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<document ID-DOI="http://doi.org/10.5281/zenodo.4762654" ID-GBIF-Dataset="564cead7-264a-4ee4-b5ee-15fd3ba81657" ID-GBIF-Taxon="182073493" ID-ISSN="1854-0392" ID-Zenodo-Dep="4762654" checkinTime="1620935982597" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Sivec, Ignac" docDate="2008" docId="03CA87B2FF95FFF7FEE22C25768CF9AE" docLanguage="en" docName="Illiesia.4.3.19-54.pdf" docOrigin="Illiesia 4 (3)" docStyle="DocumentStyle:1DEF1196F206C1C297CF0540A5C6768F.1:Illesia.2005-2008.journal_article" docStyleId="1DEF1196F206C1C297CF0540A5C6768F" docStyleName="Illesia.2005-2008.journal_article" docStyleVersion="1" docTitle="Neoperla spinaloba Stark &amp; Sivec 2008, sp. n." docType="treatment" docVersion="5" lastPageId="20" lastPageNumber="39" masterDocId="FFF3FFCAFF86FFE3FFAB2954746AFFD5" masterDocTitle="New Species And Records Of Neoperla (Plecoptera: Perlidae) From Vietnam" masterLastPageNumber="54" masterPageNumber="19" pageId="19" pageNumber="38" updateTime="1632024248002" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Species And Records Of Neoperla (Plecoptera: Perlidae) From Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>and Ignac Sivec &amp; Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivec, Ignac</mods:namePart>
<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia E-mail: isivec @ pms-lj. si</mods:affiliation>
<mods:nameIdentifier type="email">isivec@pms-lj.si</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-03-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>54</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4758856</mods:identifier>
<mods:identifier type="GBIF-Dataset">564cead7-264a-4ee4-b5ee-15fd3ba81657</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4758856</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4762654" ID-GBIF-Taxon="182073493" ID-Zenodo-Dep="4762654" LSID="urn:lsid:plazi:treatment:03CA87B2FF95FFF7FEE22C25768CF9AE" httpUri="http://treatment.plazi.org/id/03CA87B2FF95FFF7FEE22C25768CF9AE" lastPageId="20" lastPageNumber="39" pageId="19" pageNumber="38">
<subSubSection box="[329,630,1393,1417]" pageId="19" pageNumber="38" type="nomenclature">
<paragraph blockId="19.[329,630,1393,1417]" box="[329,630,1393,1417]" pageId="19" pageNumber="38">
<heading bold="true" box="[329,630,1393,1417]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="38" reason="2">
<emphasis bold="true" box="[329,630,1393,1417]" pageId="19" pageNumber="38">
<taxonomicName authority="Stark &amp; Sivec, 2008" authorityName="Stark &amp; Sivec" authorityYear="2008" box="[329,554,1393,1417]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="spinaloba" status="sp. n.">
<emphasis bold="true" box="[329,554,1393,1417]" italics="true" pageId="19" pageNumber="38">Neoperla spinaloba</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[568,630,1393,1417]" pageId="19" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[361,598,1428,1452]" pageId="19" pageNumber="38" type="description">
<paragraph blockId="19.[361,598,1428,1452]" box="[361,598,1428,1452]" pageId="19" pageNumber="38">
(
<figureCitation box="[369,492,1428,1452]" captionStart="Figs" captionStartId="19.[184,234,1255,1279]" captionTargetBox="[252,1405,417,1205]" captionTargetId="figure-383@19.[579,947,416,931]" captionTargetPageId="19" captionText="Figs. 62‐66. Neoperla spinaloba. 62. Head and pronotum, 63. Male terminalia, 64. Aedeagus, 65. Female terminalia, 66. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758886" httpUri="https://zenodo.org/record/4758886/files/figure.png" pageId="19" pageNumber="38">Figs. 62‐66</figureCitation>
,
<figureCitation box="[503,591,1428,1452]" captionStart="Figs" captionStartId="30.[184,234,1876,1900]" captionTargetBox="[184,1404,347,1770]" captionTargetPageId="30" captionText="Figs. 112‐117. Egg micrographs. 112‐114. N. song. 112. Egg, 113. Collar, 114. Lid. 115‐117. N. spinaloba. 115. Egg, 116. Collar, 117. Lid." figureDoi="http://doi.org/10.5281/zenodo.4758910" httpUri="https://zenodo.org/record/4758910/files/figure.png" pageId="19" pageNumber="38">115‐117</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="38" type="materials_examined">
<paragraph blockId="19.[184,776,1497,1971]" pageId="19" pageNumber="38">
<emphasis bold="true" box="[184,422,1497,1521]" pageId="19" pageNumber="38">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111475364" collectingDate="1995-05-08" collectingDateMax="1999-05-12" collectingDateMin="1995-04-22" collectionCode="ROM, IEBR, BPS" collectorName="D. Currie &amp; B. Hubley &amp; J. Swann" country="Vietnam" county="Sapa" elevation="1600" latitude="22.35" location="Sapa" longLatPrecision="1260" longitude="103.76667" municipality="Sapa" pageId="19" pageNumber="38" specimenCode="ROM 956022" specimenCount="26" specimenCount-female="13" specimenCount-male="13" stateProvince="Lao Cai" typeStatus="holotype">
<typeStatus box="[438,546,1497,1521]" pageId="19" pageNumber="38">Holotype</typeStatus>
<specimenCount box="[561,580,1497,1521]" pageId="19" pageNumber="38" type="male">♂</specimenCount>
and
<specimenCount box="[656,704,1497,1521]" pageId="19" pageNumber="38" type="male">8 ♂</specimenCount>
,
<specimenCount box="[727,775,1497,1521]" pageId="19" pageNumber="38" type="female">7 ♀</specimenCount>
<typeStatus box="[184,297,1533,1557]" pageId="19" pageNumber="38">paratypes</typeStatus>
from
<collectingCountry box="[390,490,1532,1556]" name="Vietnam" pageId="19" pageNumber="38">Vietnam</collectingCountry>
,
<collectingRegion box="[513,615,1532,1556]" country="Vietnam" name="Lao Cai" pageId="19" pageNumber="38">Lao Cai</collectingRegion>
,
<collectingCounty box="[638,695,1532,1556]" pageId="19" pageNumber="38">Sapa</collectingCounty>
, large waterfall on road from
<collectingMunicipality box="[444,499,1567,1591]" pageId="19" pageNumber="38">Sapa</collectingMunicipality>
to
<collectingRegion box="[531,635,1567,1591]" country="Vietnam" name="Lai Chau" pageId="19" pageNumber="38">Lai Chau</collectingRegion>
,
<date box="[644,771,1566,1591]" pageId="19" pageNumber="38" value="1995-05-08">
<collectingDate box="[644,771,1566,1591]" pageId="19" pageNumber="38" value="1995-05-08">8 May 1995</collectingDate>
</date>
,
<collectorName box="[184,305,1601,1625]" pageId="19" pageNumber="38">D. Currie</collectorName>
,
<collectorName box="[329,458,1601,1625]" pageId="19" pageNumber="38">B. Hubley</collectorName>
,
<collectorName box="[480,594,1601,1625]" pageId="19" pageNumber="38">J. Swann</collectorName>
,
<specimenCode box="[617,775,1601,1625]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="19" pageNumber="38" type="Museum">ROM 956022</specimenCode>
(
<typeStatus box="[192,301,1636,1660]" pageId="19" pageNumber="38">Holotype</typeStatus>
and
<specimenCount box="[374,420,1635,1660]" pageId="19" pageNumber="38" type="male">4 ♂</specimenCount>
,
<specimenCount box="[442,489,1635,1660]" pageId="19" pageNumber="38" type="female">6 ♀</specimenCount>
<typeStatus box="[504,617,1636,1660]" pageId="19" pageNumber="38">paratypes</typeStatus>
<collectionCode box="[632,696,1636,1660]" collectionName="Canada Entomology Department, Royal Ontario Museum" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="19" pageNumber="38" type="Museum">ROM</collectionCode>
; other
<typeStatus box="[184,297,1671,1695]" pageId="19" pageNumber="38">paratypes</typeStatus>
<collectionCode box="[325,385,1671,1694]" collectionName="IEBR" pageId="19" pageNumber="38">IEBR</collectionCode>
,
<collectionCode box="[416,465,1670,1694]" collectionName="BPS" country="USA" lsid="urn:lsid:biocol.org:col:14922" name="California Department of Food and Agriculture" pageId="19" pageNumber="38" type="Herbarium">BPS</collectionCode>
).
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111475346" collectingDate="1995-04-22" collectingDateMax="1999-05-12" collectingDateMin="1995-04-22" collectionCode="ZMB, ROM" collectorName="B. Hubley" country="Vietnam" county="Sapa" elevation="1600" latitude="22.35" location="Sapa" longLatPrecision="1260" longitude="103.76667" municipality="Sapa" pageId="19" pageNumber="38" specimenCode="ROM 992004, ROM 992006" specimenCount="12" specimenCount-female="13" specimenCount-male="12" stateProvince="Lao Cai" typeStatus="paratype">
Additional
<typeStatus box="[655,771,1671,1695]" pageId="19" pageNumber="38">paratypes</typeStatus>
:
<collectingCountry box="[184,285,1705,1729]" name="Vietnam" pageId="19" pageNumber="38">Vietnam</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:03CA87B2FF95FFF7FEE22C25768CF9AE:8EBC607FFF95FFF0FE832FFD750BF914" box="[296,353,1705,1729]" country="Vietnam" county="Sapa" latitude="22.35" longLatPrecision="1260" longitude="103.76667" municipality="Sapa" name="Sapa" pageId="19" pageNumber="38" stateProvince="Lao Cai">Sapa</location>
,
<location LSID="urn:lsid:plazi:treatment:03CA87B2FF95FFF7FEE22C25768CF9AE:8EBC607FFF95FFF0FEC72FFD760AF914" box="[364,608,1705,1729]" country="Vietnam" county="Sapa" latitude="22.35" longLatPrecision="1260" longitude="103.76667" municipality="Sapa" name="Fan Si Pan Mountain" pageId="19" pageNumber="38" stateProvince="Lao Cai">Fan Si Pan Mountain</location>
,
<quantity box="[620,703,1705,1729]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="19" pageNumber="38" unit="m" value="1600.0">
<elevation box="[620,703,1705,1729]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="19" pageNumber="38" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<date pageId="19" pageNumber="38" value="1995-04-22" valueMax="1995-04-29" valueMin="1995-04-22">
<collectingDate pageId="19" pageNumber="38" value="1995-04-22" valueMax="1995-04-29" valueMin="1995-04-22">22‐29 April 1995</collectingDate>
</date>
, L.F.
<specimenCount box="[376,418,1739,1763]" pageId="19" pageNumber="38" type="male">1 ♂</specimenCount>
(
<collectionCode box="[437,499,1739,1763]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="19" pageNumber="38" type="Museum">ZMB</collectionCode>
).
<collectingRegion box="[520,614,1739,1763]" country="Vietnam" name="Lao Cai" pageId="19" pageNumber="38">Lao Cai</collectingRegion>
,
<quantity box="[628,701,1739,1763]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="38" unit="km" value="12.0">
<locationDeviation box="[628,701,1739,1763]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="38" unit="km" value="12.0">12 km</locationDeviation>
</quantity>
along road from
<location LSID="urn:lsid:plazi:treatment:03CA87B2FF95FFF7FEE22C25768CF9AE:8EBC607FFF95FFF0FE9A2FBA7502F8D3" box="[305,360,1774,1798]" country="Vietnam" county="Sapa" latitude="22.35" longLatPrecision="1260" longitude="103.76667" municipality="Sapa" name="Sapa" pageId="19" pageNumber="38" stateProvince="Lao Cai">Sapa</location>
to
<collectingRegion box="[395,501,1774,1798]" country="Vietnam" name="Lai Chau" pageId="19" pageNumber="38">Lai Chau</collectingRegion>
,
<geoCoordinate box="[512,616,1773,1798]" degrees="22" direction="north" minutes="21" orientation="latitude" pageId="19" pageNumber="38" precision="925" value="22.35">
22
<emphasis bold="true" box="[537,547,1773,1797]" pageId="19" pageNumber="38">°</emphasis>
21’ N
</geoCoordinate>
,
<geoCoordinate box="[627,739,1773,1798]" degrees="103" direction="east" minutes="46" orientation="longitude" pageId="19" pageNumber="38" precision="925" value="103.76667">
103
<emphasis bold="true" box="[666,676,1773,1797]" pageId="19" pageNumber="38">°</emphasis>
46’ E
</geoCoordinate>
,
<date pageId="19" pageNumber="38" value="1999-04-29" valueMax="1999-05-12" valueMin="1999-04-29">
<collectingDate pageId="19" pageNumber="38" value="1999-04-29" valueMax="1999-05-12" valueMin="1999-04-29">29 April‐12 May 1999</collectingDate>
</date>
,
<collectorName box="[424,546,1808,1832]" pageId="19" pageNumber="38">B. Hubley</collectorName>
,
<specimenCode box="[562,716,1809,1833]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="19" pageNumber="38" type="Museum">ROM 992004</specimenCode>
,
<specimenCount box="[731,775,1808,1832]" pageId="19" pageNumber="38" type="male">6 ♂</specimenCount>
(pinned,
<collectionCode box="[286,352,1843,1867]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="19" pageNumber="38" type="Museum">ROM</collectionCode>
). Same site but
<date box="[527,771,1843,1867]" pageId="19" pageNumber="38" value="1999-04-30" valueMax="1999-05-12" valueMin="1999-04-30">
<collectingDate box="[527,771,1843,1867]" pageId="19" pageNumber="38" value="1999-04-30" valueMax="1999-05-12" valueMin="1999-04-30">30 April‐12 May 1999</collectingDate>
</date>
,
<collectorName box="[184,301,1877,1901]" pageId="19" pageNumber="38">B. Hubley</collectorName>
,
<specimenCode box="[311,460,1877,1902]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="19" pageNumber="38" type="Museum">ROM 992006</specimenCode>
,
<specimenCount box="[470,509,1877,1901]" pageId="19" pageNumber="38" type="male">5 ♂</specimenCount>
(pinned,
<collectionCode box="[619,685,1878,1902]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="19" pageNumber="38" type="Museum">ROM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="39" pageId="19" pageNumber="38" type="description">
<paragraph blockId="19.[184,776,1497,1971]" lastBlockId="19.[812,1404,1393,1970]" pageId="19" pageNumber="38">
<emphasis bold="true" box="[184,356,1912,1936]" pageId="19" pageNumber="38">Adult habitus.</emphasis>
Biocellate. Head with dark pigment extending from center of occiput, over ocelli and forward to M‐ line; anterior to M‐ line a dark triangular area occurs (
<figureCitation box="[1093,1177,1427,1451]" captionStart="Figs" captionStartId="19.[184,234,1255,1279]" captionTargetBox="[252,1405,417,1205]" captionTargetId="figure-383@19.[579,947,416,931]" captionTargetPageId="19" captionText="Figs. 62‐66. Neoperla spinaloba. 62. Head and pronotum, 63. Male terminalia, 64. Aedeagus, 65. Female terminalia, 66. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758886" httpUri="https://zenodo.org/record/4758886/files/figure.png" pageId="19" pageNumber="38">Fig. 62</figureCitation>
). Pronotum brown with darker rugosities.
</paragraph>
<paragraph blockId="19.[812,1404,1393,1970]" pageId="19" pageNumber="38">
<emphasis bold="true" box="[812,879,1497,1521]" pageId="19" pageNumber="38">Male.</emphasis>
Forewing length
<quantity box="[1077,1192,1497,1521]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="19" pageNumber="38" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13‐14 mm</quantity>
. Process of tergum 7 a raised plateau with posterior corners slightly projecting. Process of tergum 8 an erect, curved tongue shaped structure. Tergum 9 without sensilla patches (
<figureCitation box="[923,1011,1635,1659]" captionStart="Figs" captionStartId="19.[184,234,1255,1279]" captionTargetBox="[252,1405,417,1205]" captionTargetId="figure-383@19.[579,947,416,931]" captionTargetPageId="19" captionText="Figs. 62‐66. Neoperla spinaloba. 62. Head and pronotum, 63. Male terminalia, 64. Aedeagus, 65. Female terminalia, 66. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758886" httpUri="https://zenodo.org/record/4758886/files/figure.png" pageId="19" pageNumber="38">Fig. 63</figureCitation>
). Hemitergal lobes slender and curved at midlength. Aedeagal tube plump, poorly sclerotized, and bearing a ventroapical pair of prominent spiny lobes (
<figureCitation box="[1120,1211,1739,1763]" captionStart="Figs" captionStartId="19.[184,234,1255,1279]" captionTargetBox="[252,1405,417,1205]" captionTargetId="figure-383@19.[579,947,416,931]" captionTargetPageId="19" captionText="Figs. 62‐66. Neoperla spinaloba. 62. Head and pronotum, 63. Male terminalia, 64. Aedeagus, 65. Female terminalia, 66. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758886" httpUri="https://zenodo.org/record/4758886/files/figure.png" pageId="19" pageNumber="38">Fig. 64</figureCitation>
). Aedeagal sac cylindrical and slightly longer than tube; sac armed, except basally, with small spines and two patches of larger spines near apex.
</paragraph>
<paragraph blockId="19.[812,1404,1393,1970]" lastBlockId="20.[184,776,288,1660]" lastPageId="20" lastPageNumber="39" pageId="19" pageNumber="38">
<emphasis bold="true" box="[812,904,1877,1901]" pageId="19" pageNumber="38">Female.</emphasis>
Forewing length 16.5‐17.5 mm. Subgenital plate a small truncate tab (
<figureCitation box="[1106,1184,1912,1936]" captionStart="Figs" captionStartId="19.[184,234,1255,1279]" captionTargetBox="[252,1405,417,1205]" captionTargetId="figure-383@19.[579,947,416,931]" captionTargetPageId="19" captionText="Figs. 62‐66. Neoperla spinaloba. 62. Head and pronotum, 63. Male terminalia, 64. Aedeagus, 65. Female terminalia, 66. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758886" httpUri="https://zenodo.org/record/4758886/files/figure.png" pageId="19" pageNumber="38">Fig. 65</figureCitation>
). Vagina somewhat T‐ shaped; triangular shaped sclerite underlies spermathecal base (
<figureCitation box="[402,479,288,312]" captionStart="Figs" captionStartId="19.[184,234,1255,1279]" captionTargetBox="[252,1405,417,1205]" captionTargetId="figure-383@19.[579,947,416,931]" captionTargetPageId="19" captionText="Figs. 62‐66. Neoperla spinaloba. 62. Head and pronotum, 63. Male terminalia, 64. Aedeagus, 65. Female terminalia, 66. Vagina and spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4758886" httpUri="https://zenodo.org/record/4758886/files/figure.png" pageId="20" pageNumber="39">Fig. 66</figureCitation>
). Spermathecal stalk short, slender, spermatheca sausage shaped and curled beyond midlength; accessory gland near tip expanded apically into a large sausage shaped structure.
</paragraph>
<paragraph blockId="20.[184,776,288,1660]" pageId="20" pageNumber="39">
<emphasis bold="true" box="[184,235,461,485]" pageId="20" pageNumber="39">Egg.</emphasis>
Barrel shaped. Length ca. 0.37 mm, width ca. 0.23 mm. Collar sessile with smooth side rim; area surrounding collar with irregular, scale‐ like impressions attached to rim and to primary striae (
<figureCitation box="[192,347,599,623]" captionStart="Figs" captionStartId="30.[184,234,1876,1900]" captionTargetBox="[184,1404,347,1770]" captionTargetPageId="30" captionText="Figs. 112‐117. Egg micrographs. 112‐114. N. song. 112. Egg, 113. Collar, 114. Lid. 115‐117. N. spinaloba. 115. Egg, 116. Collar, 117. Lid." figureDoi="http://doi.org/10.5281/zenodo.4758910" httpUri="https://zenodo.org/record/4758910/files/figure.png" pageId="20" pageNumber="39">Figs. 115‐117</figureCitation>
). Chorion striate with striae closely packed; each pair of primary striae enclose a pair of secondary striae; sulci reduced, visible near lid where a single row of shallow pits occurs in each sulcus. Lid covered with small, shallow pits; follicle cell impressions absent. Micropyles subequatorial, sperm guides absent, but some micropyles associated with irregular, abnormal appearing striae.
</paragraph>
<paragraph blockId="20.[184,776,288,1660]" box="[184,385,875,899]" pageId="20" pageNumber="39">
<emphasis bold="true" box="[184,257,875,899]" pageId="20" pageNumber="39">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="39" type="etymology">
<paragraph blockId="20.[184,776,288,1660]" pageId="20" pageNumber="39">
<emphasis bold="true" box="[184,318,910,934]" pageId="20" pageNumber="39">Etymology.</emphasis>
The species name refers to the spiny aedeagal lobes.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="39" type="diagnosis">
<paragraph blockId="20.[184,776,288,1660]" pageId="20" pageNumber="39">
<emphasis bold="true" box="[184,309,979,1003]" pageId="20" pageNumber="39">Diagnosis.</emphasis>
This species is very similar in color pattern to
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[304,430,1014,1038]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis box="[304,430,1014,1038]" italics="true" pageId="20" pageNumber="39">N. nebulosa</emphasis>
</taxonomicName>
and the two species co‐ occur. The aedeagus is similar to that of
<taxonomicName authority="Wu &amp; Claassen" authorityName="Wu &amp; Claassen" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis box="[573,699,1049,1073]" italics="true" pageId="20" pageNumber="39">N. quadrata</emphasis>
Wu &amp; Claassen
</taxonomicName>
,
<taxonomicName authority="Wu" authorityName="Wu" box="[297,459,1083,1107]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="lushana">
<emphasis box="[297,412,1083,1107]" italics="true" pageId="20" pageNumber="39">N. lushana</emphasis>
Wu
</taxonomicName>
(
<emphasis box="[474,533,1084,1107]" italics="true" pageId="20" pageNumber="39">sensu</emphasis>
<bibRefCitation author="Zwick, P. &amp; I. Sivec" box="[539,763,1083,1107]" pageId="20" pageNumber="39" pagination="59 - 138" refId="ref17138" refString="Zwick, P. &amp; I. Sivec. 1980. Beitrage zur Kenntnis der Plecoptera des Himalaja. Entomologica Basiliensia, 5: 59 - 138." type="journal article" year="1980">Zwick &amp; Sivec 1980</bibRefCitation>
), and
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[234,357,1118,1142]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis box="[234,357,1118,1142]" italics="true" pageId="20" pageNumber="39">N. nebulosa</emphasis>
</taxonomicName>
(described above). The latter species is easily separated on the basis of the short aedeagal sac oriented at an angle from the tube but separation from the other species requires careful examination of aedeagal armature. The aedeagal sac of
<taxonomicName box="[661,775,1256,1280]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="lushana">
<emphasis box="[661,775,1256,1280]" italics="true" pageId="20" pageNumber="39">N. lushana</emphasis>
</taxonomicName>
has the ventromesal row of spines reduced to a few organized in a short row and these are intersected near the apex by a relatively straight transverse row of similar large spines (
<bibRefCitation author="Zwick, P. &amp; I. Sivec" box="[447,673,1394,1418]" pageId="20" pageNumber="39" pagination="59 - 138" refId="ref17138" refString="Zwick, P. &amp; I. Sivec. 1980. Beitrage zur Kenntnis der Plecoptera des Himalaja. Entomologica Basiliensia, 5: 59 - 138." type="journal article" year="1980">Zwick &amp; Sivec 1980</bibRefCitation>
). Both
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="spinaloba">
<emphasis italics="true" pageId="20" pageNumber="39">N. spinaloba</emphasis>
</taxonomicName>
and
<taxonomicName box="[368,507,1429,1453]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis box="[368,507,1429,1453]" italics="true" pageId="20" pageNumber="39">N. quadrata</emphasis>
</taxonomicName>
have a long linear ventromesal patch of slightly enlarged spines; in the latter species there are no additional enlarged spines on the sac apex or subapex (Stark unpublished) whereas in
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[337,479,1567,1591]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="20" pageNumber="39" phylum="Arthropoda" rank="species" species="spinaloba">
<emphasis box="[337,479,1567,1591]" italics="true" pageId="20" pageNumber="39">N. spinaloba</emphasis>
</taxonomicName>
a prominent subapical ventromesal patch of ca. 20 relatively large spines and a smaller patch of dorsomesal spines occurs.
</paragraph>
</subSubSection>
</treatment>
</document>
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