figureCitation id="13712A38FF80BE1644035AE9FE15F964" box="[285,430,1734,1759]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="9" pageNumber="404">Figs. 21, 23</figureCitation>
). Genital cup process ogival, long, curved, located lateral to proctiger; apex rounded, curved posteriorly, surpassing the dorsal rim in lateral view (
<figureCitation id="13712A38FF80BE1646235AC5FC7CF8B8" box="[829,967,1770,1795]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="9" pageNumber="404">Figs. 21–22</figureCitation>
). Parameres flat, mesial surface shallowly concave; anterior expansion digitiform and short, apex bifid due to small dark lobes (
<figureCitation id="13712A38FF80BE16414C5B21FABDF89C" box="[1106,1286,1806,1831]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="9" pageNumber="404">Figs. 21–22, 24</figureCitation>
); apex barely surpassing dorsal rim in lateral view (
<figureCitation id="13712A38FF80BE1647235B1DFD2AF8F0" box="[573,657,1842,1867]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="9" pageNumber="404">Fig. 22</figureCitation>
). Dorsal face of proctiger twice as wide as long; with a distal shallow concavity (
<figureCitation id="13712A38FF80BE16440C5B79FE27F8D4" box="[274,412,1878,1903]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="9" pageNumber="404">Figs. 21–22</figureCitation>
). Lateral face of proctiger with a slight constriction (
<figureCitation id="13712A38FF80BE1646FB5B79FB83F8D4" box="[997,1080,1878,1903]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="9" pageNumber="404">Fig. 21</figureCitation>
). Lateral expansion of proctiger trapezoidal to cordiform, large, three or more times wider than the anal opening, concave; apices rounded and directed posteriorly, projected dorsally, almost reaching the level of dorsal face (
<figureCitation id="13712A38FF80BE1641255BB1FB7CF80C" box="[1083,1223,1950,1975]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="9" pageNumber="404">Figs. 21–23</figureCitation>
). Ventral rim with developed lobes, not reaching the level of the posterolateral angles (
<figureCitation id="13712A38FF80BE16468E5BEDFC5FF860" box="[912,996,1986,2011]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="9" pageNumber="404">Fig. 23</figureCitation>
).
</paragraph>
<caption id="DF356635FF83BE1545895A14FAE1F9CA" httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="10" pageNumber="405" targetBox="[163,1412,190,1594]" targetPageId="10">
<paragraph id="8BF536BDFF83BE1545895A14FAE1F9CA" blockId="10.[151,1436,1595,1649]" pageId="10" pageNumber="405">
<emphasis id="B93EEAAFFF83BE1545895A14FEF4F9EA" bold="true" box="[151,335,1595,1618]" pageId="10" pageNumber="405">FIGURES 21–25</emphasis>
,
<taxonomicName id="4C4A4D3EFF83BE1544415A13FDD1F9E9" box="[351,618,1596,1618]" class="Insecta" family="Pentatomidae" genus="Paraedessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="ecuadoriensis" status="sp. nov.">
<emphasis id="B93EEAAFFF83BE1544415A13FDD1F9E9" box="[351,618,1596,1618]" italics="true" pageId="10" pageNumber="405">Paraedessa ecuadoriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57D4FF83BE15476A5A13FD7DF9E9" box="[628,710,1596,1618]" pageId="10" pageNumber="405" rank="species">
<emphasis id="B93EEAAFFF83BE15476A5A13FD7BF9EA" bold="true" box="[628,704,1596,1617]" pageId="10" pageNumber="405">sp. nov</emphasis>
.
</taxonomicNameLabel>
21–23 Male, pygophore; 21—dorsal view; 22—posterior view; 23— ventral view. (Scale = 1mm). 24—Paramere, lateral view. (Scale = 0,5mm). 25—Female, genital plates. (Scale = 1 mm).
</paragraph>
</caption>
<paragraph id="8BF536BDFF83BE1545D95AB7FCC1F942" blockId="10.[151,1436,1688,2037]" pageId="10" pageNumber="405">
Female: Gonocoxites 8 subtriangular, greatly reduced, laterally displaced and widely separated medially; posterolateral angle not notably developed or projected and contiguous with gonapophyses 8 (
<figureCitation id="13712A38FF83BE15402F5A93FA34F96E" box="[1329,1423,1724,1749]" captionStart="FIGURES 21 – 25" captionStartId="10.[151,264,1595,1618]" captionTargetBox="[163,1412,190,1594]" captionTargetId="figure@10.[158,1429,169,1598]" captionTargetPageId="10" captionText="FIGURES 21 – 25, Paraedessa ecuadoriensis sp. nov. 21 – 23 Male, pygophore; 21 — dorsal view; 22 — posterior view; 23 — ventral view. (Scale = 1 mm). 24 — Paramere, lateral view. (Scale = 0,5 mm). 25 — Female, genital plates. (Scale = 1 mm)." httpUri="https://zenodo.org/record/222980/files/figure.png" pageId="10" pageNumber="405">Fig. 25</figureCitation>
). Gonocoxites 8 overlapping gonapophyses
<quantity id="4CB29B58FF83BE15476D5ACFFD25F942" box="[627,670,1760,1785]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="10" pageNumber="405" unit="in" value="8.0">8 in</quantity>
all of its extension.
</paragraph>
</subSubSection>
<subSubSection id="C3506536FF83BE1545D95B2BFEF7F86A" pageId="10" pageNumber="405" type="discussion">
<paragraph id="8BF536BDFF83BE1545D95B2BFEF7F86A" blockId="10.[151,1436,1688,2037]" pageId="10" pageNumber="405">
<emphasis id="B93EEAAFFF83BE1545D95B2BFEF4F8A6" bold="true" box="[199,335,1796,1821]" pageId="10" pageNumber="405">Comments.</emphasis>
<taxonomicName id="4C4A4D3EFF83BE1544435B2BFD32F8A6" box="[349,649,1796,1821]" class="Insecta" family="Pentatomidae" genus="Paraedessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="ecuadoriensis" status="sp. nov.">
<emphasis id="B93EEAAFFF83BE1544435B2BFD32F8A6" box="[349,649,1796,1821]" italics="true" pageId="10" pageNumber="405">Paraedessa ecuadoriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57D4FF83BE1547895B2BFD4CF8A6" box="[663,759,1796,1821]" pageId="10" pageNumber="405" rank="species">
<emphasis id="B93EEAAFFF83BE1547895B2BFD4CF8A6" bold="true" box="[663,759,1796,1821]" pageId="10" pageNumber="405">sp. nov.</emphasis>
</taxonomicNameLabel>
is recognized by having an ogival genital cup process; parameres with bifid apex; lateral expansion of proctiger trapezoidal or cordiform; gonocoxites 8 subtriangular and greatly reduced. This species shares the lateral face of the proctiger slightly constricted with
<taxonomicName id="4C4A4D3EFF83BE1541A55B62FAC9F8DE" box="[1211,1394,1868,1893]" class="Insecta" family="Pentatomidae" genus="Paraedessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="albomaculata" status="sp. nov.">
<emphasis id="B93EEAAFFF83BE1541A55B62FAC9F8DE" box="[1211,1394,1868,1893]" italics="true" pageId="10" pageNumber="405">P. albomaculata</emphasis>
</taxonomicName>
<emphasis id="B93EEAAFFF83BE1540645B63FF73F832" bold="true" pageId="10" pageNumber="405">
<taxonomicNameLabel id="A20D57D4FF83BE1540645B63FF73F832" pageId="10" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C4A4D3EFF83BE1545C95B5EFEE6F832" box="[215,349,1904,1929]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="heymonsi">
<emphasis id="B93EEAAFFF83BE1545C95B5EFEE6F832" box="[215,349,1904,1929]" italics="true" pageId="10" pageNumber="405">P. heymonsi</emphasis>
</taxonomicName>
,
<emphasis id="B93EEAAFFF83BE1544725B5EFD35F832" box="[364,654,1904,1929]" italics="true" pageId="10" pageNumber="405">
<taxonomicName id="4C4A4D3EFF83BE1544725B5EFDBDF832" box="[364,518,1904,1929]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="paravinula">P. paravinula</taxonomicName>
,
<taxonomicName id="4C4A4D3EFF83BE15470C5B5EFD35F832" box="[530,654,1904,1929]" class="Insecta" family="Pentatomidae" genus="Paraedessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="silvicola" status="sp. nov.">P. silvicola</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20D57D4FF83BE15478B5B5FFD4BF832" box="[661,752,1904,1929]" pageId="10" pageNumber="405" rank="species">
<emphasis id="B93EEAAFFF83BE15478B5B5FFD4BF832" bold="true" box="[661,752,1904,1929]" pageId="10" pageNumber="405">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C4A4D3EFF83BE1547E15B5EFCD3F832" box="[767,872,1904,1929]" class="Insecta" family="Pentatomidae" genus="Paraedessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="stolida">
<emphasis id="B93EEAAFFF83BE1547E15B5EFCD3F832" box="[767,872,1904,1929]" italics="true" pageId="10" pageNumber="405">P. stolida</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4A4D3EFF83BE1546BE5B5EFBA4F832" box="[928,1055,1904,1929]" class="Insecta" family="Pentatomidae" genus="Paraedessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="verhoeffi">
<emphasis id="B93EEAAFFF83BE1546BE5B5EFBA4F832" box="[928,1055,1904,1929]" italics="true" pageId="10" pageNumber="405">P. verhoeffi</emphasis>
</taxonomicName>
. The species
<taxonomicName id="4C4A4D3EFF83BE1541A55B5EFAC8F832" box="[1211,1395,1904,1929]" class="Insecta" family="Pentatomidae" genus="Paraedessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="ecuadoriensis" status="sp. nov.">
<emphasis id="B93EEAAFFF83BE1541A55B5EFAC8F832" box="[1211,1395,1904,1929]" italics="true" pageId="10" pageNumber="405">P. ecuadoriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57D4FF83BE1540645B5FFF73F816" pageId="10" pageNumber="405" rank="species">
<emphasis id="B93EEAAFFF83BE1540645B5FFF73F816" bold="true" pageId="10" pageNumber="405">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C4A4D3EFF83BE1545CB5BBAFEE0F816" box="[213,347,1940,1965]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="heymonsi">
<emphasis id="B93EEAAFFF83BE1545CB5BBAFEE0F816" box="[213,347,1940,1965]" italics="true" pageId="10" pageNumber="405">P. heymonsi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4A4D3EFF83BE15448E5BBAFDB5F816" box="[400,526,1940,1965]" class="Insecta" family="Pentatomidae" genus="Paraedessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="405" phylum="Arthropoda" rank="species" species="verhoeffi">
<emphasis id="B93EEAAFFF83BE15448E5BBAFDB5F816" box="[400,526,1940,1965]" italics="true" pageId="10" pageNumber="405">P. verhoeffi</emphasis>
</taxonomicName>
have the dorsal face of proctiger with distal shallow concavity and gonocoxites 8 greatly reduced.
</paragraph>
</subSubSection>
<subSubSection id="C3506536FF83BE1545D95BF3FB18F84E" box="[199,1187,2012,2037]" pageId="10" pageNumber="405" type="distribution">
<paragraph id="8BF536BDFF83BE1545D95BF3FB18F84E" blockId="10.[151,1436,1688,2037]" box="[199,1187,2012,2037]" pageId="10" pageNumber="405">
<emphasis id="B93EEAAFFF83BE1545D95BF3FEE2F84E" bold="true" box="[199,345,2012,2037]" pageId="10" pageNumber="405">Distribution</emphasis>
(
<figureCitation id="13712A38FF83BE1544765BF3FE06F84E" box="[360,445,2012,2037]" captionStart="FIGURE 59" captionStartId="20.[151,250,1685,1707]" captionTargetBox="[233,1371,263,1673]" captionTargetId="figure@20.[212,1378,263,1676]" captionTargetPageId="20" captionText="FIGURE 59. Distribution of the species of Paraedessa." httpUri="https://zenodo.org/record/222988/files/figure.png" pageId="10" pageNumber="405">Fig. 59</figureCitation>
):
<emphasis id="B93EEAAFFF83BE1544CF5BF3FDC2F84E" bold="true" box="[465,633,2012,2037]" pageId="10" pageNumber="405">
<collectingCountry id="F35D762DFF83BE1544CF5BF3FDCFF84E" box="[465,628,2012,2037]" name="Colombia" pageId="10" pageNumber="405">COLOMBIA</collectingCountry>
:
</emphasis>
Meta;
<emphasis id="B93EEAAFFF83BE1547D65BF3FCE4F84E" bold="true" box="[712,863,2012,2037]" pageId="10" pageNumber="405">
<collectingCountry id="F35D762DFF83BE1547D65BF3FCE1F84E" box="[712,858,2012,2037]" name="Ecuador" pageId="10" pageNumber="405">ECUADOR</collectingCountry>
:
</emphasis>
Sucumbios, Napo, Orellana.
</paragraph>
</subSubSection>
</treatment>
</document>
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