303 lines
36 KiB
XML
303 lines
36 KiB
XML
<document id="65D7DD0E492E747D9C7EDAB93A3FAA46" ID-CLB-Dataset="25522" ID-DOI="10.11646/zootaxa.4671.2.12" ID-GBIF-Dataset="f94dcbe7-79ce-4dc6-8061-97e56f8bbbff" ID-ISSN="1175-5326" ID-Zenodo-Dep="3442192" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1568800056163" checkinUser="plazi" docAuthor="Kosterin, Oleg E." docDate="2019" docId="03DBB5600F2EFFA6A6A6FD684CDBFCCA" docLanguage="en" docName="zootaxa.4671.2.12.pdf.imf" docOrigin="Zootaxa 4671 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lestes Leach 1815" docType="treatment" docVersion="7" lastPageNumber="300" masterDocId="FFE2CD180F2CFFA5A661FFB24F5CFF84" masterDocTitle="Amendments and updates to F. C. Fraser’s key to Indian Lestes spp. (Odonata Lestidae) to resolve confusion of L. patricia Fraser, 1924 and L. nigriceps Fraser 1924, with notes on L. nodalis Selys 1891 and L. garoensis Lahiri, 1987" masterLastPageNumber="300" masterPageNumber="297" pageNumber="299" updateTime="1698755086446" updateUser="ExternalLinkService">
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<mods:titleInfo id="84DD41422EB6385002245FA8313736FD">
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<mods:title id="506870119154CB53CE5AAB9B35CCD1F5">Amendments and updates to F. C. Fraser’s key to Indian Lestes spp. (Odonata Lestidae) to resolve confusion of L. patricia Fraser, 1924 and L. nigriceps Fraser 1924, with notes on L. nodalis Selys 1891 and L. garoensis Lahiri, 1987</mods:title>
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<mods:name id="2078A690E55D6EE11E5AF38F50BCA4BF" type="personal">
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<mods:roleTerm id="0316EED3EAFA8FCD069478562A36DC01">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2EF477D8D85F6D5355783F388A07D18A">Kosterin, Oleg E.</mods:namePart>
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<mods:typeOfResource id="22E2A4C29A33D90678A8448926E99CF7">text</mods:typeOfResource>
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<mods:titleInfo id="9FFA9E6A696A1000575C21763D0FCC8B">
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<mods:title id="8E49A7544F20CB64BCFB91B49739CC00">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F4F5857B1630283EAA8D474764AB18E8">
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<mods:date id="EF55C4309CAB2F858C85D32143A262ED">2019</mods:date>
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<mods:number id="7A1B5BD275AA1AB7545CC384D5EE3F85">2019-09-17</mods:number>
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<mods:detail id="80BB66BF463C2613C923DB77B3FDB7E2" type="volume">
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<mods:number id="AC9FF91D4CAB5ACEF8B605B3E0ED9DA5">4671</mods:number>
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<mods:number id="3DBDA563B51E0B0B0ABC419E3E688D8A">2</mods:number>
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<mods:start id="EF0AFFFF40B13755E5A012E42A7A7D22">297</mods:start>
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<mods:end id="D0DECA80DA58A9D257C1997D278C6D51">300</mods:end>
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<mods:classification id="D881AD2F9C93F24E8EDAFEE82A25FE87">journal article</mods:classification>
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<mods:identifier id="4B167538D3B75F4D46615B6DF20C5C98" type="DOI">10.11646/zootaxa.4671.2.12</mods:identifier>
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<mods:identifier id="A70CACC6DB037140F4114EB0C26FCEBC" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03DBB5600F2EFFA6A6A6FD684CDBFCCA" ID-GBIF-Taxon="159608339" LSID="urn:lsid:plazi:treatment:03DBB5600F2EFFA6A6A6FD684CDBFCCA" httpUri="http://treatment.plazi.org/id/03DBB5600F2EFFA6A6A6FD684CDBFCCA" lastPageId="3" lastPageNumber="300" pageId="2" pageNumber="299">
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<paragraph id="8BCD04760F2EFFA7A6A6FD684E0CFB26" blockId="2.[151,1437,154,1186]" pageId="2" pageNumber="299">
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<subSubSection id="C36857FD0F2EFFA7A6A6FD684B43FD76" box="[199,1055,730,754]" pageId="2" pageNumber="299" type="discussion">For a reader’s convenience, the amended Fraser’s key to males of the species of</subSubSection>
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<emphasis id="B906D8640F2EFFA7A246FD694B2CFD76" box="[1063,1136,731,754]" italics="true" pageId="2" pageNumber="299">
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<subSubSection id="C36857FD0F2EFFA7A246FD694B30FD76" box="[1063,1132,731,754]" pageId="2" pageNumber="299" type="nomenclature">
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<taxonomicName id="4C727FF50F2EFFA7A246FD694B30FD76" ID-CoL="62WZW" authorityName="Leach" authorityYear="1815" box="[1063,1132,731,754]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">Lestes</taxonomicName>
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</subSubSection>
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<subSubSection id="C36857FD0F2EFFA7A20DFD694B2CFD76" box="[1132,1136,731,754]" pageId="2" pageNumber="299" type="discussion">,</subSubSection>
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</emphasis>
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<taxonomicName id="4C727FF50F2EFFA7A216FD684A32FD76" authority="Selys, 1862" authorityName="Selys" authorityYear="1862" box="[1143,1390,730,754]" class="Insecta" family="Lestidae" genus="Platylestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
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<emphasis id="B906D8640F2EFFA7A216FD684BB7FD76" box="[1143,1259,730,754]" italics="true" pageId="2" pageNumber="299">Platylestes</emphasis>
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Selys, 1862
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</taxonomicName>
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and
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<taxonomicName id="4C727FF50F2EFFA7A6F6FD4C4EC4FC92" authority="Fraser, 1951" authorityName="Fraser" authorityYear="1951" box="[151,408,766,790]" class="Insecta" family="Lestidae" genus="Sinhalestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
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<emphasis id="B906D8640F2EFFA7A6F6FD4C4E53FC92" box="[151,271,766,790]" italics="true" pageId="2" pageNumber="299">Sinhalestes</emphasis>
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<bibRefCitation id="EFE379870F2EFFA7A774FD4C4EC4FC92" author="Fraser, F. C." box="[277,408,766,790]" pageId="2" pageNumber="299" pagination="61 - 69" refId="ref5002" refString="Fraser, F. C. (1951) Outline of a new classification for the legion Lestes Selys (Order Odonata). Entomological News, 60 (2), 61 - 69." type="journal article" year="1951">Fraser, 1951</bibRefCitation>
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</taxonomicName>
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of
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<collectingCountry id="F36544E60F2EFFA7A7DAFD4C4EA8FC92" box="[443,500,766,790]" name="India" pageId="2" pageNumber="299">India</collectingCountry>
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,
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<collectingCountry id="F36544E60F2EFFA7A79FFD4C4D07FC92" box="[510,603,766,790]" name="Pakistan" pageId="2" pageNumber="299">Pakistan</collectingCountry>
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,
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<collectingCountry id="F36544E60F2EFFA7A404FD4C4D93FC92" box="[613,719,766,790]" name="Sri Lanka" pageId="2" pageNumber="299">Sri Lanka</collectingCountry>
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,
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<collectingCountry id="F36544E60F2EFFA7A4B8FD4C4C09FC92" box="[729,853,766,790]" name="Bangladesh" pageId="2" pageNumber="299">Bangladesh</collectingCountry>
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and
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<collectingCountry id="F36544E60F2EFFA7A5E9FD4C4CACFC92" box="[904,1008,766,790]" name="Myanmar" pageId="2" pageNumber="299">Myanmar</collectingCountry>
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is reproduced below entirely (in spite of the lack or scarce new information on some species). The genus
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<taxonomicName id="4C727FF50F2EFFA7A52EFC904C9BFCBE" authorityName="Fraser" authorityYear="1951" box="[847,967,802,826]" class="Insecta" family="Lestidae" genus="Sinhalestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
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<emphasis id="B906D8640F2EFFA7A52EFC904C9BFCBE" box="[847,967,802,826]" italics="true" pageId="2" pageNumber="299">Sinhalestes</emphasis>
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</taxonomicName>
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was introduced by
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<bibRefCitation id="EFE379870F2EFFA7A2FAFC904A70FCBE" author="Fraser, F. C." box="[1179,1324,802,826]" pageId="2" pageNumber="299" pagination="61 - 69" refId="ref5002" refString="Fraser, F. C. (1951) Outline of a new classification for the legion Lestes Selys (Order Odonata). Entomological News, 60 (2), 61 - 69." type="journal article" year="1951">Fraser (1951)</bibRefCitation>
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for
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<taxonomicName id="4C727FF50F2EFFA7A33BFC914EBFFCDA" authority="Hagen in Selys, 1862" authorityName="Hagen in Selys" authorityYear="1862" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="orientalis">
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<emphasis id="B906D8640F2EFFA7A33BFC914FA1FCDA" italics="true" pageId="2" pageNumber="299">Lestes orientalis</emphasis>
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Hagen in Selys, 1862
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</taxonomicName>
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after publication of his monograph (
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<bibRefCitation id="EFE379870F2EFFA7A503FCF44CB9FCDA" author="Fraser, F. C." box="[866,997,838,862]" pageId="2" pageNumber="299" refId="ref4932" refString="Fraser, F. C. (1933) The fauna of British India, including Ceylon and Burma. Odonata. Vol. I. Taylor and Francis, London, 423 pp." type="book" year="1933">Fraser 1933</bibRefCitation>
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).
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<taxonomicName id="4C727FF50F2EFFA7A597FCF54A74FCDA" authority="Selys, 1891" authorityName="Selys" authorityYear="1891" box="[1014,1320,838,862]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="bilineatus">
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<emphasis id="B906D8640F2EFFA7A597FCF54BFBFCDA" box="[1014,1191,838,862]" italics="true" pageId="2" pageNumber="299">Lestes bilineatus</emphasis>
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Selys, 1891
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</taxonomicName>
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and
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<taxonomicName id="4C727FF50F2EFFA7A33BFCF54EC8FC06" authority="Schmidt, 1964" authorityName="Schmidt" authorityYear="1964" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="malaisei">
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<emphasis id="B906D8640F2EFFA7A33BFCF54FADFC06" italics="true" pageId="2" pageNumber="299">Lestes malaisei</emphasis>
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Schmidt, 1964
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</taxonomicName>
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, described from
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<collectingCountry id="F36544E60F2EFFA7A423FCD84DF7FC06" box="[578,683,874,898]" name="Myanmar" pageId="2" pageNumber="299">Myanmar</collectingCountry>
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, are not included as most probably belonging to
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<taxonomicName id="4C727FF50F2EFFA7A2C9FCD84AC0FC06" authority="Fraser, 1922" authorityName="Fraser" authorityYear="1922" box="[1192,1436,874,898]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
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<emphasis id="B906D8640F2EFFA7A2C9FCD84A48FC06" box="[1192,1300,874,898]" italics="true" pageId="2" pageNumber="299">Indolestes</emphasis>
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Fraser, 1922
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</taxonomicName>
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(see, respectively,
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<bibRefCitation id="EFE379870F2EFFA7A73CFC3C4D51FC22" author="Lieftinck, M. A." box="[349,525,910,934]" pageId="2" pageNumber="299" pagination="229 - 256" refId="ref5230" refString="Lieftinck, M. A. (1960) On the identity of some little known southeast Asiatic Odonata in European museums described by De Selys Longchamps. Memorie della Societa Entomologica Italiana, 38, 229 - 256." type="journal article" year="1960">Lieftinck (1960)</bibRefCitation>
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and
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<bibRefCitation id="EFE379870F2EFFA7A423FC3C4DA4FC22" author="Kosterin, O. E." box="[578,760,910,934]" pageId="2" pageNumber="299" pagination="561 - 575" refId="ref5092" refString="Kosterin, O. E. (2018) Rediscovery of Lestes nigriceps Fraser, 1924 (Odonata: Lestidae) in eastern Cambodia. Zootaxa, 4526 (4), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4526.4.8" type="journal article" year="2018">Kosterin (2018))</bibRefCitation>
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. Both
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<taxonomicName id="4C727FF50F2EFFA7A55FFC3D4CF5FC22" authorityName="Selys Apices" authorityYear="1891" box="[830,937,910,934]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="nodalis">
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<emphasis id="B906D8640F2EFFA7A55FFC3D4CF5FC22" box="[830,937,910,934]" italics="true" pageId="2" pageNumber="299">L. nodalis</emphasis>
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</taxonomicName>
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(occurring at least in Lower
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<collectingCountry id="F36544E60F2EFFA7A280FC3C4A16FC22" box="[1249,1354,910,934]" name="Myanmar" pageId="2" pageNumber="299">Myanmar</collectingCountry>
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) and
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<taxonomicName id="4C727FF50F2EFFA7A3E6FC3D4FA2FC4D" authorityName="in Lahiri" authorityYear="1987" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="garoensis">
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<emphasis id="B906D8640F2EFFA7A3E6FC3D4FA2FC4D" italics="true" pageId="2" pageNumber="299">L. garoensis</emphasis>
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</taxonomicName>
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are included. The volume and limits of the genus
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<taxonomicName id="4C727FF50F2EFFA7A573FC004CDAFC4E" authorityName="Selys" authorityYear="1862" box="[786,902,946,970]" class="Insecta" family="Lestidae" genus="Platylestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
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<emphasis id="B906D8640F2EFFA7A573FC004CDAFC4E" box="[786,902,946,970]" italics="true" pageId="2" pageNumber="299">Platylestes</emphasis>
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</taxonomicName>
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are unclear and the attribution of the
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<taxonomicName id="4C727FF50F2EFFA7A374FC014ACFFC4E" authorityName="Hagen in Selys" authorityYear="1862" box="[1301,1427,947,970]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="praemorsus">
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<emphasis id="B906D8640F2EFFA7A374FC014ACFFC4E" box="[1301,1427,947,970]" italics="true" pageId="2" pageNumber="299">praemorsus</emphasis>
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</taxonomicName>
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- group to either
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<taxonomicName id="4C727FF50F2EFFA7A75FFC654EDDFC6A" authorityName="Leach" authorityYear="1815" box="[318,385,983,1006]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
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<emphasis id="B906D8640F2EFFA7A75FFC654EDDFC6A" box="[318,385,983,1006]" italics="true" pageId="2" pageNumber="299">Lestes</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4C727FF50F2EFFA7A7C6FC644D47FC6A" authorityName="Selys" authorityYear="1862" box="[423,539,982,1006]" class="Insecta" family="Lestidae" genus="Platylestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
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<emphasis id="B906D8640F2EFFA7A7C6FC644D47FC6A" box="[423,539,982,1006]" italics="true" pageId="2" pageNumber="299">Platylestes</emphasis>
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</taxonomicName>
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is equivocal (
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<bibRefCitation id="EFE379870F2EFFA7A4D2FC644C0EFC6A" author="Kosterin, O. E." box="[691,850,982,1006]" pageId="2" pageNumber="299" pagination="561 - 575" refId="ref5092" refString="Kosterin, O. E. (2018) Rediscovery of Lestes nigriceps Fraser, 1924 (Odonata: Lestidae) in eastern Cambodia. Zootaxa, 4526 (4), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4526.4.8" type="journal article" year="2018">Kosterin 2018</bibRefCitation>
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); here they are referred to in
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<taxonomicName id="4C727FF50F2EFFA7A2EEFC654B8EFC6A" authorityName="Leach" authorityYear="1815" box="[1167,1234,983,1006]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
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<emphasis id="B906D8640F2EFFA7A2EEFC654B8EFC6A" box="[1167,1234,983,1006]" italics="true" pageId="2" pageNumber="299">Lestes</emphasis>
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</taxonomicName>
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just in accordance with the current practice. Also I abstain from inclusion of subspecies of
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<taxonomicName id="4C727FF50F2EFFA7A5C7FC494A01FB96" authority="Hagen in Selys, 1862" authorityName="Hagen in Selys" authorityYear="1862" box="[934,1373,1018,1042]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="praemorsus">
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<emphasis id="B906D8640F2EFFA7A5C7FC494B32FB96" box="[934,1134,1019,1042]" italics="true" pageId="2" pageNumber="299">Lestes praemorsus</emphasis>
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Hagen in Selys, 1862
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</taxonomicName>
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since this matter is insufficiently resolved. The measurement ranges for
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<taxonomicName id="4C727FF50F2EFFA7A53BFBAD4BD2FBB2" authority="Fraser, 1923" authorityName="Fraser" authorityYear="1923" box="[858,1166,1054,1078]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="dorothea">
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<emphasis id="B906D8640F2EFFA7A53BFBAD4B5FFBB2" box="[858,1027,1054,1078]" italics="true" pageId="2" pageNumber="299">Lestes dorothea</emphasis>
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Fraser, 1923
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</taxonomicName>
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are given as extended by
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<bibRefCitation id="EFE379870F2EFFA7A6F6FBF04E76FBDE" author="Lahiri, A. R." box="[151,298,1090,1114]" pageId="2" pageNumber="299" pagination="1 - 402" refId="ref5140" refString="Lahiri, A. R. (1987) Studies on the odonate fauna of Meghalaya. Records of the Zoological Survey of India, Miscellaneous publication, Occasional paper No. 99, 1 - 402." type="book chapter" year="1987">Lahiri (1987)</bibRefCitation>
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. Fraser’s “thorax” is replaced by a more correct “synthorax”. Fraser’s statement “Thorax immaculate” for
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<taxonomicName id="4C727FF50F2EFFA7A6F6FBD54EEDFBFA" authority="Selys, 1891" authorityName="Selys Apices" authorityYear="1891" box="[151,433,1126,1150]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="nodalis">
|
||
<emphasis id="B906D8640F2EFFA7A6F6FBD54E73FBFA" box="[151,303,1126,1150]" italics="true" pageId="2" pageNumber="299">Lestes nodalis</emphasis>
|
||
Selys, 1891
|
||
</taxonomicName>
|
||
contradicts his own description of this species (
|
||
<bibRefCitation id="EFE379870F2EFFA7A5CCFBD44B72FBFA" author="Fraser, F. C." box="[941,1070,1126,1150]" pageId="2" pageNumber="299" refId="ref4932" refString="Fraser, F. C. (1933) The fauna of British India, including Ceylon and Burma. Odonata. Vol. I. Taylor and Francis, London, 423 pp." type="book" year="1933">Fraser 1933</bibRefCitation>
|
||
) and corrected to “Synthorax with few black spots”.
|
||
</paragraph>
|
||
<subSubSection id="C36857FD0F2EFFA6A6F6FB7F4AC1FD8B" lastPageId="3" lastPageNumber="300" pageId="2" pageNumber="299" type="key">
|
||
<paragraph id="8BCD04760F2EFFA7A6F6FB7F4AC0FB7B" blockId="2.[151,1437,1229,1933]" box="[151,1436,1229,1280]" pageId="2" pageNumber="299">1. Synthorax without metallic markings ...................................................................... 2</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A683FAB54AC0FA99" blockId="2.[151,1437,1229,1933]" box="[226,1436,1287,1309]" pageId="2" pageNumber="299">Synthorax with green metallic antehumeral stripes........................................................... 6</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A6F6FA974AC0FAD3" blockId="2.[151,1437,1229,1933]" box="[151,1436,1317,1368]" pageId="2" pageNumber="299">2. Pterostigma unicolorous................................................................................ 3</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A683FAED4AC0FAF0" blockId="2.[151,1437,1229,1933]" box="[226,1436,1375,1397]" pageId="2" pageNumber="299">Pterostigma bicolorous................................................................................ 4</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A6F6FACF4AC1FA49" blockId="2.[151,1437,1229,1933]" pageId="2" pageNumber="299">
|
||
3. No broad black dorsal stripe on synthorax (middorsal carina can be finely stressed with black); occiput reddish brown to creamy white; paraprocts extend at most to 1/3 of expanded part of cerci.................................
|
||
<taxonomicName id="4C727FF50F2EFFA7A28FFA0A4AC1FA49" ID-CoL="6Q2RW" authority="Hag. Synthorax" authorityName="Hag. Synthorax" box="[1262,1437,1464,1485]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="concinnus">
|
||
<emphasis id="B906D8640F2EFFA7A28FFA0A4A34FA49" box="[1262,1384,1464,1485]" italics="true" pageId="2" pageNumber="299">L. concinnus</emphasis>
|
||
Hag.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A683FA674AC1F983" blockId="2.[151,1437,1229,1933]" pageId="2" pageNumber="299">
|
||
Synthorax with a broad black dorsal stripe; occiput mat black; paraprocts extend to the end of expanded part of cerci..............................................................................................
|
||
<taxonomicName id="4C727FF50F2EFFA7A363FA414AC1F983" ID-CoL="6Q2RT" authority="Fras." authorityName="Fraser" authorityYear="1924" box="[1282,1437,1522,1544]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="patricia">
|
||
<emphasis id="B906D8640F2EFFA7A363FA414A3BF98C" box="[1282,1383,1523,1544]" italics="true" pageId="2" pageNumber="299">L. patricia</emphasis>
|
||
Fras.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A6F6F9BD4AC1F9C7" blockId="2.[151,1437,1229,1933]" box="[151,1437,1551,1603]" pageId="2" pageNumber="299">
|
||
4. Synthorax with about 10 small black spots on each side; pterostigma white at both ends..............
|
||
<taxonomicName id="4C727FF50F2EFFA7A2B8F99F4AC1F9C7" ID-CoL="4JSCD" authority="Ramb. Synthorax" authorityName="Ramb. Synthorax" box="[1241,1437,1581,1603]" class="Insecta" family="Lestidae" genus="Platylestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="platystylus">
|
||
<emphasis id="B906D8640F2EFFA7A2B8F99F4A0BF9C6" box="[1241,1367,1581,1602]" italics="true" pageId="2" pageNumber="299">P. platystylus</emphasis>
|
||
Ramb.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A683F9F84AC0F9F9" blockId="2.[151,1437,1229,1933]" pageId="2" pageNumber="299">Synthorax (in mature individuals) with few black spots; pterostigma with its costal third palest brown, posterior two thirds dark brown.............................................................................................. 5</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A6F6F9364AC0F93C" blockId="2.[151,1437,1229,1933]" box="[151,1436,1668,1720]" pageId="2" pageNumber="299">
|
||
5. Apices of cerci rounded, of their inner margins blunt.............................................
|
||
<taxonomicName id="4C727FF50F2EFFA7A363F9114AC0F93C" ID-CoL="6Q2RS" authority="Selys Apices" authorityName="Selys Apices" authorityYear="1891" box="[1282,1436,1698,1720]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="nodalis">
|
||
<emphasis id="B906D8640F2EFFA7A363F9114A3FF933" box="[1282,1379,1698,1720]" italics="true" pageId="2" pageNumber="299">L. nodalis</emphasis>
|
||
Selys
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A683F90D4AC1F951" blockId="2.[151,1437,1229,1933]" box="[226,1437,1727,1749]" pageId="2" pageNumber="299">
|
||
Apices of cerci and their inner inflations strongly pointed.......................................
|
||
<taxonomicName id="4C727FF50F2EFFA7A287F9724AC1F951" ID-CoL="6PQS6" authority="Lahiri" authorityName="in Lahiri" authorityYear="1987" box="[1254,1437,1727,1749]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="garoensis">
|
||
<emphasis id="B906D8640F2EFFA7A287F9724A07F951" box="[1254,1371,1728,1749]" italics="true" pageId="2" pageNumber="299">L. garoensis</emphasis>
|
||
Lahiri
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A6F6F96F4AC0F88B" blockId="2.[151,1437,1229,1933]" box="[151,1436,1757,1808]" pageId="2" pageNumber="299">6. Synthorax with a uniform narrow antehumeral stripe on each side.............................................. 7</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A683F8A54AC0F8A8" blockId="2.[151,1437,1229,1933]" box="[226,1436,1815,1837]" pageId="2" pageNumber="299">Synthorax with narrow antehumeral stripes expanded outwardly at the upper side.................................. 9</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A683F8874AC0F8CE" blockId="2.[151,1437,1229,1933]" box="[226,1436,1845,1867]" pageId="2" pageNumber="299">Synthorax with antehumeral stripes deeply scalloped or crenulate on the outer sides............................... 10</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A683F8E04AC0F8E3" blockId="2.[151,1437,1229,1933]" box="[226,1436,1874,1896]" pageId="2" pageNumber="299">Synthorax with the whole of dorsum and forepart of sides briliant metallic green................................. 10</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A6F6F8C14AC1F83A" blockId="2.[151,1437,1229,1933]" box="[151,1437,1907,1983]" lastBlockId="2.[151,1437,1961,2040]" pageId="2" pageNumber="299">
|
||
7. Vertex of head metallic green; thorax largely black; only 11 postnodal nervures to fore-wings............
|
||
<taxonomicName id="4C727FF50F2EFFA7A293F81B4AC1F83A" ID-CoL="6Q33Y" authority="Fras." authorityName="Fras." box="[1266,1437,1961,1982]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="angularis">
|
||
<emphasis id="B906D8640F2EFFA7A293F81B4A3BF83A" box="[1266,1383,1961,1982]" italics="true" pageId="2" pageNumber="299">L. angularis</emphasis>
|
||
Fras.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA7A683F8744AC0F85F" blockId="2.[151,1437,1961,2040]" box="[226,1436,1990,2012]" pageId="2" pageNumber="299">Vertex of head non-metallic; 12 to 14 postnodal nervures fore-wings............................................ 8</paragraph>
|
||
<paragraph id="8BCD04760F2EFFA6A6F6F8514AC1FF49" blockId="2.[151,1437,1961,2040]" lastBlockId="3.[151,1437,153,527]" lastPageId="3" lastPageNumber="300" pageId="2" pageNumber="299">
|
||
8. Vertex of head pale brown; dorsum of thorax pale brown, with very prominent brilliant metallic green antehumeral stripes; 12 postnodal nervures to fore-wings...........................................................
|
||
<taxonomicName id="4C727FF50F2FFFA6A289FF054AC1FF49" ID-CoL="6PR3L" authority="Ramb." authorityName="Ramb." box="[1256,1437,183,205]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="viridulus">
|
||
<emphasis id="B906D8640F2FFFA6A289FF054A0BFF48" box="[1256,1367,183,204]" italics="true" pageId="3" pageNumber="300">L. viridulus</emphasis>
|
||
Ramb.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2FFFA6A683FF674AC1FE83" blockId="3.[151,1437,153,527]" pageId="3" pageNumber="300">
|
||
Vertex of head mat black; dorsum of thorax black with an obscure narrow metallic green stripe on each side; 14 postnodal nervures to fore-wings....................................................................
|
||
<taxonomicName id="4C727FF50F2FFFA6A295FF404AC1FE83" ID-CoL="6PR3S" authority="Fras." authorityName="Fraser" authorityYear="1924" box="[1268,1437,242,263]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="nigriceps">
|
||
<emphasis id="B906D8640F2FFFA6A295FF404A3BFE83" box="[1268,1383,242,263]" italics="true" pageId="3" pageNumber="300">L. nigriceps</emphasis>
|
||
Fras.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2FFFA6A6F6FEBD4AC1FEC6" blockId="3.[151,1437,153,527]" box="[151,1437,271,322]" pageId="3" pageNumber="300">
|
||
9. Sides of thorax pale brown with a few black points................................................
|
||
<taxonomicName id="4C727FF50F2FFFA6A377FE9F4AC1FEC6" ID-CoL="6Q2RX" authority="Hag. Sides" authorityName="Hag. Sides" box="[1302,1437,300,322]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="elatus">
|
||
<emphasis id="B906D8640F2FFFA6A377FE9F4A34FEC5" box="[1302,1384,300,322]" italics="true" pageId="3" pageNumber="300">L. elatus</emphasis>
|
||
Hag.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2FFFA6A683FEFB4AC0FEDB" blockId="3.[151,1437,153,527]" box="[226,1436,329,351]" pageId="3" pageNumber="300">
|
||
Sides of thorax with two thick black bands enclosing a medial narrow brown stripe...............
|
||
<taxonomicName id="4C727FF50F2FFFA6A2A8FEF84AC0FEDB" ID-CoL="722RV" authority="Fraser" authorityName="Fraser" authorityYear="1929" box="[1225,1436,329,351]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="malabaricus">
|
||
<emphasis id="B906D8640F2FFFA6A2A8FEF84A06FEDA" box="[1225,1370,329,351]" italics="true" pageId="3" pageNumber="300">L. malabaricus</emphasis>
|
||
Fraser
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2FFFA6A6F6FED54AC0FE1E" blockId="3.[151,1437,153,527]" box="[151,1436,359,410]" pageId="3" pageNumber="300">
|
||
10. Small species, with abdomen
|
||
<quantity id="4C8AA9930F2FFFA6A461FE364D3FFE1E" box="[512,611,388,410]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="3" pageNumber="300" unit="mm" value="31.0" valueMax="32.0" valueMin="30.0">30-32 mm</quantity>
|
||
and hind-wing
|
||
<quantity id="4C8AA9930F2FFFA6A49BFE364C01FE1E" box="[762,861,388,410]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="3" pageNumber="300" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">20-21 mm</quantity>
|
||
..............................
|
||
<taxonomicName id="4C727FF50F2FFFA6A2B9FE374AC0FE1E" ID-CoL="6PQRQ" authority="Selys Larger" authorityName="Selys Larger" box="[1240,1436,388,410]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="praemorsus">
|
||
<emphasis id="B906D8640F2FFFA6A2B9FE374A3EFE1E" box="[1240,1378,389,410]" italics="true" pageId="3" pageNumber="300">L. praemorsus</emphasis>
|
||
Selys
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2FFFA6A683FE104AC1FE33" blockId="3.[151,1437,153,527]" box="[226,1437,417,439]" pageId="3" pageNumber="300">
|
||
Larger species, with abdomen
|
||
<quantity id="4C8AA9930F2FFFA6A466FE134D36FE33" box="[519,618,417,439]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.1" metricValueMin="3.6" pageId="3" pageNumber="300" unit="mm" value="38.5" valueMax="41.0" valueMin="36.0">36-41 mm</quantity>
|
||
and hind-wing
|
||
<quantity id="4C8AA9930F2FFFA6A563FE104C39FE33" box="[770,869,418,439]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.2" metricValueMin="2.4" pageId="3" pageNumber="300" unit="mm" value="28.0" valueMax="32.0" valueMin="24.0">24-32 mm</quantity>
|
||
.................................
|
||
<taxonomicName id="4C727FF50F2FFFA6A29FFE134AC1FE33" ID-CoL="6PQRX" authority="Fras." authorityName="Fraser" authorityYear="1923" box="[1278,1437,417,439]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="dorothea">
|
||
<emphasis id="B906D8640F2FFFA6A29FFE134A3BFE32" box="[1278,1383,417,438]" italics="true" pageId="3" pageNumber="300">L.dorothea</emphasis>
|
||
Fras.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2FFFA6A6F6FE0D4AC1FE76" blockId="3.[151,1437,153,527]" box="[151,1437,447,498]" pageId="3" pageNumber="300">
|
||
11. Small species, with abdomen about
|
||
<quantity id="4C8AA9930F2FFFA6A45AFE6E4D22FE76" box="[571,638,476,498]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="300" unit="mm" value="30.0">30 mm</quantity>
|
||
and hind-wing
|
||
<quantity id="4C8AA9930F2FFFA6A577FE6E4C00FE76" box="[790,860,476,498]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="300" unit="mm" value="21.0">21 mm</quantity>
|
||
; pterostigma bicolorous; palaearctic....
|
||
<taxonomicName id="4C727FF50F2FFFA6A295FE6F4AC1FE76" ID-CoL="6Q33Z" authority="Fabr. Large" authorityName="Fabr. Large" box="[1268,1437,476,498]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="barbarus">
|
||
<emphasis id="B906D8640F2FFFA6A295FE6F4A3AFE75" box="[1268,1382,476,498]" italics="true" pageId="3" pageNumber="300">L. barbarus</emphasis>
|
||
Fabr.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2FFFA6A683FE484AC1FD8B" blockId="3.[151,1437,153,527]" box="[226,1437,505,527]" pageId="3" pageNumber="300">
|
||
Large species, with abdomen
|
||
<quantity id="4C8AA9930F2FFFA6A79EFE4B4D1EFD8B" box="[511,578,505,527]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="300" unit="mm" value="52.0">52 mm</quantity>
|
||
and hind-wing
|
||
<quantity id="4C8AA9930F2FFFA6A4BBFE484C7CFD8B" box="[730,800,506,527]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="300" unit="mm" value="38.0">38 mm</quantity>
|
||
; pterostigma unicolorous, oriental...........
|
||
<taxonomicName id="4C727FF50F2FFFA6A295FE484AC1FD8B" ID-CoL="9X363" authority="Hag." authorityName="Hagen in Selys" authorityYear="1862" box="[1268,1437,505,527]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B906D8640F2FFFA6A295FE484A34FD8A" box="[1268,1384,505,527]" italics="true" pageId="3" pageNumber="300">S. orientalis</emphasis>
|
||
Hag.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36857FD0F2FFFA6A6F6FD884CDBFCCA" pageId="3" pageNumber="300" type="discussion">
|
||
<paragraph id="8BCD04760F2FFFA6A6F6FD884A10FDD6" blockId="3.[151,1437,570,847]" box="[151,1356,570,595]" pageId="3" pageNumber="300">
|
||
Naturally, using any key is insufficient to identify tropical
|
||
<taxonomicName id="4C727FF50F2FFFA6A569FD884C3EFDD6" authorityName="Fabricius" authorityYear="1793" box="[776,866,570,594]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="300" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
and the full descriptions should be consulted.
|
||
</paragraph>
|
||
<paragraph id="8BCD04760F2FFFA6A6A6FDED4C34FC82" blockId="3.[151,1437,570,847]" pageId="3" pageNumber="300">
|
||
I should make a clarification on another matter concerned in
|
||
<bibRefCitation id="EFE379870F2FFFA6A507FDEC4B49FDF3" author="Kosterin, O. E." box="[870,1045,606,631]" pageId="3" pageNumber="300" pagination="561 - 575" refId="ref5092" refString="Kosterin, O. E. (2018) Rediscovery of Lestes nigriceps Fraser, 1924 (Odonata: Lestidae) in eastern Cambodia. Zootaxa, 4526 (4), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4526.4.8" type="journal article" year="2018">Kosterin (2018)</bibRefCitation>
|
||
. Matti Hämäläinen kindly informed me that the number of 366 species described by C.H. Fraser, which I cited in
|
||
<bibRefCitation id="EFE379870F2FFFA6A5BDFD304BDBFD1F" author="Kosterin, O. E." box="[988,1159,642,667]" pageId="3" pageNumber="300" pagination="561 - 575" refId="ref5092" refString="Kosterin, O. E. (2018) Rediscovery of Lestes nigriceps Fraser, 1924 (Odonata: Lestidae) in eastern Cambodia. Zootaxa, 4526 (4), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4526.4.8" type="journal article" year="2018">Kosterin (2018)</bibRefCitation>
|
||
according to
|
||
<bibRefCitation id="EFE379870F2FFFA6A37BFD304E62FD3B" author="Wasscher, M. & Dumont, H. J." pageId="3" pageNumber="300" pagination="369 - 402" refId="ref5329" refString="Wasscher, M. & Dumont, H. J. (2013) Life and work of Michel Edmond de Selys Lonchamps (1813 - 1900), the founder of Odonatology. Odonatologica, 42 (4), 369 - 402." type="journal article" year="2013">Wasscher & Dumont (2013)</bibRefCitation>
|
||
, was strongly underestimated.
|
||
<bibRefCitation id="EFE379870F2FFFA6A4E3FD144C6EFD3A" author="Kimmins, D. E." box="[642,818,678,702]" pageId="3" pageNumber="300" pagination="173 - 227" refId="ref5038" refString="Kimmins, D. E. (1966) A list of the Odonata types described by F. C. Fraser, now in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 18 (6), 173 - 227." type="journal article" year="1966">Kimmins (1966)</bibRefCitation>
|
||
mentioned 562 species group names described by him and although some of them proposed in a subspecific rank, the number of those described as species (not necessarily considered as valid species at present) was surely more than five hundred.
|
||
</paragraph>
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<paragraph id="8BCD04760F2FFFA6A6A6FCA04CDBFCCA" blockId="3.[151,1437,570,847]" pageId="3" pageNumber="300">This work was partly supported by Russian State Scientific Project 0324-2019-0041. The author is grateful to K.D.P. Wilson and K.A. Subramanian for reviewing and improving the paper.</paragraph>
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</subSubSection>
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</treatment>
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</document> |