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<mods:title id="240FAF7AC376DE8B8D620C051A0E1C64">Taxonomic study of the genus Eloeophila Rondani, 1856 of Japan (Limoniidae Diptera)</mods:title>
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Key to Japanese species of the genus 
<taxonomicName id="4C624D0C120AFA04FD965B18FD2FF8FA" ID-CoL="87H3T" authorityName="Rondani" authorityYear="1856" box="[613,745,1822,1849]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9D120AFA04FD965B18FD2FF8FA" bold="true" box="[613,745,1822,1849]" italics="true" pageId="23" pageNumber="448">Eloeophila</emphasis>
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<paragraph id="8BDD368F120AFA04FF645B62FA5AF8BA" blockId="23.[151,1436,1892,2031]" box="[151,1436,1892,1914]" pageId="23" pageNumber="448">
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1. Most of wing cells with numerous tiny spots free from veins (
<figureCitation id="13592A0A120AFA04FCC05B62FC5FF8B9" box="[819,921,1892,1914]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="6.[151,250,1529,1553]" captionStartId-1="7.[151,250,1382,1406]" captionStartId-2="9.[151,250,1501,1525]" captionStartId-3="11.[151,250,1450,1474]" captionTargetBox-0="[170,1416,213,1483]" captionTargetBox-1="[163,1413,219,1351]" captionTargetBox-2="[164,1416,205,1460]" captionTargetBox-3="[166,1420,204,1413]" captionTargetId-0="figure-16@6.[151,1435,181,1505]" captionTargetId-1="figure-16@7.[151,1435,181,1357]" captionTargetId-2="figure-16@9.[151,1435,181,1478]" captionTargetId-3="figure-16@11.[151,1435,181,1426]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionTargetPageId-3="11" captionText-0="FIGURE 3. Eloeophila canidorsalis Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-1="FIGURE 4. Eloeophila dietziana (Alexander, 1925). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-2="FIGURE 5. Eloeophila enischnophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-3="FIGURE 6. Eloeophila hadrophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5057241" figureDoi-1="http://doi.org/10.5281/zenodo.5057243" figureDoi-2="http://doi.org/10.5281/zenodo.5057245" figureDoi-3="http://doi.org/10.5281/zenodo.5057247" httpUri-0="https://zenodo.org/record/5057241/files/figure.png" httpUri-1="https://zenodo.org/record/5057243/files/figure.png" httpUri-2="https://zenodo.org/record/5057245/files/figure.png" httpUri-3="https://zenodo.org/record/5057247/files/figure.png" pageId="23" pageNumber="448">Figs 3–6B</figureCitation>
, 
<figureCitation id="13592A0A120AFA04FC505B62FC06F8B9" box="[931,960,1892,1914]" captionStart="FIGURE 8" captionStartId="14.[151,250,1915,1939]" captionTargetBox="[188,1368,549,1869]" captionTargetId="figure-174@14.[164,1423,525,1892]" captionTargetPageId="14" captionText="FIGURE 8. Eloeophila lilliputina (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057251" httpUri="https://zenodo.org/record/5057251/files/figure.png" pageId="23" pageNumber="448">8B</figureCitation>
) …................................... 2
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<paragraph id="8BDD368F120AFA04FF645B87FA5AF854" blockId="23.[151,1436,1892,2031]" box="[151,1436,1921,1943]" pageId="23" pageNumber="448">
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– Wing cell without tiny spots free from veins (
<figureCitation id="13592A0A120AFA04FD655B87FD3BF855" box="[662,765,1921,1943]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1599,1623]" captionStartId-1="4.[151,250,1571,1595]" captionTargetBox-0="[172,1415,201,1547]" captionTargetBox-1="[161,1429,355,1536]" captionTargetId-0="figure-16@2.[151,1435,179,1574]" captionTargetId-1="figure-75@4.[151,1435,345,1548]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1. Eloeophila amamiana (Alexander, 1956). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface; H. female terminalia, lateral view. Scale bars: A–B, 2 mm; C–H, 0.1 mm. Abbreviations: ad, aedeagus; av, apodeme of vesica; cr, cercus; hv, hypogynial valve; ig, inner gonostylus; lp, lateral apodeme; ls, lower shelf; outer gonostylus; pm, paramere; s m-cu, supernumerary m-cu; sX, sternite X; tX, tergite X; vs, vesica." captionText-1="FIGURE 2. Eloeophila apicisetula Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5057237" figureDoi-1="http://doi.org/10.5281/zenodo.5057239" httpUri-0="https://zenodo.org/record/5057237/files/figure.png" httpUri-1="https://zenodo.org/record/5057239/files/figure.png" pageId="23" pageNumber="448">Figs 1–2B</figureCitation>
, 
<figureCitation id="13592A0A120AFA04FCF45B87FCE3F855" box="[775,805,1921,1942]" captionStart="FIGURE 7" captionStartId="13.[151,250,1612,1636]" captionTargetBox="[185,1394,335,1568]" captionTargetId="figure-65@13.[164,1423,309,1588]" captionTargetPageId="13" captionText="FIGURE 7. Eloeophila kintaro (Alexander, 1957). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057249" httpUri="https://zenodo.org/record/5057249/files/figure.png" pageId="23" pageNumber="448">7B</figureCitation>
, 
<figureCitation id="13592A0A120AFA04FCDC5B87FCB6F854" box="[815,880,1921,1943]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStartId-0="16.[151,250,1582,1606]" captionStartId-1="18.[151,250,1672,1696]" captionStartId-2="20.[151,250,1596,1620]" captionStartId-3="21.[151,250,1599,1623]" captionTargetBox-0="[162,1386,376,1543]" captionTargetBox-1="[183,1405,439,1637]" captionTargetBox-2="[179,1427,206,1559]" captionTargetBox-3="[199,1371,395,1540]" captionTargetId-0="figure-106@16.[151,1435,345,1559]" captionTargetId-1="figure-136@18.[164,1423,416,1648]" captionTargetId-2="figure-16@20.[151,1435,181,1572]" captionTargetId-3="figure-47@21.[189,1398,365,1576]" captionTargetPageId-0="16" captionTargetPageId-1="18" captionTargetPageId-2="20" captionTargetPageId-3="21" captionText-0="FIGURE 9. Eloeophila persalsa (Alexander, 1940). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-1="FIGURE 10. Eloeophila subaprilina (Alexander, 1919). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-2="FIGURE 11. Eloeophila tergilobellus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. gonocoxite, lateral view (upper = dorsal, left = anterior); E. outer gonostylus, ventral view; F. inner gonostylus, outer surface; G. aedeagal complex, dorsal view; H. same, lateral view (left = dorsal). Scale bars: A–B, 2 mm; C–H, 0.1 mm." captionText-3="FIGURE 12. Eloeophila ussuriana iwatensis (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. same, outer surface; F. aedeagal complex, dorsal view; G. same, lateral view (left = dorsal); H. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5057305" figureDoi-1="http://doi.org/10.5281/zenodo.5057309" figureDoi-2="http://doi.org/10.5281/zenodo.5057311" figureDoi-3="http://doi.org/10.5281/zenodo.5057315" httpUri-0="https://zenodo.org/record/5057305/files/figure.png" httpUri-1="https://zenodo.org/record/5057309/files/figure.png" httpUri-2="https://zenodo.org/record/5057311/files/figure.png" httpUri-3="https://zenodo.org/record/5057315/files/figure.png" pageId="23" pageNumber="448">9–12B</figureCitation>
)............................................ 6
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<paragraph id="8BDD368F120AFA04FF645B98FA5AF870" blockId="23.[151,1436,1892,2031]" box="[151,1436,1950,1972]" pageId="23" pageNumber="448">
<keyLead id="309395BA120AFA04FF645B98FA5AF870" box="[151,1436,1950,1972]" pageId="23" pageNumber="448">
2. Femora yellow in whole length (
<figureCitation id="13592A0A120AFA04FDE85B98FDA0F877" box="[539,614,1950,1972]" captionStart="FIGURE 3" captionStartId="6.[151,250,1529,1553]" captionTargetBox="[170,1416,213,1483]" captionTargetId="figure-16@6.[151,1435,181,1505]" captionTargetPageId="6" captionText="FIGURE 3. Eloeophila canidorsalis Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057241" httpUri="https://zenodo.org/record/5057241/files/figure.png" pageId="23" pageNumber="448">Fig. 3A</figureCitation>
) ….............................................. 
<taxonomicName id="4C624D0C120AFA04FB4B5B99FA8CF870" authority="Kato, 2021" authorityName="Kato" authorityYear="2021" box="[1208,1354,1950,1972]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="448" phylum="Arthropoda" rank="species" species="canidorsalis" status="sp. nov.">
<emphasis id="B916EA9D120AFA04FB4B5B99FA8CF870" box="[1208,1354,1950,1972]" italics="true" pageId="23" pageNumber="448">E. canidorsalis</emphasis>
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<taxonomicNameLabel id="A22557E6120AFA04FAA35B98FA5AF870" box="[1360,1436,1950,1971]" pageId="23" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BDD368F120AFA04FF645BBAFA5AF812" blockId="23.[151,1436,1892,2031]" box="[151,1436,1980,2002]" pageId="23" pageNumber="448">
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- Femora dark brown at tips (
<figureCitation id="13592A0A120AFA04FE1F5BBAFD92F811" box="[492,596,1980,2002]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="7.[151,250,1382,1406]" captionStartId-1="9.[151,250,1501,1525]" captionStartId-2="11.[151,250,1450,1474]" captionTargetBox-0="[163,1413,219,1351]" captionTargetBox-1="[164,1416,205,1460]" captionTargetBox-2="[166,1420,204,1413]" captionTargetId-0="figure-16@7.[151,1435,181,1357]" captionTargetId-1="figure-16@9.[151,1435,181,1478]" captionTargetId-2="figure-16@11.[151,1435,181,1426]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="11" captionText-0="FIGURE 4. Eloeophila dietziana (Alexander, 1925). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-1="FIGURE 5. Eloeophila enischnophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-2="FIGURE 6. Eloeophila hadrophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5057243" figureDoi-1="http://doi.org/10.5281/zenodo.5057245" figureDoi-2="http://doi.org/10.5281/zenodo.5057247" httpUri-0="https://zenodo.org/record/5057243/files/figure.png" httpUri-1="https://zenodo.org/record/5057245/files/figure.png" httpUri-2="https://zenodo.org/record/5057247/files/figure.png" pageId="23" pageNumber="448">Figs 4–6A</figureCitation>
, 
<figureCitation id="13592A0A120AFA04FDAD5BBAFDBAF811" box="[606,636,1980,2002]" captionStart="FIGURE 8" captionStartId="14.[151,250,1915,1939]" captionTargetBox="[188,1368,549,1869]" captionTargetId="figure-174@14.[164,1423,525,1892]" captionTargetPageId="14" captionText="FIGURE 8. Eloeophila lilliputina (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057251" httpUri="https://zenodo.org/record/5057251/files/figure.png" pageId="23" pageNumber="448">8A</figureCitation>
)..............................................................… 3
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3. Outer gonostylus with inner apical corner angulated at almost right angle, outer and inner margins of narrow distal part smooth (
<figureCitation id="13592A0A1205FA0BFF1A5C91FEF0FF6D" box="[233,310,151,174]" captionStart="FIGURE 4" captionStartId="7.[151,250,1382,1406]" captionTargetBox="[163,1413,219,1351]" captionTargetId="figure-16@7.[151,1435,181,1357]" captionTargetPageId="7" captionText="FIGURE 4. Eloeophila dietziana (Alexander, 1925). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057243" httpUri="https://zenodo.org/record/5057243/files/figure.png" pageId="24" pageNumber="449">Fig. 4D</figureCitation>
); mesal-apical lobe of paramere subacute at tip (
<figureCitation id="13592A0A1205FA0BFCFB5C91FC95FF6D" box="[776,851,151,174]" captionStart="FIGURE 4" captionStartId="7.[151,250,1382,1406]" captionTargetBox="[163,1413,219,1351]" captionTargetId="figure-16@7.[151,1435,181,1357]" captionTargetPageId="7" captionText="FIGURE 4. Eloeophila dietziana (Alexander, 1925). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057243" httpUri="https://zenodo.org/record/5057243/files/figure.png" pageId="24" pageNumber="449">Fig. 4G</figureCitation>
)....................... 
<taxonomicName id="4C624D0C1205FA0BFB865C9EFA5AFF6E" ID-CoL="39CBH" authority="(Alexander, 1925)" baseAuthorityName="Alexander" baseAuthorityYear="1925" box="[1141,1436,151,173]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="dietziana">
<emphasis id="B916EA9D1205FA0BFB865C9EFB21FF6F" box="[1141,1255,151,173]" italics="true" pageId="24" pageNumber="449">E. dietziana</emphasis>
(
<bibRefCitation id="EFF34B7E1205FA0BFB075C91FA53FF6E" author="Alexander, C. P." box="[1268,1429,151,173]" pageId="24" pageNumber="449" pagination="431 - 448" refId="ref17904" refString="Alexander, C. P. (1925) Undescribed species of Japanese crane-flies. Part V. Annals of the Entomological Society of America, 17, 431 - 448. https: // doi. org / 10.1093 / aesa / 17.4.431" type="journal article" year="1925">Alexander, 1925</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDD368F1205FA0BFF645CB3FA5AFF24" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645CB3FA5AFF24" pageId="24" pageNumber="449">
- Outer gonostylus with inner apical corner angulated at obtuse angle; outer and/or inner margins of narrow distal part serrate or weakly undulate (
<figureCitation id="13592A0A1205FA0BFE7E5CD4FE32FF2B" box="[397,500,210,232]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="9.[151,250,1501,1525]" captionStartId-1="11.[151,250,1450,1474]" captionTargetBox-0="[164,1416,205,1460]" captionTargetBox-1="[166,1420,204,1413]" captionTargetId-0="figure-16@9.[151,1435,181,1478]" captionTargetId-1="figure-16@11.[151,1435,181,1426]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="FIGURE 5. Eloeophila enischnophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-1="FIGURE 6. Eloeophila hadrophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5057245" figureDoi-1="http://doi.org/10.5281/zenodo.5057247" httpUri-0="https://zenodo.org/record/5057245/files/figure.png" httpUri-1="https://zenodo.org/record/5057247/files/figure.png" pageId="24" pageNumber="449">Figs 5–6D</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFE0D5CD4FDD8FF2B" box="[510,542,210,232]" captionStart="FIGURE 8" captionStartId="14.[151,250,1915,1939]" captionTargetBox="[188,1368,549,1869]" captionTargetId="figure-174@14.[164,1423,525,1892]" captionTargetPageId="14" captionText="FIGURE 8. Eloeophila lilliputina (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057251" httpUri="https://zenodo.org/record/5057251/files/figure.png" pageId="24" pageNumber="449">8D</figureCitation>
); mesal-apical lobe of paramere rounded at tip (
<figureCitation id="13592A0A1205FA0BFC195CD4FB95FF2B" box="[1002,1107,210,232]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="9.[151,250,1501,1525]" captionStartId-1="11.[151,250,1450,1474]" captionTargetBox-0="[164,1416,205,1460]" captionTargetBox-1="[166,1420,204,1413]" captionTargetId-0="figure-16@9.[151,1435,181,1478]" captionTargetId-1="figure-16@11.[151,1435,181,1426]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="FIGURE 5. Eloeophila enischnophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-1="FIGURE 6. Eloeophila hadrophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5057245" figureDoi-1="http://doi.org/10.5281/zenodo.5057247" httpUri-0="https://zenodo.org/record/5057245/files/figure.png" httpUri-1="https://zenodo.org/record/5057247/files/figure.png" pageId="24" pageNumber="449">Figs 5–6G</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFBAF5CD4FBBCFF2B" box="[1116,1146,210,232]" captionStart="FIGURE 8" captionStartId="14.[151,250,1915,1939]" captionTargetBox="[188,1368,549,1869]" captionTargetId="figure-174@14.[164,1423,525,1892]" captionTargetPageId="14" captionText="FIGURE 8. Eloeophila lilliputina (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057251" httpUri="https://zenodo.org/record/5057251/files/figure.png" pageId="24" pageNumber="449">8G</figureCitation>
)..................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E2A1205FA0BFF645CF6FA5AFE9E" pageId="24" pageNumber="449">
<paragraph id="8BDD368F1205FA0BFF645CF6FA5AFEE1" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645CF6FA5AFEE1" pageId="24" pageNumber="449">
4. Outer gonostylus with outer margin of narrow distal part serrate (
<figureCitation id="13592A0A1205FA0BFCA95CE9FC6EFEC6" box="[858,936,239,261]" captionStart="FIGURE 6" captionStartId="11.[151,250,1450,1474]" captionTargetBox="[166,1420,204,1413]" captionTargetId="figure-16@11.[151,1435,181,1426]" captionTargetPageId="11" captionText="FIGURE 6. Eloeophila hadrophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057247" httpUri="https://zenodo.org/record/5057247/files/figure.png" pageId="24" pageNumber="449">Fig. 6D</figureCitation>
); aedeagus stout, about 2.5 times as long as wide in lateral view (
<figureCitation id="13592A0A1205FA0BFE935D0BFE6EFEE0" box="[352,424,269,291]" captionStart="FIGURE 6" captionStartId="11.[151,250,1450,1474]" captionTargetBox="[166,1420,204,1413]" captionTargetId="figure-16@11.[151,1435,181,1426]" captionTargetPageId="11" captionText="FIGURE 6. Eloeophila hadrophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057247" httpUri="https://zenodo.org/record/5057247/files/figure.png" pageId="24" pageNumber="449">Fig. 6E</figureCitation>
)............................................................... 
<taxonomicName id="4C624D0C1205FA0BFB435D0BFA8FFEE1" authority="Kato, 2021" authorityName="Kato" authorityYear="2021" box="[1200,1353,269,290]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="hadrophallus" status="sp. nov.">
<emphasis id="B916EA9D1205FA0BFB435D0BFA8FFEE1" box="[1200,1353,269,290]" italics="true" pageId="24" pageNumber="449">E. hadrophallus</emphasis>
</taxonomicName>
<emphasis id="B916EA9D1205FA0BFAA35D0BFA5AFEE1" bold="true" box="[1360,1436,269,290]" pageId="24" pageNumber="449">
<taxonomicNameLabel id="A22557E61205FA0BFAA35D0BFA5AFEE1" box="[1360,1436,269,290]" pageId="24" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BDD368F1205FA0BFF645D2DFA5AFE9E" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645D2DFA5AFE9E" pageId="24" pageNumber="449">
- Outer gonostylus with outer margin of narrow distal part smooth (
<figureCitation id="13592A0A1205FA0BFCA85D2CFC6CFE83" box="[859,938,298,320]" captionStart="FIGURE 5" captionStartId="9.[151,250,1501,1525]" captionTargetBox="[164,1416,205,1460]" captionTargetId="figure-16@9.[151,1435,181,1478]" captionTargetPageId="9" captionText="FIGURE 5. Eloeophila enischnophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057245" httpUri="https://zenodo.org/record/5057245/files/figure.png" pageId="24" pageNumber="449">Figs 5D</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFC465D2CFC13FE83" box="[949,981,298,320]" captionStart="FIGURE 8" captionStartId="14.[151,250,1915,1939]" captionTargetBox="[188,1368,549,1869]" captionTargetId="figure-174@14.[164,1423,525,1892]" captionTargetPageId="14" captionText="FIGURE 8. Eloeophila lilliputina (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057251" httpUri="https://zenodo.org/record/5057251/files/figure.png" pageId="24" pageNumber="449">8D</figureCitation>
); aedeagus slender, five times longer than wide in dorsal view (
<figureCitation id="13592A0A1205FA0BFE845D41FE79FE9E" box="[375,447,327,349]" captionStart="FIGURE 5" captionStartId="9.[151,250,1501,1525]" captionTargetBox="[164,1416,205,1460]" captionTargetId="figure-16@9.[151,1435,181,1478]" captionTargetPageId="9" captionText="FIGURE 5. Eloeophila enischnophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057245" httpUri="https://zenodo.org/record/5057245/files/figure.png" pageId="24" pageNumber="449">Fig. 5E</figureCitation>
)................................................................................ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E2A1205FA0BFF645D63FA5AFE2C" pageId="24" pageNumber="449">
<paragraph id="8BDD368F1205FA0BFF645D63FA5DFE5B" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645D63FA5DFE5B" pageId="24" pageNumber="449">
5. Outer gonostylus with narrow distal part slightly curved to tip, serrate at inner margin (
<figureCitation id="13592A0A1205FA0BFBED5D63FBACFEB8" box="[1054,1130,357,379]" captionStart="FIGURE 8" captionStartId="14.[151,250,1915,1939]" captionTargetBox="[188,1368,549,1869]" captionTargetId="figure-174@14.[164,1423,525,1892]" captionTargetPageId="14" captionText="FIGURE 8. Eloeophila lilliputina (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057251" httpUri="https://zenodo.org/record/5057251/files/figure.png" pageId="24" pageNumber="449">Fig. 8D</figureCitation>
); mesal-apical lobe of paramere about three times as long as wide (
<figureCitation id="13592A0A1205FA0BFDDD5D84FDBCFE5B" box="[558,634,386,408]" captionStart="FIGURE 8" captionStartId="14.[151,250,1915,1939]" captionTargetBox="[188,1368,549,1869]" captionTargetId="figure-174@14.[164,1423,525,1892]" captionTargetPageId="14" captionText="FIGURE 8. Eloeophila lilliputina (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057251" httpUri="https://zenodo.org/record/5057251/files/figure.png" pageId="24" pageNumber="449">Fig. 8G</figureCitation>
); apodeme of vesica directed ventrally (
<figureCitation id="13592A0A1205FA0BFC085D84FB84FE5B" box="[1019,1090,386,408]" captionStart="FIGURE 8" captionStartId="14.[151,250,1915,1939]" captionTargetBox="[188,1368,549,1869]" captionTargetId="figure-174@14.[164,1423,525,1892]" captionTargetPageId="14" captionText="FIGURE 8. Eloeophila lilliputina (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057251" httpUri="https://zenodo.org/record/5057251/files/figure.png" pageId="24" pageNumber="449">Fig. 8F</figureCitation>
) … 
<taxonomicName id="4C624D0C1205FA0BFB9E5D85FA5DFE5B" ID-CoL="39CC3" authority="(Alexander, 1936)" baseAuthorityName="Alexander" baseAuthorityYear="1936" box="[1133,1435,386,408]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="lilliputina">
<emphasis id="B916EA9D1205FA0BFB9E5D85FB21FE54" box="[1133,1255,386,408]" italics="true" pageId="24" pageNumber="449">E. lilliputina</emphasis>
(Alexander, 1936)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDD368F1205FA0BFF645DA6FA5AFE2C" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645DA6FA5AFE2C" pageId="24" pageNumber="449">
- Outer gonostylus with narrow distal part distinctly curved to tip, weakly undulate at inner margin (
<figureCitation id="13592A0A1205FA0BFB665D99FB24FE76" box="[1173,1250,415,437]" captionStart="FIGURE 5" captionStartId="9.[151,250,1501,1525]" captionTargetBox="[164,1416,205,1460]" captionTargetId="figure-16@9.[151,1435,181,1478]" captionTargetPageId="9" captionText="FIGURE 5. Eloeophila enischnophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057245" httpUri="https://zenodo.org/record/5057245/files/figure.png" pageId="24" pageNumber="449">Fig. 5D</figureCitation>
); mesal-apical lobe of paramere about 1.5 times as long as wide (
<figureCitation id="13592A0A1205FA0BFD665DBBFD24FE10" box="[661,738,445,467]" captionStart="FIGURE 5" captionStartId="9.[151,250,1501,1525]" captionTargetBox="[164,1416,205,1460]" captionTargetId="figure-16@9.[151,1435,181,1478]" captionTargetPageId="9" captionText="FIGURE 5. Eloeophila enischnophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057245" httpUri="https://zenodo.org/record/5057245/files/figure.png" pageId="24" pageNumber="449">Fig. 5G</figureCitation>
); apodeme of vesica directed anteroventrally (
<figureCitation id="13592A0A1205FA0BFB6D5DBBFB23FE10" box="[1182,1253,445,467]" captionStart="FIGURE 5" captionStartId="9.[151,250,1501,1525]" captionTargetBox="[164,1416,205,1460]" captionTargetId="figure-16@9.[151,1435,181,1478]" captionTargetPageId="9" captionText="FIGURE 5. Eloeophila enischnophallus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057245" httpUri="https://zenodo.org/record/5057245/files/figure.png" pageId="24" pageNumber="449">Fig. 5F</figureCitation>
)........................................................................................... 
<taxonomicName id="4C624D0C1205FA0BFB605DDDFA8CFE2C" authority="Kato, 2021" authorityName="Kato" authorityYear="2021" box="[1171,1354,474,496]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="enischnophallus" status="sp. nov.">
<emphasis id="B916EA9D1205FA0BFB605DDDFA8CFE2C" box="[1171,1354,474,496]" italics="true" pageId="24" pageNumber="449">E. enischnophallus</emphasis>
</taxonomicName>
<emphasis id="B916EA9D1205FA0BFAA35DDCFA5AFE2C" bold="true" box="[1360,1436,474,495]" pageId="24" pageNumber="449">
<taxonomicNameLabel id="A22557E61205FA0BFAA35DDCFA5AFE2C" box="[1360,1436,474,495]" pageId="24" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E2A1205FA0BFF645DF1FA5AFDA6" pageId="24" pageNumber="449">
<paragraph id="8BDD368F1205FA0BFF645DF1FA5AFDE9" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645DF1FA5AFDE9" pageId="24" pageNumber="449">
6. Wing distinctly patterned, anterior margin with at least six dark spots, tips of veins at posterior margin usually with dark spots (
<figureCitation id="13592A0A1205FA0BFF1A5E13FEF1FDE9" box="[233,311,533,555]" captionStart="FIGURE 2" captionStartId="4.[151,250,1571,1595]" captionTargetBox="[161,1429,355,1536]" captionTargetId="figure-75@4.[151,1435,345,1548]" captionTargetPageId="4" captionText="FIGURE 2. Eloeophila apicisetula Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057239" httpUri="https://zenodo.org/record/5057239/files/figure.png" pageId="24" pageNumber="449">Figs 2B</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFEB25E13FE99FDE9" box="[321,351,533,554]" captionStart="FIGURE 7" captionStartId="13.[151,250,1612,1636]" captionTargetBox="[185,1394,335,1568]" captionTargetId="figure-65@13.[164,1423,309,1588]" captionTargetPageId="13" captionText="FIGURE 7. Eloeophila kintaro (Alexander, 1957). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057249" httpUri="https://zenodo.org/record/5057249/files/figure.png" pageId="24" pageNumber="449">7B</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFE9A5E13FE6CFDE8" box="[361,426,533,555]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="16.[151,250,1582,1606]" captionStartId-1="18.[151,250,1672,1696]" captionTargetBox-0="[162,1386,376,1543]" captionTargetBox-1="[183,1405,439,1637]" captionTargetId-0="figure-106@16.[151,1435,345,1559]" captionTargetId-1="figure-136@18.[164,1423,416,1648]" captionTargetPageId-0="16" captionTargetPageId-1="18" captionText-0="FIGURE 9. Eloeophila persalsa (Alexander, 1940). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." captionText-1="FIGURE 10. Eloeophila subaprilina (Alexander, 1919). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5057305" figureDoi-1="http://doi.org/10.5281/zenodo.5057309" httpUri-0="https://zenodo.org/record/5057305/files/figure.png" httpUri-1="https://zenodo.org/record/5057309/files/figure.png" pageId="24" pageNumber="449">9–10B</figureCitation>
).................................................................................. 7
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<paragraph id="8BDD368F1205FA0BFF645E35FA5AFDA6" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
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- Wing less patterned, anterior margin with at most four spots, tips of veins at posterior margin without dark spots (
<figureCitation id="13592A0A1205FA0BFABB5E34FA5EFD84" box="[1352,1432,562,584]" captionStart="FIGURE 1" captionStartId="2.[151,250,1599,1623]" captionTargetBox="[172,1415,201,1547]" captionTargetId="figure-16@2.[151,1435,179,1574]" captionTargetPageId="2" captionText="FIGURE 1. Eloeophila amamiana (Alexander, 1956). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface; H. female terminalia, lateral view. Scale bars: A–B, 2 mm; C–H, 0.1 mm. Abbreviations: ad, aedeagus; av, apodeme of vesica; cr, cercus; hv, hypogynial valve; ig, inner gonostylus; lp, lateral apodeme; ls, lower shelf; outer gonostylus; pm, paramere; s m-cu, supernumerary m-cu; sX, sternite X; tX, tergite X; vs, vesica." figureDoi="http://doi.org/10.5281/zenodo.5057237" httpUri="https://zenodo.org/record/5057237/files/figure.png" pageId="24" pageNumber="449">Figs 1B</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFF115E56FEE8FDA5" box="[226,302,592,614]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="20.[151,250,1596,1620]" captionStartId-1="21.[151,250,1599,1623]" captionTargetBox-0="[179,1427,206,1559]" captionTargetBox-1="[199,1371,395,1540]" captionTargetId-0="figure-16@20.[151,1435,181,1572]" captionTargetId-1="figure-47@21.[189,1398,365,1576]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="FIGURE 11. Eloeophila tergilobellus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. gonocoxite, lateral view (upper = dorsal, left = anterior); E. outer gonostylus, ventral view; F. inner gonostylus, outer surface; G. aedeagal complex, dorsal view; H. same, lateral view (left = dorsal). Scale bars: A–B, 2 mm; C–H, 0.1 mm." captionText-1="FIGURE 12. Eloeophila ussuriana iwatensis (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. same, outer surface; F. aedeagal complex, dorsal view; G. same, lateral view (left = dorsal); H. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5057311" figureDoi-1="http://doi.org/10.5281/zenodo.5057315" httpUri-0="https://zenodo.org/record/5057311/files/figure.png" httpUri-1="https://zenodo.org/record/5057315/files/figure.png" pageId="24" pageNumber="449">11–12B</figureCitation>
)........................................................................................... 10
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7. Thorax with notum and pleuron mottled with dark markings, bearing several paired spots or stripes; wing with two dark spots at tip of A 
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, basal one large (
<figureCitation id="13592A0A1205FA0BFE015E8CFD8CFD63" box="[498,586,650,672]" captionStart="FIGURE 10" captionStartId="18.[151,250,1672,1696]" captionTargetBox="[183,1405,439,1637]" captionTargetId="figure-136@18.[164,1423,416,1648]" captionTargetPageId="18" captionText="FIGURE 10. Eloeophila subaprilina (Alexander, 1919). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057309" httpUri="https://zenodo.org/record/5057309/files/figure.png" pageId="24" pageNumber="449">Fig. 10B</figureCitation>
); halter with knob dark brown (
<figureCitation id="13592A0A1205FA0BFC895E8CFC14FD63" box="[890,978,650,672]" captionStart="FIGURE 10" captionStartId="18.[151,250,1672,1696]" captionTargetBox="[183,1405,439,1637]" captionTargetId="figure-136@18.[164,1423,416,1648]" captionTargetPageId="18" captionText="FIGURE 10. Eloeophila subaprilina (Alexander, 1919). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057309" httpUri="https://zenodo.org/record/5057309/files/figure.png" pageId="24" pageNumber="449">Fig. 10A</figureCitation>
).......... 
<taxonomicName id="4C624D0C1205FA0BFBAD5E8DFA5AFD63" ID-CoL="39CDC" authority="(Alexander, 1919)" baseAuthorityName="Alexander" baseAuthorityYear="1919" box="[1118,1436,650,672]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="subaprilina">
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(
<bibRefCitation id="EFF34B7E1205FA0BFB075E8CFA53FD63" author="Alexander, C. P." box="[1268,1429,650,672]" pageId="24" pageNumber="449" pagination="327 - 348" refId="ref17810" refString="Alexander, C. P. (1919) Undescribed species of Japanese crane-flies (Tipulidae, Diptera). Annals of the Entomological Society of America, 12, 327 - 348. https: // doi. org / 10.1093 / aesa / 12.4.327" type="journal article" year="1919">Alexander, 1919</bibRefCitation>
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- Thorax with notum and pleuron almost unpatterned with dark markings except prescutal pit; wing with one or two dark spot at tips of A 
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, basal one indistinct if two spots present (
<figureCitation id="13592A0A1205FA0BFD3C5EC3FCD8FD19" box="[719,798,709,731]" captionStart="FIGURE 2" captionStartId="4.[151,250,1571,1595]" captionTargetBox="[161,1429,355,1536]" captionTargetId="figure-75@4.[151,1435,345,1548]" captionTargetPageId="4" captionText="FIGURE 2. Eloeophila apicisetula Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057239" httpUri="https://zenodo.org/record/5057239/files/figure.png" pageId="24" pageNumber="449">Figs 2B</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFCDB5EC3FC83FD19" box="[808,837,709,730]" captionStart="FIGURE 7" captionStartId="13.[151,250,1612,1636]" captionTargetBox="[185,1394,335,1568]" captionTargetId="figure-65@13.[164,1423,309,1588]" captionTargetPageId="13" captionText="FIGURE 7. Eloeophila kintaro (Alexander, 1957). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057249" httpUri="https://zenodo.org/record/5057249/files/figure.png" pageId="24" pageNumber="449">7B</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFCA35EC3FCA9FD18" box="[848,879,709,731]" captionStart="FIGURE 9" captionStartId="16.[151,250,1582,1606]" captionTargetBox="[162,1386,376,1543]" captionTargetId="figure-106@16.[151,1435,345,1559]" captionTargetPageId="16" captionText="FIGURE 9. Eloeophila persalsa (Alexander, 1940). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057305" httpUri="https://zenodo.org/record/5057305/files/figure.png" pageId="24" pageNumber="449">9B</figureCitation>
); halter entirely pale yellow (
<figureCitation id="13592A0A1205FA0BFB7B5EC3FB1EFD19" box="[1160,1240,709,731]" captionStart="FIGURE 2" captionStartId="4.[151,250,1571,1595]" captionTargetBox="[161,1429,355,1536]" captionTargetId="figure-75@4.[151,1435,345,1548]" captionTargetPageId="4" captionText="FIGURE 2. Eloeophila apicisetula Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057239" httpUri="https://zenodo.org/record/5057239/files/figure.png" pageId="24" pageNumber="449">Figs 2A</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFB115EC3FAC7FD19" box="[1250,1281,709,730]" captionStart="FIGURE 7" captionStartId="13.[151,250,1612,1636]" captionTargetBox="[185,1394,335,1568]" captionTargetId="figure-65@13.[164,1423,309,1588]" captionTargetPageId="13" captionText="FIGURE 7. Eloeophila kintaro (Alexander, 1957). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057249" httpUri="https://zenodo.org/record/5057249/files/figure.png" pageId="24" pageNumber="449">7A</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFAF85EC3FAECFD18" box="[1291,1322,709,731]" captionStart="FIGURE 9" captionStartId="16.[151,250,1582,1606]" captionTargetBox="[162,1386,376,1543]" captionTargetId="figure-106@16.[151,1435,345,1559]" captionTargetPageId="16" captionText="FIGURE 9. Eloeophila persalsa (Alexander, 1940). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057305" httpUri="https://zenodo.org/record/5057305/files/figure.png" pageId="24" pageNumber="449">9A</figureCitation>
)....... 8
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8. Wing membrane without macrotrichia in cells at wing tip (
<figureCitation id="13592A0A1205FA0BFCE65EE4FCA7FD3B" box="[789,865,738,760]" captionStart="FIGURE 9" captionStartId="16.[151,250,1582,1606]" captionTargetBox="[162,1386,376,1543]" captionTargetId="figure-106@16.[151,1435,345,1559]" captionTargetPageId="16" captionText="FIGURE 9. Eloeophila persalsa (Alexander, 1940). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057305" httpUri="https://zenodo.org/record/5057305/files/figure.png" pageId="24" pageNumber="449">Fig. 9B</figureCitation>
); each femur tipped with dark brown (
<figureCitation id="13592A0A1205FA0BFB225EE4FADBFD3B" box="[1233,1309,738,760]" captionStart="FIGURE 9" captionStartId="16.[151,250,1582,1606]" captionTargetBox="[162,1386,376,1543]" captionTargetId="figure-106@16.[151,1435,345,1559]" captionTargetPageId="16" captionText="FIGURE 9. Eloeophila persalsa (Alexander, 1940). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057305" httpUri="https://zenodo.org/record/5057305/files/figure.png" pageId="24" pageNumber="449">Fig. 9A</figureCitation>
)..................................................................................... 
<taxonomicName id="4C624D0C1205FA0BFB885F06FA5AFCD5" ID-CoL="39CCT" authority="(Alexander, 1940)" baseAuthorityName="Alexander" baseAuthorityYear="1940" box="[1147,1436,767,790]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="persalsa">
<emphasis id="B916EA9D1205FA0BFB885F06FB21FCD7" box="[1147,1255,767,789]" italics="true" pageId="24" pageNumber="449">E. persalsa</emphasis>
(
<bibRefCitation id="EFF34B7E1205FA0BFB075EF9FA53FCD5" author="Alexander, C. P." box="[1268,1429,767,790]" pageId="24" pageNumber="449" pagination="39 - 76" refId="ref18221" refString="Alexander, C. P. (1940) New or little-known Tipulidae from eastern Asia (Diptera). XLI. Philippine Journal of Science, 71, 39 - 76." type="journal article" year="1940">Alexander, 1940</bibRefCitation>
)
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</paragraph>
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- Wing membrane with macrotrichia in cells near wing tip (
<figureCitation id="13592A0A1205FA0BFCFE5F1BFC9AFCF1" box="[781,860,797,819]" captionStart="FIGURE 2" captionStartId="4.[151,250,1571,1595]" captionTargetBox="[161,1429,355,1536]" captionTargetId="figure-75@4.[151,1435,345,1548]" captionTargetPageId="4" captionText="FIGURE 2. Eloeophila apicisetula Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057239" httpUri="https://zenodo.org/record/5057239/files/figure.png" pageId="24" pageNumber="449">Figs 2B</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFC955F1BFC43FCF1" box="[870,901,797,818]" captionStart="FIGURE 7" captionStartId="13.[151,250,1612,1636]" captionTargetBox="[185,1394,335,1568]" captionTargetId="figure-65@13.[164,1423,309,1588]" captionTargetPageId="13" captionText="FIGURE 7. Eloeophila kintaro (Alexander, 1957). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057249" httpUri="https://zenodo.org/record/5057249/files/figure.png" pageId="24" pageNumber="449">7B</figureCitation>
); each femur without dark tip (
<figureCitation id="13592A0A1205FA0BFB5D5F1BFB38FCF1" box="[1198,1278,797,819]" captionStart="FIGURE 2" captionStartId="4.[151,250,1571,1595]" captionTargetBox="[161,1429,355,1536]" captionTargetId="figure-75@4.[151,1435,345,1548]" captionTargetPageId="4" captionText="FIGURE 2. Eloeophila apicisetula Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057239" httpUri="https://zenodo.org/record/5057239/files/figure.png" pageId="24" pageNumber="449">Figs 2A</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFAFB5F1BFAE1FCF0" box="[1288,1319,797,819]" captionStart="FIGURE 7" captionStartId="13.[151,250,1612,1636]" captionTargetBox="[185,1394,335,1568]" captionTargetId="figure-65@13.[164,1423,309,1588]" captionTargetPageId="13" captionText="FIGURE 7. Eloeophila kintaro (Alexander, 1957). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057249" httpUri="https://zenodo.org/record/5057249/files/figure.png" pageId="24" pageNumber="449">7A</figureCitation>
)....... 9
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<paragraph id="8BDD368F1205FA0BFF645F3CFA5AFCAE" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
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9. Pleuron weakly dark on dorsal part; outer margin of outer gonostylus without lobe near middle (
<figureCitation id="13592A0A1205FA0BFB8E5F3CFB0EFC93" box="[1149,1224,826,848]" captionStart="FIGURE 2" captionStartId="4.[151,250,1571,1595]" captionTargetBox="[161,1429,355,1536]" captionTargetId="figure-75@4.[151,1435,345,1548]" captionTargetPageId="4" captionText="FIGURE 2. Eloeophila apicisetula Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057239" httpUri="https://zenodo.org/record/5057239/files/figure.png" pageId="24" pageNumber="449">Fig. 2D</figureCitation>
).................................................................................................. 
<taxonomicName id="4C624D0C1205FA0BFB355F5EFA8FFCAF" authority="Kato, 2021" authorityName="Kato" authorityYear="2021" box="[1222,1353,855,877]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="apicisetula" status="sp. nov.">
<emphasis id="B916EA9D1205FA0BFB355F5EFA8FFCAF" box="[1222,1353,855,877]" italics="true" pageId="24" pageNumber="449">E. apicisetula</emphasis>
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<taxonomicNameLabel id="A22557E61205FA0BFAA35F5EFA5AFCAE" box="[1360,1436,856,877]" pageId="24" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
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<keyLead id="309395BA1205FA0BFF645F73FA5AFC6B" pageId="24" pageNumber="449">
- Pleuron not dark on dorsal part; outer margin of outer gonostylus with large lobe near middle of outer margin (
<figureCitation id="13592A0A1205FA0BFAFB5F73FA95FC49" box="[1288,1363,885,907]" captionStart="FIGURE 7" captionStartId="13.[151,250,1612,1636]" captionTargetBox="[185,1394,335,1568]" captionTargetId="figure-65@13.[164,1423,309,1588]" captionTargetPageId="13" captionText="FIGURE 7. Eloeophila kintaro (Alexander, 1957). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057249" httpUri="https://zenodo.org/record/5057249/files/figure.png" pageId="24" pageNumber="449">Fig. 7D</figureCitation>
).................................................................................. 
<taxonomicName id="4C624D0C1205FA0BFB7B5F95FA5AFC6B" ID-CoL="39CBY" authority="(Alexander, 1957)" baseAuthorityName="Alexander" baseAuthorityYear="1957" box="[1160,1436,914,936]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="kintaro">
<emphasis id="B916EA9D1205FA0BFB7B5F95FB21FC64" box="[1160,1255,914,936]" italics="true" pageId="24" pageNumber="449">E. kintaro</emphasis>
(
<bibRefCitation id="EFF34B7E1205FA0BFB075F94FA53FC6B" author="Alexander, C. P." box="[1268,1429,914,936]" pageId="24" pageNumber="449" pagination="349 - 388" refId="ref18400" refString="Alexander, C. P. (1957) Records and descriptions of Japanese Tipulidae (Diptera). Part V. The crane-flies of Honshu. I. Philippine Journal of Science, 85, 349 - 388." type="journal article" year="1957">Alexander, 1957</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E2A1205FA0BFF645FB6FA5AFBDE" pageId="24" pageNumber="449">
<paragraph id="8BDD368F1205FA0BFF645FB6FA5AFC21" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645FB6FA5AFC21" pageId="24" pageNumber="449">
10. Wing unpatterned except dark stigma (
<figureCitation id="13592A0A1205FA0BFDA95FA9FD74FC05" box="[602,690,943,966]" captionStart="FIGURE 11" captionStartId="20.[151,250,1596,1620]" captionTargetBox="[179,1427,206,1559]" captionTargetId="figure-16@20.[151,1435,181,1572]" captionTargetPageId="20" captionText="FIGURE 11. Eloeophila tergilobellus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. gonocoxite, lateral view (upper = dorsal, left = anterior); E. outer gonostylus, ventral view; F. inner gonostylus, outer surface; G. aedeagal complex, dorsal view; H. same, lateral view (left = dorsal). Scale bars: A–B, 2 mm; C–H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057311" httpUri="https://zenodo.org/record/5057311/files/figure.png" pageId="24" pageNumber="449">Fig. 11B</figureCitation>
); halter about as long as or longer than thorax (
<figureCitation id="13592A0A1205FA0BFB725FA9FB15FC06" box="[1153,1235,943,965]" captionStart="FIGURE 11" captionStartId="20.[151,250,1596,1620]" captionTargetBox="[179,1427,206,1559]" captionTargetId="figure-16@20.[151,1435,181,1572]" captionTargetPageId="20" captionText="FIGURE 11. Eloeophila tergilobellus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. gonocoxite, lateral view (upper = dorsal, left = anterior); E. outer gonostylus, ventral view; F. inner gonostylus, outer surface; G. aedeagal complex, dorsal view; H. same, lateral view (left = dorsal). Scale bars: A–B, 2 mm; C–H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057311" httpUri="https://zenodo.org/record/5057311/files/figure.png" pageId="24" pageNumber="449">Fig.11A</figureCitation>
); male tergite 8 with projection(s) at caudal margin (
<figureCitation id="13592A0A1205FA0BFDFC5FCBFDA5FC20" box="[527,611,973,995]" captionStart="FIGURE 11" captionStartId="20.[151,250,1596,1620]" captionTargetBox="[179,1427,206,1559]" captionTargetId="figure-16@20.[151,1435,181,1572]" captionTargetPageId="20" captionText="FIGURE 11. Eloeophila tergilobellus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. gonocoxite, lateral view (upper = dorsal, left = anterior); E. outer gonostylus, ventral view; F. inner gonostylus, outer surface; G. aedeagal complex, dorsal view; H. same, lateral view (left = dorsal). Scale bars: A–B, 2 mm; C–H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057311" httpUri="https://zenodo.org/record/5057311/files/figure.png" pageId="24" pageNumber="449">Fig. 11C</figureCitation>
)................................................................. 11
</keyLead>
</paragraph>
<paragraph id="8BDD368F1205FA0BFF645FEDFA5AFBDE" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645FEDFA5AFBDE" pageId="24" pageNumber="449">
- Wing distinctly or indistinctly patterned except stigma (
<figureCitation id="13592A0A1205FA0BFD035FECFCFBFC3C" box="[752,829,1002,1024]" captionStart="FIGURE 1" captionStartId="2.[151,250,1599,1623]" captionTargetBox="[172,1415,201,1547]" captionTargetId="figure-16@2.[151,1435,179,1574]" captionTargetPageId="2" captionText="FIGURE 1. Eloeophila amamiana (Alexander, 1956). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface; H. female terminalia, lateral view. Scale bars: A–B, 2 mm; C–H, 0.1 mm. Abbreviations: ad, aedeagus; av, apodeme of vesica; cr, cercus; hv, hypogynial valve; ig, inner gonostylus; lp, lateral apodeme; ls, lower shelf; outer gonostylus; pm, paramere; s m-cu, supernumerary m-cu; sX, sternite X; tX, tergite X; vs, vesica." figureDoi="http://doi.org/10.5281/zenodo.5057237" httpUri="https://zenodo.org/record/5057237/files/figure.png" pageId="24" pageNumber="449">Figs 1B</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFCB55FECFCB7FBC3" box="[838,881,1002,1024]" captionStart="FIGURE 12" captionStartId="21.[151,250,1599,1623]" captionTargetBox="[199,1371,395,1540]" captionTargetId="figure-47@21.[189,1398,365,1576]" captionTargetPageId="21" captionText="FIGURE 12. Eloeophila ussuriana iwatensis (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. same, outer surface; F. aedeagal complex, dorsal view; G. same, lateral view (left = dorsal); H. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057315" httpUri="https://zenodo.org/record/5057315/files/figure.png" pageId="24" pageNumber="449">12B</figureCitation>
); halter at most 0.7 times as long as thorax (
<figureCitation id="13592A0A1205FA0BFAFE5FECFA9DFC3C" box="[1293,1371,1002,1024]" captionStart="FIGURE 1" captionStartId="2.[151,250,1599,1623]" captionTargetBox="[172,1415,201,1547]" captionTargetId="figure-16@2.[151,1435,179,1574]" captionTargetPageId="2" captionText="FIGURE 1. Eloeophila amamiana (Alexander, 1956). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface; H. female terminalia, lateral view. Scale bars: A–B, 2 mm; C–H, 0.1 mm. Abbreviations: ad, aedeagus; av, apodeme of vesica; cr, cercus; hv, hypogynial valve; ig, inner gonostylus; lp, lateral apodeme; ls, lower shelf; outer gonostylus; pm, paramere; s m-cu, supernumerary m-cu; sX, sternite X; tX, tergite X; vs, vesica." figureDoi="http://doi.org/10.5281/zenodo.5057237" httpUri="https://zenodo.org/record/5057237/files/figure.png" pageId="24" pageNumber="449">Figs 1A</figureCitation>
, 
<figureCitation id="13592A0A1205FA0BFA975FECFA57FBC3" box="[1380,1425,1002,1024]" captionStart="FIGURE 12" captionStartId="21.[151,250,1599,1623]" captionTargetBox="[199,1371,395,1540]" captionTargetId="figure-47@21.[189,1398,365,1576]" captionTargetPageId="21" captionText="FIGURE 12. Eloeophila ussuriana iwatensis (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. same, outer surface; F. aedeagal complex, dorsal view; G. same, lateral view (left = dorsal); H. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057315" httpUri="https://zenodo.org/record/5057315/files/figure.png" pageId="24" pageNumber="449">12A</figureCitation>
); male tergite 8 without projection at caudal margin.......................................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E2A1205FA0BFF645823FA5AFB50" pageId="24" pageNumber="449">
<paragraph id="8BDD368F1205FA0BFF645823FA5AFB94" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645823FA5AFB94" pageId="24" pageNumber="449">
11. Male tergites 7 and 8 each with paired lobes at posterior margin (
<figureCitation id="13592A0A1205FA0BFCA65823FC6AFBF8" box="[853,940,1061,1083]" captionStart="FIGURE 11" captionStartId="20.[151,250,1596,1620]" captionTargetBox="[179,1427,206,1559]" captionTargetId="figure-16@20.[151,1435,181,1572]" captionTargetPageId="20" captionText="FIGURE 11. Eloeophila tergilobellus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. gonocoxite, lateral view (upper = dorsal, left = anterior); E. outer gonostylus, ventral view; F. inner gonostylus, outer surface; G. aedeagal complex, dorsal view; H. same, lateral view (left = dorsal). Scale bars: A–B, 2 mm; C–H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057311" httpUri="https://zenodo.org/record/5057311/files/figure.png" pageId="24" pageNumber="449">Fig. 11C</figureCitation>
); gonocoxite with one large swelling on dorsal side (
<figureCitation id="13592A0A1205FA0BFF1A5844FE99FB9B" box="[233,351,1090,1112]" captionStart="FIGURE 11" captionStartId="20.[151,250,1596,1620]" captionTargetBox="[179,1427,206,1559]" captionTargetId="figure-16@20.[151,1435,181,1572]" captionTargetPageId="20" captionText="FIGURE 11. Eloeophila tergilobellus Kato, sp. nov. A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. gonocoxite, lateral view (upper = dorsal, left = anterior); E. outer gonostylus, ventral view; F. inner gonostylus, outer surface; G. aedeagal complex, dorsal view; H. same, lateral view (left = dorsal). Scale bars: A–B, 2 mm; C–H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057311" httpUri="https://zenodo.org/record/5057311/files/figure.png" pageId="24" pageNumber="449">Figs 11C–D</figureCitation>
)...................................................................... 
<taxonomicName id="4C624D0C1205FA0BFB455845FA8FFB94" authority="Kato, 2021" authorityName="Kato" authorityYear="2021" box="[1206,1353,1090,1112]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="tergilobellus" status="sp. nov.">
<emphasis id="B916EA9D1205FA0BFB455845FA8FFB94" box="[1206,1353,1090,1112]" italics="true" pageId="24" pageNumber="449">E. tergilobellus</emphasis>
</taxonomicName>
<emphasis id="B916EA9D1205FA0BFAA35844FA5AFB94" bold="true" box="[1360,1436,1090,1111]" pageId="24" pageNumber="449">
<taxonomicNameLabel id="A22557E61205FA0BFAA35844FA5AFB94" box="[1360,1436,1090,1111]" pageId="24" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BDD368F1205FA0BFF645866FA5AFB50" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF645866FA5AFB50" pageId="24" pageNumber="449">
- Male tergite 8 with long rectangular lobe at center of posterior margin; dorsal side of gonocoxite with three lobes bearing tufts of long setae................................................................. 
<taxonomicName id="4C624D0C1205FA0BFB85587BFA5AFB50" ID-CoL="39CDR" authority="Savchenko, 1976" authorityName="Savchenko" authorityYear="1976" box="[1142,1436,1149,1171]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="verrucosa">
<emphasis id="B916EA9D1205FA0BFB85587BFB37FB51" box="[1142,1265,1149,1170]" italics="true" pageId="24" pageNumber="449">E. verrucosa</emphasis>
<bibRefCitation id="EFF34B7E1205FA0BFB04587BFA5AFB50" author="Savchenko, E. N. &amp; Krivolutskaya, G. O." box="[1271,1436,1149,1171]" pageId="24" pageNumber="449" refId="ref19194" refString="Savchenko, E. N. &amp; Krivolutskaya, G. O. (1976) Limoniidae of the south Kuril Islands and south Sakhalin. Akademiya Nauk Ukrainskoy SSR, Naukova Dumka, Kiev, 160 pp. [in Russian]" type="book" year="1976">Savchenko, 1976</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E2A1205FA0BFF64589CFA5AFACB" pageId="24" pageNumber="449">
<paragraph id="8BDD368F1205FA0BFF64589CFA5AFB0E" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF64589CFA5AFB0E" pageId="24" pageNumber="449">
12. Wing 
<quantity id="4C9A9B6A1205FA0BFEEF589DFE95FB73" box="[284,339,1179,1200]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="449" unit="mm" value="7.0">7 mm</quantity>
or longer, yellowish at basal part, with distinctly dark spot each at Rs origin, crossveins Sc-r and r-r, and supernumerary m-cu (
<figureCitation id="13592A0A1205FA0BFE9958B1FE04FB0D" box="[362,450,1207,1230]" captionStart="FIGURE 12" captionStartId="21.[151,250,1599,1623]" captionTargetBox="[199,1371,395,1540]" captionTargetId="figure-47@21.[189,1398,365,1576]" captionTargetPageId="21" captionText="FIGURE 12. Eloeophila ussuriana iwatensis (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. same, outer surface; F. aedeagal complex, dorsal view; G. same, lateral view (left = dorsal); H. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057315" httpUri="https://zenodo.org/record/5057315/files/figure.png" pageId="24" pageNumber="449">Fig. 12B</figureCitation>
); halter pale yellow (
<figureCitation id="13592A0A1205FA0BFD7E58B1FD22FB0D" box="[653,740,1207,1230]" captionStart="FIGURE 12" captionStartId="21.[151,250,1599,1623]" captionTargetBox="[199,1371,395,1540]" captionTargetId="figure-47@21.[189,1398,365,1576]" captionTargetPageId="21" captionText="FIGURE 12. Eloeophila ussuriana iwatensis (Alexander, 1936). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. same, outer surface; F. aedeagal complex, dorsal view; G. same, lateral view (left = dorsal); H. paramere, dorsal surface. Scale bars: A–B, 2 mm; C–G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5057315" httpUri="https://zenodo.org/record/5057315/files/figure.png" pageId="24" pageNumber="449">Fig. 12A</figureCitation>
)...................... 
<taxonomicName id="4C624D0C1205FA0BFBFC58BEFA5AFB0E" ID-CoL="7GS4D" authority="(Alexander, 1936)" baseAuthorityName="Alexander" baseAuthorityYear="1936" box="[1039,1436,1207,1229]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="subSpecies" species="ussuriana" subSpecies="iwatensis">
<emphasis id="B916EA9D1205FA0BFBFC58BEFB21FB0E" box="[1039,1255,1208,1229]" italics="true" pageId="24" pageNumber="449">E. ussuriana iwatensis</emphasis>
(Alexander, 1936)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDD368F1205FA0BFF6458D3FA5AFACB" blockId="24.[151,1437,151,1288]" pageId="24" pageNumber="449">
<keyLead id="309395BA1205FA0BFF6458D3FA5AFACB" pageId="24" pageNumber="449">
- Wing shorter than 
<quantity id="4C9A9B6A1205FA0BFE6A58D3FE20FB29" box="[409,486,1237,1259]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="24" pageNumber="449" unit="mm" value="6.7">6.7 mm</quantity>
, without yellowish basal part and distinctly dark spot except stigma (
<figureCitation id="13592A0A1205FA0BFB7D58D3FB1CFB28" box="[1166,1242,1237,1259]" captionStart="FIGURE 1" captionStartId="2.[151,250,1599,1623]" captionTargetBox="[172,1415,201,1547]" captionTargetId="figure-16@2.[151,1435,179,1574]" captionTargetPageId="2" captionText="FIGURE 1. Eloeophila amamiana (Alexander, 1956). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface; H. female terminalia, lateral view. Scale bars: A–B, 2 mm; C–H, 0.1 mm. Abbreviations: ad, aedeagus; av, apodeme of vesica; cr, cercus; hv, hypogynial valve; ig, inner gonostylus; lp, lateral apodeme; ls, lower shelf; outer gonostylus; pm, paramere; s m-cu, supernumerary m-cu; sX, sternite X; tX, tergite X; vs, vesica." figureDoi="http://doi.org/10.5281/zenodo.5057237" httpUri="https://zenodo.org/record/5057237/files/figure.png" pageId="24" pageNumber="449">Fig. 1B</figureCitation>
); halter dark brown except yellowish base (
<figureCitation id="13592A0A1205FA0BFE3158F4FDCAFACB" box="[450,524,1266,1288]" captionStart="FIGURE 1" captionStartId="2.[151,250,1599,1623]" captionTargetBox="[172,1415,201,1547]" captionTargetId="figure-16@2.[151,1435,179,1574]" captionTargetPageId="2" captionText="FIGURE 1. Eloeophila amamiana (Alexander, 1956). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface; H. female terminalia, lateral view. Scale bars: A–B, 2 mm; C–H, 0.1 mm. Abbreviations: ad, aedeagus; av, apodeme of vesica; cr, cercus; hv, hypogynial valve; ig, inner gonostylus; lp, lateral apodeme; ls, lower shelf; outer gonostylus; pm, paramere; s m-cu, supernumerary m-cu; sX, sternite X; tX, tergite X; vs, vesica." figureDoi="http://doi.org/10.5281/zenodo.5057237" httpUri="https://zenodo.org/record/5057237/files/figure.png" pageId="24" pageNumber="449">Fig. 1A</figureCitation>
)................................................ 
<taxonomicName id="4C624D0C1205FA0BFB9458F5FA5AFACB" ID-CoL="6F4QP" authority="(Alexander, 1956)" baseAuthorityName="Alexander" baseAuthorityYear="1956" box="[1127,1436,1266,1288]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="449" phylum="Arthropoda" rank="species" species="amamiana">
<emphasis id="B916EA9D1205FA0BFB9458F5FB21FACB" box="[1127,1255,1267,1288]" italics="true" pageId="24" pageNumber="449">E. amamiana</emphasis>
(Alexander, 1956)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>