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<mods:title id="356CDAAAB727B6379D3DC3847174421B">Late Neogene Lophophaenidae (Nassellaria, Radiolaria) from the eastern equatorial Pacific</mods:title>
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<paragraph id="8BDFDE46FFADFFC675DFE2B1FD02C59C" blockId="62.[151,709,1734,1796]" box="[151,709,1734,1760]" pageId="62" pageNumber="63">
<taxonomicName id="4C60A5C5FFADFFC675DFE2B1FD02C59C" ID-CoL="3W3V7" authority="Petrushevskaya, 1971" authorityName="Petrushevskaya" authorityYear="1971" box="[151,709,1734,1760]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="62" pageNumber="63" phylum="Radiozoa" rank="species" species="nadezdae">
<emphasis id="B9140254FFADFFC675DFE2B1FE68C59C" box="[151,431,1734,1760]" italics="true" pageId="62" pageNumber="63">Lophophaena nadezdae</emphasis>
<bibRefCitation id="EFF1A3B7FFADFFC674FEE2B1FD02C59C" author="Petrushevskaya, M. G." box="[438,709,1734,1760]" pageId="62" pageNumber="63" pagination="1 - 294" refId="ref93469" refString="Petrushevskaya, M. G. (1971) Nassellarian radiolarians in the plankton of the world oceans. Investigations of the Fauna of the Seas, 9 (17), 1 - 294. [in Russian]" type="journal article" year="1971">
<emphasis id="B9140254FFADFFC674FEE2B1FD02C59C" bold="true" box="[438,709,1734,1760]" pageId="62" pageNumber="63">Petrushevskaya, 1971</emphasis>
</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C37A8DCDFFADFFC675DFE29DFE57C478" box="[151,400,1770,1796]" pageId="62" pageNumber="63" type="description">
<paragraph id="8BDFDE46FFADFFC675DFE29DFE57C478" blockId="62.[151,709,1734,1796]" box="[151,400,1770,1796]" pageId="62" pageNumber="63">
Plate 23,
<figureCitation id="135BC2C3FFADFFC67448E29DFE9FC478" box="[256,344,1770,1796]" captionStart="FIGURE 1" captionStartId="6.[151,250,1340,1364]" captionTargetBox="[151,1430,743,1316]" captionTargetId="figure-284@6.[151,1436,741,1317]" captionTargetPageId="6" captionText="FIGURE 1. Diagram illustrating the general skeletal morphology of Lophophaenidae, and terminology used in this manuscript. Dashed lines indicate spines that are not visible from the angle shown, as they extend into the skeleton. An example specimen of Lophophaena casperi n. sp. is included for comparison to the diagrams." figureDoi="http://doi.org/10.5281/zenodo.10544060" httpUri="https://zenodo.org/record/10544060/files/figure.png" pageId="62" pageNumber="63">Figs 1A</figureCitation>
<figureCitation id="135BC2C3FFADFFC67421E29DFE4CC478" box="[361,395,1770,1796]" captionStart="FIGURE 2" captionStartId="7.[151,250,1538,1562]" captionTargetBox="[157,1428,745,1510]" captionTargetId="figure-251@7.[151,1436,741,1514]" captionTargetPageId="7" captionText="FIGURE 2. Relative abundance of specimens and taxa belonging to Plagiacanthoidea and Lophophaenidae per sample, from IODP Site U1337 in the EEP over the last ~10 Ma. The relative percentages of total taxa are represented by the blue lines and the red lines represent the percentages of total specimens (triangles = Plagiacanthoidea; circles = Lophophaenidae). The Plagiacanthoidea was among the most abundant and speciose of radiolarian groups encountered in Trubovitz et al. (2020), with at least 10% of the specimens and 20% of the taxonomic richness in every sample. Within the Plagiacanthoidea, lophophaenids consistently made up the majority of specimens (~67%) and taxa (~61%). Data are from Trubovitz et al. (2020)." figureDoi="http://doi.org/10.5281/zenodo.10544062" httpUri="https://zenodo.org/record/10544062/files/figure.png" pageId="62" pageNumber="63">2C</figureCitation>
.
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<subSubSection id="C37A8DCDFFADFFC675DFE358FBC7C4BD" pageId="62" pageNumber="63" type="reference_group">
<paragraph id="8BDFDE46FFADFFC675DFE358FC23C43B" blockId="62.[151,1024,1839,1986]" box="[151,996,1839,1863]" pageId="62" pageNumber="63">
<taxonomicName id="4C60A5C5FFADFFC675DFE358FD52C43B" authority="Ehrenberg, Benson, 1966" authorityName="Ehrenberg, Benson" authorityYear="1966" box="[151,661,1839,1863]" class="Polycystina" family="Plagiacanthidae" genus="Lithomelissa" isUncertain="true" kingdom="Chromista" order="Nassellaria" pageId="62" pageNumber="63" phylum="Radiozoa" rank="species" species="galeata">
<emphasis id="B9140254FFADFFC675DFE358FEDEC43B" box="[151,281,1839,1863]" italics="true" pageId="62" pageNumber="63">Lithomelissa</emphasis>
cf.
<emphasis id="B9140254FFADFFC67477E358FE4CC43B" box="[319,395,1839,1863]" italics="true" pageId="62" pageNumber="63">galeata</emphasis>
Ehrenberg,
<bibRefCitation id="EFF1A3B7FFADFFC6774FE358FD52C43B" author="Benson, R. N." box="[519,661,1839,1863]" pageId="62" pageNumber="63" refId="ref85564" refString="Benson, R. N. (1966) Recent Radiolaria from the Gulf of California. PhD dissertation, University of Minnesota, Minneapolis, 577 pp." type="book" year="1966">Benson, 1966</bibRefCitation>
</taxonomicName>
, pl. 24, figs. 1617 (non fig. 18).
</paragraph>
<paragraph id="8BDFDE46FFADFFC675DFE33AFCA6C41A" blockId="62.[151,1024,1839,1986]" box="[151,865,1869,1894]" pageId="62" pageNumber="63">
<taxonomicName id="4C60A5C5FFADFFC675DFE33AFE4EC419" authorityName="Petrushevskaya" authorityYear="1971" box="[151,393,1869,1893]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="62" pageNumber="63" phylum="Radiozoa" rank="species" species="nadezdae" status="sp. nov.">
<emphasis id="B9140254FFADFFC675DFE33AFE4EC419" box="[151,393,1869,1893]" italics="true" pageId="62" pageNumber="63">Lophophaena nadezdae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227BF2FFFADFFC674C7E339FE00C419" box="[399,455,1870,1893]" pageId="62" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
,
<bibRefCitation id="EFF1A3B7FFADFFC6749AE33AFD76C41A" author="Petrushevskaya, M. G." box="[466,689,1869,1894]" pageId="62" pageNumber="63" pagination="1 - 294" refId="ref93469" refString="Petrushevskaya, M. G. (1971) Nassellarian radiolarians in the plankton of the world oceans. Investigations of the Fauna of the Seas, 9 (17), 1 - 294. [in Russian]" type="journal article" year="1971">Petrushevskaya, 1971</bibRefCitation>
, pl. 60, figs. 14.
</paragraph>
<paragraph id="8BDFDE46FFADFFC675DFE31BFC64C4F8" blockId="62.[151,1024,1839,1986]" box="[151,931,1900,1924]" pageId="62" pageNumber="63">
<taxonomicName id="4C60A5C5FFADFFC675DFE31BFD08C4F8" authority="Popofsky, Van de Paverd, 1995" authorityName="Popofsky, Van de Paverd" authorityYear="1995" box="[151,719,1900,1924]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="62" pageNumber="63" phylum="Radiozoa" rank="species" species="variabilis">
<emphasis id="B9140254FFADFFC675DFE31BFE4DC4F8" box="[151,394,1900,1924]" italics="true" pageId="62" pageNumber="63">Lophophaena variabilis</emphasis>
Popofsky,
<bibRefCitation id="EFF1A3B7FFADFFC674B3E31AFD08C4F8" author="Van de Paverd, P. J." box="[507,719,1900,1924]" pageId="62" pageNumber="63" refId="ref96419" refString="Van de Paverd, P. J. (1995) Recent Polycystine Radiolaria from the Snellius-II Expedition. Ph. D. thesis, Center for Marine Earth Science (the Netherlands) and Paleontological Museum, University of Oslo, Oslo, 351 pp." type="book" year="1995">Van de Paverd, 1995</bibRefCitation>
</taxonomicName>
, pl. 65, figs. 56, 11.
</paragraph>
<paragraph id="8BDFDE46FFADFFC675DFE3FCFC3CC4DF" blockId="62.[151,1024,1839,1986]" box="[151,1019,1931,1955]" pageId="62" pageNumber="63">
<taxonomicName id="4C60A5C5FFADFFC675DFE3FCFD02C4DF" authority="Haeckel, Van de Paverd, 1995" authorityName="Haeckel, Van de Paverd" authorityYear="1995" box="[151,709,1931,1955]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="62" pageNumber="63" phylum="Radiozoa" rank="species" species="buetschlii">
<emphasis id="B9140254FFADFFC675DFE3FCFE4BC4DF" box="[151,396,1931,1955]" italics="true" pageId="62" pageNumber="63">Lophophaena buetschlii</emphasis>
Haeckel,
<bibRefCitation id="EFF1A3B7FFADFFC674B9E3FCFD02C4DF" author="Van de Paverd, P. J." box="[497,709,1931,1955]" pageId="62" pageNumber="63" refId="ref96419" refString="Van de Paverd, P. J. (1995) Recent Polycystine Radiolaria from the Snellius-II Expedition. Ph. D. thesis, Center for Marine Earth Science (the Netherlands) and Paleontological Museum, University of Oslo, Oslo, 351 pp." type="book" year="1995">Van de Paverd, 1995</bibRefCitation>
</taxonomicName>
, pl. 65, figs. 34 (non fig. 10).
</paragraph>
<paragraph id="8BDFDE46FFADFFC675DFE3DEFBC7C4BD" blockId="62.[151,1024,1839,1986]" box="[151,1024,1961,1986]" pageId="62" pageNumber="63">
<emphasis id="B9140254FFADFFC675DFE3DEFE4EC4BD" box="[151,393,1961,1985]" italics="true" pageId="62" pageNumber="63">Lophophaena nadezdae</emphasis>
Petrushevskaya, Trubovitz
<emphasis id="B9140254FFADFFC677EAE3DDFD1CC4BD" box="[674,731,1961,1985]" italics="true" pageId="62" pageNumber="63">et al.,</emphasis>
2020, supplementary data 7.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFACFFC775DFE4E1FCECC219" pageId="63" pageNumber="64" type="discussion">
<paragraph id="8BDFDE46FFACFFC775DFE4E1FCECC219" blockId="63.[151,1437,150,393]" pageId="63" pageNumber="64">
<emphasis id="B9140254FFACFFC775DFE4E1FECCC3CC" bold="true" box="[151,267,150,176]" pageId="63" pageNumber="64">Remarks.</emphasis>
Despite its abundance in our samples, this species is not commonly cited in the literature. Following
<bibRefCitation id="EFF1A3B7FFACFFC775DFE4CDFE5DC3A8" author="Petrushevskaya, M. G." box="[151,410,186,212]" pageId="63" pageNumber="64" pagination="1 - 294" refId="ref93469" refString="Petrushevskaya, M. G. (1971) Nassellarian radiolarians in the plankton of the world oceans. Investigations of the Fauna of the Seas, 9 (17), 1 - 294. [in Russian]" type="journal article" year="1971">Petrushevskaya (1971)</bibRefCitation>
, we consider this species to have an elongated spiny cephalis with indistinct neck area, and a broad thorax (~2x the cephalis width) with numerous ribs and extended spines. Illustrations published under the names
<taxonomicName id="4C60A5C5FFACFFC775ACE575FCDEC260" authority="Ehrenberg (Benson, 1966)" authorityName="Ehrenberg (Benson" authorityYear="1966" box="[228,793,258,284]" class="Polycystina" family="Plagiacanthidae" genus="Lithomelissa" isUncertain="true" kingdom="Chromista" order="Nassellaria" pageId="63" pageNumber="64" phylum="Radiozoa" rank="species" species="galeata">
<emphasis id="B9140254FFACFFC775ACE575FEB4C260" box="[228,371,258,284]" italics="true" pageId="63" pageNumber="64">Lithomelissa</emphasis>
cf.
<emphasis id="B9140254FFACFFC774D2E575FE29C260" box="[410,494,258,284]" italics="true" pageId="63" pageNumber="64">galeata</emphasis>
Ehrenberg (
<bibRefCitation id="EFF1A3B7FFACFFC7773CE575FCD6C260" author="Benson, R. N." box="[628,785,258,284]" pageId="63" pageNumber="64" refId="ref85564" refString="Benson, R. N. (1966) Recent Radiolaria from the Gulf of California. PhD dissertation, University of Minnesota, Minneapolis, 577 pp." type="book" year="1966">Benson, 1966</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C60A5C5FFACFFC7766BE575FA50C261" authority="Popofsky (Van de Paverd, 1995)" authorityName="Popofsky (Van de Paverd" authorityYear="1995" box="[803,1431,258,285]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="63" pageNumber="64" phylum="Radiozoa" rank="species" species="variabilis">
<emphasis id="B9140254FFACFFC7766BE575FBEAC260" box="[803,1069,258,284]" italics="true" pageId="63" pageNumber="64">Lophophaena variabilis</emphasis>
Popofsky (
<bibRefCitation id="EFF1A3B7FFACFFC771E0E575FA48C261" author="Van de Paverd, P. J." box="[1192,1423,258,285]" pageId="63" pageNumber="64" refId="ref96419" refString="Van de Paverd, P. J. (1995) Recent Polycystine Radiolaria from the Snellius-II Expedition. Ph. D. thesis, Center for Marine Earth Science (the Netherlands) and Paleontological Museum, University of Oslo, Oslo, 351 pp." type="book" year="1995">Van de Paverd, 1995</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C60A5C5FFACFFC7758DE551FCE8C23D" authority="Haeckel (Van de Paverd, 1995)" authorityName="Haeckel (Van de Paverd" authorityYear="1995" box="[197,815,294,321]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="63" pageNumber="64" phylum="Radiozoa" rank="species" species="buetschlii">
<emphasis id="B9140254FFACFFC7758DE551FE15C23C" box="[197,466,294,320]" italics="true" pageId="63" pageNumber="64">Lophophaena buetschlii</emphasis>
Haeckel (
<bibRefCitation id="EFF1A3B7FFACFFC77708E551FCE0C23D" author="Van de Paverd, P. J." box="[576,807,294,321]" pageId="63" pageNumber="64" refId="ref96419" refString="Van de Paverd, P. J. (1995) Recent Polycystine Radiolaria from the Snellius-II Expedition. Ph. D. thesis, Center for Marine Earth Science (the Netherlands) and Paleontological Museum, University of Oslo, Oslo, 351 pp." type="book" year="1995">Van de Paverd, 1995</bibRefCitation>
)
</taxonomicName>
best fit the description of
<taxonomicName id="4C60A5C5FFACFFC77105E551FA91C23C" authorityName="Petrushevskaya" authorityYear="1971" box="[1101,1366,294,320]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="63" pageNumber="64" phylum="Radiozoa" rank="species" species="nadezdae">
<emphasis id="B9140254FFACFFC77105E551FA91C23C" box="[1101,1366,294,320]" italics="true" pageId="63" pageNumber="64">Lophophaena nadezdae</emphasis>
</taxonomicName>
in our opinion, so these specimens are included in our synonymy.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFACFFC7758FE519FD1FC2F4" box="[199,728,366,393]" pageId="63" pageNumber="64" type="biology_ecology">
<paragraph id="8BDFDE46FFACFFC7758FE519FD1FC2F4" blockId="63.[151,1437,150,393]" box="[199,728,366,393]" pageId="63" pageNumber="64">
<emphasis id="B9140254FFACFFC7758FE519FEDDC2F4" bold="true" box="[199,282,366,392]" pageId="63" pageNumber="64">Range.</emphasis>
Late Miocene—Recent, EEP (
<tableCitation id="C6E2EBFDFFACFFC7773BE519FD00C2F4" box="[627,711,366,392]" captionStart="TABLE 1" captionStartId="84.[151,248,834,860]" captionTargetPageId="84" captionText="TABLE 1. Occurrence chart of Late NeogeneRecent lophophaenids in the EEP." httpUri="http://table.plazi.org/id/DF1F8ECEFFC7FFAC75DFE735FB46C021" pageId="63" pageNumber="64" tableUuid="DF1F8ECEFFC7FFAC75DFE735FB46C021">Table 1</tableCitation>
).
</paragraph>
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