595 lines
126 KiB
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595 lines
126 KiB
XML
<document id="405072B487A8F77054A15A9EF5065DAD" ID-DOI="10.5852/ejt.2021.747.1333" ID-GBIF-Dataset="aca67cc8-96ca-4f24-b8af-2b532ecd355c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4836744" ID-ZooBank="3F0B6EAF-C616-4865-811A-414A094B590C" IM.bibliography_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1619605517524" checkinUser="plazi" docAuthor="Moctezuma, Victor & Halffter, Gonzalo" docDate="2021" docId="03CA87F2FFCDFFF4FF154720BA053DD3" docLanguage="en" docName="ejt-747_1-71.pdf" docOrigin="European Journal of Taxonomy 747" docSource="http://zoobank.org/3f0b6eaf-c616-4865-811a-414a094b590c" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phanaeus Macleay 1819" docType="key" docVersion="11" lastPageNumber="54" masterDocId="FFF3FF8AFFFEFFC1FFA8420BBF723A3A" masterDocTitle="Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species" masterLastPageNumber="71" masterPageNumber="1" pageNumber="52" updateTime="1733497921341" updateUser="jonas" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="86552FCEBCF758561575E7683F978FA5">Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species</mods:title>
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<mods:namePart id="85C8EE4EC48D08F9FCB95EA28F30CA74">Moctezuma, Victor</mods:namePart>
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<mods:affiliation id="F8477806900466913938A84CEE8258EC">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="6BF321AE79062382207843D53F7E0277" type="email">abadonjvpm@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="464C6EAC23968EB16C9B403F60F4FD50">Halffter, Gonzalo</mods:namePart>
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<mods:affiliation id="E82905BB1BF640707E1B52D4F82FBD65">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:nameIdentifier id="7421E1E9B8310098CC33C983B322F326" type="email">gonzalohalffter@inecol.mx</mods:nameIdentifier>
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<mods:title id="A7A95D23A6A362B1F121E1122D70E1DB">European Journal of Taxonomy</mods:title>
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<mods:date id="52FB46FD321DD6A1F5AB543FDA35EDB5">2021</mods:date>
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<treatment id="03CA87F2FFCDFFF4FF154720BA053DD3" ID-DOI="http://doi.org/10.5281/zenodo.4726243" ID-GBIF-Taxon="181753780" ID-Zenodo-Dep="4726243" LSID="urn:lsid:plazi:treatment:03CA87F2FFCDFFF4FF154720BA053DD3" httpUri="http://treatment.plazi.org/id/03CA87F2FFCDFFF4FF154720BA053DD3" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52" scope_family="Scarabaeidae" scope_order="Coleoptera">
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<subSubSection id="C379656FFFCDFFF2FF154720BCE53F7D" box="[189,919,1322,1351]" pageId="51" pageNumber="52" type="nomenclature">
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<paragraph id="8BDC36E4FFCDFFF2FF154720BCE53F7D" blockId="51.[189,919,1322,1351]" box="[189,919,1322,1351]" pageId="51" pageNumber="52">
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<emphasis id="B917EAF6FFCDFFF2FF154720BCE53F7D" bold="true" box="[189,919,1322,1351]" italics="true" pageId="51" pageNumber="52">
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Key to species of the
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<taxonomicName id="4C634D67FFCDFFF2FE784721BDAF3F7D" ID-CoL="6MRP" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[464,733,1322,1351]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">Phanaeus endymion</taxonomicName>
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species group
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</emphasis>
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</paragraph>
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<subSubSection id="C379656FFFCDFFF4FF154754BA053DD3" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52" type="key">
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<key id="FFF261D5FFCDFFF4FF154754BA053DD3" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52">
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<paragraph id="8BDC36E4FFCDFFF2FF154754BA053F85" blockId="51.[189,1399,1375,1507]" pageId="51" pageNumber="52">
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<keyLead id="309295D1FFCDFFF2FF154754BA053F85" pageId="51" pageNumber="52">1. Sutural margin of each elytron upturned to form a sharp ridge, which is progressively more elevated posteriorly and prolonged into a small, sharp tooth at apical angle; elytral margin slightly excised adjacent to this tooth.......................................................................................................................... 2</keyLead>
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</paragraph>
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<paragraph id="8BDC36E4FFCDFFF2FF6947C2BA053FD8" blockId="51.[189,1399,1375,1507]" box="[193,1399,1480,1507]" pageId="51" pageNumber="52">
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<keyLead id="309295D1FFCDFFF2FF6947C2BA053FD8" box="[193,1399,1480,1507]" pageId="51" pageNumber="52">– Sutural margin of elytra simple (running straight, not upturned)...................................................... 3</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30972E41FFCDFFF2FF154401BA053C8B" pageId="51" pageNumber="52">
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<paragraph id="8BDC36E4FFCDFFF2FF154401BA053C7D" blockId="51.[189,1399,1546,1713]" pageId="51" pageNumber="52">
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<keyLead id="309295D1FFCDFFF2FF154401BA053C7D" pageId="51" pageNumber="52">
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2. Male with dentiform keel in the middle of anterior pronotal margin, lateral lines of pronotal triangle straight. Southern
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<collectingRegion id="49A7F806FFCDFFF2FE6A4426BD2C3C7D" box="[450,606,1581,1607]" country="Mexico" name="Mexico" pageId="51" pageNumber="52">Mexico State</collectingRegion>
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and
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<collectingRegion id="49A7F806FFCDFFF2FD3E4426BD8A3C7D" box="[662,760,1581,1607]" country="Mexico" name="Morelos" pageId="51" pageNumber="52">Morelos</collectingRegion>
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(
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<figureCitation id="13582A61FFCDFFF2FCA04426BC2F3C72" box="[776,861,1581,1608]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="51" pageNumber="52">Fig. 15</figureCitation>
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)....................
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<taxonomicName id="4C634D67FFCDFFF2FC564426BA053C7D" ID-CoL="4FSJM" authority="Edmonds, 1979" authorityName="Edmonds" authorityYear="1979" box="[1022,1399,1580,1607]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="halffterorum">
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<emphasis id="B917EAF6FFCDFFF2FC564426BBC53C7D" bold="true" box="[1022,1207,1580,1607]" italics="true" pageId="51" pageNumber="52">P. halffterorum</emphasis>
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<bibRefCitation id="EFF24B15FFCDFFF2FB164426BA053C7D" author="Edmonds" box="[1214,1399,1581,1607]" firstAuthor="Edmonds" pageId="51" pageNumber="52" pagination="99 - 105" refId="ref37459" refString="Edmonds W. D. 1979. A new species of Phanaeus from Mexico (Coleoptera: Scarabaeidae). Pan-Pacific Entomologist 55: 99 - 105." type="journal article" year="1979">Edmonds, 1979</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BDC36E4FFCDFFF2FF69445ABA053C8B" blockId="51.[189,1399,1546,1713]" pageId="51" pageNumber="52">
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<keyLead id="309295D1FFCDFFF2FF69445ABA053C8B" pageId="51" pageNumber="52">
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– Male with keel absent in the middle of anterior pronotal margin, lateral lines of pronotal triangle curved. Sierra Madre del Sur, central
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<collectingRegion id="49A7F806FFCDFFF2FD0D4478BC7D3CB7" box="[677,783,1651,1677]" country="Mexico" name="Guerrero" pageId="51" pageNumber="52">Guerrero</collectingRegion>
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(
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<figureCitation id="13582A61FFCDFFF2FCB74478BC063CB4" box="[799,884,1651,1678]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="51" pageNumber="52">Fig. 15</figureCitation>
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) ................................................................... .............................................................
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<taxonomicName id="4C634D67FFCDFFF2FD1B449CBA053C8B" ID-CoL="4FSJD" authority="Moctezuma, Sanchez-Huerta & Halffter, 2017" authorityName="Moctezuma, Sanchez-Huerta & Halffter" authorityYear="2017" box="[691,1399,1686,1713]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="bravoensis">
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<emphasis id="B917EAF6FFCDFFF2FD1B449CBC233C8B" bold="true" box="[691,849,1686,1713]" italics="true" pageId="51" pageNumber="52">P. bravoensis</emphasis>
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<bibRefCitation id="EFF24B15FFCDFFF2FCF0449CBA053C8B" author="Moctezuma, Sanchez-Huerta & Halffter" box="[856,1399,1686,1713]" firstAuthor="Moctezuma" pageId="51" pageNumber="52" pagination="113 - 135" refId="ref40712" refString="Moctezuma V., Sanchez-Huerta J. L. & Halffter G. 2017. Two new species of the Phanaeus endymion species group (Coleoptera, Scarabaeidae, Scarabaeinae). ZooKeys 702: 113 - 135. https: // doi. org / 10.3897 / zookeys. 702.14728" type="journal article" year="2017">Moctezuma, Sánchez-Huerta & Halffter, 2017</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30972E41FFCDFFF2FF1544D3BA053DD2" pageId="51" pageNumber="52">
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<paragraph id="8BDC36E4FFCDFFF2FF1544D3BA053D45" blockId="51.[189,1399,1752,2025]" pageId="51" pageNumber="52">
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<keyLead id="309295D1FFCDFFF2FF1544D3BA053D45" pageId="51" pageNumber="52">
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3. Pronotal disc of male evenly and densely but finely granulate, granules in most specimens larger and becoming squamose along lateral margins of disc and extending onto posterolateral angles; sides of pronotum strongly roughened (
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<figureCitation id="13582A61FFCDFFF2FD1A4515BC623D02" box="[690,784,1822,1848]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="51" pageNumber="52">Figs 2L</figureCitation>
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,
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<figureCitation id="13582A61FFCDFFF2FCB34515BC003D02" box="[795,882,1822,1848]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="51" pageNumber="52">14A, C</figureCitation>
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). Female pronotum slightly roughened; with distinctly impressed midline (
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<figureCitation id="13582A61FFCDFFF2FDE9454ABDC23D66" box="[577,688,1857,1884]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="51" pageNumber="52">Figs 14B</figureCitation>
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,
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<figureCitation id="13582A61FFCDFFF2FD134549BD833D66" box="[699,753,1858,1884]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="51" pageNumber="52">18Q</figureCitation>
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). Sierra Madre del Sur and Sierra Norte, central
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<collectingRegion id="49A7F806FFCDFFF2FAB64549BA043D66" box="[1310,1398,1858,1884]" country="Mexico" name="Oaxaca" pageId="51" pageNumber="52">Oaxaca</collectingRegion>
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||
(
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<figureCitation id="13582A61FFCDFFF2FF5A456EBE353DBA" box="[242,327,1893,1920]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="51" pageNumber="52">Fig. 15</figureCitation>
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) .................................................................................................
|
||
<taxonomicName id="4C634D67FFCDFFF2FB8D456EBA053D45" ID-CoL="4FSK2" authority="Edmonds, 2006" authorityName="Edmonds" authorityYear="2006" box="[1061,1399,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="zapotecus">
|
||
<emphasis id="B917EAF6FFCDFFF2FB8D456EBBC53D45" bold="true" box="[1061,1207,1893,1919]" italics="true" pageId="51" pageNumber="52">P. zapotecus</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCDFFF2FB16456EBA053D45" author="Edmonds" box="[1214,1399,1893,1919]" firstAuthor="Edmonds" pageId="51" pageNumber="52" pagination="31 - 37" refId="ref37655" refString="Edmonds W. D. 2006. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico. Zootaxa 1171 (3): 31 - 37. https: // doi. org / 10.11646 / zootaxa. 1171.1.3" type="journal article" year="2006">Edmonds, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCDFFF2FF694583BA053DD2" blockId="51.[189,1399,1752,2025]" pageId="51" pageNumber="52">
|
||
<keyLead id="309295D1FFCDFFF2FF694583BA053DD2" pageId="51" pageNumber="52">– Pronotal disc of male lacking distinct granulation, or, if granules present, these are minute and restricted along lateral margins of disc; sides of pronotum smooth. Female pronotum smooth; with superficially impressed to completely effaced midline...................................................................... 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCAFFF5FF154302BA053BE6" pageId="52" pageNumber="53">
|
||
<paragraph id="8BDC36E4FFCAFFF5FF154302BA053B57" blockId="52.[189,1401,265,476]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF154302BA053B57" pageId="52" pageNumber="53">
|
||
4. Elytral striae deeply punctate; all strial punctures forming a distinct fossa, giving a completely roughened surface to striae (
|
||
<figureCitation id="13582A61FFCAFFF5FD9E4325BDEA3B72" box="[566,664,302,328]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="52" pageNumber="53">Figs 1K</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FD0C4325BDB83B72" box="[676,714,302,328]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="52" pageNumber="53">2G</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FD7F4325BD9A3B72" box="[727,744,302,328]" captionStart="Fig" captionStartId="34.[189,232,1997,2023]" captionTargetBox="[211,1370,749,1965]" captionTargetId="figure-104@34.[189,1399,749,1965]" captionTargetPageId="34" captionText="Fig. 9. Phanaeus pacificus sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836782" httpUri="https://zenodo.org/record/4836782/files/figure.png" pageId="52" pageNumber="53">9</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FD5C4325BC553B72" box="[756,807,302,328]" captionStart="Fig" captionStartId="56.[189,231,1678,1704]" captionTargetBox="[306,1282,265,1646]" captionTargetId="figure-158@56.[306,1282,265,1646]" captionTargetPageId="56" captionText="Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836804" httpUri="https://zenodo.org/record/4836804/files/figure.png" pageId="52" pageNumber="53">19L</figureCitation>
|
||
). Pacific slope of the Sierra Madre de Chiapas and
|
||
<collectingCountry id="F3747674FFCAFFF5FF424359BE063B56" box="[234,372,338,364]" name="El Salvador" pageId="52" pageNumber="53">El Salvador</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFCAFFF5FE2C4359BEAB3B57" box="[388,473,338,365]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="52" pageNumber="53">Fig. 15</figureCitation>
|
||
).............................................................................................
|
||
<taxonomicName id="4C634D67FFCAFFF5FB254358BA673B56" ID-CoL="874FG" authority="Moctezuma & Halffter, 2021" authorityName="Moctezuma & Halffter" authorityYear="2021" box="[1165,1301,338,365]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pacificus" status="sp. nov.">
|
||
<emphasis id="B917EAF6FFCAFFF5FB254358BA673B56" bold="true" box="[1165,1301,338,365]" italics="true" pageId="52" pageNumber="53">P. pacificus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFCAFFF5FAB44358BA053B57" box="[1308,1399,339,365]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCAFFF5FF694373BA053BE6" blockId="52.[189,1401,265,476]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF694373BA053BE6" pageId="52" pageNumber="53">– Elytral striae distinctly punctate to impunctate. If strial punctures distinctly impressed, forming a distinct fossa giving a completely roughened surface to first and second striae, or strial punctures forming a distinct fossa giving a partially roughened surface to basal half of striae ....................... 5</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCAFFF5FF15400FBA04388B" pageId="52" pageNumber="53">
|
||
<paragraph id="8BDC36E4FFCAFFF5FF15400FBA053879" blockId="52.[189,1399,516,690]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF15400FBA053879" pageId="52" pageNumber="53">
|
||
5. Pronotum of female with anteromedial concavity bounded anteriorly by three variable on shape tubercles (
|
||
<figureCitation id="13582A61FFCAFFF5FECC4022BEB63879" box="[356,452,553,579]" captionStart="Fig" captionStartId="13.[189,232,1925,1951]" captionTargetBox="[218,1398,491,1881]" captionTargetId="figure-72@13.[189,1398,491,1889]" captionTargetPageId="13" captionText="Fig. 4. Phanaeus endymion Harold, 1863. A. ♂, blue morph (typical; VMC). B. ♂, green morph (VMC). C. ♀, blue morph (typical; VMC). D. Lectotype, ♂ (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836760" httpUri="https://zenodo.org/record/4836760/files/figure.png" pageId="52" pageNumber="53">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FE784022BD703879" box="[464,514,553,579]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="52" pageNumber="53">11B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FDA74022BD303879" box="[527,578,553,579]" captionStart="Fig" captionStartId="45.[189,232,1998,2024]" captionTargetBox="[220,1360,1157,1946]" captionTargetId="figure-204@45.[189,1399,1157,1960]" captionTargetPageId="45" captionText="Fig. 13. Phanaeus rzedowskii sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836792" httpUri="https://zenodo.org/record/4836792/files/figure.png" pageId="52" pageNumber="53">13B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FDE64022BDB03879" box="[590,706,553,579]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="52" pageNumber="53">18E, N, P</figureCitation>
|
||
).......................................................................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCAFFF5FF694045BA04388B" blockId="52.[189,1399,516,690]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF694045BA04388B" pageId="52" pageNumber="53">
|
||
– Pronotum of female evenly convex, lacking anteromedial concavity even in largest specimens, bearing three round, smooth tubercles in transverse line near anterior margin (
|
||
<figureCitation id="13582A61FFCAFFF5FBD64079BBAC38B6" box="[1150,1246,626,652]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[198,1364,1148,1905]" captionTargetId="figure-267@10.[189,1399,1148,1928]" captionTargetPageId="10" captionText="Fig. 3. Phanaeus chiapanecus sp. nov. A. Holotype, ♂, blue morph (typical; TAMU). B. ♀, green morph (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836756" httpUri="https://zenodo.org/record/4836756/files/figure.png" pageId="52" pageNumber="53">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FB434079BA7D38B7" box="[1259,1295,626,653]" captionStart="Fig" captionStartId="19.[189,232,1962,1988]" captionTargetBox="[189,1373,555,1930]" captionTargetId="figure-106@19.[189,1399,555,1931]" captionTargetPageId="19" captionText="Fig. 5. Phanaeus funereus Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Jiří Hájek, Národní Muzeum, Prague, Czech Republic) (NMPC)." figureDoi="http://doi.org/10.5281/zenodo.4836764" httpUri="https://zenodo.org/record/4836764/files/figure.png" pageId="52" pageNumber="53">5B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FAB44079BA3238B6" box="[1308,1344,626,652]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="52" pageNumber="53">6B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FAE54078BA0038B7" box="[1357,1394,627,653]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="52" pageNumber="53">7B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FF43409CBE7D388B" box="[235,271,663,689]" captionStart="Fig" captionStartId="31.[189,232,1962,1988]" captionTargetBox="[199,1366,462,1913]" captionTargetId="figure-52@31.[189,1399,462,1925]" captionTargetPageId="31" captionText="Fig. 8. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Lectotype ♂ (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836778" httpUri="https://zenodo.org/record/4836778/files/figure.png" pageId="52" pageNumber="53">8B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FEB3409CBE3C388B" box="[283,334,663,689]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="52" pageNumber="53">10B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FEF3409CBEFC388B" box="[347,398,663,689]" captionStart="Fig" captionStartId="43.[189,232,1962,1988]" captionTargetBox="[319,1279,265,1926]" captionTargetId="figure-38@43.[306,1281,265,1926]" captionTargetPageId="43" captionText="Fig. 12. Phanaeus pyrois Bates, 1887. A. ♂, red morph (VCM). B. ♂, blue morph (VMC). C. ♀, green morph (TAMU). D. Lectotype, ♂ (by Mario Cupello, Universidade Federal do Paraná, Brazil) (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.4836790" httpUri="https://zenodo.org/record/4836790/files/figure.png" pageId="52" pageNumber="53">12C</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FE33409CBD00388B" box="[411,626,663,690]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="52" pageNumber="53">18C, F, I–K, M, O</figureCitation>
|
||
)................................................................................................... 10
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCAFFF5FF1540D1BA053E7A" pageId="52" pageNumber="53">
|
||
<paragraph id="8BDC36E4FFCAFFF5FF1540D1BA053959" blockId="52.[189,1399,730,1088]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF1540D1BA053959" pageId="52" pageNumber="53">
|
||
6. Pronotum of female with concavity bounded anteriorly by a raised U- or V-shaped process; middle pronotal tubercle dentiform or rounded, always more developed and frontally projected than lateral tubercles; pronotal concavity interrupted posteriorly by a small rounded bump or strongly developed dentiform tubercle (
|
||
<figureCitation id="13582A61FFCAFFF5FE7A4143BD403959" box="[466,562,840,867]" captionStart="Fig" captionStartId="13.[189,232,1925,1951]" captionTargetBox="[218,1398,491,1881]" captionTargetId="figure-72@13.[189,1398,491,1889]" captionTargetPageId="13" captionText="Fig. 4. Phanaeus endymion Harold, 1863. A. ♂, blue morph (typical; VMC). B. ♂, green morph (VMC). C. ♀, blue morph (typical; VMC). D. Lectotype, ♂ (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836760" httpUri="https://zenodo.org/record/4836760/files/figure.png" pageId="52" pageNumber="53">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FD964142BD033959" box="[574,625,841,867]" captionStart="Fig" captionStartId="45.[189,232,1998,2024]" captionTargetBox="[220,1360,1157,1946]" captionTargetId="figure-204@45.[189,1399,1157,1960]" captionTargetPageId="45" captionText="Fig. 13. Phanaeus rzedowskii sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836792" httpUri="https://zenodo.org/record/4836792/files/figure.png" pageId="52" pageNumber="53">13B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FDD64142BDA23959" box="[638,720,841,867]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="52" pageNumber="53">18E, P</figureCitation>
|
||
). Endophallite copulatrix variable................................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCAFFF5FF694165BA053E7A" blockId="52.[189,1399,730,1088]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF694165BA053E7A" pageId="52" pageNumber="53">
|
||
– Pronotum of female with concavity bounded anteriorly by three isolated, round, almost aligned tubercles; middle tubercle sometimes dentiform and more strongly developed than lateral tubercles (
|
||
<figureCitation id="13582A61FFCAFFF5FEC541BCBE9639EB" box="[365,484,951,977]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="52" pageNumber="53">Figs 11B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FE5241BCBD4339EB" box="[506,561,951,977]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="52" pageNumber="53">18N</figureCitation>
|
||
). Right and left lobes of endophallite copulatrix similar in size; right lobe obtusely triangular in shape, weakly developed superiorly; left lobe obtusely lobed, strongly developed; central ridge distinctly developed (
|
||
<figureCitation id="13582A61FFCAFFF5FCD6460BBC913E21" box="[894,995,1024,1051]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="52" pageNumber="53">Fig. 1M</figureCitation>
|
||
). Caribbean
|
||
<collectingCountry id="F3747674FFCAFFF5FBD4460BBB9C3E20" box="[1148,1262,1024,1050]" name="Honduras" pageId="52" pageNumber="53">Honduras</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3747674FFCAFFF5FA84460BBA053E20" box="[1324,1399,1024,1050]" name="Belize" pageId="52" pageNumber="53">Belize</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFCAFFF5FF5D462EBE383E7A" box="[245,330,1061,1088]" captionStart="Fig" captionStartId="51.[189,232,1233,1259]" captionTargetBox="[193,1395,270,1197]" captionTargetId="figure-241@51.[189,1398,265,1199]" captionTargetPageId="51" captionText="Fig. 16. Distribution of the Phanaeus endymion species group in Central America (in part)." figureDoi="http://doi.org/10.5281/zenodo.4836798" httpUri="https://zenodo.org/record/4836798/files/figure.png" pageId="52" pageNumber="53">Fig. 16</figureCitation>
|
||
) ..........................................................................................
|
||
<taxonomicName id="4C634D67FFCAFFF5FC59462DBA7F3E05" ID-CoL="874DL" authority="Arnaud, 2002" authorityName="Arnaud" authorityYear="2002" baseAuthorityName="Arnaud" baseAuthorityYear="2001" box="[1009,1293,1061,1088]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="porioni" status="stat. nov.">
|
||
<emphasis id="B917EAF6FFCAFFF5FC59462DBB143E05" bold="true" box="[1009,1126,1061,1088]" italics="true" pageId="52" pageNumber="53">P. porioni</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCAFFF5FBC4462EBA7F3E05" author="Arnaud P." box="[1132,1293,1061,1087]" journalOrPublisher="Besoiro" pageId="52" pageNumber="53" pagination="2 - 5" part="8" refId="ref35833" refString="Arnaud P. 2002 a. Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae). Besoiro 8: 2 - 5." title="Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae)" type="journal article" year="2002">Arnaud, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFCAFFF5FABC462DBA053E7A" box="[1300,1399,1062,1088]" pageId="52" pageNumber="53">stat. rev.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCAFFF5FF154662BA053F00" pageId="52" pageNumber="53">
|
||
<paragraph id="8BDC36E4FFCAFFF5FF154662BA053ECB" blockId="52.[189,1399,1128,1339]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF154662BA053ECB" pageId="52" pageNumber="53">
|
||
7. Pronotal disc of male with superficially impressed midline (
|
||
<figureCitation id="13582A61FFCAFFF5FC694663BB503EB8" box="[961,1058,1128,1154]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="52" pageNumber="53">Fig. 2K</figureCitation>
|
||
). Right lobe of endophallite copulatrix more developed than left lobe; right lobe obtusely triangular in shape, sharply acute frontally, rounded superiorly; left lobe strongly developed, obtusely lobed; central ridge strongly developed (
|
||
<figureCitation id="13582A61FFCAFFF5FEDB46DCBEA23ECB" box="[371,464,1239,1265]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="52" pageNumber="53">Fig. 1O</figureCitation>
|
||
). Sierra Madre del Sur, southern
|
||
<collectingRegion id="49A7F806FFCAFFF5FCFD46DCBCA53ECB" box="[853,983,1239,1265]" country="Mexico" name="Michoacan de Ocampo" pageId="52" pageNumber="53">Michoacán</collectingRegion>
|
||
(
|
||
<figureCitation id="13582A61FFCAFFF5FC4F46DCBB4E3EC8" box="[999,1084,1239,1266]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="52" pageNumber="53">Fig. 15</figureCitation>
|
||
) ......
|
||
<taxonomicName id="4C634D67FFCAFFF5FBD146DCBA673ECB" ID-CoL="874G3" authority="Moctezuma & Halffter, 2021" authorityName="Moctezuma & Halffter" authorityYear="2021" box="[1145,1301,1238,1265]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="rzedowskii" status="sp. nov.">
|
||
<emphasis id="B917EAF6FFCAFFF5FBD146DCBA673ECB" bold="true" box="[1145,1301,1238,1265]" italics="true" pageId="52" pageNumber="53">P. rzedowskii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFCAFFF5FAB446DCBA053ECB" box="[1308,1399,1239,1265]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCAFFF5FF6946F7BA053F00" blockId="52.[189,1399,1128,1339]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF6946F7BA053F00" pageId="52" pageNumber="53">
|
||
– Pronotal disc of male with completely effaced midline (
|
||
<figureCitation id="13582A61FFCAFFF5FC3446F0BC8F3F2C" box="[924,1021,1275,1302]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="52" pageNumber="53">Fig. 2B</figureCitation>
|
||
). Central ridge of endophallite copulatrix distinctly developed (
|
||
<figureCitation id="13582A61FFCAFFF5FDCE472BBC7B3F00" box="[614,777,1312,1338]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="52" pageNumber="53">Fig. 1D, G, Q</figureCitation>
|
||
) ................................................................................ 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCAFFF5FF154768BA053C9E" pageId="52" pageNumber="53">
|
||
<paragraph id="8BDC36E4FFCAFFF5FF154768BA053C0F" blockId="52.[189,1401,1379,1700]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF154768BA053C0F" pageId="52" pageNumber="53">
|
||
8. Pronotum of male uniformly dark blue, dark metallic blue-green or dark metallic green (
|
||
<figureCitation id="13582A61FFCAFFF5FAB84768BA003F47" box="[1296,1394,1379,1405]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="52" pageNumber="53">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCAFFF5FF444783BE453F98" box="[236,311,1416,1442]" captionStart="Fig" captionStartId="13.[189,232,1925,1951]" captionTargetBox="[218,1398,491,1881]" captionTargetId="figure-72@13.[189,1398,491,1889]" captionTargetPageId="13" captionText="Fig. 4. Phanaeus endymion Harold, 1863. A. ♂, blue morph (typical; VMC). B. ♂, green morph (VMC). C. ♀, blue morph (typical; VMC). D. Lectotype, ♂ (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836760" httpUri="https://zenodo.org/record/4836760/files/figure.png" pageId="52" pageNumber="53">4A–B</figureCitation>
|
||
). Weakly developed keel close to anterior margin of pronotum, or keel completely effaced (
|
||
<figureCitation id="13582A61FFCAFFF5FF5D47A6BE3F3FF2" box="[245,333,1453,1480]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="52" pageNumber="53">Fig. 2b</figureCitation>
|
||
). Right lobe of endophallite copulatrix more developed than left lobe; right lobe obtusely triangular in shape; left lobe obtusely lobed inferiorly, obtusely triangular superiorly; central ridge less developed than central column (
|
||
<figureCitation id="13582A61FFCAFFF5FD3F47FDBD873C2B" box="[663,757,1526,1553]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="52" pageNumber="53">Fig. 1D</figureCitation>
|
||
). Lowlands and midlands of the Gulf of Mexico slope, Yucatán Peninsula,
|
||
<collectingCountry id="F3747674FFCAFFF5FE794410BD6D3C0F" box="[465,543,1563,1589]" name="Belize" pageId="52" pageNumber="53">Belize</collectingCountry>
|
||
, northern
|
||
<collectingCountry id="F3747674FFCAFFF5FD3D4410BC663C0F" box="[661,788,1563,1589]" name="Guatemala" pageId="52" pageNumber="53">Guatemala</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFCAFFF5FC8C4410BC0B3C0C" box="[804,889,1563,1590]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="52" pageNumber="53">Fig. 15</figureCitation>
|
||
).........................
|
||
<taxonomicName id="4C634D67FFCAFFF5FBE94417BA053C0F" ID-CoL="4FSJL" authority="Harold, 1863" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[1089,1399,1563,1590]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFCAFFF5FBE94417BBA63C0C" bold="true" box="[1089,1236,1563,1590]" italics="true" pageId="52" pageNumber="53">P. endymion</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCAFFF5FB744410BA053C0F" author="Harold" box="[1244,1399,1563,1589]" firstAuthor="Harold" pageId="52" pageNumber="53" pagination="161 - 176" refId="ref38961" refString="Harold E. 1863. Note sur les especes mexicaines du genre Phanaeus et descriptions de quelques especes nouvelles de coleopteres mexicains. Annales de la Societe entomologique de France 3: 161 - 176." type="journal article" year="1863">Harold, 1863</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCAFFF5FF69444ABA053C9E" blockId="52.[189,1401,1379,1700]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF69444ABA053C9E" pageId="52" pageNumber="53">– Pronotum of male uniformly bright metallic green, bright metallic green-blue, sometimes showing a red or golden sheen. Keel always absent on anterior margin of pronotum. Endophallite copulatrix variable .............................................................................................................................................. 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCAFFF5FF1544C6BA053DD2" pageId="52" pageNumber="53">
|
||
<paragraph id="8BDC36E4FFCAFFF5FF1544C6BA053D6F" blockId="52.[189,1399,1741,2025]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF1544C6BA053D6F" pageId="52" pageNumber="53">
|
||
9. Anterior metasternal angle obtuse in lateral view. Lateral metasternal angles well defined and slightly curved. Left lobe of endophallite copulatrix almost completely lobed (
|
||
<figureCitation id="13582A61FFCAFFF5FB2F44FABB973D36" box="[1159,1253,1777,1804]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="52" pageNumber="53">Fig. 1Q</figureCitation>
|
||
). Temperate mountains of Los Chimalapas, eastern
|
||
<collectingRegion id="49A7F806FFCAFFF5FD1B451DBC793D0A" box="[691,779,1814,1840]" country="Mexico" name="Oaxaca" pageId="52" pageNumber="53">Oaxaca</collectingRegion>
|
||
and western
|
||
<collectingRegion id="49A7F806FFCAFFF5FC0F451DBB743D0A" box="[935,1030,1814,1840]" country="Mexico" name="Chiapas" pageId="52" pageNumber="53">Chiapas</collectingRegion>
|
||
(
|
||
<figureCitation id="13582A61FFCAFFF5FBBD451DBB183D0B" box="[1045,1130,1814,1841]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="52" pageNumber="53">Fig. 15</figureCitation>
|
||
).................................. .................................................................................................
|
||
<taxonomicName id="4C634D67FFCAFFF5FC104530BA053D6F" ID-CoL="4FSK3" authority="Moctezuma & Halffter, 2017" authorityName="Moctezuma & Halffter" authorityYear="2017" box="[952,1399,1851,1878]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="zoque">
|
||
<emphasis id="B917EAF6FFCAFFF5FC104530BB693D6F" bold="true" box="[952,1051,1851,1877]" italics="true" pageId="52" pageNumber="53">P. zoque</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCAFFF5FB8A4537BA053D6F" author="Moctezuma & Halffter" box="[1058,1399,1851,1878]" firstAuthor="Moctezuma" pageId="52" pageNumber="53" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma & Halffter, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCAFFF5FF69456BBA053DD2" blockId="52.[189,1399,1741,2025]" pageId="52" pageNumber="53">
|
||
<keyLead id="309295D1FFCAFFF5FF69456BBA053DD2" pageId="52" pageNumber="53">
|
||
– Anterior metasternal angle almost right angled but with rounded apex in lateral view. Lateral metasternal angles evanescent. Left lobe of endophallite copulatrix straight posteroinferiorly (
|
||
<figureCitation id="13582A61FFCAFFF5FF5B45A2BE223DFE" box="[243,336,1961,1988]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="52" pageNumber="53">Fig. 1G</figureCitation>
|
||
).
|
||
<collectingRegion id="49A7F806FFCAFFF5FECE45A2BECB3DF9" box="[358,441,1961,1987]" country="Mexico" name="Jalisco" pageId="52" pageNumber="53">Jalisco</collectingRegion>
|
||
, northern
|
||
<collectingRegion id="49A7F806FFCAFFF5FD8745A2BDC63DF9" box="[559,692,1961,1987]" country="Mexico" name="Michoacan de Ocampo" pageId="52" pageNumber="53">Michoacán</collectingRegion>
|
||
,
|
||
<collectingRegion id="49A7F806FFCAFFF5FD6845A2BC693DF9" box="[704,795,1961,1987]" country="Mexico" name="Nayarit" pageId="52" pageNumber="53">Nayarit</collectingRegion>
|
||
, southern
|
||
<collectingRegion id="49A7F806FFCAFFF5FC3B45A2BC993DF9" box="[915,1003,1961,1987]" country="Mexico" name="Sinaloa" pageId="52" pageNumber="53">Sinaloa</collectingRegion>
|
||
(
|
||
<figureCitation id="13582A61FFCAFFF5FC5345A2BB223DFE" box="[1019,1104,1961,1988]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="52" pageNumber="53">Fig. 15</figureCitation>
|
||
) ..................................... ..................................................................
|
||
<taxonomicName id="4C634D67FFCAFFF5FD7F45C4BA053DD2" ID-CoL="4FSJP" authority="Moctezuma, Sanchez-Huerta & Halffter, 2017" authorityName="Moctezuma, Sanchez-Huerta & Halffter" authorityYear="2017" box="[727,1399,1998,2025]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="huichol">
|
||
<emphasis id="B917EAF6FFCAFFF5FD7F45C4BC233DD3" bold="true" box="[727,849,1998,2025]" italics="true" pageId="52" pageNumber="53">P. huichol</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCAFFF5FCF045C4BA053DD2" author="Moctezuma, Sanchez-Huerta & Halffter" box="[856,1399,1998,2025]" firstAuthor="Moctezuma" pageId="52" pageNumber="53" pagination="113 - 135" refId="ref40712" refString="Moctezuma V., Sanchez-Huerta J. L. & Halffter G. 2017. Two new species of the Phanaeus endymion species group (Coleoptera, Scarabaeidae, Scarabaeinae). ZooKeys 702: 113 - 135. https: // doi. org / 10.3897 / zookeys. 702.14728" type="journal article" year="2017">Moctezuma, Sánchez-Huerta & Halffter, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCBFFF4FF154302BA043BA2" pageId="53" pageNumber="54">
|
||
<paragraph id="8BDC36E4FFCBFFF4FF154302BA043B70" blockId="53.[189,1398,265,409]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF154302BA043B70" pageId="53" pageNumber="54">
|
||
10. Elytral interstriae evenly convex and glossy midlongitudinally; striae impressed basally as distinct fossae (
|
||
<figureCitation id="13582A61FFCBFFF4FEEC433BBEE23B70" box="[324,400,304,330]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[198,1364,1148,1905]" captionTargetId="figure-267@10.[189,1399,1148,1928]" captionTargetPageId="10" captionText="Fig. 3. Phanaeus chiapanecus sp. nov. A. Holotype, ♂, blue morph (typical; TAMU). B. ♀, green morph (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836756" httpUri="https://zenodo.org/record/4836756/files/figure.png" pageId="53" pageNumber="54">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FE35433BBEDC3B70" box="[413,430,304,330]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="53" pageNumber="54">6</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FE12433ABEB93B71" box="[442,459,305,331]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="53" pageNumber="54">7</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FE70433BBD4F3B71" box="[472,573,304,331]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="53" pageNumber="54">18C, I–J</figureCitation>
|
||
) ...........................................................................................................11
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCBFFF4FF694353BA043BA2" blockId="53.[189,1398,265,409]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF694353BA043BA2" pageId="53" pageNumber="54">
|
||
– Elytral interstriae distinctly flattened and uniformly dull; striae not strongly impressed basally (
|
||
<figureCitation id="13582A61FFCBFFF4FF5B4375BE4C3BA3" box="[243,318,382,409]" captionStart="Fig" captionStartId="19.[189,232,1962,1988]" captionTargetBox="[189,1373,555,1930]" captionTargetId="figure-106@19.[189,1399,555,1931]" captionTargetPageId="19" captionText="Fig. 5. Phanaeus funereus Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Jiří Hájek, Národní Muzeum, Prague, Czech Republic) (NMPC)." figureDoi="http://doi.org/10.5281/zenodo.4836764" httpUri="https://zenodo.org/record/4836764/files/figure.png" pageId="53" pageNumber="54">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FEE34375BE293BA2" box="[331,347,382,408]" captionStart="Fig" captionStartId="31.[189,232,1962,1988]" captionTargetBox="[199,1366,462,1913]" captionTargetId="figure-52@31.[189,1399,462,1925]" captionTargetPageId="31" captionText="Fig. 8. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Lectotype ♂ (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836778" httpUri="https://zenodo.org/record/4836778/files/figure.png" pageId="53" pageNumber="54">8</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FEC14375BEFB3BA2" box="[361,393,382,408]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="53" pageNumber="54">10</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FE3D4375BEC73BA2" box="[405,437,382,408]" captionStart="Fig" captionStartId="43.[189,232,1962,1988]" captionTargetBox="[319,1279,265,1926]" captionTargetId="figure-38@43.[306,1281,265,1926]" captionTargetPageId="43" captionText="Fig. 12. Phanaeus pyrois Bates, 1887. A. ♂, red morph (VCM). B. ♂, blue morph (VMC). C. ♀, green morph (TAMU). D. Lectotype, ♂ (by Mario Cupello, Universidade Federal do Paraná, Brazil) (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.4836790" httpUri="https://zenodo.org/record/4836790/files/figure.png" pageId="53" pageNumber="54">12</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FE694375BD2F3BA2" box="[449,605,382,409]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="53" pageNumber="54">18F, K, M, O</figureCitation>
|
||
)...................................................................................................... 14
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCBFFF4FF1543C8BA043811" pageId="53" pageNumber="54">
|
||
<paragraph id="8BDC36E4FFCBFFF4FF1543C8BA05383E" blockId="53.[189,1399,451,556]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF1543C8BA05383E" pageId="53" pageNumber="54">
|
||
11. Almost completely bright black dorsally (
|
||
<figureCitation id="13582A61FFCBFFF4FD7D43C8BC463BE7" box="[725,820,451,477]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Figs 2E</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FCE843CFBC233BE4" box="[832,849,452,478]" captionStart="Fig" captionStartId="27.[189,232,1925,1951]" captionTargetBox="[219,1380,610,1882]" captionTargetId="figure-108@27.[189,1399,610,1883]" captionTargetPageId="27" captionText="Fig. 7. Phanaeus malyi Arnaud, 2002.A. ♂ (TAMU). B. ♀ (TAMU). C. Colombian specimen previously labelled as a paralectotype of P. blanchardi Olsoufieff, 1924 (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836774" httpUri="https://zenodo.org/record/4836774/files/figure.png" pageId="53" pageNumber="54">7</figureCitation>
|
||
). Pacific slope of the Cordillera de Talamanca,
|
||
<collectingCountry id="F3747674FFCBFFF4FF4243E1BE18383E" box="[234,362,490,516]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3747674FFCBFFF4FE0B43E0BD72383F" box="[419,512,491,517]" name="Panama" pageId="53" pageNumber="54">Panama</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFCBFFF4FDB843E1BD17383F" box="[528,613,490,517]" captionStart="Fig" captionStartId="51.[189,232,1233,1259]" captionTargetBox="[193,1395,270,1197]" captionTargetId="figure-241@51.[189,1398,265,1199]" captionTargetPageId="51" captionText="Fig. 16. Distribution of the Phanaeus endymion species group in Central America (in part)." figureDoi="http://doi.org/10.5281/zenodo.4836798" httpUri="https://zenodo.org/record/4836798/files/figure.png" pageId="53" pageNumber="54">Fig. 16</figureCitation>
|
||
) .....................................................................
|
||
<taxonomicName id="4C634D67FFCBFFF4FBD843E0BA05383E" ID-CoL="4FSJS" authority="Arnaud, 2002" authorityName="Arnaud" authorityYear="2002" box="[1136,1399,490,517]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="malyi">
|
||
<emphasis id="B917EAF6FFCBFFF4FBD843E0BBA2383F" bold="true" box="[1136,1232,490,517]" italics="true" pageId="53" pageNumber="54">P. malyi</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCBFFF4FB7D43E1BA05383E" author="Arnaud P." box="[1237,1399,490,516]" journalOrPublisher="Besoiro" pageId="53" pageNumber="54" pagination="2 - 5" part="8" refId="ref35833" refString="Arnaud P. 2002 a. Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae). Besoiro 8: 2 - 5." title="Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae)" type="journal article" year="2002">Arnaud, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCBFFF4FF694019BA043811" blockId="53.[189,1399,451,556]" box="[193,1398,529,556]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF694019BA043811" box="[193,1398,529,556]" pageId="53" pageNumber="54">
|
||
– Pronotum typically bright metallic green, blue-green or dark metallic blue (
|
||
<figureCitation id="13582A61FFCBFFF4FBF6401ABB963816" box="[1118,1252,529,556]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Figs 2A, D</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FB58401ABA723811" box="[1264,1280,529,555]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[198,1364,1148,1905]" captionTargetId="figure-267@10.[189,1399,1148,1928]" captionTargetPageId="10" captionText="Fig. 3. Phanaeus chiapanecus sp. nov. A. Holotype, ♂, blue morph (typical; TAMU). B. ♀, green morph (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836756" httpUri="https://zenodo.org/record/4836756/files/figure.png" pageId="53" pageNumber="54">3</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FAA5401ABA6E3816" box="[1293,1308,529,556]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="53" pageNumber="54">6</figureCitation>
|
||
)...... 12
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCBFFF4FF15405CBA043937" pageId="53" pageNumber="54">
|
||
<paragraph id="8BDC36E4FFCBFFF4FF15405CBA053885" blockId="53.[189,1399,598,781]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF15405CBA053885" pageId="53" pageNumber="54">
|
||
12. Distinctly developed keel on central anterior margin of pronotum of male; right lobe of endophallite copulatrix tapering superiorly and straight apically (
|
||
<figureCitation id="13582A61FFCBFFF4FCED4076BCED38A2" box="[837,927,637,664]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="53" pageNumber="54">Fig. 1C</figureCitation>
|
||
). Sierra Madre del Sur, southern
|
||
<collectingRegion id="49A7F806FFCBFFF4FAB64075BA0438A2" box="[1310,1398,638,664]" country="Mexico" name="Oaxaca" pageId="53" pageNumber="54">Oaxaca</collectingRegion>
|
||
(
|
||
<figureCitation id="13582A61FFCBFFF4FF5A40AEBE353885" box="[242,327,676,703]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="53" pageNumber="54">Fig. 15</figureCitation>
|
||
) ...............................................................
|
||
<taxonomicName id="4C634D67FFCBFFF4FC8F40AEBA053885" ID-CoL="4FSJK" authority="Moctezuma, Deloya & Halffter, 2019" authorityName="Moctezuma, Deloya & Halffter" authorityYear="2019" box="[807,1399,676,703]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="edmondsi">
|
||
<emphasis id="B917EAF6FFCBFFF4FC8F40AEBCCA3885" bold="true" box="[807,952,676,703]" italics="true" pageId="53" pageNumber="54">P. edmondsi</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCBFFF4FC1740AEBA053885" author="Moctezuma, Deloya & Halffter" box="[959,1399,677,703]" firstAuthor="Moctezuma" pageId="53" pageNumber="54" pagination="249 - 254" refId="ref40763" refString="Moctezuma V., Deloya C., Sanchez-Huerta J. L. & Halffter G. 2019. A new species of the Phanaeus endymion species group (Coleoptera: Scarabaeidae: Scarabaeinae), with comments on ecology and distribution. Annales de la Societe entomologique de France (N. S.) 55: 249 - 254. https: // doi. org / 10.1080 / 00379271.2019.1577170" type="journal article" year="2019">Moctezuma, Deloya & Halffter, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCBFFF4FF6940C7BA043937" blockId="53.[189,1399,598,781]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF6940C7BA043937" pageId="53" pageNumber="54">
|
||
– Always with keel absent on central anterior margin of pronotum of male (
|
||
<figureCitation id="13582A61FFCBFFF4FBE240C7BBBE38DC" box="[1098,1228,716,742]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Fig. 2A, D</figureCitation>
|
||
). Endophallite copulatrix variable ........................................................................................................................... 13
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCBFFF4FF154133BA053E2C" pageId="53" pageNumber="54">
|
||
<paragraph id="8BDC36E4FFCBFFF4FF154133BA05399A" blockId="53.[189,1399,824,1046]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF154133BA05399A" pageId="53" pageNumber="54">
|
||
13. Larger species, frequently attaining
|
||
<quantity id="4C9B9B01FFCBFFF4FD354133BC6C3968" box="[669,798,824,850]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="53" pageNumber="54" unit="mm" value="19.5" valueMax="20.0" valueMin="19.0">19–20 mm</quantity>
|
||
in length. Posterolateral angles of male widened, slightly projected posteriorly (
|
||
<figureCitation id="13582A61FFCBFFF4FDE54154BDDC3943" box="[589,686,863,889]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FD134154BDB93943" box="[699,715,863,889]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[198,1364,1148,1905]" captionTargetId="figure-267@10.[189,1399,1148,1928]" captionTargetPageId="10" captionText="Fig. 3. Phanaeus chiapanecus sp. nov. A. Holotype, ♂, blue morph (typical; TAMU). B. ♀, green morph (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836756" httpUri="https://zenodo.org/record/4836756/files/figure.png" pageId="53" pageNumber="54">3</figureCitation>
|
||
). Central highlands of
|
||
<collectingRegion id="49A7F806FFCBFFF4FC714154BB4A3943" box="[985,1080,863,889]" country="Mexico" name="Chiapas" pageId="53" pageNumber="54">Chiapas</collectingRegion>
|
||
(
|
||
<figureCitation id="13582A61FFCBFFF4FBEF4154BBEE3940" box="[1095,1180,863,890]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="53" pageNumber="54">Fig. 15</figureCitation>
|
||
) ........................... ........................................................................................................................
|
||
<taxonomicName id="4C634D67FFCBFFF4FBC9418DBA67399A" ID-CoL="874FN" authority="Moctezuma & Halffter, 2021" authorityName="Moctezuma & Halffter" authorityYear="2021" box="[1121,1301,901,928]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="chiapanecus" status="sp. nov.">
|
||
<emphasis id="B917EAF6FFCBFFF4FBC9418DBA67399A" bold="true" box="[1121,1301,901,928]" italics="true" pageId="53" pageNumber="54">P. chiapanecus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFCBFFF4FAB4418DBA05399A" box="[1308,1399,902,928]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCBFFF4FF6941A6BA053E2C" blockId="53.[189,1399,824,1046]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF6941A6BA053E2C" pageId="53" pageNumber="54">
|
||
– Smaller species, never attaining
|
||
<quantity id="4C9B9B01FFCBFFF4FDD341A6BDA139FD" box="[635,723,941,967]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="53" pageNumber="54" unit="mm" value="19.0">19 mm</quantity>
|
||
in length. Posterolateral angles of male sharply acute, strongly projected posteriorly (
|
||
<figureCitation id="13582A61FFCBFFF4FDF941DFBDC339D4" box="[593,689,980,1006]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Figs 2D</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FD1441DFBDBF39D5" box="[700,717,980,1007]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="53" pageNumber="54">6</figureCitation>
|
||
). Inner slope of the Sierra Madre de Chiapas, south-central
|
||
<collectingCountry id="F3747674FFCBFFF4FF4141F0BE1A3E2F" box="[233,360,1019,1045]" name="Guatemala" pageId="53" pageNumber="54">Guatemala</collectingCountry>
|
||
and southeastern Chiapas (
|
||
<figureCitation id="13582A61FFCBFFF4FD0241F0BD8C3E2C" box="[682,766,1019,1046]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="53" pageNumber="54">Fig. 15</figureCitation>
|
||
) ...................................................
|
||
<taxonomicName id="4C634D67FFCBFFF4FB2141F0BA673E2F" ID-CoL="874F5" authority="Moctezuma & Halffter, 2021" authorityName="Moctezuma & Halffter" authorityYear="2021" box="[1161,1301,1018,1045]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="jackenioi" status="sp. nov.">
|
||
<emphasis id="B917EAF6FFCBFFF4FB2141F0BA673E2F" bold="true" box="[1161,1301,1018,1045]" italics="true" pageId="53" pageNumber="54">P. jackenioi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFCBFFF4FAB441F7BA053E2C" box="[1308,1399,1020,1046]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCBFFF4FF15464BBA043E92" pageId="53" pageNumber="54">
|
||
<paragraph id="8BDC36E4FFCBFFF4FF15464BBA043E61" blockId="53.[189,1398,1088,1193]" box="[189,1398,1088,1115]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF15464BBA043E61" box="[189,1398,1088,1115]" pageId="53" pageNumber="54">
|
||
14. Pronotal disc of male completely bright metallic (
|
||
<figureCitation id="13582A61FFCBFFF4FC99464BBCFB3E60" box="[817,905,1088,1114]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Figs 2J</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FC3D464BBCC13E60" box="[917,947,1088,1114]" captionStart="Fig" captionStartId="43.[189,232,1962,1988]" captionTargetBox="[319,1279,265,1926]" captionTargetId="figure-38@43.[306,1281,265,1926]" captionTargetPageId="43" captionText="Fig. 12. Phanaeus pyrois Bates, 1887. A. ♂, red morph (VCM). B. ♂, blue morph (VMC). C. ♀, green morph (TAMU). D. Lectotype, ♂ (by Mario Cupello, Universidade Federal do Paraná, Brazil) (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.4836790" httpUri="https://zenodo.org/record/4836790/files/figure.png" pageId="53" pageNumber="54">12</figureCitation>
|
||
)....................................................... 15
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCBFFF4FF694663BA043E92" blockId="53.[189,1398,1088,1193]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF694663BA043E92" pageId="53" pageNumber="54">
|
||
– Pronotal disc of male distinctly black, without metallic reflection (
|
||
<figureCitation id="13582A61FFCBFFF4FC5C466CBBFE3EB8" box="[1012,1164,1127,1154]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Figs 2E, J, L</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FB31466CBB903EBB" box="[1177,1250,1127,1154]" captionStart="Fig" captionStartId="19.[189,232,1962,1988]" captionTargetBox="[189,1373,555,1930]" captionTargetId="figure-106@19.[189,1399,555,1931]" captionTargetPageId="19" captionText="Fig. 5. Phanaeus funereus Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Jiří Hájek, Národní Muzeum, Prague, Czech Republic) (NMPC)." figureDoi="http://doi.org/10.5281/zenodo.4836764" httpUri="https://zenodo.org/record/4836764/files/figure.png" pageId="53" pageNumber="54">5A, C</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FB46466CBA663EBB" box="[1262,1300,1127,1153]" captionStart="Fig" captionStartId="31.[189,232,1962,1988]" captionTargetBox="[199,1366,462,1913]" captionTargetId="figure-52@31.[189,1399,462,1925]" captionTargetPageId="31" captionText="Fig. 8. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Lectotype ♂ (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836778" httpUri="https://zenodo.org/record/4836778/files/figure.png" pageId="53" pageNumber="54">8A</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FA88466CBA263EBB" box="[1312,1364,1127,1153]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="53" pageNumber="54">10A</figureCitation>
|
||
) .. ........................................................................................................................................................ 16
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCBFFF4FF1546D8BA053F8B" pageId="53" pageNumber="54">
|
||
<paragraph id="8BDC36E4FFCBFFF4FF1546D8BA053F06" blockId="53.[189,1400,1235,1457]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF1546D8BA053F06" pageId="53" pageNumber="54">
|
||
15. Pronotal disc typically bright metallic green, rarely dark metallic blue. Pronotal process of female with central tubercle more developed and anteriorly projected than lateral tubercles.
|
||
<collectingCountry id="F3747674FFCBFFF4FABE46F1BA0A3F2E" box="[1302,1400,1274,1300]" name="Ecuador" pageId="53" pageNumber="54">Ecuador</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFCBFFF4FF5B472ABE3F3F06" box="[243,333,1313,1340]" captionStart="Fig" captionStartId="54.[191,234,1247,1273]" captionTargetBox="[193,1393,268,1195]" captionTargetId="figure-243@54.[189,1396,265,1198]" captionTargetPageId="54" captionText="Fig. 17. Distribution of the Phanaeus endymion species group in South America. Phanaeus malyi Arnaud, 2001 was reported from Colombia without accurate locality. Phanaeus arletteae Arnaud, 2018 was modified from Arnaud (2018)." figureDoi="http://doi.org/10.5281/zenodo.4836800" httpUri="https://zenodo.org/record/4836800/files/figure.png" pageId="53" pageNumber="54">Figs 17</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FEF24729BEFF3F06" box="[346,397,1314,1340]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="53" pageNumber="54">18A</figureCitation>
|
||
)..............................................................................................
|
||
<taxonomicName id="4C634D67FFCBFFF4FBE24729BA053F06" authority="Arnaud, 2002" authorityName="Arnaud" authorityYear="2002" box="[1098,1399,1313,1340]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="arletteae">
|
||
<emphasis id="B917EAF6FFCBFFF4FBE24729BBBD3F06" bold="true" box="[1098,1231,1313,1340]" italics="true" pageId="53" pageNumber="54">P. arletteae</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCBFFF4FB7D472ABA053F06" author="Arnaud P." box="[1237,1399,1313,1340]" journalOrPublisher="Besoiro" pageId="53" pageNumber="54" pagination="2 - 5" part="8" refId="ref35833" refString="Arnaud P. 2002 a. Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae). Besoiro 8: 2 - 5." title="Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae)" type="journal article" year="2002">Arnaud, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCBFFF4FF694742BA053F8B" blockId="53.[189,1400,1235,1457]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF694742BA053F8B" pageId="53" pageNumber="54">
|
||
– Pronotal disc typically bright metallic red, green, or dark metallic blue (
|
||
<figureCitation id="13582A61FFCBFFF4FB894743BB053F59" box="[1057,1143,1352,1379]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Figs 2J</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FB294742BBD03F59" box="[1153,1186,1353,1379]" captionStart="Fig" captionStartId="43.[189,232,1962,1988]" captionTargetBox="[319,1279,265,1926]" captionTargetId="figure-38@43.[306,1281,265,1926]" captionTargetPageId="43" captionText="Fig. 12. Phanaeus pyrois Bates, 1887. A. ♂, red morph (VCM). B. ♂, blue morph (VMC). C. ♀, green morph (TAMU). D. Lectotype, ♂ (by Mario Cupello, Universidade Federal do Paraná, Brazil) (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.4836790" httpUri="https://zenodo.org/record/4836790/files/figure.png" pageId="53" pageNumber="54">12</figureCitation>
|
||
). Pronotal process of female with tubercles almost aligned, central tubercle not projecting anteriorly (
|
||
<figureCitation id="13582A61FFCBFFF4FB1D4764BA563FB0" box="[1205,1316,1391,1418]" captionStart="Fig" captionStartId="43.[189,232,1962,1988]" captionTargetBox="[319,1279,265,1926]" captionTargetId="figure-38@43.[306,1281,265,1926]" captionTargetPageId="43" captionText="Fig. 12. Phanaeus pyrois Bates, 1887. A. ♂, red morph (VCM). B. ♂, blue morph (VMC). C. ♀, green morph (TAMU). D. Lectotype, ♂ (by Mario Cupello, Universidade Federal do Paraná, Brazil) (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.4836790" httpUri="https://zenodo.org/record/4836790/files/figure.png" pageId="53" pageNumber="54">Figs 12C</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FA99477BBA153FB0" box="[1329,1383,1392,1418]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="53" pageNumber="54">18O</figureCitation>
|
||
).
|
||
<collectingCountry id="F3747674FFCBFFF4FF43479CBE163F8B" box="[235,356,1431,1457]" name="Nicaragua" pageId="53" pageNumber="54">Nicaragua</collectingCountry>
|
||
and north Caribbean
|
||
<collectingCountry id="F3747674FFCBFFF4FDC9479CBD903F8B" box="[609,738,1431,1457]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFCBFFF4FD59479CBC343F8B" box="[753,838,1430,1457]" captionStart="Fig" captionStartId="51.[189,232,1233,1259]" captionTargetBox="[193,1395,270,1197]" captionTargetId="figure-241@51.[189,1398,265,1199]" captionTargetPageId="51" captionText="Fig. 16. Distribution of the Phanaeus endymion species group in Central America (in part)." figureDoi="http://doi.org/10.5281/zenodo.4836798" httpUri="https://zenodo.org/record/4836798/files/figure.png" pageId="53" pageNumber="54">Fig. 16</figureCitation>
|
||
).........................................
|
||
<taxonomicName id="4C634D67FFCBFFF4FBD6479CBA053F8B" ID-CoL="4FSJX" authority="Bates, 1887" authorityName="Bates, Det. W. D. Edmonds" authorityYear="1887" box="[1150,1399,1430,1457]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pyrois">
|
||
<emphasis id="B917EAF6FFCBFFF4FBD6479CBB963F8A" bold="true" box="[1150,1252,1430,1457]" italics="true" pageId="53" pageNumber="54">P. pyrois</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCBFFF4FB44479CBA053F8B" author="Bates" box="[1260,1399,1431,1457]" firstAuthor="Bates" pageId="53" pageNumber="54" refId="ref36135" refString="Bates H. W. 1887. Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. Taylor & Francis, London." type="book" year="1887">Bates, 1887</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCBFFF4FF1547D7BA053CA8" pageId="53" pageNumber="54">
|
||
<paragraph id="8BDC36E4FFCBFFF4FF1547D7BA043C27" blockId="53.[189,1399,1500,1682]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF1547D7BA043C27" pageId="53" pageNumber="54">
|
||
16 Pronotum distinctly bright metallic red frontolaterally. Head bright metallic red with green-yellow metallic sheen (
|
||
<figureCitation id="13582A61FFCBFFF4FE0C4408BD503C27" box="[420,546,1539,1565]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Figs 2F, H</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FD864408BD4C3C27" box="[558,574,1539,1565]" captionStart="Fig" captionStartId="31.[189,232,1962,1988]" captionTargetBox="[199,1366,462,1913]" captionTargetId="figure-52@31.[189,1399,462,1925]" captionTargetPageId="31" captionText="Fig. 8. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Lectotype ♂ (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836778" httpUri="https://zenodo.org/record/4836778/files/figure.png" pageId="53" pageNumber="54">8</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FDE44408BD183C27" box="[588,618,1539,1565]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="53" pageNumber="54">10</figureCitation>
|
||
).................................................................................................... 17
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCBFFF4FF694421BA053CA8" blockId="53.[189,1399,1500,1682]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF694421BA053CA8" pageId="53" pageNumber="54">
|
||
– Pronotum distinctly dull metallic red frontolaterally. Head dull metallic red without green-yellow metallic sheen (
|
||
<figureCitation id="13582A61FFCBFFF4FE1A445ABD643C51" box="[434,534,1617,1643]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFCBFFF4FD81445ABD483C56" box="[553,570,1617,1644]" captionStart="Fig" captionStartId="19.[189,232,1962,1988]" captionTargetBox="[189,1373,555,1930]" captionTargetId="figure-106@19.[189,1399,555,1931]" captionTargetPageId="19" captionText="Fig. 5. Phanaeus funereus Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Jiří Hájek, Národní Muzeum, Prague, Czech Republic) (NMPC)." figureDoi="http://doi.org/10.5281/zenodo.4836764" httpUri="https://zenodo.org/record/4836764/files/figure.png" pageId="53" pageNumber="54">5</figureCitation>
|
||
). Pacific slope of the Andes, north-central
|
||
<collectingCountry id="F3747674FFCBFFF4FBF3445ABBCF3C51" box="[1115,1213,1617,1643]" name="Ecuador" pageId="53" pageNumber="54">Ecuador</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3747674FFCBFFF4FAAB445ABA053C51" box="[1283,1399,1617,1643]" name="Colombia" pageId="53" pageNumber="54">Colombia</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFCBFFF4FF5B4473BE3A3CA8" box="[243,328,1656,1682]" captionStart="Fig" captionStartId="54.[191,234,1247,1273]" captionTargetBox="[193,1393,268,1195]" captionTargetId="figure-243@54.[189,1396,265,1198]" captionTargetPageId="54" captionText="Fig. 17. Distribution of the Phanaeus endymion species group in South America. Phanaeus malyi Arnaud, 2001 was reported from Colombia without accurate locality. Phanaeus arletteae Arnaud, 2018 was modified from Arnaud (2018)." figureDoi="http://doi.org/10.5281/zenodo.4836800" httpUri="https://zenodo.org/record/4836800/files/figure.png" pageId="53" pageNumber="54">Fig. 17</figureCitation>
|
||
) .....................................................................................
|
||
<taxonomicName id="4C634D67FFCBFFF4FC6E4473BA7F3CA8" ID-CoL="874G8" authority="Balthasar, 1939" authorityName="Balthasar" authorityYear="1939" box="[966,1293,1655,1682]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="funereus" status="stat. nov.">
|
||
<emphasis id="B917EAF6FFCBFFF4FC6E4473BB3D3CAB" bold="true" box="[966,1103,1655,1682]" italics="true" pageId="53" pageNumber="54">P. funereus</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCBFFF4FBFE4473BA7F3CA8" author="Balthasar" box="[1110,1293,1656,1682]" firstAuthor="Balthasar" pageId="53" pageNumber="54" pagination="238 - 247" refId="ref36059" refString="Balthasar V. 1939. Neue Phanaeus - Arten. Folia Zoologica et Hydrobiologica 9: 238 - 247." type="journal article" year="1939">Balthasar, 1939</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFCBFFF4FABC4473BA053CA8" box="[1300,1399,1656,1682]" pageId="53" pageNumber="54">stat. rev.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30972E41FFCBFFF4FF1544B6BA053DD3" pageId="53" pageNumber="54">
|
||
<paragraph id="8BDC36E4FFCBFFF4FF1544B6BA053D77" blockId="53.[189,1401,1725,2025]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF1544B6BA053D77" pageId="53" pageNumber="54">
|
||
17. Larger species, frequently attaining from
|
||
<quantity id="4C9B9B01FFCBFFF4FD6744B6BC163CE2" box="[719,868,1725,1752]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="53" pageNumber="54" unit="mm" value="23.0" valueMax="24.0" valueMin="22.0">22 to 24 mm</quantity>
|
||
in length. Carinate, distinctly developed keel on central anterior margin of pronotum of male (
|
||
<figureCitation id="13582A61FFCBFFF4FC9244EFBCE53CC4" box="[826,919,1764,1790]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Fig. 2F</figureCitation>
|
||
). Pronotal posterolateral angles of male strongly developed, projected laterally (
|
||
<figureCitation id="13582A61FFCBFFF4FD114500BC643D1F" box="[697,790,1803,1829]" captionStart="Fig" captionStartId="31.[189,232,1962,1988]" captionTargetBox="[199,1366,462,1913]" captionTargetId="figure-52@31.[189,1399,462,1925]" captionTargetPageId="31" captionText="Fig. 8. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Lectotype ♂ (by Christophe Rivier, Muséum national d’histoire naturelle, Paris, France) (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4836778" httpUri="https://zenodo.org/record/4836778/files/figure.png" pageId="53" pageNumber="54">Fig. 8A</figureCitation>
|
||
). Pacific slope of the Andes, south-central
|
||
<collectingCountry id="F3747674FFCBFFF4FAAB4500BA053D1F" box="[1283,1399,1803,1829]" name="Colombia" pageId="53" pageNumber="54">Colombia</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="F3747674FFCBFFF4FE2F4539BE9B3D76" box="[391,489,1842,1868]" name="Ecuador" pageId="53" pageNumber="54">Ecuador</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFCBFFF4FE514539BD3F3D76" box="[505,589,1842,1868]" captionStart="Fig" captionStartId="54.[191,234,1247,1273]" captionTargetBox="[193,1393,268,1195]" captionTargetId="figure-243@54.[189,1396,265,1198]" captionTargetPageId="54" captionText="Fig. 17. Distribution of the Phanaeus endymion species group in South America. Phanaeus malyi Arnaud, 2001 was reported from Colombia without accurate locality. Phanaeus arletteae Arnaud, 2018 was modified from Arnaud (2018)." figureDoi="http://doi.org/10.5281/zenodo.4836800" httpUri="https://zenodo.org/record/4836800/files/figure.png" pageId="53" pageNumber="54">Fig. 17</figureCitation>
|
||
)................................................
|
||
<taxonomicName id="4C634D67FFCBFFF4FC114538BA7F3D76" ID-CoL="874GF" authority="Balthasar, 1939" authorityName="Balthasar" authorityYear="1939" box="[953,1293,1842,1869]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="olsoufieffi" status="stat. nov.">
|
||
<emphasis id="B917EAF6FFCBFFF4FC114538BB3D3D77" bold="true" box="[953,1103,1842,1869]" italics="true" pageId="53" pageNumber="54">P. olsoufieffi</emphasis>
|
||
<bibRefCitation id="EFF24B15FFCBFFF4FBFE4539BA7F3D76" author="Balthasar" box="[1110,1293,1842,1868]" firstAuthor="Balthasar" pageId="53" pageNumber="54" pagination="238 - 247" refId="ref36059" refString="Balthasar V. 1939. Neue Phanaeus - Arten. Folia Zoologica et Hydrobiologica 9: 238 - 247." type="journal article" year="1939">Balthasar, 1939</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFCBFFF4FABC4538BA053D77" box="[1300,1399,1843,1869]" pageId="53" pageNumber="54">stat. rev.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFCBFFF4FF694551BA053DD3" blockId="53.[189,1401,1725,2025]" pageId="53" pageNumber="54">
|
||
<keyLead id="309295D1FFCBFFF4FF694551BA053DD3" pageId="53" pageNumber="54">
|
||
– Smaller species, rarely attaining
|
||
<quantity id="4C9B9B01FFCBFFF4FDD44552BDA63D4E" box="[636,724,1881,1908]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="53" pageNumber="54" unit="mm" value="20.0">20 mm</quantity>
|
||
in length. Always with keel absent on central anterior margin of pronotum of male (
|
||
<figureCitation id="13582A61FFCBFFF4FDF9458BBDC03DA0" box="[593,690,1920,1946]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="53" pageNumber="54">Fig. 2H</figureCitation>
|
||
). Pronotal posterolateral angles of male weakly developed, widened, and slightly projected posteriorly (
|
||
<figureCitation id="13582A61FFCBFFF4FC8D45ACBCEE3DFB" box="[805,924,1959,1985]" captionStart="Fig" captionStartId="37.[191,234,1996,2022]" captionTargetBox="[210,1374,1195,1954]" captionTargetId="figure-230@37.[189,1399,1195,1963]" captionTargetPageId="37" captionText="Fig. 10. Phanaeus panamensis sp. nov. A. Holotype, ♂ (TAMU). B. ♀ (TAMU)." figureDoi="http://doi.org/10.5281/zenodo.4836786" httpUri="https://zenodo.org/record/4836786/files/figure.png" pageId="53" pageNumber="54">Fig. 10A</figureCitation>
|
||
). Caribbean
|
||
<collectingCountry id="F3747674FFCBFFF4FBED45ACBBBD3DFB" box="[1093,1231,1959,1985]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3747674FFCBFFF4FAB445A3BA0B3DF8" box="[1308,1401,1960,1986]" name="Panama" pageId="53" pageNumber="54">Panama</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFCBFFF4FF5B45C5BE3A3DD3" box="[243,328,1998,2025]" captionStart="Fig" captionStartId="51.[189,232,1233,1259]" captionTargetBox="[193,1395,270,1197]" captionTargetId="figure-241@51.[189,1398,265,1199]" captionTargetPageId="51" captionText="Fig. 16. Distribution of the Phanaeus endymion species group in Central America (in part)." figureDoi="http://doi.org/10.5281/zenodo.4836798" httpUri="https://zenodo.org/record/4836798/files/figure.png" pageId="53" pageNumber="54">Fig. 16</figureCitation>
|
||
) ...........................................................................................................
|
||
<taxonomicName id="4C634D67FFCBFFF4FBC145C4BA663DD2" ID-CoL="874GT" authority="Moctezuma & Halffter, 2021" authorityName="Moctezuma & Halffter" authorityYear="2021" box="[1129,1300,1998,2025]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
|
||
<emphasis id="B917EAF6FFCBFFF4FBC145C4BA663DD2" bold="true" box="[1129,1300,1998,2025]" italics="true" pageId="53" pageNumber="54">P. panamensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFCBFFF4FAB445C4BA053DD3" box="[1308,1399,1999,2025]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |