198 lines
26 KiB
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198 lines
26 KiB
XML
<document id="B439D83A3C0FEFAF2FDD7FD0EF5E99F4" ID-CLB-Dataset="20176" ID-DOI="10.11646/zootaxa.5116.2.5" ID-GBIF-Dataset="dc6e6f10-3b06-448e-bf5f-8bc181884366" ID-ISSN="1175-5326" ID-Zenodo-Dep="6367323" ID-ZooBank="2558C8B0-FABB-4104-B103-56C0830AF277" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647595653746" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V., Chigray, Ivan A., Ntatsopoulos, Konstantinos & Papadopoulou, Anna" docDate="2022" docId="03D0A51AFFD9FFB89C97AACFCCD296FB" docLanguage="en" docName="zootaxa.5116.2.5.pdf" docOrigin="Zootaxa 5116 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blaps subalpina Menetries 1832" docType="treatment" docVersion="7" lastPageNumber="286" masterDocId="FFE9DD62FFCAFFAB9C00A841CF16931E" masterDocTitle="A key to Russian and Eastern European species of Blaps Fabricius, 1775 (Coleoptera: Tenebrionidae: Blaptinae) with the description of a new species from the North Caucasus supported by morphological and molecular data" masterLastPageNumber="291" masterPageNumber="267" pageNumber="286" updateTime="1699318281145" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B889B41A05B6AE8DC32207D1C33C3B22">
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<mods:title id="A07E5220B94C2ECDE3E30427AB9752B2">A key to Russian and Eastern European species of Blaps Fabricius, 1775 (Coleoptera: Tenebrionidae: Blaptinae) with the description of a new species from the North Caucasus supported by morphological and molecular data</mods:title>
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<mods:name id="9A58356269644C42206683914AAB0294" type="personal">
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<mods:roleTerm id="D2507D1757C28B740E8C36C38A40F241">Author</mods:roleTerm>
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<mods:namePart id="36D6FB845A433F495B1346F4C7C29E4E">Nabozhenko, Maxim V.</mods:namePart>
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<mods:affiliation id="A90732B98D299A5EDB9474FBCBFC7B95">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000, Russia. & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000, Russia & A. N. Severtsov Institute of ecology and evolution of the Russian Academy of Sciences, Leninsky Prospect 33, 119071 Moscow, Russia Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb., 1, Saint Petersburg 199034 Russia. chigray 93 @ bk. ru; https: // orcid. org / 0000 - 0002 - 3830 - 6860</mods:affiliation>
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<mods:namePart id="21940EC39B4159872D56DD9CCD85250C">Chigray, Ivan A.</mods:namePart>
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<mods:nameIdentifier id="E6FE7408CF9A9FBE82B6520998F647BD" type="ORCID">0000-0002-3830-6860</mods:nameIdentifier>
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<mods:nameIdentifier id="B4D2E0DD7D07AC19DC0E325BA66E6BCD" type="email">chigray93@bk.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name id="2FEE7C52290439B0ED9D2E4254AA8DE9" type="personal">
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<mods:role id="679927AA3406FBF4CEFC42ED56EE1EAA">
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<mods:roleTerm id="EEBA25AAA017FD4E6BB77094FD8974CD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5EA5EF8EF30A01FDD82E147129C5AC7C">Ntatsopoulos, Konstantinos</mods:namePart>
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<mods:nameIdentifier id="8B80AC80D10960B010F08F93453ED1BC" type="ORCID">0000-0003-0673-2416</mods:nameIdentifier>
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<mods:affiliation id="F083C196F49DC5778CDAB84BE0C72B40">Department of Biological Sciences, University of Cyprus, 1678 Nicosia P. O. Box 20537, Cyprus. & ntatsopoulos. konstantinos @ ucy. ac. cy; https: // orcid. org / 0000 - 0003 - 0673 - 2416</mods:affiliation>
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<mods:nameIdentifier id="A08BD9D1F9737F03F4B7D132335C1A3B" type="email">ntatsopoulos.konstantinos@ucy.ac.cy</mods:nameIdentifier>
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<mods:namePart id="1719C54367589108D942F4F3CFEC5D7E">Papadopoulou, Anna</mods:namePart>
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<mods:nameIdentifier id="4C4031EE517C3EB76E1C2AC43DFAC49E" type="ORCID">0000-0002-4656-4894</mods:nameIdentifier>
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<mods:affiliation id="4B8BB33F643A9CAD6B63E16931065700">Department of Biological Sciences, University of Cyprus, 1678 Nicosia P. O. Box 20537, Cyprus. & papadopoulou. g. anna @ ucy. ac. cy; https: // orcid. org / 0000 - 0002 - 4656 - 4894</mods:affiliation>
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<mods:nameIdentifier id="75DEA03CAAB2D98B9E6D74FC8833E88E" type="email">papadopoulou.g.anna@ucy.ac.cy</mods:nameIdentifier>
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<mods:typeOfResource id="10AAD1D73C1A18BC79FA8888910501AC">text</mods:typeOfResource>
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<mods:title id="DFD1CA6F8B4DF006DA8E104D7865C472">Zootaxa</mods:title>
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<mods:part id="27E3150F6C4D1DD89F7B6A7B1F75394B">
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<mods:date id="5966F241D63DA38501C08CA711A391C6">2022</mods:date>
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<mods:number id="25B3C04E81982E3A1E03AA9B7C2BABC0">2022-03-18</mods:number>
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<mods:detail id="958A9BB59B4E509C563E566B12BA2511" type="volume">
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<mods:number id="44C9D067775333B194885FE5426D01DB">5116</mods:number>
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<mods:detail id="37698F05516D8B75875C22615DEDEF2A" type="issue">
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<mods:number id="79A5A246227856BCD3B059623741ED0E">2</mods:number>
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<mods:start id="F1D053EE8546734B58B508B1CAFBEFA5">267</mods:start>
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<mods:end id="7F275BB87E5198200101D596E517C4D2">291</mods:end>
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<mods:classification id="59A7F5DF46883834C84994E007F76052">journal article</mods:classification>
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<mods:identifier id="B77793698FB37ABAD305126F0C70F6C1" type="CLB-Dataset">20176</mods:identifier>
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<mods:identifier id="CE4796200F3A05F19B0D9B351FC0356D" type="DOI">10.11646/zootaxa.5116.2.5</mods:identifier>
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<treatment id="03D0A51AFFD9FFB89C97AACFCCD296FB" ID-DOI="http://doi.org/10.5281/zenodo.6367327" ID-GBIF-Taxon="194098881" ID-Zenodo-Dep="6367327" LSID="urn:lsid:plazi:treatment:03D0A51AFFD9FFB89C97AACFCCD296FB" httpUri="http://treatment.plazi.org/id/03D0A51AFFD9FFB89C97AACFCCD296FB" lastPageNumber="286" pageId="19" pageNumber="286">
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<subSubSection id="C3634787FFD9FFB89C97AACFCDD691B7" box="[151,704,654,681]" pageId="19" pageNumber="286" type="nomenclature">
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<paragraph id="8BC6140CFFD9FFB89C97AACFCDD691B7" blockId="19.[151,704,654,717]" box="[151,704,654,681]" pageId="19" pageNumber="286">
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<heading id="D08EA360FFD9FFB89C97AACFCDD691B7" bold="true" box="[151,704,654,681]" fontSize="11" level="1" pageId="19" pageNumber="286" reason="1">
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<emphasis id="B90DC81EFFD9FFB89C97AACFCDD691B7" bold="true" box="[151,704,654,681]" pageId="19" pageNumber="286">
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<taxonomicName id="4C796F8FFFD9FFB89C97AACFCD3E91B6" authority="Menetries, 1832" authorityName="Menetries" authorityYear="1832" box="[151,552,654,681]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="subalpina">
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<emphasis id="B90DC81EFFD9FFB89C97AACFCE4C91B7" bold="true" box="[151,346,654,681]" italics="true" pageId="19" pageNumber="286">Blaps subalpina</emphasis>
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<bibRefCitation id="EFE869FDFFD9FFB89D61AACFCD3E91B6" author="Menetries, E." box="[353,552,654,681]" pageId="19" pageNumber="286" refId="ref14478" refString="Menetries, E. (1832) Catalogue raissonne des objets de Zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse. St. Petersbourg, L'Imprimerie de l'Academie Imperiale des Sciences, 272 pp., 5 pls. https: // doi. org / 10.5962 / bhl. title. 51784" type="book" year="1832">Ménétriés, 1832</bibRefCitation>
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</taxonomicName>
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stat. resurr.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3634787FFD9FFB89C97AAF2CCD296FB" pageId="19" pageNumber="286" type="description">
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<paragraph id="8BC6140CFFD9FFB89C97AAF2CE4A91D3" blockId="19.[151,704,654,717]" box="[151,348,691,717]" pageId="19" pageNumber="286">
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(
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<figureCitation id="13420889FFD9FFB89C9FAAF2CFE091D3" box="[159,246,691,717]" captionStart="FIGURE 10" captionStartId="7.[151,250,1931,1955]" captionTargetBox="[154,1428,1285,1906]" captionTargetId="figure-553@7.[151,1437,1285,1907]" captionTargetPageId="7" captionText="FIGURE 10. B. subalpina, habitus. A, B = ♂; C = ♀; D = prothoracic hypomeron; A, C = dorsal view; B = ventral view." figureDoi="http://doi.org/10.5281/zenodo.6367347" httpUri="https://zenodo.org/record/6367347/files/figure.png" pageId="19" pageNumber="286">Figs 10</figureCitation>
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,
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<figureCitation id="13420889FFD9FFB89D02AAF2CE4591D3" box="[258,339,691,717]" captionStart="FIGURE 23" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[162,1430,186,1912]" captionTargetId="figure-23@20.[154,1433,181,1920]" captionTargetPageId="20" captionText="FIGURE 23. Blaps spp., habitus. A = non-rugose male of B. verrucosa from Baku, Azerbaijan, (the type locality of B. scabriuscula) ZM MSU; B = rugose male of B. verrucosa from Georgia (Kazbegi), ZIN; C = lectotype labels of B. subalpina, D = lectotype of B. subalpina, male (ZIN); E = lectotype labels of B. pterotapha; F = lectotype of B. pterotapha, female (ZIN)." figureDoi="http://doi.org/10.5281/zenodo.6367379" httpUri="https://zenodo.org/record/6367379/files/figure.png" pageId="19" pageNumber="286">23C, D</figureCitation>
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)
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</paragraph>
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<paragraph id="8BC6140CFFD9FFB89C97AABACC61962F" blockId="19.[151,1437,763,1509]" pageId="19" pageNumber="286">
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This species was described from fortress Groznaya (now Groznyi city, Chechen Republic,
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<collectingCountry id="F36E549CFFD9FFB898C6AABACA049008" box="[1222,1298,763,790]" name="Russia" pageId="19" pageNumber="286">Russia</collectingCountry>
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) or its vast surroundings; type locality: “Caucase, près de Grosnaïa”; the
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<typeStatus id="54C2AAAEFFD9FFB89F73AB5ECCCC9027" box="[883,986,799,825]" pageId="19" pageNumber="286" type="lectotype">lectotype</typeStatus>
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(ZIN) was designated and studied by
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<bibRefCitation id="EFE869FDFFD9FFB89C97AB02CD769043" author="Abdurakhmanov, G. M. & Nabozhenko, M. V." box="[151,608,835,861]" pageId="19" pageNumber="286" refId="ref12648" refString="Abdurakhmanov, G. M. & Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov & Nabozhenko (2011)</bibRefCitation>
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. Since Allard (1880, 1882), many authors have interpreted this name as a subspecies of
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<taxonomicName id="4C796F8FFFD9FFB89D54AB26CEE8909F" authorityName="Marsham" authorityYear="1802" box="[340,510,871,897]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="lethifera">
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<emphasis id="B90DC81EFFD9FFB89D54AB26CEE8909F" box="[340,510,871,897]" italics="true" pageId="19" pageNumber="286">Blaps lethifera</emphasis>
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</taxonomicName>
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based on its smooth elytra.
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<bibRefCitation id="EFE869FDFFD9FFB89F4EAB26CA0E909F" author="Abdurakhmanov, G. M. & Nabozhenko, M. V." box="[846,1304,871,897]" pageId="19" pageNumber="286" refId="ref12648" refString="Abdurakhmanov, G. M. & Nabozhenko, M. V. (2011) Keys and catalogue to darkling beetles (Coleoptera: Tenebrionidae s. str.) of the Caucasus and south of European part of Russia. KMK Scientific Press Ltd, Moscow, 361 pp. [in Russian]" type="book" year="2011">Abdurakhmanov & Nabozhenko (2011)</bibRefCitation>
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interpreted this taxon as a subspecies of
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<taxonomicName id="4C796F8FFFD9FFB89DE4ABCACC6A90BB" authority="Menetries, 1832" authorityName="Menetries" authorityYear="1832" box="[484,892,907,933]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="scabriuscula">
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<emphasis id="B90DC81EFFD9FFB89DE4ABCACDAA90BB" box="[484,700,907,933]" italics="true" pageId="19" pageNumber="286">Blaps scabriuscula</emphasis>
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<bibRefCitation id="EFE869FDFFD9FFB89EC4ABCACC6A90BB" author="Menetries, E." box="[708,892,907,933]" pageId="19" pageNumber="286" refId="ref14478" refString="Menetries, E. (1832) Catalogue raissonne des objets de Zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse. St. Petersbourg, L'Imprimerie de l'Academie Imperiale des Sciences, 272 pp., 5 pls. https: // doi. org / 10.5962 / bhl. title. 51784" type="book" year="1832">Ménétriés, 1832</bibRefCitation>
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</taxonomicName>
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and proposed the synonymy
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<taxonomicName id="4C796F8FFFD9FFB898CDABCACA9590BB" authorityName="Menetries" authorityYear="1832" box="[1229,1411,907,933]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="subalpina">
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<emphasis id="B90DC81EFFD9FFB898CDABCACA9590BB" box="[1229,1411,907,933]" italics="true" pageId="19" pageNumber="286">Blaps subalpina</emphasis>
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</taxonomicName>
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<emphasis id="B90DC81EFFD9FFB8998CABCACA8A90BB" bold="true" box="[1420,1436,907,933]" pageId="19" pageNumber="286">=</emphasis>
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<taxonomicName id="4C796F8FFFD9FFB89C97ABEECD3790D7" authority="Motschulsky, 1839" authorityName="Motschulsky" authorityYear="1839" box="[151,545,943,969]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="montana">
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<emphasis id="B90DC81EFFD9FFB89C97ABEECE5790D6" box="[151,321,943,969]" italics="true" pageId="19" pageNumber="286">Blaps montana</emphasis>
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<bibRefCitation id="EFE869FDFFD9FFB89D48ABEECD3790D7" author="Motschulsky, V. de" box="[328,545,943,969]" pageId="19" pageNumber="286" pagination="44 - 93" refId="ref14537" refString="Motschulsky, V. de (1839) Insectes du Caucase et des Provinces Transcaucausiennes. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 12, 44 - 93, pls. I, II, V, VI." type="journal article" year="1839">Motschulsky, 1839</bibRefCitation>
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</taxonomicName>
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. In fact,
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<taxonomicName id="4C796F8FFFD9FFB89E84ABEECC0490D7" authorityName="Menetries" authorityYear="1832" box="[644,786,943,969]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="subalpina">
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<emphasis id="B90DC81EFFD9FFB89E84ABEECC0490D7" box="[644,786,943,969]" italics="true" pageId="19" pageNumber="286">B. subalpina</emphasis>
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</taxonomicName>
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is a separate species (not subspecies), which differs from
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<taxonomicName id="4C796F8FFFD9FFB89C97AB92CE3090F2" authorityName="Adams" authorityYear="1817" box="[151,294,979,1004]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="verrucosa">
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<emphasis id="B90DC81EFFD9FFB89C97AB92CE3090F2" box="[151,294,979,1004]" italics="true" pageId="19" pageNumber="286">B. verrucosa</emphasis>
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</taxonomicName>
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(
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<emphasis id="B90DC81EFFD9FFB89D37AB92CE5190F3" bold="true" box="[311,327,979,1005]" pageId="19" pageNumber="286">=</emphasis>
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<taxonomicName id="4C796F8FFFD9FFB89D4EAB92CED990F2" authorityName="Motschulsky" authorityYear="1839" box="[334,463,979,1004]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="montana">
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<emphasis id="B90DC81EFFD9FFB89D4EAB92CED990F2" box="[334,463,979,1004]" italics="true" pageId="19" pageNumber="286">B. montana</emphasis>
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</taxonomicName>
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) by the shape of the narrow pronotum, narrowed to apical margin, smooth elytra, little wider mucro in males and the absence of the mucro in females. In addition, both species differ in their bionomics.
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<taxonomicName id="4C796F8FFFD9FFB89C97AC5ACE5B972A" authorityName="Adams" authorityYear="1817" box="[151,333,1051,1077]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="verrucosa">
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<emphasis id="B90DC81EFFD9FFB89C97AC5ACE5B972A" box="[151,333,1051,1077]" italics="true" pageId="19" pageNumber="286">Blaps verrucosa</emphasis>
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</taxonomicName>
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is cavernicolous species, which inhabits caves, niches and deep cracks in the rocks. High mountain populations (usually more or strongly rugose) of
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<taxonomicName id="4C796F8FFFD9FFB89ECBAC7ECC4A9746" authorityName="Adams" authorityYear="1817" box="[715,860,1087,1112]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="verrucosa">
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<emphasis id="B90DC81EFFD9FFB89ECBAC7ECC4A9746" box="[715,860,1087,1112]" italics="true" pageId="19" pageNumber="286">B. verrucosa</emphasis>
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</taxonomicName>
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prefer wet meadow mountain habitats with rocks, while lower mountain Transcaucasian populations (for example, in Gobustan desert) occur in rocks or clusters of huge boulders.
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<taxonomicName id="4C796F8FFFD9FFB89D68ACC6CD0997BF" authorityName="Menetries" authorityYear="1832" box="[360,543,1159,1185]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="subalpina">
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<emphasis id="B90DC81EFFD9FFB89D68ACC6CD0997BF" box="[360,543,1159,1185]" italics="true" pageId="19" pageNumber="286">Blaps subalpina</emphasis>
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</taxonomicName>
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inhabits xerophytic mountain steppes and slopes only on the North Caucasus (
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<collectingRegion id="49BDDAEEFFD9FFB89C9FACEACE1A97DB" box="[159,268,1195,1221]" country="Russia" name="Dagestan" pageId="19" pageNumber="286">Dagestan</collectingRegion>
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,
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<collectingRegion id="49BDDAEEFFD9FFB89D17ACEACE9F97D8" box="[279,393,1195,1222]" country="Russia" name="Chechnya" pageId="19" pageNumber="286">Chechnya</collectingRegion>
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|
) and can often be found in phrygana-like steppes at the foot of
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<taxonomicName id="4C796F8FFFD9FFB8984FACEACBEB97DB" box="[1103,1277,1195,1221]" class="Magnoliopsida" family="Fabaceae" genus="Astragalus" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="286" phylum="Tracheophyta" rank="species" species="undetermined">
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<emphasis id="B90DC81EFFD9FFB8984FACEACBDE97DB" box="[1103,1224,1195,1221]" italics="true" pageId="19" pageNumber="286">Astragalus</emphasis>
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|
spp.
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|
</taxonomicName>
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|
(milkvetches;
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|
<taxonomicName id="4C796F8FFFD9FFB89C97AC8ECE1297F4" box="[151,260,1231,1258]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="286" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
|
|
),
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|
<taxonomicName id="4C796F8FFFD9FFB89D17AC8ECE9097F7" box="[279,390,1231,1257]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="19" pageNumber="286" phylum="Tracheophyta" rank="species" species="undetermined">
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|
<emphasis id="B90DC81EFFD9FFB89D17AC8ECE4797F7" box="[279,337,1231,1257]" italics="true" pageId="19" pageNumber="286">Stipa</emphasis>
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|
spp.
|
|
</taxonomicName>
|
|
(feather grasses;
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|
<taxonomicName id="4C796F8FFFD9FFB89E4AAC8ECDBE97F7" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[586,680,1231,1257]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="19" pageNumber="286" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
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|
) and in rodents burrows. These two species are allopatric in natural non-transformed landscapes. However, both species tend to a synanthropic lifestyle and are sometimes sympatric in ruins of abandoned mountain villages in
|
|
<collectingRegion id="49BDDAEEFFD9FFB89E60AD56CDDE962F" box="[608,712,1303,1329]" country="Russia" name="Dagestan" pageId="19" pageNumber="286">Dagestan</collectingRegion>
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|
and
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|
<collectingRegion id="49BDDAEEFFD9FFB89EFFAD56CC65962F" box="[767,883,1303,1329]" country="Russia" name="Chechnya" pageId="19" pageNumber="286">Chechnya</collectingRegion>
|
|
.
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|
</paragraph>
|
|
<paragraph id="8BC6140CFFD9FFB89CC7AD7ACB579683" blockId="19.[151,1437,763,1509]" pageId="19" pageNumber="286">
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|
<taxonomicName id="4C796F8FFFD9FFB89CC7AD7ACE94964A" authorityName="Adams" authorityYear="1817" box="[199,386,1339,1365]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="verrucosa">
|
|
<emphasis id="B90DC81EFFD9FFB89CC7AD7ACE94964A" box="[199,386,1339,1365]" italics="true" pageId="19" pageNumber="286">Blaps verrucosa</emphasis>
|
|
</taxonomicName>
|
|
is a species demonstrating great morphological variation.
|
|
<location id="8EA642D7FFD9FFB89833AD7ACBF7964B" LSID="urn:lsid:plazi:treatment:03D0A51AFFD9FFB89C97AACFCCD296FB:8EA642D7FFD9FFB89833AD7ACBF7964B" box="[1075,1249,1339,1365]" country="Azerbaijan" name="High mountain" pageId="19" pageNumber="286" stateProvince="Qobustan">High mountain</location>
|
|
populations are strongly rugose and slender (
|
|
<figureCitation id="13420889FFD9FFB89DE1AD1ECD5E9667" box="[481,584,1375,1401]" captionStart="FIGURE 23" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[162,1430,186,1912]" captionTargetId="figure-23@20.[154,1433,181,1920]" captionTargetPageId="20" captionText="FIGURE 23. Blaps spp., habitus. A = non-rugose male of B. verrucosa from Baku, Azerbaijan, (the type locality of B. scabriuscula) ZM MSU; B = rugose male of B. verrucosa from Georgia (Kazbegi), ZIN; C = lectotype labels of B. subalpina, D = lectotype of B. subalpina, male (ZIN); E = lectotype labels of B. pterotapha; F = lectotype of B. pterotapha, female (ZIN)." figureDoi="http://doi.org/10.5281/zenodo.6367379" httpUri="https://zenodo.org/record/6367379/files/figure.png" pageId="19" pageNumber="286">Fig. 23B</figureCitation>
|
|
), while almost smooth (but with coarse elytral punctation) specimens have been found in subdesert
|
|
<collectingRegion id="49BDDAEEFFD9FFB89DA9ADC2CD029683" box="[425,532,1411,1437]" country="Azerbaijan" name="Qobustan" pageId="19" pageNumber="286">Gobustan</collectingRegion>
|
|
populations around
|
|
<collectingRegion id="49BDDAEEFFD9FFB89EFAADC2CC2E9683" box="[762,824,1411,1437]" country="Azerbaijan" name="Baku" pageId="19" pageNumber="286">Baku</collectingRegion>
|
|
,
|
|
<collectingCountry id="F36E549CFFD9FFB89F41ADC2CCAA9683" box="[833,956,1411,1437]" name="Azerbaijan" pageId="19" pageNumber="286">Azerbaijan</collectingCountry>
|
|
(
|
|
<figureCitation id="13420889FFD9FFB89FCBADC2CB229683" box="[971,1076,1411,1437]" captionStart="FIGURE 23" captionStartId="20.[151,250,1943,1967]" captionTargetBox="[162,1430,186,1912]" captionTargetId="figure-23@20.[154,1433,181,1920]" captionTargetPageId="20" captionText="FIGURE 23. Blaps spp., habitus. A = non-rugose male of B. verrucosa from Baku, Azerbaijan, (the type locality of B. scabriuscula) ZM MSU; B = rugose male of B. verrucosa from Georgia (Kazbegi), ZIN; C = lectotype labels of B. subalpina, D = lectotype of B. subalpina, male (ZIN); E = lectotype labels of B. pterotapha; F = lectotype of B. pterotapha, female (ZIN)." figureDoi="http://doi.org/10.5281/zenodo.6367379" httpUri="https://zenodo.org/record/6367379/files/figure.png" pageId="19" pageNumber="286">Fig. 23A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BC6140CFFD9FFB89CC7ADE6CCD296FB" blockId="19.[151,1437,763,1509]" pageId="19" pageNumber="286">
|
|
Given that the species status of
|
|
<taxonomicName id="4C796F8FFFD9FFB89E33ADE6CDFF96DF" authorityName="Menetries" authorityYear="1832" box="[563,745,1447,1473]" class="Insecta" family="Tenebrionidae" genus="Blaps" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="286" phylum="Arthropoda" rank="species" species="subalpina">
|
|
<emphasis id="B90DC81EFFD9FFB89E33ADE6CDFF96DF" box="[563,745,1447,1473]" italics="true" pageId="19" pageNumber="286">Blaps subalpina</emphasis>
|
|
</taxonomicName>
|
|
is also corroborated by molecular data (
|
|
<figureCitation id="13420889FFD9FFB898B6ADE6CA1F96DF" box="[1206,1289,1447,1473]" captionStart="FIGURE 22" captionStartId="15.[151,250,1093,1117]" captionTargetBox="[151,1429,433,1070]" captionTargetId="figure-325@15.[151,1436,433,1070]" captionTargetPageId="15" captionText="FIGURE 22. ML phylogenetic tree reconstruction of Blaps species from the Northern Caucasus in IQ-TREE. Numbers above branches represent bootstrap support values (values <50% and the ones corresponding to short intraspecific branches are omitted)." figureDoi="http://doi.org/10.5281/zenodo.6367377" httpUri="https://zenodo.org/record/6367377/files/figure.png" pageId="19" pageNumber="286">Fig. 22</figureCitation>
|
|
) (at least for specimens from
|
|
<collectingRegion id="49BDDAEEFFD9FFB89D50AD8ACEAB96FB" box="[336,445,1483,1509]" country="Russia" name="Dagestan" pageId="19" pageNumber="286">Dagestan</collectingRegion>
|
|
), the species rank for this taxon is resurrected.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |