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<document id="0B0C96440FD07DC10D857F3D9A229135" ID-CLB-Dataset="287311" ID-DOI="https://doi.org/10.5852/ejt.2023.881.2175" ID-GBIF-Dataset="225ee515-e526-4cc0-b86b-85e8f2e4c83a" ID-ISSN="2118-9773" ID-Zenodo-Dep="8178280" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="admin" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="admin" checkinTime="1690201937279" checkinUser="felipe" docAuthor="Moonlight, Peter. W., Jara-Muñoz, Orlando A., Purvis, David A., Delves, Jay, Allen, Josh P. &amp; Reynel, Carlos" docDate="2023" docId="03B26B4BFEB6FEEFFE24F9EDAC54EFF4" docLanguage="en" docName="ejt-881_1-334.imf" docOrigin="European Journal of Taxonomy 881" docSource="http://dx.doi.org/10.5852/ejt.2023.881.2175" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia yuracyacuensis Moonlight 2023, sp. nov." docType="treatment" docVersion="14" lastPageNumber="279" masterDocId="FF8B1333FFA5FFF9FFD9FFBFAD27EA79" masterDocTitle="The genus Begonia (Begoniaceae) in Peru" masterLastPageNumber="334" masterPageNumber="1" pageNumber="276" updateTime="1732576113334" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C1A09774342E335A7FC44850BC829C64">The genus Begonia (Begoniaceae) in Peru</mods:title>
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<mods:namePart id="648F4346782F382267CC422914C4C148">Moonlight, Peter. W.</mods:namePart>
<mods:affiliation id="3B7112D56CFF2324FD88B33E66BDD685">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK. &amp; Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland.</mods:affiliation>
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<mods:namePart id="C4C6783AAACDB3635C8C0C8DB5570FA7">Jara-Muñoz, Orlando A.</mods:namePart>
<mods:affiliation id="1C83D332CA4882FD4545D06D1FC8F0ED">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá D. C., Colombia.</mods:affiliation>
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<mods:namePart id="507DC5A499DF62AFFF2E2DBE0B241543">Purvis, David A.</mods:namePart>
<mods:affiliation id="8C3369ABD94F2C19C0959DB14158EADB">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
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<mods:namePart id="39B5CD1B1FDA312B88B216225F6FDCC6">Delves, Jay</mods:namePart>
<mods:affiliation id="36C7B1EE86BABB19EA395F911C0D2CBF">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh, EH 3 5 LR, Scotland, UK.</mods:affiliation>
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<mods:nameIdentifier id="4295C06A20A79B32373927161AB5D873" type="email">myrmeciaman@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9C6101F0F11E3E9E9848905A05318EAA">Reynel, Carlos</mods:namePart>
<mods:affiliation id="786A5B2A35C1229D511BEC8B1F058153">Facultad de Ciencias Forestales, Universidad Nacional Agraria-La Molina, Lima 12, Peru.</mods:affiliation>
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<treatment id="03B26B4BFEB6FEEFFE24F9EDAC54EFF4" ID-DOI="http://doi.org/10.5281/zenodo.11978166" ID-GBIF-Taxon="221057797" ID-Zenodo-Dep="11978166" LSID="urn:lsid:plazi:treatment:03B26B4BFEB6FEEFFE24F9EDAC54EFF4" httpUri="http://treatment.plazi.org/id/03B26B4BFEB6FEEFFE24F9EDAC54EFF4" lastPageId="278" lastPageNumber="279" pageId="275" pageNumber="276" scope_class="Equisetopsida" scope_family="Begoniaceae" scope_order="Cucurbitales" scope_subClass="Magnoliidae" scope_superOrder="Rosanae">
<subSubSection id="C30189D6FEB6FEEAFE24F9EDA910EC14" box="[509,1079,1618,1645]" pageId="275" pageNumber="276" type="nomenclature">
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69.
<taxonomicName id="4C1BA1DEFEB6FEEAFDF0F9EDAEF2EC15" ID-CoL="CFGWK" authority="Moonlight" authorityName="Moonlight" authorityYear="2023" box="[553,981,1618,1644]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="275" pageNumber="276" phylum="Tracheophyta" rank="species" species="yuracyacuensis" status="sp. nov.">
<emphasis id="B96F064FFEB6FEEAFDF0F9EDAE75EC15" bold="true" box="[553,850,1618,1644]" italics="true" pageId="275" pageNumber="276">Begonia yuracyacuensis</emphasis>
Moonlight
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<taxonomicNameLabel id="A25CBB34FEB6FEEAFC04F9ECA910EC14" box="[989,1079,1619,1645]" pageId="275" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C30189D6FEB6FEEAFD9EF9CAAE59ECCA" pageId="275" pageNumber="276" type="description">
<paragraph id="8BA4DA5DFEB6FEEAFD9EF9CAAECAECF6" blockId="275.[509,1079,1618,1715]" box="[583,1005,1653,1679]" pageId="275" pageNumber="276">urn:lsid:ipni.org:names:77323297-1</paragraph>
<paragraph id="8BA4DA5DFEB6FEEAFD6CF927AE59ECCA" blockId="275.[509,1079,1618,1715]" box="[693,894,1688,1715]" pageId="275" pageNumber="276">
<figureCitation id="1320C6D8FEB6FEEAFD6CF927AE34ECCB" box="[693,787,1688,1714]" captionStart="Fig" captionStartId="11.[189,232,1822,1848]" captionTargetBox="[189,1398,915,1782]" captionTargetId="figure-11@11.[189,1398,915,1782]" captionTargetPageId="11" captionText="Fig. 6. Styles and stigmas in Peruvian Begonia. AC. Bifid styles. D. Reniform styles. EF. Irregularly multifid styles. A. Three styles and stigmas of B. bracteosa A.DC. B. Three styles and stigmas of B. glabra Aubl. C. Single style and stigma of B. yuracyacuensis Moonlight sp. nov. D. Three styles and stigmas of B. pedemontana Moonlight sp. nov. E. Three styles and stigmas of B. obtecticaulis Irmsch. F. Single styles and stigma of B. pleiopetala A.DC. Photographs taken by D.A. Purvis &amp; P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh." figureDoi="http://doi.org/10.5281/zenodo.8178292" httpUri="https://zenodo.org/record/8178292/files/figure.png" pageId="275" pageNumber="276">Figs 6C</figureCitation>
,
<figureCitation id="1320C6D8FEB6FEEAFCF9F926AE72ECCA" box="[800,853,1689,1715]" captionStart="Fig" captionStartId="263.[189,232,1857,1883]" captionTargetBox="[194,1392,1194,1813]" captionTargetId="figure-326@263.[189,1398,1184,1819]" captionTargetPageId="263" captionText="Fig. 84. Distribution of Begonia sect. Ruizopavonia A.DC. in Peru and surrounding countries. A. B. glauca (Klotzsch) Ruiz &amp; Pav. ex A.DC. (red) and B. yuracyacuensis Moonlight sp. nov. (blue). B. B. nunezii Moonlight sp. nov. (red) and B. peruviana A.DC. (blue). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178482" httpUri="https://zenodo.org/record/8178482/files/figure.png" pageId="275" pageNumber="276">84A</figureCitation>
,
<figureCitation id="1320C6D8FEB6FEEAFCB8F926AE59ECCA" box="[865,894,1689,1715]" captionStart="Fig" captionStartId="277.[189,232,1822,1848]" captionTargetBox="[257,1330,265,1783]" captionTargetId="figure-11@277.[257,1330,265,1783]" captionTargetPageId="277" captionText="Fig. 87. Begonia yuracyacuensis Moonlight sp. nov. A. Habit. B. Stipule. C. Leaf, adaxial surface. D. Leaf, abaxial surface. E. Shade leaf, adaxial surface. F. Cross section of ovary. G. Pistillate flower, side view. H. Pistillate flower, front view. I. Pistils, side view. J. Bracteole. K. Staminate flower, side view. L. Staminate flower, front view. M. Androecium, side view. Photographs taken by D.A. Purvis and P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh (Accession 20160139, grown from seeds collected as part of P.W. Moonlight &amp; A. Daza 199)." figureDoi="http://doi.org/10.5281/zenodo.8178489" httpUri="https://zenodo.org/record/8178489/files/figure.png" pageId="275" pageNumber="276">87</figureCitation>
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<emphasis id="B96F064FFEB6FEEAFF64F965AC10EC8D" bold="true" box="[189,311,1754,1780]" pageId="275" pageNumber="276">Diagnosis</emphasis>
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Most similar to
<taxonomicName id="4C1BA1DEFEB6FEEAFE58F8B6AF39ED5B" baseAuthorityName="de Candolle" baseAuthorityYear="1859" box="[385,542,1801,1826]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="275" pageNumber="276" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B96F064FFEB6FEEAFE58F8B6AF39ED5B" box="[385,542,1801,1826]" italics="true" pageId="275" pageNumber="276">B. peruviana</emphasis>
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but differing in its inflorescences that branch up to 4 times (vs 6 to 8 times); its subequal, obdeltoid fruit wings with straight sides (vs unequal wings, the largest obdeltoid with rounded sides, the smallest ridgelike); its broadly ovate bracts and bracteoles (vs elliptic bracts and linear to ovate bracteoles); its larger staminate tepals (1014 ×
<quantity id="4CE377B8FEB6FEEAFC4EF8CDA932EDF4" box="[919,1045,1906,1933]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="275" pageNumber="276" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
vs 25.5 ×
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); and its styles, which are thickened at the bifurcation (vs not thickened).
</paragraph>
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<emphasis id="B96F064FFEB1FEEDFF64FEB6AC62EB5A" bold="true" box="[189,325,265,291]" pageId="276" pageNumber="277">Etymology</emphasis>
</heading>
</paragraph>
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The name refers to the río Yuracyacu, which flows through the
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de Padre Abad. This means the white river in Quechua.
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<typeStatus id="54A064FFFEB1FEEDFF64FE22ADDCEBCE" box="[189,251,413,439]" pageId="276" pageNumber="277">Type</typeStatus>
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<materialsCitation id="3B73D000FEB1FEEDFF64FE73AF35E854" ID-GBIF-Occurrence="4517462470" collectingDate="1962-08-01" collectionCode="USM, MO" collectorName="M. E. Mathias &amp; D. Taylor" country="Peru" county="Prov. Coronel Portillo" elevation="300" latitude="-9.066667" location="Boqueron de Padre Abad" longLatPrecision="1300" longitude="-75.683334" municipality="Vicinity of Aguaytia" pageId="276" pageNumber="277" specimenCode="MO-1641350" specimenCount="1" stateProvince="San Martin Region" typeStatus="holotype">
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:
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<collectingMunicipality id="6BC04027FEB1FEEDFC89FE74A966EB9F" box="[848,1089,459,486]" pageId="276" pageNumber="277">Vicinity of Aguaytía</collectingMunicipality>
,
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; [
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,
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a.s.l.
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;
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<collectorName id="26EEBF8BFEB1FEEDFCC3FE4FAE98E870" box="[794,959,495,521]" pageId="276" pageNumber="277">M.E. Mathias</collectorName>
&amp;
<collectorName id="26EEBF8BFEB1FEEDFC32FE4FA979E870" box="[1003,1118,495,521]" pageId="276" pageNumber="277">D. Taylor</collectorName>
6096
</emphasis>
;
<typeStatus id="54A064FFFEB1FEEDFB6FFE50A806E870" box="[1206,1313,495,521]" pageId="276" pageNumber="277">holotype</typeStatus>
:
<collectionCode id="ED0A4298FEB1FEEDFAF6FE50A855E870" box="[1327,1394,495,521]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="276" pageNumber="277">USM</collectionCode>
; isotype:
<collectionCode id="ED0A4298FEB1FEEDFEF8FDACAC77E854" box="[289,336,531,557]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="276" pageNumber="277" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFEBBFDADAF25E854" box="[354,514,530,557]" pageId="276" pageNumber="277">MO-1641350</specimenCode>
].
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</paragraph>
<paragraph id="8BA4DA5DFEB1FEEDFF64FDEBACE3E817" blockId="276.[189,452,596,622]" box="[189,452,596,622]" pageId="276" pageNumber="277">
<heading id="D0EC6D31FEB1FEEDFF64FDEBACE3E817" bold="true" box="[189,452,596,622]" fontSize="11" level="3" pageId="276" pageNumber="277" reason="3">
<emphasis id="B96F064FFEB1FEEDFF64FDEBACE3E817" bold="true" box="[189,452,596,622]" pageId="276" pageNumber="277">Specimens examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFEB1FEEDFF64FD3BA9F3EE31" blockId="276.[187,1399,642,1096]" pageId="276" pageNumber="277">
<materialsCitation id="3B73D000FEB1FEEDFF64FD3BAFEEE89C" ID-GBIF-Occurrence="4517462491" collectingDate="1996-11-01" collectionCode="CPUN, F, US" collectorName="I. Sanchez Vega &amp; M. O. Dillon" country="Peru" county="Prov. Rioja" elevation="890" latitude="-5.9" location="cerca del Poblado Palestina" longLatPrecision="1305" longitude="-77.35" municipality="Dist. Nuevo Cajamarca" pageId="276" pageNumber="277" specimenCode="V0086778F, US00673143" specimenCount="1" stateProvince="San Martin Region">
<collectingCountry id="F30C9ACDFEB1FEEDFF64FD3BAC2FE8E7" box="[189,264,644,670]" name="Peru" pageId="276" pageNumber="277">PERU</collectingCountry>
<emphasis id="B96F064FFEB1FEEDFEC7FD3CAF8FE8E4" bold="true" box="[286,680,642,669]" pageId="276" pageNumber="277">
<collectingRegion id="49DF14BFFEB1FEEDFEC7FD3CAF37E8E4" box="[286,528,642,669]" country="Peru" name="San Martin" pageId="276" pageNumber="277">San Martín Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FEB1FEEDFDC2FD3CAF8FE8E4" box="[539,680,643,669]" pageId="276" pageNumber="277">Prov. Rioja</collectingCounty>
</emphasis>
<collectingMunicipality id="6BC04027FEB1FEEDFD67FD3CAEF7E8E4" box="[702,976,643,670]" pageId="276" pageNumber="277">Dist. Nuevo Cajamarca</collectingMunicipality>
,
<location id="8EC48C86FEB1FEEDFC00FD3BA830E8E4" LSID="urn:lsid:plazi:treatment:03B26B4BFEB6FEEFFE24F9EDAC54EFF4:8EC48C86FEB1FEEDFC00FD3BA830E8E4" box="[985,1303,643,670]" country="Peru" county="Prov. Rioja" latitude="-5.9" longLatPrecision="1305" longitude="-77.35" municipality="Dist. Nuevo Cajamarca" name="cerca del Poblado Palestina" pageId="276" pageNumber="277" stateProvince="San Martin Region">cerca del Poblado Palestina</location>
,
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;
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,
<geoCoordinate id="EE2FBC9AFEB1FEEDFDCFFD1AAFAFE8B8" box="[534,648,677,705]" degrees="77" direction="west" minutes="21" orientation="longitude" pageId="276" pageNumber="277" precision="925" value="-77.35">77°21 W</geoCoordinate>
;
<elevation id="00363D6EFEB1FEEDFD4AFD18AE3DE8B9" box="[659,794,678,705]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="276" pageNumber="277" unit="m" value="890.0">
<quantity id="4CE377B8FEB1FEEDFD4AFD18AFFBE8B8" box="[659,732,679,705]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="276" pageNumber="277" unit="m" value="890.0">890 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFEB1FEEDFCFEFD18AE90E8B9" box="[807,951,678,705]" pageId="276" pageNumber="277" value="1996-11-01">
<collectingDate id="EFE10575FEB1FEEDFCFEFD18AE90E8B9" box="[807,951,678,705]" pageId="276" pageNumber="277" value="1996-11-01">1 Nov. 1996</collectingDate>
</date>
;
<emphasis id="B96F064FFEB1FEEDFC18FD18A84AE8B8" box="[961,1389,678,705]" italics="true" pageId="276" pageNumber="277">
<collectorName id="26EEBF8BFEB1FEEDFC18FD18A95EE8B9" box="[961,1145,678,704]" pageId="276" pageNumber="277">I. Sánchez Vega</collectorName>
&amp;
<collectorName id="26EEBF8BFEB1FEEDFB44FD18A80BE8B9" box="[1181,1324,678,705]" pageId="276" pageNumber="277">M.O. Dillon</collectorName>
8389
</emphasis>
;
<collectionCode id="ED0A4298FEB1FEEDFF64FD75AC29E89D" box="[189,270,714,740]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14355" lsid="urn:lsid:biocol.org:col:14355" name="Universidad Nacional de Cajamarca" pageId="276" pageNumber="277" type="Herbarium">CPUN</collectionCode>
,
<collectionCode id="ED0A4298FEB1FEEDFEC2FD74AC0CE89C" box="[283,299,715,741]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="276" pageNumber="277" type="Herbarium">F</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFEE7FD75ACEDE89D" box="[318,458,714,740]" pageId="276" pageNumber="277">V0086778F</specimenCode>
],
<collectionCode id="ED0A4298FEB1FEEDFE3DFD75AF2EE89D" box="[484,521,714,740]" pageId="276" pageNumber="277">US</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFDC4FD75AF9FE89D" box="[541,696,714,740]" pageId="276" pageNumber="277">US00673143</specimenCode>
].
</materialsCitation>
<materialsCitation id="3B73D000FEB1FEEDFD33FD75AE67E952" ID-GBIF-Occurrence="4517462700" collectingDate="1946-08-08" collectionCode="G, MO" collectorName="G. Woytkowski" county="Prov. Coronel Portillo" elevation="300" latitude="-9.066667" location="Boqueron de Padre Abad" longLatPrecision="1300" longitude="-75.683334" municipality="Dist. Padre Abad" pageId="276" pageNumber="277" specimenCode="MO-1641413" specimenCount="1" stateProvince="Ucayali Region">
<emphasis id="B96F064FFEB1FEEDFD33FD75A9F0E89D" bold="true" box="[746,1239,714,740]" pageId="276" pageNumber="277">
<collectingRegion id="49DF14BFFEB1FEEDFD33FD75AE97E89D" box="[746,944,714,740]" pageId="276" pageNumber="277">Ucayali Region</collectingRegion>
:
<collectingCounty id="62C5A2D1FEB1FEEDFC64FD75A9F0E89D" box="[957,1239,714,740]" pageId="276" pageNumber="277">Prov. Coronel Portillo</collectingCounty>
</emphasis>
<collectingMunicipality id="6BC04027FEB1FEEDFB2AFD75AC25E97E" pageId="276" pageNumber="277">Dist. Padre Abad</collectingMunicipality>
,
<location id="8EC48C86FEB1FEEDFED4FD51AF1FE97E" LSID="urn:lsid:plazi:treatment:03B26B4BFEB6FEEFFE24F9EDAC54EFF4:8EC48C86FEB1FEEDFED4FD51AF1FE97E" box="[269,568,749,776]" country="Palestine" county="Prov. Coronel Portillo" latitude="-9.066667" longLatPrecision="1300" longitude="-75.683334" municipality="Dist. Padre Abad" name="Boqueron de Padre Abad" pageId="276" pageNumber="277" stateProvince="Ucayali Region">Boquerón de Padre Abad</location>
;
<geoCoordinate id="EE2FBC9AFEB1FEEDFD9CFD53AFBCE971" box="[581,667,748,776]" degrees="9" direction="south" minutes="04" orientation="latitude" pageId="276" pageNumber="277" precision="925" value="-9.066667">9°04 S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFEB1FEEDFD70FD53AE39E971" box="[681,798,748,776]" degrees="75" direction="west" minutes="41" orientation="longitude" pageId="276" pageNumber="277" precision="925" value="-75.683334">75°41 W</geoCoordinate>
];
<elevation id="00363D6EFEB1FEEDFCEBFD51AE9AE97E" box="[818,957,749,776]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="276" pageNumber="277" unit="m" value="300.0">
<quantity id="4CE377B8FEB1FEEDFCEBFD51AE5AE971" box="[818,893,750,776]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="276" pageNumber="277" unit="m" value="300.0">300 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFEB1FEEDFC12FD51A947E97E" box="[971,1120,749,776]" pageId="276" pageNumber="277" value="1946-08-08">
<collectingDate id="EFE10575FEB1FEEDFC12FD51A947E97E" box="[971,1120,749,776]" pageId="276" pageNumber="277" value="1946-08-08">8 Aug. 1946</collectingDate>
</date>
;
<emphasis id="B96F064FFEB1FEEDFBB4FD51A849E971" box="[1133,1390,749,776]" italics="true" pageId="276" pageNumber="277">
<collectorName id="26EEBF8BFEB1FEEDFBB4FD51A83BE97E" box="[1133,1308,749,776]" pageId="276" pageNumber="277">G. Woytkowski</collectorName>
34341
</emphasis>
;
<collectionCode id="ED0A4298FEB1FEEDFF64FCAEADF5E952" box="[189,210,785,811]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="276" pageNumber="277" type="Herbarium">G</collectionCode>
[2: 1 mixed sheet],
<collectionCode id="ED0A4298FEB1FEEDFE62FCAEACCDE952" box="[443,490,785,811]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="276" pageNumber="277" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFE22FCAEAFBCE952" box="[507,667,785,811]" pageId="276" pageNumber="277">MO-1641413</specimenCode>
, mixed sheet]
</materialsCitation>
<materialsCitation id="3B73D000FEB1FEEDFC81FCAEAEB2E936" ID-GBIF-Occurrence="4517461969" collectingDate="1980-09-10" collectionCode="U, US, USM" collectorName="P. J. M. Maas &amp; L. Cobb &amp; E. H. M. Grothe" county="Prov. Coronel Portillo" elevation="500" latitude="-9.066667" location="Boqueron de Padre Abad" longLatPrecision="1300" longitude="-75.683334" municipality="Dist. Padre Abad" pageId="276" pageNumber="277" specimenCode="US00967205" specimenCount="1" stateProvince="Ucayali Region">
ibid.;
<elevation id="00363D6EFEB1FEEDFC45FCAEA901E952" box="[924,1062,785,812]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="276" pageNumber="277" unit="m" value="500.0">
<quantity id="4CE377B8FEB1FEEDFC45FCAEAEC1E955" box="[924,998,785,812]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="276" pageNumber="277" unit="m" value="500.0">500 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFEB1FEEDFBEAFCAEA9F7E952" box="[1075,1232,785,811]" pageId="276" pageNumber="277" value="1980-09-10">
<collectingDate id="EFE10575FEB1FEEDFBEAFCAEA9F7E952" box="[1075,1232,785,811]" pageId="276" pageNumber="277" value="1980-09-10">10 Sep. 1980</collectingDate>
</date>
;
<emphasis id="B96F064FFEB1FEEDFB02FCADAF10E936" italics="true" pageId="276" pageNumber="277">
<collectorName id="26EEBF8BFEB1FEEDFB02FCADA855E952" box="[1243,1394,786,811]" pageId="276" pageNumber="277">P.J.M. Maas</collectorName>
,
<collectorName id="26EEBF8BFEB1FEEDFF64FC8AAC3BE937" box="[189,284,820,846]" pageId="276" pageNumber="277">L. Cobb</collectorName>
&amp;
<collectorName id="26EEBF8BFEB1FEEDFE9AFC8AACD3E937" box="[323,500,820,846]" pageId="276" pageNumber="277">E.H.M. Grothe</collectorName>
4574
</emphasis>
;
<collectionCode id="ED0A4298FEB1FEEDFD9EFC8AAF78E936" box="[583,607,821,847]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15513" name="Nationaal Herbarium Nederland" pageId="276" pageNumber="277" type="Herbarium">U</collectionCode>
,
<collectionCode id="ED0A4298FEB1FEEDFDB2FC8AAFB7E936" box="[619,656,821,847]" pageId="276" pageNumber="277">US</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFD7BFC8BAE1AE936" box="[674,829,820,847]" pageId="276" pageNumber="277">US00967205</specimenCode>
],
<collectionCode id="ED0A4298FEB1FEEDFC8CFC8AAEB2E936" box="[853,917,821,847]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="276" pageNumber="277">USM</collectionCode>
</materialsCitation>
<materialsCitation id="3B73D000FEB1FEEDFC77FC8BAFF9E90B" ID-GBIF-Occurrence="4517461527" collectingDate="1946-08-17" collectionCode="MO, US, USM" collectorName="R. Ferreyra" county="Prov. Coronel Portillo" elevation="500" latitude="-9.066667" location="Boqueron de Padre Abad" longLatPrecision="1300" longitude="-75.683334" municipality="Dist. Padre Abad" pageId="276" pageNumber="277" specimenCode="MO-2264390, US00967391" specimenCount="1" stateProvince="Ucayali Region">
ibid.;
<date id="FFA5FC9DFEB1FEEDFC2AFC8AA9B1E937" box="[1011,1174,820,847]" pageId="276" pageNumber="277" value="1946-08-17">
<collectingDate id="EFE10575FEB1FEEDFC2AFC8AA9B1E937" box="[1011,1174,820,847]" pageId="276" pageNumber="277" value="1946-08-17">17 Aug. 1946</collectingDate>
</date>
;
<emphasis id="B96F064FFEB1FEEDFB7BFC8AA849E936" box="[1186,1390,821,847]" italics="true" pageId="276" pageNumber="277">
<collectorName id="26EEBF8BFEB1FEEDFB7BFC8AA809E937" box="[1186,1326,821,846]" pageId="276" pageNumber="277">R. Ferreyra</collectorName>
1147
</emphasis>
;
<collectionCode id="ED0A4298FEB1FEEDFF64FCE7ADCBE90B" box="[189,236,856,882]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="276" pageNumber="277" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFF24FCE7ACBBE90B" box="[253,412,856,882]" pageId="276" pageNumber="277">MO-2264390</specimenCode>
],
<collectionCode id="ED0A4298FEB1FEEDFE6CFCE7ACFDE90B" box="[437,474,856,882]" pageId="276" pageNumber="277">US</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFE32FCE7AFA1E90B" box="[491,646,856,882]" pageId="276" pageNumber="277">US00967391</specimenCode>
],
<collectionCode id="ED0A4298FEB1FEEDFD47FCE7AFF9E90B" box="[670,734,856,882]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="276" pageNumber="277">USM</collectionCode>
</materialsCitation>
<materialsCitation id="3B73D000FEB1FEEDFD2FFCE7AE71E9EF" ID-GBIF-Occurrence="4517461678" collectingDate="1950-10-02" collectionCode="MO, MOL, US, USM" collectorName="R. Ferreyra" county="Prov. Coronel Portillo" elevation="525" latitude="-9.066667" location="Boqueron de Padre Abad" longLatPrecision="1300" longitude="-75.683334" municipality="Dist. Padre Abad" pageId="276" pageNumber="277" specimenCode="MO-2264395, US00967204" specimenCount="1" stateProvince="Ucayali Region">
ibid.;
<elevation id="00363D6EFEB1FEEDFCE3FCE7AED9E90B" box="[826,1022,856,883]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="276" pageNumber="277" unit="m" value="525.0" valueMax="550.0" valueMin="500.0">
<quantity id="4CE377B8FEB1FEEDFCE3FCE7AE98E90A" box="[826,959,856,883]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="276" pageNumber="277" unit="m" value="525.0" valueMax="550.0" valueMin="500.0">500550 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFEB1FEEDFBD2FCE7A9BFE90A" box="[1035,1176,856,883]" pageId="276" pageNumber="277" value="1950-10-02">
<collectingDate id="EFE10575FEB1FEEDFBD2FCE7A9BFE90A" box="[1035,1176,856,883]" pageId="276" pageNumber="277" value="1950-10-02">2 Oct. 1950</collectingDate>
</date>
;
<emphasis id="B96F064FFEB1FEEDFB7AFCE6A849E90B" box="[1187,1390,856,882]" italics="true" pageId="276" pageNumber="277">
<collectorName id="26EEBF8BFEB1FEEDFB7AFCE6A809E90B" box="[1187,1326,857,882]" pageId="276" pageNumber="277">R. Ferreyra</collectorName>
8110
</emphasis>
;
<collectionCode id="ED0A4298FEB1FEEDFF64FCC3ADCBE9EF" box="[189,236,892,918]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="276" pageNumber="277" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFF24FCC4ACBAE9EF" box="[253,413,891,918]" pageId="276" pageNumber="277">MO-2264395</specimenCode>
],
<collectionCode id="ED0A4298FEB1FEEDFE6CFCC3ACDEE9EF" box="[437,505,892,918]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="276" pageNumber="277" type="Herbarium">MOL</collectionCode>
,
<collectionCode id="ED0A4298FEB1FEEDFDDDFCC3AF0EE9EF" box="[516,553,892,918]" pageId="276" pageNumber="277">US</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFDE2FCC4AFF2E9EC" box="[571,725,891,917]" pageId="276" pageNumber="277">US00967204</specimenCode>
],
<collectionCode id="ED0A4298FEB1FEEDFD37FCC3AE09E9EF" box="[750,814,892,918]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="276" pageNumber="277">USM</collectionCode>
[2]
</materialsCitation>
<materialsCitation id="3B73D000FEB1FEEDFCB6FCC4A90EE9C0" ID-GBIF-Occurrence="4517462168" collectingDate="2016-02-08" collectionCode="E, MO, MOL" collectorName="P. W. Moonlight &amp; A. Daza" county="Prov. Coronel Portillo" elevation="403" latitude="-9.066944" location="Boqueron de Padre Abad" longLatPrecision="21" longitude="-75.67944" municipality="Dist. Padre Abad" pageId="276" pageNumber="277" specimenCode="E00885477" specimenCount="1" stateProvince="Ucayali Region">
ibid.;
<geoCoordinate id="EE2FBC9AFEB1FEEDFC6AFCC3A913E9EF" box="[947,1076,890,918]" degrees="9" direction="south" minutes="04" orientation="latitude" pageId="276" pageNumber="277" precision="15" seconds="01" value="-9.066944">9°0401″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFEB1FEEDFB99FCC5A9FCE9EF" box="[1088,1243,890,918]" degrees="75" direction="west" minutes="40" orientation="longitude" pageId="276" pageNumber="277" precision="15" seconds="46" value="-75.67944">75°4046″ W</geoCoordinate>
;
<elevation id="00363D6EFEB1FEEDFB3EFCC3A857E9EC" box="[1255,1392,891,918]" metricMagnitude="2" metricUnit="m" metricValue="4.029999999999999" pageId="276" pageNumber="277" unit="m" value="403.0">
<quantity id="4CE377B8FEB1FEEDFB3EFCC3A816E9EF" box="[1255,1329,892,918]" metricMagnitude="2" metricUnit="m" metricValue="4.029999999999999" pageId="276" pageNumber="277" unit="m" value="403.0">403 m</quantity>
a.s.l.
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;
<date id="FFA5FC9DFEB1FEEDFF64FC20AC68E9C0" box="[189,335,927,953]" pageId="276" pageNumber="277" value="2016-02-08">
<collectingDate id="EFE10575FEB1FEEDFF64FC20AC68E9C0" box="[189,335,927,953]" pageId="276" pageNumber="277" value="2016-02-08">8 Feb. 2016</collectingDate>
</date>
;
<emphasis id="B96F064FFEB1FEEDFE87FC1FAFF4E9C0" box="[350,723,927,953]" italics="true" pageId="276" pageNumber="277">
<collectorName id="26EEBF8BFEB1FEEDFE87FC1FAF32E9C0" box="[350,533,927,953]" pageId="276" pageNumber="277">P.W. Moonlight</collectorName>
&amp;
<collectorName id="26EEBF8BFEB1FEEDFDE7FC20AFB9E9C0" box="[574,670,927,953]" pageId="276" pageNumber="277">A. Daza</collectorName>
199
</emphasis>
;
<collectionCode id="ED0A4298FEB1FEEDFD3DFC1FAFD1E9C3" box="[740,758,928,954]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="276" pageNumber="277" type="Herbarium">E</collectionCode>
[
<specimenCode id="DBBD7226FEB1FEEDFCD1FC20AEB7E9C3" box="[776,912,927,954]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="276" pageNumber="277" type="Herbarium">E00885477</specimenCode>
],
<collectionCode id="ED0A4298FEB1FEEDFC70FC20AEFCE9C0" box="[937,987,927,953]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="276" pageNumber="277" type="Herbarium">MO</collectionCode>
,
<collectionCode id="ED0A4298FEB1FEEDFC30FC20A90EE9C0" box="[1001,1065,927,953]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="276" pageNumber="277" type="Herbarium">MOL</collectionCode>
</materialsCitation>
<materialsCitation id="3B73D000FEB1FEEDFB9CFC20AF1DE9A4" ID-GBIF-Occurrence="4517461703" collectingDate="1946-08-08" collectionCode="USM" collectorName="F. Woytkowski" county="Prov. Coronel Portillo" elevation="470" latitude="-9.066944" location="Boqueron de Padre Abad" longLatPrecision="21" longitude="-75.67944" municipality="Dist. Padre Abad" pageId="276" pageNumber="277" specimenCount="1" stateProvince="Ucayali Region">
ibid.;
<elevation id="00363D6EFEB1FEEDFB52FC20A83FE9C0" box="[1163,1304,927,954]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="276" pageNumber="277" unit="m" value="470.0">
<quantity id="4CE377B8FEB1FEEDFB52FC20A9F0E9C3" box="[1163,1239,927,954]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="276" pageNumber="277" unit="m" value="470.0">470 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFEB1FEEDFAFEFC20ADDBE9A5" pageId="276" pageNumber="277" value="1946-08-08">
<collectingDate id="EFE10575FEB1FEEDFAFEFC20ADDBE9A5" pageId="276" pageNumber="277" value="1946-08-08">8 Aug. 1946</collectingDate>
</date>
;
<emphasis id="B96F064FFEB1FEEDFED0FC7CACCFE9A4" box="[265,488,962,989]" italics="true" pageId="276" pageNumber="277">
<collectorName id="26EEBF8BFEB1FEEDFED0FC7CAC94E9A5" box="[265,435,962,988]" pageId="276" pageNumber="277">F. Woytkowski</collectorName>
341
</emphasis>
;
<collectionCode id="ED0A4298FEB1FEEDFE23FC7CAF1DE9A4" box="[506,570,963,989]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="276" pageNumber="277">USM</collectionCode>
</materialsCitation>
<materialsCitation id="3B73D000FEB1FEEDFD8FFC7DAC4AEE5D" ID-GBIF-Occurrence="4517461502" collectingDate="2007-10-05" collectionCode="MOL, US" collectorName="J. G. Graham" elevation="425" latitude="-9.4372225" location="Cuenca del rio Iparia" longLatPrecision="21" longitude="-74.558334" municipality="Dist. Iparia" pageId="276" pageNumber="277" specimenCode="US01008606" specimenCount="1">
<collectingMunicipality id="6BC04027FEB1FEEDFD8FFC7DAFF8E9A4" box="[598,735,962,989]" pageId="276" pageNumber="277">Dist. Iparía</collectingMunicipality>
,
<location id="8EC48C86FEB1FEEDFD35FC7CAECCE9A4" LSID="urn:lsid:plazi:treatment:03B26B4BFEB6FEEFFE24F9EDAC54EFF4:8EC48C86FEB1FEEDFD35FC7CAECCE9A4" box="[748,1003,962,989]" latitude="-9.4372225" longLatPrecision="21" longitude="-74.558334" municipality="Dist. Iparia" name="Cuenca del rio Iparia" pageId="276" pageNumber="277">Cuenca del río Iparía</location>
,
<locationDeviation id="9A16C37FFEB1FEEDFC20FC7DA833E9A5" box="[1017,1300,962,989]" pageId="276" pageNumber="277">afluente del río Ucayali</locationDeviation>
,
<location id="8EC48C86FEB1FEEDFAFBFC7CAC5BEE79" LSID="urn:lsid:plazi:treatment:03B26B4BFEB6FEEFFE24F9EDAC54EFF4:8EC48C86FEB1FEEDFAFBFC7CAC5BEE79" latitude="-9.4372225" longLatPrecision="21" longitude="-74.558334" municipality="Dist. Iparia" name="reserva Comunal el Sira" pageId="276" pageNumber="277">reserva Comunal el Sira</location>
;
<geoCoordinate id="EE2FBC9AFEB1FEEDFE5EFC5BAF21EE79" box="[391,518,996,1024]" degrees="9" direction="south" minutes="26" orientation="latitude" pageId="276" pageNumber="277" precision="15" seconds="14" value="-9.4372225">9°2614″ S</geoCoordinate>
,
<geoCoordinate id="EE2FBC9AFEB1FEEDFDCBFC5BAF8BEE78" box="[530,684,996,1025]" degrees="74" direction="west" minutes="33" orientation="longitude" pageId="276" pageNumber="277" precision="15" seconds="30" value="-74.558334">74°3330″ W</geoCoordinate>
;
<elevation id="00363D6EFEB1FEEDFD6FFC59AE5FEE79" box="[694,888,998,1025]" metricMagnitude="2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="276" pageNumber="277" unit="m" value="425.0" valueMax="450.0" valueMin="400.0">
<quantity id="4CE377B8FEB1FEEDFD6FFC59AE1DEE78" box="[694,826,998,1025]" metricMagnitude="2" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="276" pageNumber="277" unit="m" value="425.0" valueMax="450.0" valueMin="400.0">400450 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA5FC9DFEB1FEEDFC5DFC59A929EE79" box="[900,1038,998,1025]" pageId="276" pageNumber="277" value="2007-10-05">
<collectingDate id="EFE10575FEB1FEEDFC5DFC59A929EE79" box="[900,1038,998,1025]" pageId="276" pageNumber="277" value="2007-10-05">5 Oct. 2007</collectingDate>
</date>
;
<emphasis id="B96F064FFEB1FEEDFBC0FC59A9D3EE79" box="[1049,1268,998,1024]" italics="true" pageId="276" pageNumber="277">
<collectorName id="26EEBF8BFEB1FEEDFBC0FC59A993EE79" box="[1049,1204,998,1024]" pageId="276" pageNumber="277">J.G. Graham</collectorName>
4898
</emphasis>
;
<collectionCode id="ED0A4298FEB1FEEDFADAFC59A860EE79" box="[1283,1351,998,1024]" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="276" pageNumber="277" type="Herbarium">MOL</collectionCode>
,
<collectionCode id="ED0A4298FEB1FEEDFA88FC59A851EE79" box="[1361,1398,998,1024]" pageId="276" pageNumber="277">US</collectionCode>
[
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]
</materialsCitation>
.
<materialsCitation id="3B73D000FEB1FEEDFE49FBB6A9F3EE31" ID-GBIF-Occurrence="4517461314" collectingDate="2021-03-22" collectionCode="E" collectorName="P. W. Moonlight" pageId="276" pageNumber="277" specimenCount="1">
<emphasis id="B96F064FFEB1FEEDFE49FBB6AF32EE5A" bold="true" box="[400,533,1033,1059]" pageId="276" pageNumber="277">Cultivated</emphasis>
• Grown in the Royal Botanic Garden Edinburgh from
<emphasis id="B96F064FFEB1FEEDFB7BFBB5AC77EE3E" italics="true" pageId="276" pageNumber="277">P.W. Moonlight &amp; A. Daza 199</emphasis>
RBGE Living Accession 20160139;
<date id="FFA5FC9DFEB1FEEDFCDFFB92AE80EE3E" box="[774,935,1069,1096]" pageId="276" pageNumber="277" value="2021-03-22">
<collectingDate id="EFE10575FEB1FEEDFCDFFB92AE80EE3E" box="[774,935,1069,1096]" pageId="276" pageNumber="277" value="2021-03-22">22 Mar. 2021</collectingDate>
</date>
;
<emphasis id="B96F064FFEB1FEEDFC6AFB91A98CEE3E" box="[947,1195,1069,1095]" italics="true" pageId="276" pageNumber="277">
<collectorName id="26EEBF8BFEB1FEEDFC6AFB91A94FEE3E" box="[947,1128,1069,1095]" pageId="276" pageNumber="277">P.W. Moonlight</collectorName>
1952
</emphasis>
;
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.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FEB1FEEDFF64FBD0AEE4ED90" pageId="276" pageNumber="277" type="description">
<paragraph id="8BA4DA5DFEB1FEEDFF64FBD0AC68EEF0" blockId="276.[189,335,1135,1161]" box="[189,335,1135,1161]" pageId="276" pageNumber="277">
<heading id="D0EC6D31FEB1FEEDFF64FBD0AC68EEF0" bold="true" box="[189,335,1135,1161]" fontSize="11" level="3" pageId="276" pageNumber="277" reason="3">
<emphasis id="B96F064FFEB1FEEDFF64FBD0AC68EEF0" bold="true" box="[189,335,1135,1161]" pageId="276" pageNumber="277">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFEB1FEEDFF64FB21AEE4ED90" blockId="276.[187,1401,1181,2025]" pageId="276" pageNumber="277">
Caulescent herb, to at least
<quantity id="4CE377B8FEB1FEEDFDC9FB21AF67EEC1" box="[528,576,1182,1208]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="276" pageNumber="277" unit="m" value="1.0">1 m</quantity>
high.
<emphasis id="B96F064FFEB1FEEDFD48FB21AFEDEEC1" box="[657,714,1182,1208]" italics="true" pageId="276" pageNumber="277">Stem</emphasis>
erect, frequently branching; internodes to
<quantity id="4CE377B8FEB1FEEDFB0BFB22A828EEC1" box="[1234,1295,1181,1208]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="276" pageNumber="277" unit="cm" value="5.0">5 cm</quantity>
long, to
<quantity id="4CE377B8FEB1FEEDFF64FB7EAC26EEA5" box="[189,257,1217,1244]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="276" pageNumber="277" unit="mm" value="6.0">6 mm</quantity>
thick, succulent, green to red at the nodes, glabrous.
<emphasis id="B96F064FFEB1FEEDFCA8FB7EAEE8EEA2" box="[881,975,1217,1243]" italics="true" pageId="276" pageNumber="277">Stipules</emphasis>
early deciduous, lanceolate, 814 ×
<quantity id="4CE377B8FEB1FEEDFF64FB5AAC07EE86" box="[189,288,1253,1279]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="276" pageNumber="277" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
, apex acuminate, translucent, pale green, glabrous, margin entire, aciliate.
<emphasis id="B96F064FFEB1FEEDFBA7FB5AA9F7EE87" box="[1150,1232,1253,1278]" italics="true" pageId="276" pageNumber="277">Leaves</emphasis>
&gt; 5, alternate, basifixed; petiole
<quantity id="4CE377B8FEB1FEEDFE48FAB7AF3EEF5A" box="[401,537,1288,1315]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.7" metricValueMin="0.5" pageId="276" pageNumber="277" unit="cm" value="1.1" valueMax="1.7" valueMin="0.5">0.51.7 cm</quantity>
long, pale green to red, glabrous; blade subsymmetric, oblong, to 12 ×
<quantity id="4CE377B8FEB1FEEDFF64FA94AC28EF3F" box="[189,271,1323,1350]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="276" pageNumber="277" unit="cm" value="6.5">6.5 cm</quantity>
, succulent, apex short acuminate, base inequilateral, cuneate to rounded on the narrow side of the bade, rounded on the broad side of the blade, margin serrulate, ciliate, upper surface dark green, often with red or purple spots on young leaves, glabrous, though young leaves often with a single hair at the centre of each coloured spot, lower surface pale green, glabrous, veins pinnate with 912 secondary veins on the larger side, 812 secondary veins on the smaller side.
<emphasis id="B96F064FFEB1FEEDFCA4FA05A902EFAD" box="[893,1061,1466,1492]" italics="true" pageId="276" pageNumber="277">Inflorescences</emphasis>
&gt; 3, bisexual, axillary, erect, cymose, branching up to 4 times, bearing up to 8 staminate flowers and 8 pistillate flowers, protandrous; peduncle to
<quantity id="4CE377B8FEB1FEEDFE95F9BEACBAEC62" box="[332,413,1537,1563]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="276" pageNumber="277" unit="cm" value="2.8">2.8 cm</quantity>
long, pale green, glabrous, bracts deciduous, ovate, ca 4 ×
<quantity id="4CE377B8FEB1FEEDFB8AF9BEA9BDEC62" box="[1107,1178,1537,1563]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="276" pageNumber="277" unit="mm" value="3.0">3 mm</quantity>
, translucent, white to pink, glabrous, apex acute, margin serrulate towards the apex, ciliate.
<emphasis id="B96F064FFEB1FEEDFBCDF99AA9C0EC47" box="[1044,1255,1572,1599]" italics="true" pageId="276" pageNumber="277">Staminate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FEB1FEEDFF64F9F7AC35EC1B" box="[189,274,1608,1634]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="276" pageNumber="277" unit="mm" value="13.0">13 mm</quantity>
long, glabrous; tepals 2, spreading, broadly ovate, 1014 ×
<quantity id="4CE377B8FEB1FEEDFC33F9F7A949EC1B" box="[1002,1134,1608,1635]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="276" pageNumber="277" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
, apex rounded, white, glabrous, margin entire, aciliate; stamens ca 35, projecting, yellow, filaments
<quantity id="4CE377B8FEB1FEEDFB9FF9D4A9BAECFF" box="[1094,1181,1643,1670]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="276" pageNumber="277" unit="mm" value="0.2">0.2 mm</quantity>
long, free, anthers elliptic, 11.5 ×
<quantity id="4CE377B8FEB1FEEDFEA0F930ACF5ECD0" box="[377,466,1679,1705]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="276" pageNumber="277" unit="mm" value="0.8">0.8 mm</quantity>
long, dehiscing via lateral slits, connectives extending to
<quantity id="4CE377B8FEB1FEEDFBB3F930A9E2ECD0" box="[1130,1221,1679,1706]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="276" pageNumber="277" unit="mm" value="0.5">0.5 mm</quantity>
, symmetrically basifixed.
<emphasis id="B96F064FFEB1FEEDFEEEF90DAF27ECB5" box="[311,512,1714,1740]" italics="true" pageId="276" pageNumber="277">Pistillate flowers</emphasis>
: pedicels to
<quantity id="4CE377B8FEB1FEEDFD43F90DAFC9ECB4" box="[666,750,1714,1741]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="276" pageNumber="277" unit="mm" value="13.0">13 mm</quantity>
long; bracteoles 2, directly beneath the ovary, broadly ovate, 57 ×
<quantity id="4CE377B8FEB1FEEDFE82F969ACE7EC89" box="[347,448,1750,1776]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="276" pageNumber="277" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
, apex rounded, translucent, white flushed pink, glabrous, margin serrulate at the apex, ciliate, tepals 2, equal, deciduous in fruit, spreading, ovate, 811 ×
<quantity id="4CE377B8FEB1FEEDFB9FF946A99AED6D" box="[1094,1213,1785,1812]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="276" pageNumber="277" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">812 mm</quantity>
, apex rounded, white, glabrous, margin entire, aciliate; ovary body ovoid, 89 ×
<quantity id="4CE377B8FEB1FEEDFC63F8A2A939ED4E" box="[954,1054,1821,1847]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="276" pageNumber="277" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
, white, glabrous, sub-equally 3-winged, the largest wing obdeltoid, 812 ×
<quantity id="4CE377B8FEB1FEEDFD38F8FFAE43ED22" box="[737,868,1856,1883]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="276" pageNumber="277" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">1013 mm</quantity>
, the smallest obdeltoid, to 12 ×
<quantity id="4CE377B8FEB1FEEDFB3EF8FFA80AED22" box="[1255,1325,1856,1883]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="276" pageNumber="277" unit="mm" value="8.0">8 mm</quantity>
wide; 3-locular, placentae branches divided, bearing ovules on both surfaces; styles 3, yellow, free, ca
<quantity id="4CE377B8FEB1FEEDFAEDF8DBA85EED07" box="[1332,1401,1892,1918]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="276" pageNumber="277" unit="mm" value="4.0">4 mm</quantity>
long, once-divided, thickened above the division, stigmatic papillae in a spirally twisted band.
<emphasis id="B96F064FFEB1FEEDFACDF837AC34EDBC" italics="true" pageId="276" pageNumber="277">Fruiting pedicel</emphasis>
to
<quantity id="4CE377B8FEB1FEEDFEE3F814ACAAEDBC" box="[314,397,1963,1989]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="276" pageNumber="277" unit="mm" value="18.0">18 mm</quantity>
long.
<emphasis id="B96F064FFEB1FEEDFE00F813AF70EDBC" box="[473,599,1963,1989]" italics="true" pageId="276" pageNumber="277">Fruit body</emphasis>
ovoid, to 12 ×
<quantity id="4CE377B8FEB1FEEDFCD6F814AE71EDBC" box="[783,854,1963,1989]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="276" pageNumber="277" unit="mm" value="7.0">7 mm</quantity>
, drying brown, wings same shape as in ovary, the largest expanding to 16 ×
<quantity id="4CE377B8FEB1FEEDFDC3F871AF57ED90" box="[538,624,1998,2025]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="276" pageNumber="277" unit="mm" value="13.0">13 mm</quantity>
, the smallest to 14 ×
<quantity id="4CE377B8FEB1FEEDFCB1F871AE99ED90" box="[872,958,1998,2025]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="276" pageNumber="277" unit="mm" value="16.0">16 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF648AD5FEB0FEECFF64F8A1AEDBED91" ID-DOI="http://doi.org/10.5281/zenodo.8178489" ID-Zenodo-Dep="8178489" httpUri="https://zenodo.org/record/8178489/files/figure.png" pageId="277" pageNumber="278" startId="277.[189,232,1822,1848]" targetBox="[257,1330,265,1783]" targetPageId="277">
<paragraph id="8BA4DA5DFEB0FEECFF64F8A1AEDBED91" blockId="277.[189,1399,1822,2025]" pageId="277" pageNumber="278">
<emphasis id="B96F064FFEB0FEECFF64F8A1AC39ED41" bold="true" box="[189,286,1822,1848]" pageId="277" pageNumber="278">Fig. 87.</emphasis>
<taxonomicName id="4C1BA1DEFEB0FEECFEF1F8A0AFF3ED41" authority="Moonlight" authorityName="Moonlight" authorityYear="2023" box="[296,724,1822,1848]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="277" pageNumber="278" phylum="Tracheophyta" rank="species" species="yuracyacuensis" status="sp. nov.">
<emphasis id="B96F064FFEB0FEECFEF1F8A0AF6AED41" box="[296,589,1823,1848]" italics="true" pageId="277" pageNumber="278">Begonia yuracyacuensis</emphasis>
Moonlight
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FEB0FEECFD07F8A0AE1BED40" box="[734,828,1823,1849]" pageId="277" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B96F064FFEB0FEECFC9FF8A1AE7CED41" bold="true" box="[838,859,1822,1848]" pageId="277" pageNumber="278">A</emphasis>
. Habit.
<emphasis id="B96F064FFEB0FEECFC18F8A1AEF2ED41" bold="true" box="[961,981,1822,1848]" pageId="277" pageNumber="278">B</emphasis>
. Stipule.
<emphasis id="B96F064FFEB0FEECFB95F8A1A946ED41" bold="true" box="[1100,1121,1822,1848]" pageId="277" pageNumber="278">C</emphasis>
. Leaf, adaxial surface.
<emphasis id="B96F064FFEB0FEECFF64F8FDADF5ED25" bold="true" box="[189,210,1858,1884]" pageId="277" pageNumber="278">D</emphasis>
. Leaf, abaxial surface.
<emphasis id="B96F064FFEB0FEECFE30F8FDACDBED25" bold="true" box="[489,508,1858,1884]" pageId="277" pageNumber="278">E</emphasis>
. Shade leaf, adaxial surface.
<emphasis id="B96F064FFEB0FEECFC81F8FDAE4EED25" bold="true" box="[856,873,1858,1884]" pageId="277" pageNumber="278">F</emphasis>
. Cross section of ovary.
<emphasis id="B96F064FFEB0FEECFB48F8FEA980ED22" bold="true" box="[1169,1191,1857,1883]" pageId="277" pageNumber="278">G</emphasis>
. Pistillate flower, side view.
<emphasis id="B96F064FFEB0FEECFEE7F8DAAC74ED06" bold="true" box="[318,339,1893,1919]" pageId="277" pageNumber="278">H</emphasis>
. Pistillate flower, front view.
<emphasis id="B96F064FFEB0FEECFD6FF8DAAFE6ED06" bold="true" box="[694,705,1893,1919]" pageId="277" pageNumber="278">I</emphasis>
. Pistils, side view.
<emphasis id="B96F064FFEB0FEECFC73F8DAAE90ED06" bold="true" box="[938,951,1893,1919]" pageId="277" pageNumber="278">J</emphasis>
. Bracteole.
<emphasis id="B96F064FFEB0FEECFB93F8DAA978ED06" bold="true" box="[1098,1119,1893,1919]" pageId="277" pageNumber="278">K</emphasis>
. Staminate flower, side view.
<emphasis id="B96F064FFEB0FEECFEDBF837AC31EDDB" bold="true" box="[258,278,1928,1954]" pageId="277" pageNumber="278">L</emphasis>
. Staminate flower, front view. M. Androecium, side view. Photographs taken by D.A. Purvis and P.W. Moonlight in the living collections of the Royal Botanic Garden Edinburgh (Accession 20160139, grown from seeds collected as part of
<emphasis id="B96F064FFEB0FEECFDA5F870AECCED91" box="[636,1003,1998,2025]" italics="true" pageId="277" pageNumber="278">P.W. Moonlight &amp; A. Daza 199</emphasis>
).
</paragraph>
</caption>
<subSubSection id="C30189D6FEB3FEEFFF64FEB6ACACE86D" pageId="278" pageNumber="279" type="conservation">
<paragraph id="8BA4DA5DFEB3FEEFFF64FEB6AF4AEB5A" blockId="278.[189,621,264,291]" box="[189,621,264,291]" pageId="278" pageNumber="279">
<heading id="D0EC6D31FEB3FEEFFF64FEB6AF4AEB5A" bold="true" box="[189,621,264,291]" fontSize="11" level="3" pageId="278" pageNumber="279" reason="3">
<emphasis id="B96F064FFEB3FEEFFF64FEB6AF4AEB5A" bold="true" box="[189,621,264,291]" pageId="278" pageNumber="279">Proposed conservation assessment</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFEB3FEEFFF64FE84ACACE86D" blockId="278.[189,1400,314,532]" pageId="278" pageNumber="279">
Known from localities approximately
<quantity id="4CE377B8FEB3FEEFFD58FE85AFFDEB2D" box="[641,730,314,341]" metricMagnitude="5" metricUnit="m" metricValue="1.25" pageId="278" pageNumber="279" unit="km" value="125.0">125 km</quantity>
apart in
<collectingRegion id="49DF14BFFEB3FEEFFC9CFE85AEDAEB2D" box="[837,1021,314,340]" country="Peru" name="Ucayali" pageId="278" pageNumber="279">Ucayali Region</collectingRegion>
<collectingCountry id="F30C9ACDFEB3FEEFFBDCFE84A91CEB2C" box="[1029,1083,315,341]" name="Peru" pageId="278" pageNumber="279">Peru</collectingCountry>
and a third locality in
<collectingRegion id="49DF14BFFEB3FEEFFA93FE85AC49EB03" country="Peru" name="San Martin" pageId="278" pageNumber="279">San Martín Region</collectingRegion>
. These three localities are low lying, extremely humid hills. Similar localities elsewhere are poorly explored in
<collectingCountry id="F30C9ACDFEB3FEEFFE09FE38AF2EEBD8" box="[464,521,391,417]" name="Peru" pageId="278" pageNumber="279">Peru</collectingCountry>
, so it is likely found elsewhere. However, because it is only known from three localities, we assess
<taxonomicName id="4C1BA1DEFEB3FEEFFE21FE11AFF5EBBE" authorityName="Moonlight" authorityYear="2023" box="[504,722,430,455]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="278" pageNumber="279" phylum="Tracheophyta" rank="species" species="yuracyacuensis" status="sp. nov.">
<emphasis id="B96F064FFEB3FEEFFE21FE11AFF5EBBE" box="[504,722,430,455]" italics="true" pageId="278" pageNumber="279">B. yuracyacuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FEB3FEEFFD00FE12AE11EBBE" box="[729,822,429,455]" pageId="278" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
as Vulnerable (VU D2), owing to the proximity of two of its collections to major roads and the widescale deforestation in lowland montane forests in Amazonian
<collectingCountry id="F30C9ACDFEB3FEEFFE94FE45ACA0E86D" box="[333,391,506,532]" name="Peru" pageId="278" pageNumber="279">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FEB3FEEFFF64FD81AF84EE4B" pageId="278" pageNumber="279" type="discussion">
<paragraph id="8BA4DA5DFEB3FEEFFF64FD81AC23E821" blockId="278.[189,260,574,600]" box="[189,260,574,600]" pageId="278" pageNumber="279">
<heading id="D0EC6D31FEB3FEEFFF64FD81AC23E821" bold="true" box="[189,260,574,600]" fontSize="11" level="3" pageId="278" pageNumber="279" reason="3">
<emphasis id="B96F064FFEB3FEEFFF64FD81AC23E821" bold="true" box="[189,260,574,600]" pageId="278" pageNumber="279">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFEB3FEEFFF64FDD0A91DE8C9" blockId="278.[189,1398,623,688]" pageId="278" pageNumber="279">
The label of the
<typeStatus id="54A064FFFEB3FEEFFE5DFDCFAC90E8F3" box="[388,439,624,650]" pageId="278" pageNumber="279">type</typeStatus>
collection states it was collected in
<collectingRegion id="49DF14BFFEB3FEEFFCB3FDCFA976E8F0" box="[874,1105,623,650]" country="Peru" name="San Martin" pageId="278" pageNumber="279">San Martín Region</collectingRegion>
. The locality description refers however to the
<collectingRegion id="49DF14BFFEB3FEEFFE67FD29AF15E8C9" box="[446,562,662,688]" country="Paraguay" name="Boqueron" pageId="278" pageNumber="279">Boquerón</collectingRegion>
de Padre Abad, which is in
<collectingRegion id="49DF14BFFEB3FEEFFCA2FD29A910E8C9" box="[891,1079,662,688]" country="Peru" name="Ucayali" pageId="278" pageNumber="279">Ucayali Region</collectingRegion>
.
</paragraph>
<paragraph id="8BA4DA5DFEB3FEEFFF64FD65AC97E88D" blockId="278.[189,432,730,756]" box="[189,432,730,756]" pageId="278" pageNumber="279">
<heading id="D0EC6D31FEB3FEEFFF64FD65AC97E88D" bold="true" box="[189,432,730,756]" fontSize="11" level="3" pageId="278" pageNumber="279" reason="3">
<emphasis id="B96F064FFEB3FEEFFF64FD65AC97E88D" bold="true" box="[189,432,730,756]" pageId="278" pageNumber="279">Identification notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFEB3FEEFFF64FCB3AF84EE4B" blockId="278.[188,1400,780,1074]" pageId="278" pageNumber="279">
When in flower and fruit, it is trivial to distinguish the new species from
<taxonomicName id="4C1BA1DEFEB3FEEFFBF1FCB2A9E4E95F" baseAuthorityName="de Candolle" baseAuthorityYear="1859" box="[1064,1219,781,806]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="278" pageNumber="279" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B96F064FFEB3FEEFFBF1FCB2A9E4E95F" box="[1064,1219,781,806]" italics="true" pageId="278" pageNumber="279">B. peruviana</emphasis>
</taxonomicName>
based upon its fewer-branched inflorescences (up to 4 times branching vs up to 8 times branching) and much larger staminate and pistillate flowers (see Diagnosis). The two species are very difficult to distinguish when sterile. The stipules of the new species tend to be broader (
<quantity id="4CE377B8FEB3FEEFFCB7FCC0AEF6E9E0" box="[878,977,895,921]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="278" pageNumber="279" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
wide vs
<quantity id="4CE377B8FEB3FEEFFBE3FCC0A9BBE9E0" box="[1082,1180,895,921]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="278" pageNumber="279" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
wide) and the leaf margins of both species can be serrulate but
<taxonomicName id="4C1BA1DEFEB3FEEFFD1CFC19AE79E9C6" baseAuthorityName="de Candolle" baseAuthorityYear="1859" box="[709,862,934,959]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="278" pageNumber="279" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B96F064FFEB3FEEFFD1CFC19AE79E9C6" box="[709,862,934,959]" italics="true" pageId="278" pageNumber="279">B. peruviana</emphasis>
</taxonomicName>
more frequently has serrate or doubly-serrate margins. Finally, the two species differ in their habits.
<taxonomicName id="4C1BA1DEFEB3FEEFFC8BFC73AEC9E99C" baseAuthorityName="de Candolle" baseAuthorityYear="1859" box="[850,1006,972,997]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="278" pageNumber="279" phylum="Tracheophyta" rank="species" species="peruviana">
<emphasis id="B96F064FFEB3FEEFFC8BFC73AEC9E99C" box="[850,1006,972,997]" italics="true" pageId="278" pageNumber="279">B. peruviana</emphasis>
</taxonomicName>
tends to scramble through other plants and reaches up to at least
<quantity id="4CE377B8FEB3FEEFFDE9FC4DAF7BEE75" box="[560,604,1010,1036]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="278" pageNumber="279" unit="m" value="3.0">3 m</quantity>
in height. In contrast,
<taxonomicName id="4C1BA1DEFEB3FEEFFC84FC4DA913EE72" authorityName="Moonlight" authorityYear="2023" box="[861,1076,1010,1035]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="278" pageNumber="279" phylum="Tracheophyta" rank="species" species="yuracyacuensis" status="sp. nov.">
<emphasis id="B96F064FFEB3FEEFFC84FC4DA913EE72" box="[861,1076,1010,1035]" italics="true" pageId="278" pageNumber="279">B. yuracyacuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25CBB34FEB3FEEFFBE2FC4DA9B3EE75" box="[1083,1172,1010,1036]" pageId="278" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
grows horizontally from vertical cliffs and is self-supporting.
</paragraph>
</subSubSection>
<subSubSection id="C30189D6FEB3FEEFFF64FBE3AC54EFF4" pageId="278" pageNumber="279" type="distribution">
<paragraph id="8BA4DA5DFEB3FEEFFF64FBE3ACD2EE0F" blockId="278.[189,501,1116,1142]" box="[189,501,1116,1142]" pageId="278" pageNumber="279">
<heading id="D0EC6D31FEB3FEEFFF64FBE3ACD2EE0F" bold="true" box="[189,501,1116,1142]" fontSize="11" level="3" pageId="278" pageNumber="279" reason="3">
<emphasis id="B96F064FFEB3FEEFFF64FBE3ACD2EE0F" bold="true" box="[189,501,1116,1142]" pageId="278" pageNumber="279">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4DA5DFEB3FEEFFF64FB31AC54EFF4" blockId="278.[187,1399,1166,1421]" pageId="278" pageNumber="279">
Endemic to
<collectingCountry id="F30C9ACDFEB3FEEFFE91FB31AC59EED1" box="[328,382,1166,1192]" name="Peru" pageId="278" pageNumber="279">Peru</collectingCountry>
and known from
<collectingRegion id="49DF14BFFEB3FEEFFD91FB31AFEDEED1" box="[584,714,1166,1192]" country="Peru" name="San Martin" pageId="278" pageNumber="279">San Martín</collectingRegion>
and
<collectingRegion id="49DF14BFFEB3FEEFFCD9FB31AE7CEED1" box="[768,859,1166,1192]" country="Peru" name="Ucayali" pageId="278" pageNumber="279">Ucayali</collectingRegion>
Regions (
<figureCitation id="1320C6D8FEB3FEEFFC09FB31A91BEED1" box="[976,1084,1166,1192]" captionStart="Fig" captionStartId="263.[189,232,1857,1883]" captionTargetBox="[194,1392,1194,1813]" captionTargetId="figure-326@263.[189,1398,1184,1819]" captionTargetPageId="263" captionText="Fig. 84. Distribution of Begonia sect. Ruizopavonia A.DC. in Peru and surrounding countries. A. B. glauca (Klotzsch) Ruiz &amp; Pav. ex A.DC. (red) and B. yuracyacuensis Moonlight sp. nov. (blue). B. B. nunezii Moonlight sp. nov. (red) and B. peruviana A.DC. (blue). Black lines indicate country borders, grey lines indicate major administrative divisions, blue lines indicate rivers, and shading indicates elevation." figureDoi="http://doi.org/10.5281/zenodo.8178482" httpUri="https://zenodo.org/record/8178482/files/figure.png" pageId="278" pageNumber="279">Fig. 84A</figureCitation>
). Known from an elevation of
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a.s.l. and collected by the authors within the Boquerón de Padre Abad, a ca
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a.s.l. deep gorge in Amazonian
<collectingCountry id="F30C9ACDFEB3FEEFFE28FB64AF0DEE8C" box="[497,554,1243,1269]" name="Peru" pageId="278" pageNumber="279">Peru</collectingCountry>
, where the río Yuracyacu cuts through a north-south ridge to the west of the Cordillera Azul. This is the most easterly ridge of the Andes in central
<collectingCountry id="F30C9ACDFEB3FEEFFBFBFABEA97FEF62" box="[1058,1112,1281,1307]" name="Peru" pageId="278" pageNumber="279">Peru</collectingCountry>
and therefore one of the wettest locations in
<collectingCountry id="F30C9ACDFEB3FEEFFE72FA98ACC2EF38" box="[427,485,1319,1345]" name="Peru" pageId="278" pageNumber="279">Peru</collectingCountry>
. The new species is found growing under and around waterfalls on the walls of the gorge. It has been collected growing in association with
<taxonomicName id="4C1BA1DEFEB3FEEFFC6EFAF1A95BEF1E" box="[951,1148,1357,1383]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="278" pageNumber="279" phylum="Tracheophyta" rank="species" species="albomaculata">
<emphasis id="B96F064FFEB3FEEFFC6EFAF1A95BEF1E" box="[951,1148,1357,1383]" italics="true" pageId="278" pageNumber="279">B. albomaculata</emphasis>
</taxonomicName>
, which is a distantly related species.
</paragraph>
</subSubSection>
</treatment>
</document>