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<mods:title id="3854FFD1A52FA2E187A1D549B12B599D">Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena</mods:title>
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<mods:namePart id="FA4CC80D4AAFF547655F6E3B956989BC">Souza, Leonardo Santos De</mods:namePart>
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<mods:namePart id="CE4404FA6133464565190C0D2A5EEE79">Araújo, Tarciso Almeida De</mods:namePart>
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<emphasis id="B917EA8EBA77DE24FF4FFF407134FF2D" bold="true" box="[151,374,151,178]" italics="true" pageId="25" pageNumber="476">Aclis marplatensis</emphasis>
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<figureCitation id="13582A19BA77DE24FF4FFF6C70A5FF4A" box="[151,231,187,213]" captionStart="FIGURE 13" captionStartId="24.[151,250,1908,1932]" captionTargetBox="[206,1382,181,1884]" captionTargetId="figure-23@24.[206,1382,181,1884]" captionTargetPageId="24" captionText="FIGURE 13. Aclis marplatensis (Castellanos &amp; Landoni, 1984). A, B. MZSP 85401. C. MZSP 88368. D. MNRJ 15963. E–I. MNRJ 15964. A–C, E. Shell in frontal view; rectangle in E indicates detail in H. D. Shell in lateral view. F. Detail of body whorl in frontal view. G. Detail of apical whorls. H, I. Detail of teleoconch surface; rectangle in H indicates detail in I. Scale bars: A–E. 1 mm; F. 500 μm; G. 100 μm; H, I. 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10954455" httpUri="https://zenodo.org/record/10954455/files/figure.png" pageId="25" pageNumber="476">Fig. 13</figureCitation>
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<emphasis id="B917EA8EBA77DE24FF4FFED771EAFE87" box="[151,424,256,280]" italics="true" pageId="25" pageNumber="476">Turritellopsis marplatensis</emphasis>
<bibRefCitation id="EFF24B6DBA77DE24FE74FED77346FE87" author="Castellanos, Z. J. A. &amp; Landoni, N." box="[428,772,256,280]" pageId="25" pageNumber="476" pagination="291 - 299" refId="ref24249" refString="Castellanos, Z. J. A. &amp; Landoni, N. (1984) Nueva contribuicion al conocimiento de los micromoluscos de la plataforma continental argentina. Revista del Museo de La Plata, Nueva Serie, Seccion Zoologia, 13 (149), 291 - 299." type="journal article" year="1984">Castellanos &amp; Landoni, 1984: 293</bibRefCitation>
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<figureCitation id="13582A19BA77DE24FCD5FED7732BFE87" box="[781,873,256,280]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="8.[151,250,1858,1882]" captionStartId-1="9.[151,250,1891,1915]" captionStartId-2="10.[151,250,1646,1670]" captionTargetBox-0="[151,1435,309,1835]" captionTargetBox-1="[151,1435,181,1867]" captionTargetBox-2="[189,1397,957,1623]" captionTargetId-0="figure-96@8.[151,1435,309,1835]" captionTargetId-1="figure-17@9.[151,1435,181,1867]" captionTargetId-2="figure-402@10.[189,1397,957,1623]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 2. Aclis sarissa Watson, 1881. A–C, E. Lectotype, NHMUK 1887.2.9.1576, herein designated; D. Paralectotype, NHMUK 1887.2.9.1576; F–I. Paralectotype, NMW 1955.158.26255; J–K. Holotype of Aclis carolinensis, USNM 83743. A, D, F, J. Shell in frontal view; B, G, K. Shell in lateral view. C, H. Shell in dorsal view. E, I. Body whorl in frontal view. Scale bars: A–D, F–H, I–J. 1 mm; E, I. 500 μm. Credits: A–E. NHMUK photographic unit; F–I. NMW." captionText-1="FIGURE 3. Aclis sarissa Watson, 1881. A–B, H–K. MORG 49199. C–D. CZUFS GAS-00102. E–G. CZUFS GAS-00104. A, C, E. Shell in frontal view; square in A indicates detail in I. B, D, F. Shell in lateral view. G–H. Detail of the body whorl in frontal view. I–J. Detail of teleoconch surface; rectangle in I indicates detail in J. K. Detail of apical whorls; arrow indicates the transition protoconch–teleoconch. Scale bars: A–F. 1 mm; G–H. 500 μm; I–J. 50 μm; K. 200 μm." captionText-2="FIGURE 4. Aclis watsoni Barros, Lima &amp; Francisco, 2007. A–B. CZUFS GAS-00105. A. C. Shell in frontal view. B. Shell in lateral view. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10954424" figureDoi-1="http://doi.org/10.5281/zenodo.10954428" figureDoi-2="http://doi.org/10.5281/zenodo.10954432" httpUri-0="https://zenodo.org/record/10954424/files/figure.png" httpUri-1="https://zenodo.org/record/10954428/files/figure.png" httpUri-2="https://zenodo.org/record/10954432/files/figure.png" pageId="25" pageNumber="476">Figs. 2-4</figureCitation>
: 
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<bibRefCitation id="EFF24B6DBA77DE24FCA8FED77423FE88" author="Absalao, R. S. &amp; Pimenta, A. D. &amp; Costa, P. M. S." box="[880,1121,255,280]" pageId="25" pageNumber="476" pagination="57 - 68" refId="ref23241" refString="Absalao, R. S., Pimenta, A. D. &amp; Costa, P. M. S. (1996) Novas ocorrencias de gastropodes no litoral do Rio de Janeiro (Brasil). Neritica, 10, 57 - 68." type="journal article" year="1996">
Absalão 
<emphasis id="B917EA8EBA77DE24FC10FED773B8FE87" box="[968,1018,256,280]" italics="true" pageId="25" pageNumber="476">et al.</emphasis>
(1996: 59
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, 
<figureCitation id="13582A19BA77DE24FBB1FED7748BFE87" box="[1129,1225,256,280]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="8.[151,250,1858,1882]" captionStartId-1="9.[151,250,1891,1915]" captionTargetBox-0="[151,1435,309,1835]" captionTargetBox-1="[151,1435,181,1867]" captionTargetId-0="figure-96@8.[151,1435,309,1835]" captionTargetId-1="figure-17@9.[151,1435,181,1867]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 2. Aclis sarissa Watson, 1881. A–C, E. Lectotype, NHMUK 1887.2.9.1576, herein designated; D. Paralectotype, NHMUK 1887.2.9.1576; F–I. Paralectotype, NMW 1955.158.26255; J–K. Holotype of Aclis carolinensis, USNM 83743. A, D, F, J. Shell in frontal view; B, G, K. Shell in lateral view. C, H. Shell in dorsal view. E, I. Body whorl in frontal view. Scale bars: A–D, F–H, I–J. 1 mm; E, I. 500 μm. Credits: A–E. NHMUK photographic unit; F–I. NMW." captionText-1="FIGURE 3. Aclis sarissa Watson, 1881. A–B, H–K. MORG 49199. C–D. CZUFS GAS-00102. E–G. CZUFS GAS-00104. A, C, E. Shell in frontal view; square in A indicates detail in I. B, D, F. Shell in lateral view. G–H. Detail of the body whorl in frontal view. I–J. Detail of teleoconch surface; rectangle in I indicates detail in J. K. Detail of apical whorls; arrow indicates the transition protoconch–teleoconch. Scale bars: A–F. 1 mm; G–H. 500 μm; I–J. 50 μm; K. 200 μm." figureDoi-0="http://doi.org/10.5281/zenodo.10954424" figureDoi-1="http://doi.org/10.5281/zenodo.10954428" httpUri-0="https://zenodo.org/record/10954424/files/figure.png" httpUri-1="https://zenodo.org/record/10954428/files/figure.png" pageId="25" pageNumber="476">Figs. 2–3</figureCitation>
); 
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. pl. 12, Fig. 95); 
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Absalão 
<emphasis id="B917EA8EBA77DE24FDEAFEC8721CFEA9" box="[562,606,286,310]" italics="true" pageId="25" pageNumber="476">et al</emphasis>
. (2006: 235)
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; 
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<bibRefCitation id="EFF24B6DBA77DE24FD37FEC973D1FEA9" author="Rios, E." box="[751,915,286,310]" pageId="25" pageNumber="476" refId="ref27447" refString="Rios, E. (2009) Compendium of Brazilian Seashells. Evangraf, Rio Grande, 668 pp." type="book" year="2009">Rios (2009: 114</bibRefCitation>
</treatmentCitation>
, Fig. 278); 
<treatmentCitation id="0AC2108DBA77DE24FBD0FEC9754DFEA9" author="Signorelli, J. H. &amp; Urteaga, D. &amp; Teso, V." box="[1032,1295,286,310]" page="65" pageId="25" pageNumber="476" year="2015">
<bibRefCitation id="EFF24B6DBA77DE24FBD0FEC9754DFEA9" author="Signorelli, J. H. &amp; Urteaga, D. &amp; Teso, V." box="[1032,1295,286,310]" pageId="25" pageNumber="476" pagination="45 - 69" refId="ref27575" refString="Signorelli, J. H., Urteaga, D. &amp; Teso, V. (2015) Zulma Ageitos de Castellanos: Publications and status of described taxa. Zootaxa, 4034 (1), 45 - 69. https: // doi. org / 10.11646 / zootaxa. 4034.1.2" type="journal article" year="2015">
Signorelli 
<emphasis id="B917EA8EBA77DE24FBAAFEC874DDFEA9" box="[1138,1183,286,310]" italics="true" pageId="25" pageNumber="476">et al</emphasis>
. (2015: 65
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, 
<figureCitation id="13582A19BA77DE24FAC2FEC97515FEA9" box="[1306,1367,286,310]" captionStart="FIGURE 6" captionStartId="13.[151,250,1785,1809]" captionTargetBox="[151,1435,993,1762]" captionTargetId="figure-349@13.[151,1435,993,1762]" captionTargetPageId="13" captionText="FIGURE 6. Aclis sp. 1: MNRJ 33505. A. Shell in frontal view. B. Apical whorls: double arrows indicate microscopic and sigmoid axial line at the protoconch surface, single bottom arrow indicate microscopic spiral striae at the teleoconch surface. C. Body whorl. Scale bars: A. 1 mm; B. 200 μm; C. 500 μm." figureDoi="http://doi.org/10.5281/zenodo.10954438" httpUri="https://zenodo.org/record/10954438/files/figure.png" pageId="25" pageNumber="476">Fig. 6</figureCitation>
).
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<emphasis id="B917EA8EBA77DE24FF4FFEB27105FEE1" bold="true" box="[151,327,356,383]" pageId="25" pageNumber="476">Type material.</emphasis>
(see 
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Signorelli 
<emphasis id="B917EA8EBA77DE24FE27FEB27278FEE1" box="[511,570,356,382]" italics="true" pageId="25" pageNumber="476">et al.</emphasis>
2015
</bibRefCitation>
for notes and illustrations of the type material). 
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<typeStatus id="54D8883EBA77DE24FB76FEB37559FEE1" box="[1198,1307,356,382]" pageId="25" pageNumber="476" type="holotype">Holotype</typeStatus>
: 
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. 
<materialsCitation id="3B0B3CC1BA77DE24FF33FE5E73FFFE3C" box="[235,957,392,419]" country="Argentina" location="Argentina" pageId="25" pageNumber="476" specimenCode="MACN-In 28294-1" specimenCount="1" typeStatus="paratype">
<typeStatus id="54D8883EBA77DE24FF33FE5E711EFE3C" box="[235,348,393,419]" pageId="25" pageNumber="476" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="F374760CBA77DE24FEBDFE5F719BFE3D" box="[357,473,392,418]" name="Argentina" pageId="25" pageNumber="476">Argentina</collectingCountry>
: from the type locality: 
<specimenCode id="DBC59EE7BA77DE24FD3CFE5E73FFFE3C" box="[740,957,393,419]" collectionCode="MACN-In" pageId="25" pageNumber="476">MACN-In 28294-1</specimenCode>
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[1].
</paragraph>
<paragraph id="8BDC369CBA77DE24FF1FFE7A7579FE59" blockId="25.[151,1437,356,2039]" box="[199,1339,428,455]" pageId="25" pageNumber="476">
<materialsCitation id="3B0B3CC1BA77DE24FF1FFE7A74B2FE59" box="[199,1264,428,455]" collectorName="N. Landoni" country="Argentina" location="Mar del Plata" pageId="25" pageNumber="476" specimenCount="1" stateProvince="Buenos Aires" typeStatus="holotype">
<emphasis id="B917EA8EBA77DE24FF1FFE7A7124FE59" bold="true" box="[199,358,428,455]" pageId="25" pageNumber="476">
<typeStatus id="54D8883EBA77DE24FF1FFE7A7140FE58" box="[199,258,429,455]" pageId="25" pageNumber="476">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F374760CBA77DE24FEB3FE7B719DFE59" box="[363,479,428,454]" name="Argentina" pageId="25" pageNumber="476">Argentina</collectingCountry>
: 
<collectingRegion id="49A7F87EBA77DE24FE31FE7A72AEFE59" box="[489,748,428,455]" country="Argentina" name="Buenos Aires" pageId="25" pageNumber="476">Buenos Aires province</collectingRegion>
: 
<location id="8EBC6047BA77DE24FD2FFE7A73D3FE59" LSID="urn:lsid:plazi:treatment:03CA878ABA77DE27FF4FFF407501FD86:8EBC6047BA77DE24FD2FFE7A73D3FE59" box="[759,913,428,455]" country="Argentina" name="Mar del Plata" pageId="25" pageNumber="476" stateProvince="Buenos Aires">Mar del Plata</location>
, Chapadmalal; 
<collectorName id="2696534ABA77DE24FBE6FE7A74FFFE59" box="[1086,1213,428,455]" pageId="25" pageNumber="476">N. Landoni</collectorName>
leg.
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, 1976.
</paragraph>
<paragraph id="8BDC369CBA77DE24FF1FFE077482FBD0" blockId="25.[151,1437,356,2039]" pageId="25" pageNumber="476">
<emphasis id="B917EA8EBA77DE24FF1FFE0771F6FE75" bold="true" box="[199,436,464,490]" pageId="25" pageNumber="476">Material examined.</emphasis>
<materialsCitation id="3B0B3CC1BA77DE24FE67FE0775DEFE74" box="[447,1436,464,491]" collectionCode="IBUFRJ" collectorName="Royal Charlote Bank Revizee Central V" country="Brazil" location="Bahia state" pageId="25" pageNumber="476" specimenCode="IBUFRJ 14050" specimenCount="1" stateProvince="Bahia">
<collectingCountry id="F374760CBA77DE24FE67FE07724AFE75" box="[447,520,464,490]" name="Brazil" pageId="25" pageNumber="476">Brazil</collectingCountry>
: 
<collectingRegion id="49A7F87EBA77DE24FDCEFE0772D8FE74" box="[534,666,464,491]" country="Brazil" name="Bahia" pageId="25" pageNumber="476">Bahia state</collectingRegion>
: 
<collectorName id="2696534ABA77DE24FD7FFE077439FE74" box="[679,1147,464,491]" pageId="25" pageNumber="476">Royal Charlote Bank Revizee Central V</collectorName>
sta. 07r: 
<specimenCode id="DBC59EE7BA77DE24FB36FE0675DEFE74" box="[1262,1436,464,491]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 14050</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FF4FFE22734FFD90" box="[151,781,500,527]" collectionCode="IBUFRJ" collectorName="Revizee Central V" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 14050" specimenCount="1" stateProvince="Bahia state">
[1†]; Bank, 
<collectorName id="2696534ABA77DE24FEF9FE2371B1FD90" box="[289,499,500,527]" pageId="25" pageNumber="476">Revizee Central V</collectorName>
sta. 07r: 
<specimenCode id="DBC59EE7BA77DE24FDB9FE22734FFD90" box="[609,781,500,527]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 14050</specimenCode>
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[1†]. 
<materialsCitation id="3B0B3CC1BA77DE24FC8CFE23723EFDAC" collectionCode="IBUFRJ" collectorName="Revizee Central V" country="Brazil" location="Espirito Santo state" pageId="25" pageNumber="476" specimenCode="IBUFRJ 14008" specimenCount="1" stateProvince="Espirito Santo">
<collectingRegion id="49A7F87EBA77DE24FC8CFE237474FD90" box="[852,1078,500,527]" country="Brazil" name="Espirito Santo" pageId="25" pageNumber="476">Espírito Santo state</collectingRegion>
: South edge of Abrolhos Shelf, 
<collectorName id="2696534ABA77DE24FF4FFDCF7124FDAC" box="[151,358,536,563]" pageId="25" pageNumber="476">Revizee Central V</collectorName>
sta. 25a: 
<specimenCode id="DBC59EE7BA77DE24FE0AFDCE723EFDAC" box="[466,636,537,563]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 14008</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FD5CFDCE7103FDC8" collectionCode="IBUFRJ" collectorName="Revizee Central V" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 13998" specimenCount="1" stateProvince="Espirito Santo state">
[1†]; Vitória-Trindade Chain: Jaseur Bank, 
<collectorName id="2696534ABA77DE24FBB4FDCF7579FDAC" box="[1132,1339,536,563]" pageId="25" pageNumber="476">Revizee Central V</collectorName>
sta. 23r: 
<specimenCode id="DBC59EE7BA77DE24FF4FFDEA7103FDC8" box="[151,321,573,599]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 13998</specimenCode>
</materialsCitation>
[1†]; 
<materialsCitation id="3B0B3CC1BA77DE24FE5DFDEB7384FDC8" box="[389,966,572,599]" collectionCode="MNRJ" country="Brazil" location="Rio de Janeiro state" pageId="25" pageNumber="476" specimenCode="MNRJ 15962" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingRegion id="49A7F87EBA77DE24FE5DFDEB7227FDC8" box="[389,613,572,599]" country="Brazil" name="Rio de Janeiro" pageId="25" pageNumber="476">Rio de Janeiro state</collectingRegion>
: HAB 11 sta. D2: 
<specimenCode id="DBC59EE7BA77DE24FCF7FDEA7384FDC8" box="[815,966,572,599]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 15962</specimenCode>
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<materialsCitation id="3B0B3CC1BA77DE24FC15FDEA74E3FDC9" box="[973,1185,572,599]" collectionCode="MNRJ" country="Brazil" isTailEnumeration="true" location="Rio de Janeiro state" pageId="25" pageNumber="476" specimenCode="MNRJ 16969" specimenCount="1" stateProvince="Rio de Janeiro state">
[3†]; 
<specimenCode id="DBC59EE7BA77DE24FBD2FDEA74E3FDC9" box="[1034,1185,572,599]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 16969</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FB70FDEA7172FDE4" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 15963" specimenCount="1" stateProvince="Rio de Janeiro state">
[1†]; HAB 11 sta. E3: 
<specimenCode id="DBC59EE7BA77DE24FF4FFDB67172FDE4" box="[151,304,608,635]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 15963</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FEE1FDB67253FDE4" box="[313,529,608,635]" collectionCode="MNRJ" country="Brazil" isTailEnumeration="true" pageId="25" pageNumber="476" specimenCode="MNRJ 16851" specimenCount="1" stateProvince="Rio de Janeiro state">
[3†]; 
<specimenCode id="DBC59EE7BA77DE24FEA0FDB67253FDE4" box="[376,529,608,635]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 16851</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FDC1FDB673F4FDE4" box="[537,950,608,635]" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 15964" specimenCount="1" stateProvince="Rio de Janeiro state">
[1†]; HAB 11 sta. E2: 
<specimenCode id="DBC59EE7BA77DE24FCC5FDB673F4FDE4" box="[797,950,608,635]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 15964</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FC67FDB674D5FDE4" box="[959,1175,608,635]" collectionCode="MNRJ" country="Brazil" isTailEnumeration="true" pageId="25" pageNumber="476" specimenCode="MNRJ 16583" specimenCount="1" stateProvince="Rio de Janeiro state">
[1†]; 
<specimenCode id="DBC59EE7BA77DE24FC26FDB674D5FDE4" box="[1022,1175,608,635]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 16583</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FB47FDB67172FD00" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 15965" specimenCount="1" stateProvince="Rio de Janeiro state">
[3†]; HAB 11 sta. A1: 
<specimenCode id="DBC59EE7BA77DE24FF4FFD527172FD00" box="[151,304,644,671]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 15965</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FEE1FD527296FD00" box="[313,724,644,671]" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 15966" specimenCount="1" stateProvince="Rio de Janeiro state">
[1†]; HAB 11 sta. F2: 
<specimenCode id="DBC59EE7BA77DE24FDE3FD527296FD00" box="[571,724,644,671]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 15966</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FD05FD5273F7FD00" box="[733,949,644,671]" collectionCode="MNRJ" country="Brazil" isTailEnumeration="true" pageId="25" pageNumber="476" specimenCode="MNRJ 16582" specimenCount="1" stateProvince="Rio de Janeiro state">
[1†]; 
<specimenCode id="DBC59EE7BA77DE24FCC4FD5273F7FD00" box="[796,949,644,671]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 16582</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FC65FD5274D7FD01" box="[957,1173,644,671]" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 16966" specimenCount="1" stateProvince="Rio de Janeiro state">
[1†]; 
<specimenCode id="DBC59EE7BA77DE24FC24FD5274D7FD01" box="[1020,1173,644,671]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 16966</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FB46FD52716DFD5C" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 15968" specimenCount="1" stateProvince="Rio de Janeiro state">
[1†]; HAB 11 sta. G3: 
<specimenCode id="DBC59EE7BA77DE24FF4FFD7E716DFD5C" box="[151,303,680,707]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 15968</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FEEEFD7E716CFD78" collectingDate="1993-05" collectionCode="MNRJ" collectorName="B Astro Garoupa" country="Brazil" elevation="38" latitude="-22.408333" location="Santana Archipelago" longLatPrecision="21" longitude="-41.708336" municipality="Macae" pageId="25" pageNumber="476" specimenCode="MNRJ 22141" specimenCount="1" stateProvince="Rio de Janeiro state">
[1†]; 
<collectingMunicipality id="6BB8ACE6BA77DE24FEABFD7E7182FD5C" box="[371,448,681,707]" pageId="25" pageNumber="476">Macaé</collectingMunicipality>
, 
<location id="8EBC6047BA77DE24FE14FD7E72FAFD5D" LSID="urn:lsid:plazi:treatment:03CA878ABA77DE27FF4FFF407501FD86:8EBC6047BA77DE24FE14FD7E72FAFD5D" box="[460,696,680,707]" country="Brazil" latitude="-22.408333" longLatPrecision="21" longitude="-41.708336" municipality="Macae" name="Santana Archipelago" pageId="25" pageNumber="476" stateProvince="Rio de Janeiro state">Santana Archipelago</location>
; 
<geoCoordinate id="EE57505BBA77DE24FD1BFD7D7304FD5A" box="[707,838,682,709]" degrees="22" direction="south" minutes="24" orientation="latitude" pageId="25" pageNumber="476" precision="15" seconds="30" value="-22.408333">22°24′30″S</geoCoordinate>
, 
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, 
<quantity id="4C9B9B79BA77DE24FC31FD7E7467FD5C" box="[1001,1061,681,707]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="476" unit="m" value="38.0">
<elevation id="004ED1AFBA77DE24FC31FD7E7467FD5C" box="[1001,1061,681,707]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="476" unit="m" value="38.0">38 m</elevation>
</quantity>
; S/ 
<collectorName id="2696534ABA77DE24FB90FD7E754AFD5C" box="[1096,1288,681,707]" pageId="25" pageNumber="476">
B 
<emphasis id="B917EA8EBA77DE24FBBAFD7E754AFD5C" box="[1122,1288,681,707]" italics="true" pageId="25" pageNumber="476">Astro Garoupa</emphasis>
</collectorName>
col., 
<date id="FFDD105CBA77DE24FA9FFD7E75DBFD5C" box="[1351,1433,681,707]" bridgedPair="/" pageId="25" pageNumber="476" value="1993-05">
<collectingDate id="EF99E9B4BA77DE24FA9FFD7E75DBFD5C" box="[1351,1433,681,707]" bridgedPair="/" pageId="25" pageNumber="476" value="1993-05">v/1993</collectingDate>
</date>
: 
<specimenCode id="DBC59EE7BA77DE24FF4FFD1A716CFD78" box="[151,302,717,743]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 22141</specimenCode>
</materialsCitation>
[5†]: off 
<materialsCitation id="3B0B3CC1BA77DE24FE4DFD1B71D9FC94" collectingDate="1997-11" collectionCode="MNRJ" collectorName="V Professor Wladimir Besnard" country="Brazil" elevation="225" latitude="-23.817835" location="Rio de Janeiro" longLatPrecision="1" longitude="-42.780167" pageId="25" pageNumber="476" specimenCode="MNRJ 28195" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingRegion id="49A7F87EBA77DE24FE4DFD1B727AFD79" box="[405,568,716,742]" country="Brazil" name="Rio de Janeiro" pageId="25" pageNumber="476">Rio de Janeiro</collectingRegion>
; 
<geoCoordinate id="EE57505BBA77DE24FD9AFD1972ABFD76" box="[578,745,718,745]" degrees="23" direction="south" minutes="49" orientation="latitude" pageId="25" pageNumber="476" precision="1" seconds="04.20" value="-23.817835">23°49′04.20″S</geoCoordinate>
, 
<geoCoordinate id="EE57505BBA77DE24FD2AFD1973E3FD76" box="[754,929,718,745]" degrees="42" direction="west" minutes="46" orientation="longitude" pageId="25" pageNumber="476" precision="1" seconds="48.60" value="-42.780167">42°46′48.60″W</geoCoordinate>
, 
<quantity id="4C9B9B79BA77DE24FC73FD1B73B6FD78" box="[939,1012,716,743]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="25" pageNumber="476" unit="m" value="225.0">
<elevation id="004ED1AFBA77DE24FC73FD1B73B6FD78" box="[939,1012,716,743]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="25" pageNumber="476" unit="m" value="225.0">225 m</elevation>
</quantity>
; R/ 
<collectorName id="2696534ABA77DE24FBC1FD1A752FFD79" box="[1049,1389,716,743]" pageId="25" pageNumber="476">
V Professor 
<emphasis id="B917EA8EBA77DE24FB79FD1B752FFD79" box="[1185,1389,716,742]" italics="true" pageId="25" pageNumber="476">Wladimir Besnard</emphasis>
</collectorName>
col. 
<date id="FFDD105CBA77DE24FF4FFD2770B5FC95" box="[151,247,752,778]" bridgedPair="/" pageId="25" pageNumber="476" value="1997-11">
<collectingDate id="EF99E9B4BA77DE24FF4FFD2770B5FC95" box="[151,247,752,778]" bridgedPair="/" pageId="25" pageNumber="476" value="1997-11">Xi/1997</collectingDate>
</date>
; 
<specimenCode id="DBC59EE7BA77DE24FEDAFD2671D9FC94" box="[258,411,752,779]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 28195</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FE7AFD267369FC94" box="[418,811,752,779]" collectionCode="MNRJ" country="Brazil" location="Campos Basin" pageId="25" pageNumber="476" specimenCode="MNRJ 28198" specimenCount="1" stateProvince="Rio de Janeiro">
[7†]: 
<location id="8EBC6047BA77DE24FE38FD2672C4FC95" LSID="urn:lsid:plazi:treatment:03CA878ABA77DE27FF4FFF407501FD86:8EBC6047BA77DE24FE38FD2672C4FC95" box="[480,646,752,779]" country="Brazil" name="Campos Basin" pageId="25" pageNumber="476" stateProvince="Rio de Janeiro">Campos Basin</location>
; 
<specimenCode id="DBC59EE7BA77DE24FD4AFD267369FC94" box="[658,811,753,779]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 28198</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FCEAFD26754EFC94" box="[818,1292,752,779]" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 60185" specimenCount="1" stateProvince="Rio de Janeiro">
[1†]; Revizee Sul sta. 6653: 
<specimenCode id="DBC59EE7BA77DE24FBACFD26754EFC94" box="[1140,1292,752,779]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 60185</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FACBFD2671E3FCB0" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 60254" specimenCount="1" stateProvince="Rio de Janeiro">
[1]; PADCT sta. 6627: 
<specimenCode id="DBC59EE7BA77DE24FED2FCC271E3FCB0" box="[266,417,788,815]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 60254</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FE70FCC273DAFCB0" box="[424,920,788,815]" collectionCode="IBUFRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 2428" specimenCount="1" stateProvince="Rio de Janeiro">
[1]; Cabo Frio VII sta. 6147;: 
<specimenCode id="DBC59EE7BA77DE24FD24FCC273DAFCB0" box="[764,920,789,815]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 2428</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FC46FCC275DEFCB0" box="[926,1436,788,815]" collectionCode="IBUFRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 2429" specimenCount="1" stateProvince="Rio de Janeiro">
[1†]; Cabo Frio VII sta. 6165;: 
<specimenCode id="DBC59EE7BA77DE24FAD8FCC275DEFCB0" box="[1280,1436,789,815]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 2429</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FF4FFCEF71C6FCCC" box="[151,388,824,851]" collectionCode="IBUFRJ" country="Brazil" isTailEnumeration="true" pageId="25" pageNumber="476" specimenCode="IBUFRJ 14813" specimenCount="1" stateProvince="Rio de Janeiro">
[6†]; 
<specimenCode id="DBC59EE7BA77DE24FF0FFCEE71C6FCCC" box="[215,388,825,851]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 14813</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FE55FCEE73D2FCCC" box="[397,912,824,851]" collectionCode="IBUFRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 2430" specimenCount="1" stateProvince="Rio de Janeiro">
[4†]; Cabo Frio VII sta. 6199; 
<specimenCode id="DBC59EE7BA77DE24FD29FCEE73D2FCCC" box="[753,912,825,851]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 2430</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FC41FCEE75DEFCCC" box="[921,1436,824,851]" collectionCode="IBUFRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 2431" specimenCount="1" stateProvince="Rio de Janeiro">
[2†]; Cabo Frio VII sta. 6198; 
<specimenCode id="DBC59EE7BA77DE24FB25FCEE75DEFCCC" box="[1277,1436,825,851]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 2431</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FF4FFC8A738FFCE8" box="[151,973,860,887]" collectingDate="1993-05-20" collectionCode="IBUFRJ" collectorName="B Astro Garoupa" country="Brazil" location="Cabiunas" municipality="Macae" pageId="25" pageNumber="476" specimenCode="IBUFRJ 7223" specimenCount="1" stateProvince="Rio de Janeiro">
[1†]; 
<collectingMunicipality id="6BB8ACE6BA77DE24FF0BFC8A7162FCE8" box="[211,288,861,887]" pageId="25" pageNumber="476">Macaé</collectingMunicipality>
, 
<location id="8EBC6047BA77DE24FEF3FC8B71D5FCE9" LSID="urn:lsid:plazi:treatment:03CA878ABA77DE27FF4FFF407501FD86:8EBC6047BA77DE24FEF3FC8B71D5FCE9" box="[299,407,860,886]" country="Brazil" municipality="Macae" name="Cabiunas" pageId="25" pageNumber="476" stateProvince="Rio de Janeiro">Cabiúnas</location>
; S/ 
<collectorName id="2696534ABA77DE24FE60FC8A7234FCE8" box="[440,630,861,887]" pageId="25" pageNumber="476">
B 
<emphasis id="B917EA8EBA77DE24FE09FC8A7234FCE8" box="[465,630,861,887]" italics="true" pageId="25" pageNumber="476">Astro Garoupa</emphasis>
</collectorName>
col., 
<date id="FFDD105CBA77DE24FD6AFC8A736AFCE8" box="[690,808,861,887]" pageId="25" pageNumber="476" value="1993-05-20">
<collectingDate id="EF99E9B4BA77DE24FD6AFC8A736AFCE8" box="[690,808,861,887]" pageId="25" pageNumber="476" value="1993-05-20">20/v/1993</collectingDate>
</date>
: 
<specimenCode id="DBC59EE7BA77DE24FCE9FC8A738FFCE8" box="[817,973,861,887]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 7223</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FC0AFC8A739CFC04" collectingDate="1993-05" collectionCode="IBUFRJ" collectorName="B Astro Garoupa" country="Brazil" elevation="15" latitude="-22.358334" location="Santana Archipelago" longLatPrecision="855" longitude="-41.683334" municipality="Macae" pageId="25" pageNumber="476" specimenCode="IBUFRJ 7410" specimenCount="1" stateProvince="Rio de Janeiro">
[1†]; 
<collectingMunicipality id="6BB8ACE6BA77DE24FBD7FC8A741EFCE8" box="[1039,1116,861,887]" pageId="25" pageNumber="476">Macaé</collectingMunicipality>
, 
<location id="8EBC6047BA77DE24FBBEFC8A7512FCE9" LSID="urn:lsid:plazi:treatment:03CA878ABA77DE27FF4FFF407501FD86:8EBC6047BA77DE24FBBEFC8A7512FCE9" box="[1126,1360,860,887]" country="Brazil" latitude="-22.358334" longLatPrecision="855" longitude="-41.683334" municipality="Macae" name="Santana Archipelago" pageId="25" pageNumber="476" stateProvince="Rio de Janeiro">Santana Archipelago</location>
, sta. 9; 
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, 
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, 
<quantity id="4C9B9B79BA77DE24FE4AFC57718FFC04" box="[402,461,896,923]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="476" unit="m" value="15.0">
<elevation id="004ED1AFBA77DE24FE4AFC57718FFC04" box="[402,461,896,923]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="476" unit="m" value="15.0">15 m</elevation>
</quantity>
; S/ 
<collectorName id="2696534ABA77DE24FE37FC5672EEFC04" box="[495,684,897,923]" pageId="25" pageNumber="476">
B 
<emphasis id="B917EA8EBA77DE24FDDFFC5672EEFC04" box="[519,684,897,923]" italics="true" pageId="25" pageNumber="476">Astro Garoupa</emphasis>
</collectorName>
col., 
<date id="FFDD105CBA77DE24FD30FC567378FC04" box="[744,826,897,923]" bridgedPair="/" pageId="25" pageNumber="476" value="1993-05">
<collectingDate id="EF99E9B4BA77DE24FD30FC567378FC04" box="[744,826,897,923]" bridgedPair="/" pageId="25" pageNumber="476" value="1993-05">v/1993</collectingDate>
</date>
: 
<specimenCode id="DBC59EE7BA77DE24FC9BFC56739CFC04" box="[835,990,897,923]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 7410</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FC3CFC56708DFC21" collectionCode="IBUFRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 7463" specimenCount="1" stateProvince="Rio de Janeiro">
[11†]; Cabo Frio VII sta. 6194; 
<specimenCode id="DBC59EE7BA77DE24FAE7FC56708DFC21" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 7463</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FF00FC7272DBFC20" box="[216,665,932,959]" collectionCode="IBUFRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 7570" specimenCount="1" stateProvince="Rio de Janeiro">
[1†]; Geomar XII sta. 76; 
<specimenCode id="DBC59EE7BA77DE24FE20FC7372DBFC20" box="[504,665,932,959]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 7570</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FD7AFC727438FC21" box="[674,1146,932,959]" collectionCode="IBUFRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 7693" specimenCount="1" stateProvince="Rio de Janeiro">
[1†]; Geomar XII sta. 125; 
<specimenCode id="DBC59EE7BA77DE24FC04FC727438FC21" box="[988,1146,932,959]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 7693</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FB5BFC727103FC7C" collectionCode="IBUFRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="IBUFRJ 13344" specimenCount="1" stateProvince="Rio de Janeiro">
[1†]; Revizee V sta. 52f: 
<specimenCode id="DBC59EE7BA77DE24FF4FFC1E7103FC7C" box="[151,321,969,995]" collectionCode="IBUFRJ" pageId="25" pageNumber="476">IBUFRJ 13344</specimenCode>
</materialsCitation>
[2†]; 
<materialsCitation id="3B0B3CC1BA77DE24FE5BFC1E7388FC7D" box="[387,970,968,995]" collectionCode="MNRJ" country="Brazil" location="Sao Paulo state" pageId="25" pageNumber="476" specimenCode="MNRJ 28196" specimenCount="1" stateProvince="Sao Paulo">
<collectingRegion id="49A7F87EBA77DE24FE5BFC1E7273FC7C" box="[387,561,968,995]" country="Brazil" name="Sao Paulo" pageId="25" pageNumber="476">São Paulo state</collectingRegion>
: Revizee sul sta. 6656: 
<specimenCode id="DBC59EE7BA77DE24FCEBFC1E7388FC7D" box="[819,970,968,995]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 28196</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FC08FC1E75DEFC7C" box="[976,1436,968,995]" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 28197" specimenCount="1" stateProvince="Sao Paulo state">
[1†]; Revizee sul sta. 6680: 
<specimenCode id="DBC59EE7BA77DE24FADDFC1E75DEFC7C" box="[1285,1436,969,995]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 28197</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FF4FFC3B7205FB99" box="[151,583,1004,1031]" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 60260" specimenCount="1" stateProvince="Sao Paulo state">
[5†]; PADCT sta. 6631: 
<specimenCode id="DBC59EE7BA77DE24FE76FC3A7205FB99" box="[430,583,1004,1031]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 60260</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FD97FC3A745EFB99" box="[591,1052,1004,1031]" collectionCode="MNRJ" country="Brazil" pageId="25" pageNumber="476" specimenCode="MNRJ 60268" specimenCount="1" stateProvince="Sao Paulo state">
[1]; Revizee sul sta. 6652: 
<specimenCode id="DBC59EE7BA77DE24FC5AFC3A745EFB99" box="[898,1052,1004,1031]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="25" pageNumber="476">MNRJ 60268</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FBFCFC3A720FFBB4" collectingDate="2001-02-15" collectionCode="MZSP" country="Brazil" location="Caraguatatuba" pageId="25" pageNumber="476" specimenCode="MZSP 42037" specimenCount="1" stateProvince="Sao Paulo state">
[2]; 
<location id="8EBC6047BA77DE24FB8DFC3B74B8FB99" LSID="urn:lsid:plazi:treatment:03CA878ABA77DE27FF4FFF407501FD86:8EBC6047BA77DE24FB8DFC3B74B8FB99" box="[1109,1274,1004,1030]" country="Brazil" name="Caraguatatuba" pageId="25" pageNumber="476" stateProvince="Sao Paulo state">Caraguatatuba</location>
; 23°53′097″S, 45°30′843″W, 
<date id="FFDD105CBA77DE24FEE2FBC771F0FBB4" box="[314,434,1040,1067]" pageId="25" pageNumber="476" value="2001-02-15">
<collectingDate id="EF99E9B4BA77DE24FEE2FBC771F0FBB4" box="[314,434,1040,1067]" pageId="25" pageNumber="476" value="2001-02-15">15/ii/2001</collectingDate>
</date>
: 
<specimenCode id="DBC59EE7BA77DE24FE62FBC6720FFBB4" box="[442,589,1041,1067]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="476" type="Museum">MZSP 42037</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FD8AFBC67541FBB4" box="[594,1283,1040,1069]" collectionCode="MZSP" country="Brazil" elevation="47" latitude="-23.783333" location="Ubatuba" longLatPrecision="1254" longitude="-44.966667" pageId="25" pageNumber="476" specimenCode="MZSP 85269" specimenCount="1" stateProvince="Sao Paulo state">
[2]; 
<location id="8EBC6047BA77DE24FD58FBC772A0FBB5" LSID="urn:lsid:plazi:treatment:03CA878ABA77DE27FF4FFF407501FD86:8EBC6047BA77DE24FD58FBC772A0FBB5" box="[640,738,1040,1066]" country="Brazil" latitude="-23.783333" longLatPrecision="1254" longitude="-44.966667" name="Ubatuba" pageId="25" pageNumber="476" stateProvince="Sao Paulo state">Ubatuba</location>
, sta. 4854; 
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, 
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, 
<quantity id="4C9B9B79BA77DE24FBF4FBC67425FBB4" box="[1068,1127,1041,1067]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="25" pageNumber="476" unit="m" value="47.0">
<elevation id="004ED1AFBA77DE24FBF4FBC67425FBB4" box="[1068,1127,1041,1067]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="25" pageNumber="476" unit="m" value="47.0">47 m</elevation>
</quantity>
: 
<specimenCode id="DBC59EE7BA77DE24FBA8FBC67541FBB4" box="[1136,1283,1040,1067]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="476" type="Museum">MZSP 85269</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FAD0FBC673E8FBD0" collectingDate="1986-10-10" collectionCode="MZSP" collectorName="V Veliger II" country="Brazil" elevation="38" latitude="-23.883333" location="Ubatuba" longLatPrecision="1254" longitude="-45.15" pageId="25" pageNumber="476" specimenCode="MZSP 85401" specimenCount="1" stateProvince="Sao Paulo state">
[3]; 
<location id="8EBC6047BA77DE24FAEEFBC775DBFBB5" LSID="urn:lsid:plazi:treatment:03CA878ABA77DE27FF4FFF407501FD86:8EBC6047BA77DE24FAEEFBC775DBFBB5" box="[1334,1433,1040,1066]" country="Brazil" latitude="-23.883333" longLatPrecision="1254" longitude="-45.15" name="Ubatuba" pageId="25" pageNumber="476" stateProvince="Sao Paulo state">Ubatuba</location>
; 
<geoCoordinate id="EE57505BBA77DE24FF4FFBE170B0FBCE" box="[151,242,1078,1105]" degrees="23" direction="south" minutes="53" orientation="latitude" pageId="25" pageNumber="476" precision="925" value="-23.883333">23°53′S</geoCoordinate>
, 
<geoCoordinate id="EE57505BBA77DE24FF26FBE17120FBCE" box="[254,354,1078,1105]" degrees="45" direction="west" minutes="09" orientation="longitude" pageId="25" pageNumber="476" precision="925" value="-45.15">45°09′W</geoCoordinate>
, 
<quantity id="4C9B9B79BA77DE24FEB5FBE271EBFBD0" box="[365,425,1077,1103]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="476" unit="m" value="38.0">
<elevation id="004ED1AFBA77DE24FEB5FBE271EBFBD0" box="[365,425,1077,1103]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="476" unit="m" value="38.0">38 m</elevation>
</quantity>
; R/ 
<collectorName id="2696534ABA77DE24FE08FBE27214FBD1" box="[464,598,1076,1103]" pageId="25" pageNumber="476">
V 
<emphasis id="B917EA8EBA77DE24FE32FBE37214FBD1" box="[490,598,1076,1102]" italics="true" pageId="25" pageNumber="476">Véliger II</emphasis>
</collectorName>
col., 
<date id="FFDD105CBA77DE24FD4CFBE37348FBD1" box="[660,778,1076,1102]" pageId="25" pageNumber="476" value="1986-10-10">
<collectingDate id="EF99E9B4BA77DE24FD4CFBE37348FBD1" box="[660,778,1076,1102]" pageId="25" pageNumber="476" value="1986-10-10">10/x/1986</collectingDate>
</date>
: 
<specimenCode id="DBC59EE7BA77DE24FCCDFBE273E8FBD0" box="[789,938,1076,1103]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="476" type="Museum">MZSP 85401</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA77DE24FC69FBE274C1FBD1" box="[945,1155,1076,1103]" collectingDate="1986-10-10" collectionCode="MZSP" collectorName="V Veliger II" country="Brazil" elevation="38" isTailEnumeration="true" latitude="-23.883333" location="Ubatuba" longLatPrecision="1254" longitude="-45.15" pageId="25" pageNumber="476" specimenCode="MZSP 88368" specimenCount="1" stateProvince="Sao Paulo state">
[18]; 
<specimenCode id="DBC59EE7BA77DE24FC36FBE274C1FBD1" box="[1006,1155,1076,1103]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="476" type="Museum">MZSP 88368</specimenCode>
</materialsCitation>
[40].
</paragraph>
<paragraph id="8BDC369CBA77DE24FF1FFB8F7424FA0C" blockId="25.[151,1437,356,2039]" pageId="25" pageNumber="476">
<emphasis id="B917EA8EBA77DE24FF1FFB8F71FAFBED" bold="true" box="[199,440,1112,1138]" pageId="25" pageNumber="476">Revised description.</emphasis>
Shell vitreous or whitish, tall, conical, with an obtuse apex, reaching about 
<quantity id="4C9B9B79BA77DE24FADFFB8E751FFBEC" box="[1287,1373,1113,1139]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="25" pageNumber="476" unit="mm" value="4.1">4.1 mm</quantity>
long, 
<quantity id="4C9B9B79BA77DE24FF4FFBAA70AFFB08" box="[151,237,1149,1175]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="476" unit="mm" value="1.2">1.2 mm</quantity>
wide. Protoconch globose, subcylindrical, about 2.0 whorls, 400 μm high, smooth, transition to teleoconch marked by a gentle incremental scar. Teleoconch reaching about eight whorls, regularly convex outline; suture slightly deep, well impressed, sloping; surface glossy, presenting faint microscopic spiral striae in subsutural zone of each whorl; incremental scars strongly demarcated, appearing at irregular intervals; presence of spiral keels, emerging faintly in the first teleoconch whorl, in number of two in the initial whorls, reaching five in the body whorl of 
<typeStatus id="54D8883EBA77DE24FF6FFAE7715FFAD5" box="[183,285,1328,1354]" pageId="25" pageNumber="476" type="holotype">holotype</typeStatus>
. Last whorl about 40% of total length, broad; base rounded. Aperture high, elliptical in shape, about 60% of body whorl length, slightly expanded laterally, rounded and slightly advanced anteriorly, acute posteriorly; outer lip thin, orthocline, rectilinear; inner lip thin, reflected. Umbilical fissure present.
</paragraph>
</subSubSection>
<subSubSection id="C3796517BA77DE24FF1FFA4A7104FA60" pageId="25" pageNumber="476" type="description">
<paragraph id="8BDC369CBA77DE24FF1FFA4A7104FA60" blockId="25.[151,1437,356,2039]" pageId="25" pageNumber="476">
<emphasis id="B917EA8EBA77DE24FF1FFA4A713CFA28" bold="true" box="[199,382,1437,1463]" pageId="25" pageNumber="476">Measurements.</emphasis>
<typeStatus id="54D8883EBA77DE24FE5CFA4B71AEFA29" box="[388,492,1436,1462]" pageId="25" pageNumber="476" type="holotype">Holotype</typeStatus>
(MACN-In 28924): 8 whorls; SL= 
<quantity id="4C9B9B79BA77DE24FCA5FA4A73A6FA28" box="[893,996,1437,1463]" metricMagnitude="-3" metricUnit="m" metricValue="4.13" pageId="25" pageNumber="476" unit="mm" value="4.13">4.13 mm</quantity>
; BWL= 
<quantity id="4C9B9B79BA77DE24FB9CFA4B74E9FA28" box="[1092,1195,1436,1463]" metricMagnitude="-3" metricUnit="m" metricValue="1.59" pageId="25" pageNumber="476" unit="mm" value="1.59">1.59 mm</quantity>
; AL= 
<quantity id="4C9B9B79BA77DE24FB28FA4A7515FA28" box="[1264,1367,1437,1463]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="25" pageNumber="476" unit="mm" value="0.91">0.91 mm</quantity>
; SW= 
<quantity id="4C9B9B79BA77DE24FF4FFA1670BCFA44" box="[151,254,1473,1499]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="25" pageNumber="476" unit="mm" value="1.19">1.19 mm</quantity>
; AW= 
<quantity id="4C9B9B79BA77DE24FE93FA1771F0FA44" box="[331,434,1472,1499]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="25" pageNumber="476" unit="mm" value="0.67">0.67 mm</quantity>
. MNRJ 15964: 8 whorls; SL= 
<quantity id="4C9B9B79BA77DE24FCD2FA177333FA44" box="[778,881,1472,1499]" metricMagnitude="-3" metricUnit="m" metricValue="3.62" pageId="25" pageNumber="476" unit="mm" value="3.62">3.62 mm</quantity>
; BWL= 
<quantity id="4C9B9B79BA77DE24FC09FA16747AFA44" box="[977,1080,1473,1499]" metricMagnitude="-3" metricUnit="m" metricValue="1.41" pageId="25" pageNumber="476" unit="mm" value="1.41">1.41 mm</quantity>
; AL= 
<quantity id="4C9B9B79BA77DE24FBA6FA1674A7FA44" box="[1150,1253,1473,1499]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="25" pageNumber="476" unit="mm" value="0.83">0.83 mm</quantity>
; SW= 
<quantity id="4C9B9B79BA77DE24FAE9FA1675DAFA44" box="[1329,1432,1473,1499]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="25" pageNumber="476" unit="mm" value="1.03">1.03 mm</quantity>
; AW= 
<quantity id="4C9B9B79BA77DE24FF02FA337103FA60" box="[218,321,1508,1535]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="25" pageNumber="476" unit="mm" value="0.64">0.64 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796517BA77DE27FF1FF9DE7501FD86" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476" type="discussion">
<paragraph id="8BDC369CBA77DE24FF1FF9DE73D8F9F4" blockId="25.[151,1437,356,2039]" pageId="25" pageNumber="476">
<emphasis id="B917EA8EBA77DE24FF1FF9DE7179F9BC" bold="true" box="[199,315,1545,1571]" pageId="25" pageNumber="476">Remarks.</emphasis>
The present species was originally described in the genus 
<taxonomicName id="4C634D1FBA77DE24FC66F9DF754EF9BC" authority="G. O. Sars, 1878" authorityName="G. O. Sars" authorityYear="1878" box="[958,1292,1544,1571]" class="Gastropoda" family="Mathildidae" genus="Turritellopsis" kingdom="Animalia" pageId="25" pageNumber="476" phylum="Mollusca" rank="genus">
<emphasis id="B917EA8EBA77DE24FC66F9DF7417F9BD" box="[958,1109,1544,1570]" italics="true" pageId="25" pageNumber="476">Turritellopsis</emphasis>
G.O. 
<bibRefCitation id="EFF24B6DBA77DE24FB4FF9DE754EF9BC" author="Sars, G. O." box="[1175,1292,1545,1571]" pageId="25" pageNumber="476" pagination="1 - 34" refId="ref27534" refString="Sars, G. O. (1878) Bidrag til Kundskaben om Norges Artktische Fauna. I. Mollusca Regionis Articae Norvegiae. Brogger, Christiania, 466 pp., 1 map, pls. 1 - 34." type="book chapter" year="1878">Sars, 1878</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFF24B6DBA77DE24FAC1F9DF711AF9D8" author="Castellanos, Z. J. A. &amp; Landoni, N." pageId="25" pageNumber="476" pagination="291 - 299" refId="ref24249" refString="Castellanos, Z. J. A. &amp; Landoni, N. (1984) Nueva contribuicion al conocimiento de los micromoluscos de la plataforma continental argentina. Revista del Museo de La Plata, Nueva Serie, Seccion Zoologia, 13 (149), 291 - 299." type="journal article" year="1984">Castellanos &amp; Landoni 1984</bibRefCitation>
), which was included in 
<taxonomicName id="4C634D1FBA77DE24FDA0F9FB7341F9D9" authorityName="Loven" authorityYear="1847" box="[632,771,1580,1606]" class="Gastropoda" family="Turritellidae" kingdom="Animalia" pageId="25" pageNumber="476" phylum="Mollusca" rank="family">Turritellidae</taxonomicName>
until 
<bibRefCitation id="EFF24B6DBA77DE24FC90F9FA73A6F9D9" author="Waren, A." box="[840,996,1580,1607]" pageId="25" pageNumber="476" pagination="197 - 245" refId="ref28221" refString="Waren, A. (1996) New and little known mollusca from Iceland and Scandinavia. Part 3. Sarsia, 81, 197 - 245. https: // doi. org / 10.1080 / 00364827.1996.10413622" type="journal article" year="1996">Warén (1996)</bibRefCitation>
transferred it to 
<taxonomicName id="4C634D1FBA77DE24FB7FF9FB756CF9D9" authorityName="Dall" authorityYear="1889" box="[1191,1326,1580,1606]" class="Gastropoda" family="Mathildidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="25" pageNumber="476" phylum="Mollusca" rank="family">Mathildidae</taxonomicName>
based on anatomical characteristics studied by him and by 
<bibRefCitation id="EFF24B6DBA77DE24FD18F98673D5F9F4" author="Haszprunar, G." box="[704,919,1617,1643]" pageId="25" pageNumber="476" pagination="367 - 441" refId="ref25243" refString="Haszprunar, G. (1988) On the origin and evolution of major gastropod groups, with special reference to the Streptoneura. Journal of Molluscan Studies, 54, 367 - 441. https: // doi. org / 10.1093 / mollus / 54.4.367" type="journal article" year="1988">Haszprunar (1988)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDC369CBA77DE24FF1FF9A3751DF869" blockId="25.[151,1437,356,2039]" pageId="25" pageNumber="476">
The type species of 
<taxonomicName id="4C634D1FBA77DE24FE61F9A37212F911" authorityName="G. O. Sars" authorityYear="1878" box="[441,592,1652,1678]" class="Gastropoda" family="Mathildidae" genus="Turritellopsis" kingdom="Animalia" pageId="25" pageNumber="476" phylum="Mollusca" rank="genus">
<emphasis id="B917EA8EBA77DE24FE61F9A37212F911" box="[441,592,1652,1678]" italics="true" pageId="25" pageNumber="476">Turritellopsis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D1FBA77DE24FDBAF9A373BFF910" authority=", Dall, 1919" authorityName=", Dall" authorityYear="1919" box="[610,1021,1652,1679]" class="Gastropoda" family="Mathildidae" genus="Turritellopsis" kingdom="Animalia" pageId="25" pageNumber="476" phylum="Mollusca" rank="species" species="stimpsoni">
<emphasis id="B917EA8EBA77DE24FDBAF9A37332F911" box="[610,880,1652,1678]" italics="true" pageId="25" pageNumber="476">Turritellopsis stimpsoni</emphasis>
, 
<bibRefCitation id="EFF24B6DBA77DE24FC5BF9A373BFF910" author="Dall, W. H." box="[899,1021,1652,1679]" pageId="25" pageNumber="476" pagination="293 - 371" refId="ref24543" refString="Dall, W. H. (1919) Descriptions of new species of Mollusca from the North Pacific Ocean in the collection of the United States National Museum. Proceedings of the United States National Museum, 56 (2295), 293 - 371. https: // doi. org / 10.5479 / si. 00963801.56 - 2295.293" type="journal article" year="1919">Dall, 1919</bibRefCitation>
</taxonomicName>
[= 
<taxonomicName id="4C634D1FBA77DE24FBFAF9A37098F92C" authority="(Stimpson, 1851)" baseAuthorityName="Stimpson" baseAuthorityYear="1851" class="Gastropoda" family="Mathildidae" genus="Turritellopsis" kingdom="Animalia" pageId="25" pageNumber="476" phylum="Mollusca" rank="species" species="acicula">
<emphasis id="B917EA8EBA77DE24FBFAF9A37554F911" box="[1058,1302,1652,1678]" italics="true" pageId="25" pageNumber="476">Turritellopsis acicula</emphasis>
(Stimpson, 1851)
</taxonomicName>
] was revised and figured by 
<bibRefCitation id="EFF24B6DBA77DE24FDFFF94E7284F92D" author="Waren, A." box="[551,710,1688,1715]" pageId="25" pageNumber="476" pagination="197 - 245" refId="ref28221" refString="Waren, A. (1996) New and little known mollusca from Iceland and Scandinavia. Part 3. Sarsia, 81, 197 - 245. https: // doi. org / 10.1080 / 00364827.1996.10413622" type="journal article" year="1996">Warén (1996)</bibRefCitation>
. The shell has convex whorls with sharp spiral keels, similar to those present in 
<taxonomicName id="4C634D1FBA77DE24FE95F96B725FF949" authorityName="Souza &amp; Araújo &amp; Pimenta" authorityYear="2024" baseAuthorityName="Castellanos &amp; Landoni" baseAuthorityYear="1984" box="[333,541,1724,1750]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="476" phylum="Mollusca" rank="species" species="marplatensis" status="comb. nov.">
<emphasis id="B917EA8EBA77DE24FE95F96B725FF949" box="[333,541,1724,1750]" italics="true" pageId="25" pageNumber="476">Aclis marplatensis</emphasis>
</taxonomicName>
<emphasis id="B917EA8EBA77DE24FDFBF96A72E1F948" bold="true" box="[547,675,1725,1751]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A22457F5BA77DE24FDFBF96A72E1F948" box="[547,675,1725,1751]" pageId="25" pageNumber="476" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13582A19BA77DE24FD6AF96B7378F948" box="[690,826,1724,1751]" captionStart="FIGURE 13" captionStartId="24.[151,250,1908,1932]" captionTargetBox="[206,1382,181,1884]" captionTargetId="figure-23@24.[206,1382,181,1884]" captionTargetPageId="24" captionText="FIGURE 13. Aclis marplatensis (Castellanos &amp; Landoni, 1984). A, B. MZSP 85401. C. MZSP 88368. D. MNRJ 15963. E–I. MNRJ 15964. A–C, E. Shell in frontal view; rectangle in E indicates detail in H. D. Shell in lateral view. F. Detail of body whorl in frontal view. G. Detail of apical whorls. H, I. Detail of teleoconch surface; rectangle in H indicates detail in I. Scale bars: A–E. 1 mm; F. 500 μm; G. 100 μm; H, I. 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10954455" httpUri="https://zenodo.org/record/10954455/files/figure.png" pageId="25" pageNumber="476">Fig. 13A–E</figureCitation>
). However, 
<taxonomicName id="4C634D1FBA77DE24FC67F96A7404F949" authorityName=", Dall" authorityYear="1919" box="[959,1094,1725,1750]" class="Gastropoda" family="Mathildidae" genus="Turritellopsis" kingdom="Animalia" pageId="25" pageNumber="476" phylum="Mollusca" rank="species" species="stimpsoni">
<emphasis id="B917EA8EBA77DE24FC67F96A7404F949" box="[959,1094,1725,1750]" italics="true" pageId="25" pageNumber="476">T. stimpsoni</emphasis>
</taxonomicName>
presents the intervals between the spiral keels sculptured by microscopic axial threads (
<bibRefCitation id="EFF24B6DBA77DE24FCD5F93673D8F965" author="Waren, A." box="[781,922,1760,1787]" pageId="25" pageNumber="476" pagination="197 - 245" refId="ref28221" refString="Waren, A. (1996) New and little known mollusca from Iceland and Scandinavia. Part 3. Sarsia, 81, 197 - 245. https: // doi. org / 10.1080 / 00364827.1996.10413622" type="journal article" year="1996">Warén 1996</bibRefCitation>
: Fig. 28A–C) and lacks the microscopic spiral striae adjacent to the sutures. 
<taxonomicName id="4C634D1FBA77DE24FE2AF8D37287F881" authorityName="Souza &amp; Araújo &amp; Pimenta" authorityYear="2024" baseAuthorityName="Castellanos &amp; Landoni" baseAuthorityYear="1984" box="[498,709,1796,1822]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="476" phylum="Mollusca" rank="species" species="marplatensis">
<emphasis id="B917EA8EBA77DE24FE2AF8D37287F881" box="[498,709,1796,1822]" italics="true" pageId="25" pageNumber="476">Aclis marplatensis</emphasis>
</taxonomicName>
has no axial sculpture and has the microscopic spiral striae in the intervals between the suture and the subsutural spiral cord (
<figureCitation id="13582A19BA77DE24FCB1F8FF73A8F8DC" box="[873,1002,1832,1859]" captionStart="FIGURE 13" captionStartId="24.[151,250,1908,1932]" captionTargetBox="[206,1382,181,1884]" captionTargetId="figure-23@24.[206,1382,181,1884]" captionTargetPageId="24" captionText="FIGURE 13. Aclis marplatensis (Castellanos &amp; Landoni, 1984). A, B. MZSP 85401. C. MZSP 88368. D. MNRJ 15963. E–I. MNRJ 15964. A–C, E. Shell in frontal view; rectangle in E indicates detail in H. D. Shell in lateral view. F. Detail of body whorl in frontal view. G. Detail of apical whorls. H, I. Detail of teleoconch surface; rectangle in H indicates detail in I. Scale bars: A–E. 1 mm; F. 500 μm; G. 100 μm; H, I. 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10954455" httpUri="https://zenodo.org/record/10954455/files/figure.png" pageId="25" pageNumber="476">Fig. 13H, I</figureCitation>
), which leads to the new combination proposed here. The 
<typeStatus id="54D8883EBA77DE24FEA2F89B719EF8F9" box="[378,476,1868,1894]" pageId="25" pageNumber="476" type="holotype">holotype</typeStatus>
of the species was recently illustrated with SEM images (
<bibRefCitation id="EFF24B6DBA77DE24FBABF89B7528F8F8" author="Signorelli, J. H. &amp; Urteaga, D. &amp; Teso, V." box="[1139,1386,1868,1895]" pageId="25" pageNumber="476" pagination="45 - 69" refId="ref27575" refString="Signorelli, J. H., Urteaga, D. &amp; Teso, V. (2015) Zulma Ageitos de Castellanos: Publications and status of described taxa. Zootaxa, 4034 (1), 45 - 69. https: // doi. org / 10.11646 / zootaxa. 4034.1.2" type="journal article" year="2015">
Signorelli 
<emphasis id="B917EA8EBA77DE24FB34F89A7564F8F9" box="[1260,1318,1868,1894]" italics="true" pageId="25" pageNumber="476">et al.</emphasis>
2015
</bibRefCitation>
: fig. 6K–L). Based on these images, it is possible to assure that there are no axial threads in the 
<typeStatus id="54D8883EBA77DE24FB7EF8A7754AF815" box="[1190,1288,1904,1930]" pageId="25" pageNumber="476" type="holotype">holotype</typeStatus>
compared to what was described and figured by 
<bibRefCitation id="EFF24B6DBA77DE24FDFFF843733DF830" author="Castellanos, Z. J. A. &amp; Landoni, N." box="[551,895,1940,1967]" pageId="25" pageNumber="476" pagination="291 - 299" refId="ref24249" refString="Castellanos, Z. J. A. &amp; Landoni, N. (1984) Nueva contribuicion al conocimiento de los micromoluscos de la plataforma continental argentina. Revista del Museo de La Plata, Nueva Serie, Seccion Zoologia, 13 (149), 291 - 299." type="journal article" year="1984">Castellanos &amp; Landoni (1984)</bibRefCitation>
. The first record of the present species in 
<collectingCountry id="F374760CBA77DE24FA80F84375DEF831" box="[1368,1436,1940,1966]" name="Brazil" pageId="25" pageNumber="476">Brazil</collectingCountry>
was reported by 
<bibRefCitation id="EFF24B6DBA77DE24FE8FF86F7208F84D" author="Absalao, R. S. &amp; Pimenta, A. D. &amp; Costa, P. M. S." box="[343,586,1976,2002]" pageId="25" pageNumber="476" pagination="57 - 68" refId="ref23241" refString="Absalao, R. S., Pimenta, A. D. &amp; Costa, P. M. S. (1996) Novas ocorrencias de gastropodes no litoral do Rio de Janeiro (Brasil). Neritica, 10, 57 - 68." type="journal article" year="1996">
Absalão 
<emphasis id="B917EA8EBA77DE24FE64F86E71B4F84D" box="[444,502,1976,2002]" italics="true" pageId="25" pageNumber="476">et al.</emphasis>
(1996)
</bibRefCitation>
(as “ 
<taxonomicName id="4C634D1FBA77DE24FD52F86F73F9F84D" authorityName="Castellanos &amp; Landoni" authorityYear="1984" box="[650,955,1976,2002]" class="Gastropoda" family="Mathildidae" genus="Turritellopsis" kingdom="Animalia" pageId="25" pageNumber="476" phylum="Mollusca" rank="species" species="marplatensis">
<emphasis id="B917EA8EBA77DE24FD52F86F73F9F84D" box="[650,955,1976,2002]" italics="true" pageId="25" pageNumber="476">Turritellopsis marplatensis</emphasis>
</taxonomicName>
”). These authors described that the shells from 
<collectingCountry id="F374760CBA77DE24FF0DF80B715BF869" box="[213,281,2012,2038]" name="Brazil" pageId="25" pageNumber="476">Brazil</collectingCountry>
do not present microscopic axial sculpture, but only microscopic spiral striae as described above.
</paragraph>
<paragraph id="8BDC369CBA74DE27FF1FFF407443FE82" blockId="26.[151,1437,151,537]" pageId="26" pageNumber="477">
A revised description of the shell is provided here to elucidate the shell morphology of the species. The development of the spiral keels is very homogeneous in 
<taxonomicName id="4C634D1FBA74DE27FCD5FF6C73FFFF4A" authorityName="Castellanos &amp; Landoni" authorityYear="1984" box="[781,957,187,213]" class="Gastropoda" family="Mathildidae" genus="Turritellopsis" kingdom="Animalia" pageId="26" pageNumber="477" phylum="Mollusca" rank="species" species="marplatensis">
<emphasis id="B917EA8EBA74DE27FCD5FF6C73FFFF4A" box="[781,957,187,213]" italics="true" pageId="26" pageNumber="477">A. marplatensis</emphasis>
</taxonomicName>
, with all of them showing a similar degree of elevation from the teleoconch surface and does not vary among the specimens studied. This pattern seems robust, taking into consideration that this is one of the most abundant species studied.
</paragraph>
<paragraph id="8BDC369CBA74DE27FF1FFEF07233FE4E" blockId="26.[151,1437,151,537]" pageId="26" pageNumber="477">
<taxonomicName id="4C634D1FBA74DE27FF1FFEF071D4FEDE" authorityName="Souza &amp; Araújo &amp; Pimenta" authorityYear="2024" baseAuthorityName="Castellanos &amp; Landoni" baseAuthorityYear="1984" box="[199,406,295,321]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="26" pageNumber="477" phylum="Mollusca" rank="species" species="marplatensis">
<emphasis id="B917EA8EBA74DE27FF1FFEF071D4FEDE" box="[199,406,295,321]" italics="true" pageId="26" pageNumber="477">Aclis marplatensis</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="4C634D1FBA74DE27FD18FEF07371FEDE" authorityName="Bartsch" authorityYear="1947" box="[704,819,295,321]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="26" pageNumber="477" phylum="Mollusca" rank="species" species="eolis">
<emphasis id="B917EA8EBA74DE27FD18FEF07371FEDE" box="[704,819,295,321]" italics="true" pageId="26" pageNumber="477">Aclis eolis</emphasis>
</taxonomicName>
(
<figureCitation id="13582A19BA74DE27FC99FEF07388FEDE" box="[833,970,295,321]" captionStart="FIGURE 19" captionStartId="33.[151,250,1761,1785]" captionTargetBox="[171,1417,453,1737]" captionTargetId="figure-143@33.[171,1417,453,1737]" captionTargetPageId="33" captionText="FIGURE 19. A–D. Aclis eolis Bartsch, 1947: Holotype, USNM 573617. A. Shell in frontal view. B. Shell in lateral view. C. Detail of apical whorls. D. Detail of the body whorl in frontal view. E–F. Aclis hendersoni Dall, 1927: E, F. Holotype, USNM 333465; E. Shell in frontal view. F. Protoconch in apical view. G–H. Aclis georgiana Dall, 1927: Holotype, USNM 108020. G. Shell in frontal view. H. Protoconch in apical view. Scale bars: A, B, E, G. 1 mm; C, F, H. 200 μm, D. 500 μm. Credits: E–H. USNM." figureDoi="http://doi.org/10.5281/zenodo.10954467" httpUri="https://zenodo.org/record/10954467/files/figure.png" pageId="26" pageNumber="477">Fig. 19A–D</figureCitation>
), 
<taxonomicName id="4C634D1FBA74DE27FC05FEF07560FEDE" authority="Bartsch, 1911" authorityName="Bartsch" authorityYear="1911" box="[989,1314,295,321]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="26" pageNumber="477" phylum="Mollusca" rank="species" species="floridana">
<emphasis id="B917EA8EBA74DE27FC05FEF074C1FEDE" box="[989,1155,295,321]" italics="true" pageId="26" pageNumber="477">Aclis floridana</emphasis>
<bibRefCitation id="EFF24B6DBA74DE27FB51FEF07560FEDE" author="Bartsch, P." box="[1161,1314,295,321]" pageId="26" pageNumber="477" pagination="435 - 438" refId="ref23751" refString="Bartsch, P. (1911) New mollusks of the genus Aclis from the North Atlantic. Proceedings of the United States National Museum, 40 (1829), 435 - 438. https: // doi. org / 10.5479 / si. 00963801.40 - 1829.435" type="journal article" year="1911">Bartsch, 1911</bibRefCitation>
</taxonomicName>
(see 
<bibRefCitation id="EFF24B6DBA74DE27FA83FEF0717FFEFA" author="Penas, A. &amp; Rolan, E." pageId="26" pageNumber="477" pagination="15 - 64" refId="ref26541" refString="Penas, A. &amp; Rolan, E. (2013) Revision of the genera Murchisonella and Pseudoaclisina (Gastropoda, Heterobranchia, Murchisonellidae). Vita Malacologica, 11, 15 - 64." type="journal article" year="2013">Peñas &amp; Rolán 2013</bibRefCitation>
: pl. 2, 
<figureCitation id="13582A19BA74DE27FE54FE9C7244FEFA" box="[396,518,331,357]" captionStart="FIGURE 2" captionStartId="8.[151,250,1858,1882]" captionTargetBox="[151,1435,309,1835]" captionTargetId="figure-96@8.[151,1435,309,1835]" captionTargetPageId="8" captionText="FIGURE 2. Aclis sarissa Watson, 1881. A–C, E. Lectotype, NHMUK 1887.2.9.1576, herein designated; D. Paralectotype, NHMUK 1887.2.9.1576; F–I. Paralectotype, NMW 1955.158.26255; J–K. Holotype of Aclis carolinensis, USNM 83743. A, D, F, J. Shell in frontal view; B, G, K. Shell in lateral view. C, H. Shell in dorsal view. E, I. Body whorl in frontal view. Scale bars: A–D, F–H, I–J. 1 mm; E, I. 500 μm. Credits: A–E. NHMUK photographic unit; F–I. NMW." figureDoi="http://doi.org/10.5281/zenodo.10954424" httpUri="https://zenodo.org/record/10954424/files/figure.png" pageId="26" pageNumber="477">Fig. 2A, B</figureCitation>
for illustrations of the 
<typeStatus id="54D8883EBA74DE27FCD7FE9C737DFEFA" box="[783,831,331,357]" pageId="26" pageNumber="477">type</typeStatus>
material), 
<taxonomicName id="4C634D1FBA74DE27FC65FE9C74B8FEFA" authority="Dall, 1927" authorityName="Dall" authorityYear="1927" box="[957,1274,331,357]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="26" pageNumber="477" phylum="Mollusca" rank="species" species="hendersoni">
<emphasis id="B917EA8EBA74DE27FC65FE9C743BFEFA" box="[957,1145,331,357]" italics="true" pageId="26" pageNumber="477">Aclis hendersoni</emphasis>
<bibRefCitation id="EFF24B6DBA74DE27FB5AFE9C74B8FEFA" author="Dall, W. H." box="[1154,1274,331,357]" pageId="26" pageNumber="477" pagination="1 - 134" refId="ref24604" refString="Dall, W. H. (1927) Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steamer ' Albatross' in 1885 and 1886. Proceedings of the United States National Museum, 70 (2667), 1 - 134. https: // doi. org / 10.5479 / si. 00963801.70 - 2667.1" type="journal article" year="1927">Dall, 1927</bibRefCitation>
</taxonomicName>
(
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), all from 
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, 
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, and from 
<taxonomicName id="4C634D1FBA74DE27FDC8FEB87318FE16" authority="Watson, 1897" authorityName="Watson" authorityYear="1897" box="[528,858,367,393]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="26" pageNumber="477" phylum="Mollusca" rank="species" species="trilineata">
<emphasis id="B917EA8EBA74DE27FDC8FEB872FAFE16" box="[528,696,367,393]" italics="true" pageId="26" pageNumber="477">Aclis trilineata</emphasis>
<bibRefCitation id="EFF24B6DBA74DE27FD67FEB87318FE16" author="Watson, R. B." box="[703,858,367,393]" pageId="26" pageNumber="477" pagination="233 - 329" refId="ref28420" refString="Watson, R. B. (1897) On the marine Mollusca of Madeira; with descriptions of thirty-five new species, and an index-list of all the known sea-dwelling species of that island. Journal of the Linnean Society of London, Zoology, 26 (19), 233 - 329, pls. 19 - 20. https: // doi. org / 10.1111 / j. 1096 - 3642.1897. tb 00033. x" type="journal article" year="1897">Watson, 1897</bibRefCitation>
</taxonomicName>
(see 
<bibRefCitation id="EFF24B6DBA74DE27FC4FFEB87410FE16" author="Bogi, C. &amp; Renda, W. &amp; Giacobbe, S." box="[919,1106,367,393]" pageId="26" pageNumber="477" pagination="94 - 97" refId="ref23883" refString="Bogi, C., Renda, W. &amp; Giacobbe, S. (2016) Aclis trilineata Watson, 1897 (Gastropoda, Aclididae) from the Southern Tyrrhenian Sea. Bollettino Malacologico, 52 (2), 94 - 97." type="journal article" year="2016">
Bogi 
<emphasis id="B917EA8EBA74DE27FC0DFEB8744DFE16" box="[981,1039,367,393]" italics="true" pageId="26" pageNumber="477">et al.</emphasis>
2016
</bibRefCitation>
: 
<figureCitation id="13582A19BA74DE27FB86FEB87490FE16" box="[1118,1234,367,393]" captionStart="FIGURE 1" captionStartId="6.[151,250,1295,1319]" captionTargetBox="[151,1435,181,1271]" captionTargetId="figure-23@6.[151,1435,181,1271]" captionTargetPageId="6" captionText="FIGURE 1. Terminology of some of the shell features used in descriptions in the present study. A. Microscopic spiral striae of the teleoconch indicated by arrows. B. Spiral keels of the teleoconch indicated by arrows. C. Umbilical fissure indicated by arrow. D. Umbilicus indicated by arrow." figureDoi="http://doi.org/10.5281/zenodo.10954422" httpUri="https://zenodo.org/record/10954422/files/figure.png" pageId="26" pageNumber="477">Fig. 1A–F</figureCitation>
, for illustrations), from Madeira and western Mediterranean, by the more pronounced spiral keels of the teleoconch and by the less expanded aperture anteriorly and laterally.
</paragraph>
<paragraph id="8BDC369CBA74DE27FF1FFE0D7501FD86" blockId="26.[151,1437,151,537]" pageId="26" pageNumber="477">
<emphasis id="B917EA8EBA74DE27FF1FFE0D71B4FE6B" bold="true" box="[199,502,474,500]" pageId="26" pageNumber="477">Geographic Distribution.</emphasis>
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: 
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, 
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, Vitória-Trindade Chain, 
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, 
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(
<bibRefCitation id="EFF24B6DBA74DE27FF47FE28717CFD86" author="Absalao, R. S. &amp; Pimenta, A. D. &amp; Costa, P. M. S." box="[159,318,511,537]" pageId="26" pageNumber="477" pagination="57 - 68" refId="ref23241" refString="Absalao, R. S., Pimenta, A. D. &amp; Costa, P. M. S. (1996) Novas ocorrencias de gastropodes no litoral do Rio de Janeiro (Brasil). Neritica, 10, 57 - 68." type="journal article" year="1996">Absalão 1996</bibRefCitation>
; present study); 
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: 
<collectingMunicipality id="6BB8ACE6BA74DE27FDABFE2872E0FD86" box="[627,674,511,537]" pageId="26" pageNumber="477">Mar</collectingMunicipality>
del plata (
<bibRefCitation id="EFF24B6DBA74DE27FCCFFE28741CFD86" author="Castellanos, Z. J. A. &amp; Landoni, N." box="[791,1118,511,537]" pageId="26" pageNumber="477" pagination="291 - 299" refId="ref24249" refString="Castellanos, Z. J. A. &amp; Landoni, N. (1984) Nueva contribuicion al conocimiento de los micromoluscos de la plataforma continental argentina. Revista del Museo de La Plata, Nueva Serie, Seccion Zoologia, 13 (149), 291 - 299." type="journal article" year="1984">Castellanos &amp; Landoni 1984</bibRefCitation>
). 
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