233 lines
31 KiB
XML
233 lines
31 KiB
XML
<document id="4A8EE8915478EEE44E3AB01ECC5C5BD8" ID-CLB-Dataset="31855" ID-DOI="10.11646/zootaxa.4331.1.1" ID-GBIF-Dataset="14c29731-224f-43a4-ae52-f76823177ce5" ID-ISSN="1175-5326" ID-Zenodo-Dep="1009725" ID-ZooBank="FB50725E-4D03-4507-9CAF-EB939C7F548F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507876567978" checkinUser="plazi" docAuthor="Asami, Takahiro" docDate="2017" docId="03CA87D8FFB8FF90FF0BFD74FB9DFE77" docLanguage="en" docName="zootaxa.4331.1.1.pdf" docOrigin="Zootaxa 4331 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dicharax robustus Asami 2017, n. sp." docType="treatment" docVersion="9" lastPageNumber="71" masterDocId="FFF3FFA0FFFDFFD6FF9CFFC0FF80FFFF" masterDocTitle="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus" masterLastPageNumber="124" masterPageNumber="1" pageNumber="70" updateTime="1698469440756" updateUser="ExternalLinkService">
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<mods:title id="0064D90B8D46EDF778B5A33FE2B22BEB">Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus</mods:title>
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<mods:namePart id="40157308B6CE16F61AA0C70FB10180E3">Asami, Takahiro</mods:namePart>
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<mods:title id="CB12F3816E055D984D24ECA183C67BDD">Zootaxa</mods:title>
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<mods:part id="CE293C885D2B55D16F7A766238D57440">
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<mods:date id="BBF3B8ED191BE8CF3922D4F03F6499F8">2017</mods:date>
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<mods:number id="3CBF1CD309E7212A191AE054BF748EB9">2017-10-11</mods:number>
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<mods:number id="17CF0DCC185AA6746E0ABADFDE531071">4331</mods:number>
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<mods:identifier id="12EEDF44CA95D84658DC2E1CFB1B1EA8" type="DOI">10.11646/zootaxa.4331.1.1</mods:identifier>
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<mods:identifier id="0E458165853FEFCE8C7733DE5D6CD414" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CA87D8FFB8FF90FF0BFD74FB9DFE77" ID-DOI="http://doi.org/10.5281/zenodo.6004452" ID-GBIF-Taxon="135435143" ID-Zenodo-Dep="6004452" LSID="urn:lsid:plazi:treatment:03CA87D8FFB8FF90FF0BFD74FB9DFE77" httpUri="http://treatment.plazi.org/id/03CA87D8FFB8FF90FF0BFD74FB9DFE77" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
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<subSubSection id="C3796545FFB8FF93FF0BFD74FE30FD10" pageId="69" pageNumber="70" type="nomenclature">
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<paragraph id="8BDC36CEFFB8FF93FF0BFD74FC83FD31" blockId="69.[151,771,692,751]" box="[151,771,692,718]" pageId="69" pageNumber="70">
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<heading id="D09481A2FFB8FF93FF0BFD74FC83FD31" bold="true" box="[151,771,692,718]" fontSize="11" level="1" pageId="69" pageNumber="70" reason="1">
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<emphasis id="B917EADCFFB8FF93FF0BFD74FC83FD31" bold="true" box="[151,771,692,718]" pageId="69" pageNumber="70">
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<taxonomicName id="4C634D4DFFB8FF93FF0BFD74FD34FD31" ID-CoL="35L5Z" authority="Pall-Gergely & Hunyadi" authorityName="Asami" authorityYear="2017" box="[151,692,692,718]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="69" pageNumber="70" phylum="Mollusca" rank="species" species="robustus" status="sp. nov.">
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||
<emphasis id="B917EADCFFB8FF93FF0BFD74FEF8FD31" bold="true" box="[151,376,692,718]" italics="true" pageId="69" pageNumber="70">Dicharax robustus</emphasis>
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||
Páll-Gergely & Hunyadi
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||
</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="A22457A7FFB8FF93FD5CFD74FC83FD31" box="[704,771,692,718]" pageId="69" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDC36CEFFB8FF93FF0BFD17FE30FD10" blockId="69.[151,771,692,751]" box="[151,432,726,751]" pageId="69" pageNumber="70">
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<figureCitation id="13582A4BFFB8FF93FF0BFD17FEA2FD10" box="[151,290,727,751]" captionStart="FIGURE 19" captionStartId="30.[151,250,1940,1962]" captionTargetBox="[151,1436,769,1919]" captionTargetId="figure@30.[151,1436,769,1919]" captionTargetPageId="30" captionText="FIGURE 19. Shells of Dicharax species. A. Dicharax anthostoma (Möllendorff, 1885) (lectotype, SMF 39225). B. Dicharax anthostoma (syntype of Alycaeus pentagonus, MCZ 167215). C. Dicharax robustus Páll-Gergely & Hunyadi, n. sp. (holotype, HNHM 99704). Scale represents 5 mm. All photos: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009763/files/figure.png" pageId="69" pageNumber="70">FIGUrEs 19C</figureCitation>
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,
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||
<figureCitation id="13582A4BFFB8FF93FEB2FD17FE02FD10" box="[302,386,727,751]" captionStart="FIGURE 29" captionStartId="46.[151,250,1943,1965]" captionTargetBox="[152,1436,250,1861]" captionTargetId="figure@46.[151,1436,250,1861]" captionTargetPageId="46" captionText="FIGURE 29. Protoconchs of Dicharax species. A – B. Dicharax robustus Páll-Gergely & Hunyadi, n. sp. (20000319, spec. 1, HNHM 99726, paratype from the type locality). C – D. Dicharax cristatus (Möllendorff, 1886) (20100508 A, coll. PGB). E – F. Dicharax depressus (Bavay & Dautzenberg, 1912), MAA 1, Specimen 2. All images: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009783/files/figure.png" pageId="69" pageNumber="70">29A–B</figureCitation>
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,
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||
<figureCitation id="13582A4BFFB8FF93FE11FD16FE2CFD10" box="[397,428,726,751]" captionStart="FIGURE 45" captionStartId="70.[151,250,1564,1586]" captionTargetBox="[152,1436,447,1542]" captionTargetId="figure@70.[151,1436,447,1542]" captionTargetPageId="70" captionText="FIGURE 45. R 2 ribs Dicharax robustus Páll-Gergely & Hunyadi, n. sp. (20000319, spec. 1, HNHM 99726 paratype from the type locality). All images: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009815/files/figure.png" pageId="69" pageNumber="70">45</figureCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3796545FFB8FF93FF0BFCDEFE6FFC38" pageId="69" pageNumber="70" type="materials_examined">
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<paragraph id="8BDC36CEFFB8FF93FF0BFCDEFE6FFC38" blockId="69.[151,1437,798,2011]" pageId="69" pageNumber="70">
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<materialsCitation id="3B0B3C93FFB8FF93FF0BFCDEFE6BFC38" ID-GBIF-Occurrence="1671715149" collectingDate="2011-03-18" collectionCode="COLL, NOT, SHI" country="China" elevation="1778" latitude="24.564034" location="Ex" longLatPrecision="1" longitude="102.84357" municipality="Ex" pageId="69" pageNumber="70" specimenCode="HNHM 99704, HNHM 99726" specimenCount="7" stateProvince="YUNNaN" typeStatus="holotype">
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<emphasis id="B917EADCFFB8FF93FF0BFCDEFEC6FCC8" bold="true" box="[151,326,798,823]" pageId="69" pageNumber="70">
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<typeStatus id="54D8886CFFB8FF93FF0BFCDEFF52FCC8" box="[151,210,798,823]" pageId="69" pageNumber="70">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54D8886CFFB8FF93FED1FCDFFE35FCC8" box="[333,437,799,823]" pageId="69" pageNumber="70" type="holotype">HOLOTypE</typeStatus>
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<specimenCode id="DBC59EB5FFB8FF93FE21FCDFFDE4FCC8" box="[445,612,799,823]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="69" pageNumber="70">HNHM 99704</specimenCode>
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,
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<specimenCode id="DBC59EB5FFB8FF93FDF3FCDFFC97FCC8" box="[623,791,798,823]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="69" pageNumber="70">HNHM 99726</specimenCode>
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/1 (fIGUrEd
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<typeStatus id="54D8886CFFB8FF93FC0FFCDFFC77FCC8" box="[915,1015,799,823]" pageId="69" pageNumber="70" type="paratype">paraTypE</typeStatus>
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,
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<collectingMunicipality id="6BB8ACB4FFB8FF93FB9EFCDFFB9DFCC9" box="[1026,1053,799,822]" pageId="69" pageNumber="70">Ex</collectingMunicipality>
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<collectionCode id="ED72AE0BFFB8FF93FBB8FCDFFBD3FCC8" box="[1060,1107,799,823]" pageId="69" pageNumber="70">COLL</collectionCode>
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. K. OHara), OK/19
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<typeStatus id="54D8886CFFB8FF93FAADFCDFFA1CFCC8" box="[1329,1436,799,823]" pageId="69" pageNumber="70" type="paratype">paraTypEs</typeStatus>
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+ 1 jUvENILE sHELL (
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||
<collectionCode id="ED72AE0BFFB8FF93FEF1FC83FE15FCA4" box="[365,405,835,859]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13781" name="Nottingham City Natural History Museum" pageId="69" pageNumber="70">NOT</collectionCode>
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<typeStatus id="54D8886CFFB8FF93FE02FC83FD84FCA4" box="[414,516,835,859]" pageId="69" pageNumber="70" type="paratype">paraTypE</typeStatus>
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), PGB/2
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<typeStatus id="54D8886CFFB8FF93FDF1FC83FD58FCA4" box="[621,728,835,859]" pageId="69" pageNumber="70" type="paratype">paraTypEs</typeStatus>
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(
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||
<location id="8EBC6015FFB8FF93FD75FC83FC86FCA4" LSID="urn:lsid:plazi:treatment:03CA87D8FFB8FF90FF0BFD74FB9DFE77:8EBC6015FFB8FF93FD75FC83FC86FCA4" box="[745,774,835,859]" country="China" latitude="24.564034" longLatPrecision="1" longitude="102.84357" municipality="Ex" name="Ex" pageId="69" pageNumber="70" stateProvince="YUNNaN">Ex</location>
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<collectionCode id="ED72AE0BFFB8FF93FC93FC83FCBEFCA4" box="[783,830,835,859]" pageId="69" pageNumber="70">COLL</collectionCode>
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||
. K. OHara),
|
||
<emphasis id="B917EADCFFB8FF93FC57FC82FBBBFCA4" bold="true" box="[971,1083,834,859]" pageId="69" pageNumber="70">20000319</emphasis>
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||
<collectingCountry id="F374765EFFB8FF93FBD8FC83FB0AFCA4" box="[1092,1162,835,859]" name="China" pageId="69" pageNumber="70">CHINa</collectingCountry>
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||
,
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||
<collectingRegion id="49A7F82CFFB8FF93FB0BFC83FB70FCA5" box="[1175,1264,835,858]" country="China" name="Yunnan" pageId="69" pageNumber="70">YUNNaN</collectingRegion>
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||
, KUNmING
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<collectionCode id="ED72AE0BFFB8FF93FAECFC83FA18FCA4" box="[1392,1432,835,859]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14032" name="Shihezi Agricultural College" pageId="69" pageNumber="70">SHI</collectionCode>
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, YUqIqU, BIjIaNsHaN, GUaNyINsI (TEmpLE), apprOxImaTE GPS daTa:
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||
<geoCoordinate id="EE575009FFB8FF93FCEFFCA6FB83FC80" box="[883,1027,870,895]" direction="north" orientation="latitude" pageId="69" pageNumber="70" precision="1" value="24.271183">24°16.271'N</geoCoordinate>
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,
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<geoCoordinate id="EE575009FFB8FF93FB93FCA6FB2AFC80" box="[1039,1194,870,895]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="1" value="102.828766">102°49.726'E</geoCoordinate>
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, LEG. K. OHara; HA/3
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<typeStatus id="54D8886CFFB8FF93FF0BFC4BFE86FC5C" box="[151,262,907,931]" pageId="69" pageNumber="70" type="paratype">paraTypEs</typeStatus>
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,
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<emphasis id="B917EADCFFB8FF93FE8CFC4AFEDDFC5C" bold="true" box="[272,349,906,931]" pageId="69" pageNumber="70">2011/5</emphasis>
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<collectingCountry id="F374765EFFB8FF93FEF8FC4BFE2AFC5C" box="[356,426,907,931]" name="China" pageId="69" pageNumber="70">CHINa</collectingCountry>
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,
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||
<collectingRegion id="49A7F82CFFB8FF93FE29FC4BFD8EFC5D" box="[437,526,907,930]" country="China" name="Yunnan" pageId="69" pageNumber="70">YUNNaN</collectingRegion>
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||
, YUxI
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||
<collectionCode id="ED72AE0BFFB8FF93FDC9FC4BFDFDFC5C" box="[597,637,907,931]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14032" name="Shihezi Agricultural College" pageId="69" pageNumber="70">SHI</collectionCode>
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||
, CHENGjIaNG XIaN, LUCHONG CUN,
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||
<elevation id="004ED1FDFFB8FF93FC62FC4BFBD5FC5D" box="[1022,1109,907,931]" metricMagnitude="3" metricUnit="m" metricValue="1.778" pageId="69" pageNumber="70" unit="m" value="1778.0">
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||
<quantity id="4C9B9B2BFFB8FF93FC62FC4BFBD5FC5D" box="[1022,1109,907,931]" metricMagnitude="3" metricUnit="m" metricValue="1.778" pageId="69" pageNumber="70" unit="m" value="1778.0">1778 m</quantity>
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||
</elevation>
|
||
,
|
||
<geoCoordinate id="EE575009FFB8FF93FBFDFC4BFB71FC5C" box="[1121,1265,907,931]" direction="north" orientation="latitude" pageId="69" pageNumber="70" precision="1" value="24.564034">24°33.842'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE575009FFB8FF93FB61FC4AFA18FC5C" box="[1277,1432,906,931]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="1" value="102.84357">102°50.614'E</geoCoordinate>
|
||
, LEG. A. HUNyadI,
|
||
<collectingDate id="EF99E9E6FFB8FF93FEC4FC6FFE6BFC38" box="[344,491,943,967]" pageId="69" pageNumber="70" value="2011-03-18">
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||
<date id="FFDD100EFFB8FF93FEC4FC6FFE6BFC38" box="[344,491,943,967]" pageId="69" pageNumber="70" value="2011-03-18">18 Mar 2011</date>
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||
</collectingDate>
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||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796545FFB8FF93FF5BFC12FC35FC14" box="[199,949,978,1003]" pageId="69" pageNumber="70" type="etymology">
|
||
<paragraph id="8BDC36CEFFB8FF93FF5BFC12FC35FC14" blockId="69.[151,1437,798,2011]" box="[199,949,978,1003]" pageId="69" pageNumber="70">
|
||
<emphasis id="B917EADCFFB8FF93FF5BFC12FECEFC14" bold="true" box="[199,334,978,1003]" pageId="69" pageNumber="70">Etymology.</emphasis>
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THIs NEw spECIEs Is NamEd afTEr ITs rObUsT appEaraNCE.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3796545FFB8FF93FF5BFC36FC0FFBA8" pageId="69" pageNumber="70" type="diagnosis">
|
||
<paragraph id="8BDC36CEFFB8FF93FF5BFC36FC0FFBA8" blockId="69.[151,1437,798,2011]" pageId="69" pageNumber="70">
|
||
<emphasis id="B917EADCFFB8FF93FF5BFC36FEC1FBF0" bold="true" box="[199,321,1014,1039]" pageId="69" pageNumber="70">Diagnosis.</emphasis>
|
||
A mEdIUm sIzEd spECIEs wITH ELEvaTEd spIrE, LONG R2, mOdEraTELy LONG R3, bLUNT swELLING ON R3, aNd rELaTIvELy LONG dIsTaNCE bETwEEN THE TwO LIps Of THE pErIsTOmE. THE CrOss sECTION Of rIbs Of R2 arE T-sHapEd, THEIr TOps aLmOsT rEaCH EaCH OTHEr fOrmING TUNNELs bETwEEN THE rIbs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796545FFB8FF93FF5BFBA2FED6F8B5" pageId="69" pageNumber="70" type="description">
|
||
<paragraph id="8BDC36CEFFB8FF93FF5BFBA2FE38F920" blockId="69.[151,1437,798,2011]" pageId="69" pageNumber="70">
|
||
<emphasis id="B917EADCFFB8FF93FF5BFBA2FED9FB84" bold="true" box="[199,345,1122,1147]" pageId="69" pageNumber="70">Description.</emphasis>
|
||
SHELL (
|
||
<figureCitation id="13582A4BFFB8FF93FE34FBA3FDADFB84" box="[424,557,1123,1147]" captionStart="FIGURE 19" captionStartId="30.[151,250,1940,1962]" captionTargetBox="[151,1436,769,1919]" captionTargetId="figure@30.[151,1436,769,1919]" captionTargetPageId="30" captionText="FIGURE 19. Shells of Dicharax species. A. Dicharax anthostoma (Möllendorff, 1885) (lectotype, SMF 39225). B. Dicharax anthostoma (syntype of Alycaeus pentagonus, MCZ 167215). C. Dicharax robustus Páll-Gergely & Hunyadi, n. sp. (holotype, HNHM 99704). Scale represents 5 mm. All photos: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009763/files/figure.png" pageId="69" pageNumber="70">FIGUrE 19C</figureCitation>
|
||
) Off-wHITE, aLTHOUGH mOsT avaILabLE sHELLs Had brOwNIsH-rEddIsH dIrT bETwEEN THEIr rIbs; spIrE ELEvaTEd, sHELL OUTLINE rOUNdEd frOm dOrsaL vIEw, bOdy wHOrL rOUNdEd; prOTOCONCH (
|
||
<figureCitation id="13582A4BFFB8FF93FB63FB47FF2AFB3C" captionStart="FIGURE 29" captionStartId="46.[151,250,1943,1965]" captionTargetBox="[152,1436,250,1861]" captionTargetId="figure@46.[151,1436,250,1861]" captionTargetPageId="46" captionText="FIGURE 29. Protoconchs of Dicharax species. A – B. Dicharax robustus Páll-Gergely & Hunyadi, n. sp. (20000319, spec. 1, HNHM 99726, paratype from the type locality). C – D. Dicharax cristatus (Möllendorff, 1886) (20100508 A, coll. PGB). E – F. Dicharax depressus (Bavay & Dautzenberg, 1912), MAA 1, Specimen 2. All images: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009783/files/figure.png" pageId="69" pageNumber="70">FIGUrEs 29A– B</figureCitation>
|
||
) CONsIsTs Of apprOxImaTELy 2 wHOrLs, bUT bOUNdary bETwEEN prOTOCONCH aNd R1 NOT vIsIbLE CLEarLy IN aNy Of ExamINEd spECImENs, aLTHOUGH wE ExamINEd INTaCT (NOT wEaTHErEd) sHELLs; prOTOCONCH NOrmaLLy ELEvaTEd (NOT HIGHEr Or LOwEr THaN wHaT wOULd bE ExpECTEd frOm THE OvEraLL sHELL sHapE), IT Is maTTE, wITHOUT aNy NOTabLE sCULpTUrE; R1 apprOxImaTELy 1.5 wHOrLs IN LENGTH, rEGULarLy rIbbEd; IN frEsH sHELLs rIbs sHarp aNd sTrONGLy CUrvEd TOwards apErTUrE; TraNsITION bETwEEN R1 aNd R2 vEry CONspICUOUs bECaUsE R2 mUCH mOrE dENsEr rIbbEd THaN prECEdING arEa; R2 vEry dENsELy rIbbEd (
|
||
<figureCitation id="13582A4BFFB8FF93FDE3FA9FFCB8FA88" box="[639,824,1374,1399]" captionStart="FIGURE 45" captionStartId="70.[151,250,1564,1586]" captionTargetBox="[152,1436,447,1542]" captionTargetId="figure@70.[151,1436,447,1542]" captionTargetPageId="70" captionText="FIGURE 45. R 2 ribs Dicharax robustus Páll-Gergely & Hunyadi, n. sp. (20000319, spec. 1, HNHM 99726 paratype from the type locality). All images: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009815/files/figure.png" pageId="69" pageNumber="70">FIGUrEs 45A–C</figureCitation>
|
||
); CrOss sECTIONaL vIEw Of R2 was ExamINEd IN TwO spECImENs (
|
||
<figureCitation id="13582A4BFFB8FF93FE81FA43FE32FA64" box="[285,434,1410,1435]" captionStart="FIGURE 45" captionStartId="70.[151,250,1564,1586]" captionTargetBox="[152,1436,447,1542]" captionTargetId="figure@70.[151,1436,447,1542]" captionTargetPageId="70" captionText="FIGURE 45. R 2 ribs Dicharax robustus Páll-Gergely & Hunyadi, n. sp. (20000319, spec. 1, HNHM 99726 paratype from the type locality). All images: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009815/files/figure.png" pageId="69" pageNumber="70">FIGUrEs 45D</figureCitation>
|
||
,
|
||
<figureCitation id="13582A4BFFB8FF93FE5EFA42FE76FA64" box="[450,502,1410,1435]" captionStart="FIGURE 68" captionStartId="101.[151,250,1819,1841]" captionTargetBox="[160,1433,429,1796]" captionTargetId="figure@101.[151,1436,409,1797]" captionTargetPageId="101" captionText="FIGURE 68. Schematic diagram of cross sections of R 2 of Dicharax and Metalycaeus species. The aperture opens leftwards. A. Dicharax bison Páll-Gergely & Hunyadi n. sp. B. Dicharax robustus Páll-Gergely & Hunyadi, n. sp. C. Dicharax fimbriatus (Bavay & Dautzenberg, 1912). D. Dicharax depressus (Bavay & Dautzenberg, 1912). E. Dicharax (?) alticola Páll- Gergely & Hunyadi n. sp. F. Dicharax (?) immaculatus Páll-Gergely n. sp. G. Dicharax (?) moellendorffi (Kobelt & Möllendorff, 1886). H. Metalycaeus muciferus (Heude, 1885). I. Metalycaeus oharai Páll-Gergely & Hunyadi n. sp. J. Metalycaeus (?) okuboi Páll-Gergely & Hunyadi, n. sp. Abbreviations: a: anterior crust; p: posterior crust; t: tunnel." httpUri="https://zenodo.org/record/1009859/files/figure.png" pageId="69" pageNumber="70">68B</figureCitation>
|
||
): aNTErIOr CrUsT fOrms a sIGNIfICaNTLy ELEvaTEd, sTrONG HOLLOw rIb, wHICH Has THE aNTErIOr aNd pOsTErIOr prOjECTIONs Of abOUT THE samE LENGTH; THIs rEsULTs IN T-sHapEd CrOss sECTIONaL vIEw; aNTErIOr CrUsT fOLds OvEr THE mUCH wEakEr pOsTErIOr CrUsT, wHICH Is NOT HOLLOw; mICrOTUNNEL NarrOw COmparEd TO rIbs, ITs CrOss sECTIONaL vIEw dEfOrmEd TEardrOp-sHapEd; R2 aNd R3 apprOxImaTELy 110 dEGrEEs TOGETHEr; R2 sOmEwHaT LONGEr THaN R3; R2 aNd R3 sEparaTEd by CONspICUOUs CONsTrICTION; R3 raTHEr IrrEGULarLy wrINkLEd; wrINkLEs apprOxImaTELy as dENsE as rIbs aT ENd Of R1; R3 wITH a bLUNT swELLING ON ITs mIddLE; apErTUrE rOUNdEd wITH sLIGHT UppEr INCIsION; pErIsTOmE rOUNdEd, NOT CrENULaTEd; INNEr LIp vEry sLIGHTLy THICkENEd, ExpaNdEd aNd prOTrUdING; pOsTErIOr LIp aLsO sLIGHTLy ExpaNdEd, rELaTIvELy sHarp, aTTaCHEd TO OUTEr LIp aT COLUmELLar rEGION, bUT bOUNdary bETwEEN TwO LIps CLEarLy vIsIbLE HErE as wELL; dIsTaNCE bETwEEN INNEr aNd OUTEr LIps bECOmEs LarGEr aT THE paLaTaL rEGION; UmbILICUs NarrOw.
|
||
</paragraph>
|
||
<paragraph id="8BDC36CEFFB8FF93FF5BF92AFCDFF8FC" blockId="69.[151,1437,798,2011]" box="[199,863,1770,1795]" pageId="69" pageNumber="70">
|
||
<emphasis id="B917EADCFFB8FF93FF5BF92AFE63F8FC" bold="true" box="[199,483,1770,1795]" pageId="69" pageNumber="70">Measurements (in mm).</emphasis>
|
||
D = 4.4–4.7, H = 3.3–3.6 (N = 4).
|
||
</paragraph>
|
||
<paragraph id="8BDC36CEFFB8FF93FF5BF8CEFED6F8B5" blockId="69.[151,1437,798,2011]" pageId="69" pageNumber="70">
|
||
<emphasis id="B917EADCFFB8FF93FF5BF8CEFED6F8D8" bold="true" box="[199,342,1806,1831]" pageId="69" pageNumber="70">Operculum.</emphasis>
|
||
THE OUTEr sUrfaCE Of THE OpErCULa Of sEvEraL spECImENs was ExamINEd: IT Is mULTIspIraL wITHOUT ELEvaTEd LamELLa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796545FFB8FF93FF5BF896FB49F890" box="[199,1225,1878,1903]" pageId="69" pageNumber="70" type="materials_examined">
|
||
<paragraph id="8BDC36CEFFB8FF93FF5BF896FB49F890" blockId="69.[151,1437,798,2011]" box="[199,1225,1878,1903]" pageId="69" pageNumber="70">
|
||
<emphasis id="B917EADCFFB8FF93FF5BF896FE5DF890" bold="true" box="[199,477,1878,1903]" pageId="69" pageNumber="70">Variation in specimens.</emphasis>
|
||
NO NOTabLE varIabILITy was fOUNd wITHIN aNd bETwEEN pOpULaTIONs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796545FFB8FF90FF5BF8BAFCB8FEBF" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="diagnosis">
|
||
<paragraph id="8BDC36CEFFB8FF90FF5BF8BAFCB8FEBF" blockId="69.[151,1437,798,2011]" lastBlockId="70.[151,1437,151,392]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
|
||
<emphasis id="B917EADCFFB8FF93FF5BF8BAFE4DF86C" bold="true" box="[199,461,1914,1939]" pageId="69" pageNumber="70">Differential diagnosis.</emphasis>
|
||
THIs NEw spECIEs Is mOrE sTrONGLy bUILT OvEraLL THaN
|
||
<taxonomicName id="4C634D4DFFB8FF93FB8BF8BBFB5CF86C" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[1047,1244,1915,1939]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="69" pageNumber="70" phylum="Mollusca" rank="species" species="alticola" status="sp. nov.">
|
||
<emphasis id="B917EADCFFB8FF93FB8BF8BBFB5CF86C" box="[1047,1244,1915,1939]" italics="true" pageId="69" pageNumber="70">Dicharax alticola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457A7FFB8FF93FB7FF8BAFAA1F86C" box="[1251,1313,1914,1939]" pageId="69" pageNumber="70" rank="species">
|
||
<emphasis id="B917EADCFFB8FF93FB7FF8BAFAA1F86C" bold="true" box="[1251,1313,1914,1939]" pageId="69" pageNumber="70">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
MOrEOvEr, IT Has a LarGEr sHELL, a mOrE ELEvaTEd spIrE, dENsEr rIbs ON R2, aNd THE swELLING ON R3 Is sHarpEr, aNd sITUaTEd CLOsEr TO THE apErTUrE.
|
||
<taxonomicName id="4C634D4DFFB8FF93FED0F803FD9EF824" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[332,542,1987,2011]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="69" pageNumber="70" phylum="Mollusca" rank="species" species="robustus" status="sp. nov.">
|
||
<emphasis id="B917EADCFFB8FF93FED0F803FD9EF824" box="[332,542,1987,2011]" italics="true" pageId="69" pageNumber="70">Dicharax robustus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457A7FFB8FF93FDBBF802FDE8F824" box="[551,616,1986,2011]" pageId="69" pageNumber="70" rank="species">
|
||
<emphasis id="B917EADCFFB8FF93FDBBF802FDE8F824" bold="true" box="[551,616,1986,2011]" pageId="69" pageNumber="70">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Has a mOrE ELEvaTEd spIrE, THICkEr pErIsTOmE, sHOrTEr R3 aNd LONGEr R2 THaN THE NEIGHbOUrING pOpULaTIONs Of
|
||
<taxonomicName id="4C634D4DFFBBFF90FDD7FF59FD51FF50" baseAuthorityName="Mollendorff" baseAuthorityYear="1886" box="[587,721,152,176]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="cristatus">
|
||
<emphasis id="B917EADCFFBBFF90FDD7FF59FD51FF50" box="[587,721,152,176]" italics="true" pageId="70" pageNumber="71">D. cristatus</emphasis>
|
||
</taxonomicName>
|
||
. MOrEOvEr, THE swELLING ON R2 Is sITUaTEd mUCH CLOsEr TO THE pErIsTOmE IN
|
||
<taxonomicName id="4C634D4DFFBBFF90FEB2FF7DFE33FF2B" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[302,435,188,212]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="robustus" status="sp. nov.">
|
||
<emphasis id="B917EADCFFBBFF90FEB2FF7DFE33FF2B" box="[302,435,188,212]" italics="true" pageId="70" pageNumber="71">D. robustus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457A7FFBBFF90FE20FF7CFE7EFF2A" box="[444,510,188,213]" pageId="70" pageNumber="71" rank="species">
|
||
<emphasis id="B917EADCFFBBFF90FE20FF7CFE7EFF2A" bold="true" box="[444,510,188,213]" pageId="70" pageNumber="71">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
THE mOsT sImILar spECIEs TO
|
||
<taxonomicName id="4C634D4DFFBBFF90FCD3FF7DFC53FF2B" authorityName="Asami" authorityYear="2017" box="[847,979,188,212]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="robustus">
|
||
<emphasis id="B917EADCFFBBFF90FCD3FF7DFC53FF2B" box="[847,979,188,212]" italics="true" pageId="70" pageNumber="71">D. robustus</emphasis>
|
||
</taxonomicName>
|
||
N. sp Is THE BUrmEsE
|
||
<taxonomicName id="4C634D4DFFBBFF90FB48FF7CFA1CFF2B" box="[1236,1436,188,212]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="vestitus">
|
||
<emphasis id="B917EADCFFBBFF90FB48FF7CFA1CFF2B" box="[1236,1436,188,212]" italics="true" pageId="70" pageNumber="71">Dicharax vestitus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFF24B3FFFBBFF90FF01FF1FFECDFF07" author="BLaNfOrd" box="[157,333,223,248]" pageId="70" pageNumber="71" refString="Blanford, W. T. (1862) Contributions to Indian Malacology, No. III. Descriptions of new operculated land-shells from Pegu, Arakan and the Khasi Hills. Journal of the Asiatic Society of Bengal, 31, 135 - 144. Avaliable from: http: // www. biodiversitylibrary. org / item / 114386 # page / 181 / mode / 1 up (Accessed 10 Oct. 2017)" type="journal article" year="1862">BLaNfOrd, 1862</bibRefCitation>
|
||
) (maTErIaL ExamINEd: MOdITOUNG, NHMUK 1906.04.0 4.53,
|
||
<typeStatus id="54D8886CFFBBFF90FC63FF20FBE9FF07" box="[1023,1129,224,248]" pageId="70" pageNumber="71" type="holotype">HOLOTypE</typeStatus>
|
||
), THaT dIffErs frOm THE NEw spECIEs by HavE HavING smOOTHEr R1, sHOrTEr R2, aNd THE OUTEr pErIsTOmE Is NOT ExpaNdEd, bUT TUrNs IN THE dIrECTION Of THE INNEr pErIsTOmE. SEE UNdEr
|
||
<taxonomicName id="4C634D4DFFBBFF90FD8DFEE8FD72FEBF" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[529,754,296,320]" class="Gastropoda" family="Alycaeidae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="70" pageNumber="71" phylum="Mollusca" rank="species" species="okuboi" status="sp. nov.">
|
||
<emphasis id="B917EADCFFBBFF90FD8DFEE8FD72FEBF" box="[529,754,296,320]" italics="true" pageId="70" pageNumber="71">Metalycaeus okuboi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22457A7FFBBFF90FD65FEE7FCB8FEBF" box="[761,824,295,320]" pageId="70" pageNumber="71" rank="species">
|
||
<emphasis id="B917EADCFFBBFF90FD65FEE7FCB8FEBF" bold="true" box="[761,824,295,320]" pageId="70" pageNumber="71">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796545FFBBFF90FF5BFE8CFB9DFE77" pageId="70" pageNumber="71" type="distribution">
|
||
<paragraph id="8BDC36CEFFBBFF90FF5BFE8CFB9DFE77" blockId="70.[151,1437,151,392]" pageId="70" pageNumber="71">
|
||
<emphasis id="B917EADCFFBBFF90FF5BFE8CFEE0FE9A" bold="true" box="[199,352,332,357]" pageId="70" pageNumber="71">Distribution.</emphasis>
|
||
THIs NEw spECIEs Is kNOwN frOm TwO LOCaLITIEs IN THE
|
||
<collectingRegion id="49A7F82CFFBBFF90FC65FE8DFBC8FE9B" box="[1017,1096,333,356]" country="Sri Lanka" name="Eastern" pageId="70" pageNumber="71">EasTErN</collectingRegion>
|
||
parT Of THE CHINEsE
|
||
<collectingRegion id="49A7F82CFFBBFF90FADBFE8DFF7FFE77" country="China" name="Yunnan" pageId="70" pageNumber="71">YUNNaN PrOvINCE</collectingRegion>
|
||
. THE dIsTaNCE bETwEEN THE TwO sITEs Is apprOxImaTELy
|
||
<quantity id="4C9B9B2BFFBBFF90FCC4FEB0FC1EFE77" box="[856,926,367,392]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="70" pageNumber="71" unit="km" value="32.0">32 km</quantity>
|
||
(
|
||
<figureCitation id="13582A4BFFBBFF90FC37FEB0FB8FFE77" box="[939,1039,368,392]" captionStart="FIGURE 7" captionStartId="15.[151,250,1122,1144]" captionTargetBox="[151,1436,193,1101]" captionTargetId="figure@15.[151,1436,193,1101]" captionTargetPageId="15" captionText="FIGURE 7. Distribution of narrow range or single-site endemic Dicharax species. Filled circle. Dicharax tangmaiensis (Chen & Zhang, 2001). Filled triangle, top up. Dicharax (?) alticola Páll-Gergely & Hunyadi n. sp. Filled diamond. Dicharax (?) immaculatus Páll-Gergely n. sp. Empty diamond. Dicharax microcostatus Páll-Gergely n. sp., Dicharax micropolitus Páll- Gergely & Hunyadi, n. sp. and Dicharax bison Páll-Gergely & Hunyadi n. sp. Filled triangle, top down. Dicharax robustus Páll-Gergely & Hunyadi, n. sp. Empty triangle, top down. Dicharax draco Páll-Gergely & Hunyadi n. sp. Empty square. Dicharax imitator Páll-Gergely & Hunyadi n. sp. Empty triangle, top up. Dicharax (?) moellendorffi (Kobelt & Möllendorff, 1886). Filled square. Dicharax (?) ellipticus n. sp. Cross. Dicharax (?) abdoui Páll-Gergely n. sp. Empty circle shows the position of Tchen K’eou (Chengkou), which is the type locality of Alycaeus pentagonus Heude, 1886 (synonym of Alycaeus anthostoma Möllendorff, 1885), Dicharax diminutus (Heude, 1885), Dicharax (?) fargesianus (Heude, 1885), Dicharax planorbulus (Heude, 1885), Metalycaeus muciferus (Heude, 1885), Alycaeus dolomiticus Heude, 1890 (synonym of Alycaeus rathouisianus Heude, 1882), Dicharax (?) dolichodeiros (Heude, 1890), Metalycaeus (?) elevatus (Heude, 1886), Metalycaeus (?) expansus (Heude, 1890)." httpUri="https://zenodo.org/record/1009739/files/figure.png" pageId="70" pageNumber="71">FIGUrE 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |