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<document id="8FCF89157B903C88277940D6ECFCAE6D" ID-DOI="10.1080/00222930010015861" ID-ISSN="1464-5262" ID-Zenodo-Dep="5299861" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629999714736" checkinUser="felipe" docAuthor="Ohtsuka, Susumu, Nishida, Shuhei &amp; Nakaguchi, Kazumitsu" docDate="2002" docId="03B0E830FF9AFFC0579BF823B8F423DD" docLanguage="en" docName="JNatHist.36.5.531-564.pdf" docOrigin="Journal of Natural History 36 (5)" docSource="http://www.tandfonline.com/doi/abs/10.1080/00222930010015861" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Macandrewella serratipes Ohtsuka &amp; Nishida &amp; Nakaguchi 2002, new species" docType="treatment" docVersion="4" lastPageNumber="559" masterDocId="FF899048FF83FFDE560FF906B8052201" masterDocTitle="Three new species of the genus Macandrewella (Copepoda: Calanoida: Scolecitrichidae) from the Paci ® c Ocean, with notes on distribution and feeding habits" masterLastPageNumber="564" masterPageNumber="531" pageNumber="554" updateTime="1699135089635" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B905734EFDBC1E2713BA666E01B1D569">Three new species of the genus Macandrewella (Copepoda: Calanoida: Scolecitrichidae) from the Paci ® c Ocean, with notes on distribution and feeding habits</mods:title>
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<mods:namePart id="B125ED000B6D00D597B3C4C7BED5ED93">Ohtsuka, Susumu</mods:namePart>
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<mods:namePart id="CF87A9729E467677E6CD37967E6A8AF1">Nishida, Shuhei</mods:namePart>
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<mods:title id="D87675AD7C8D08C2DD16819E6033512D">Journal of Natural History</mods:title>
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<paragraph id="8BA65926FF9AFFC7579BF823BB3B233C" blockId="25.[404,830,293,349]" box="[404,830,293,317]" pageId="25" pageNumber="554">
<heading id="D0EEEE4AFF9AFFC7579BF823BB3B233C" box="[404,830,293,317]" centered="true" fontSize="10" level="2" pageId="25" pageNumber="554" reason="3">
<taxonomicName id="4C1922A5FF9AFFC7579BF823BAB7233C" authority="Ohtsuka &amp; Nishida &amp; Nakaguchi, 2002" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[404,690,293,317]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="25" pageNumber="554" phylum="Arthropoda" rank="species" species="serratipes" status="sp. nov.">
<emphasis id="B96D8534FF9AFFC7579BF823BAB7233C" bold="true" box="[404,690,293,317]" italics="true" pageId="25" pageNumber="554">Macandrewella serratipes</emphasis>
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<taxonomicNameLabel id="A25E384FFF9AFFC754B5F823BB3B233C" box="[698,830,293,317]" pageId="25" pageNumber="554" rank="species">new species</taxonomicNameLabel>
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</paragraph>
<paragraph id="8BA65926FF9AFFC7541CF843BABA235C" blockId="25.[404,830,293,349]" box="[531,703,325,349]" pageId="25" pageNumber="554">(®gures 19, 20)</paragraph>
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<paragraph id="8BA65926FF9AFFC756BCF872BB28238D" blockId="25.[140,1095,372,939]" box="[179,813,372,396]" pageId="25" pageNumber="554">
<emphasis id="B96D8534FF9AFFC756BCF872B924238D" box="[179,289,372,396]" italics="true" pageId="25" pageNumber="554">Synonym.</emphasis>
<taxonomicName id="4C1922A5FF9AFFC75733F872BB20238D" authority="sensu Wilson, 1950" authorityName="sensu Wilson" authorityYear="1950" box="[316,805,372,396]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="25" pageNumber="554" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9AFFC75733F872BA8D238D" box="[316,648,372,396]" italics="true" pageId="25" pageNumber="554">Macandrewella chelipes sensu</emphasis>
<bibRefCitation id="EF8824D7FF9AFFC7549EF872BB20238D" author="WILSON, C. B." box="[657,805,372,396]" pageId="25" pageNumber="554" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilson, 1950</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA65926FF9AFFC756BCF892BA8D202D" blockId="25.[140,1095,372,939]" pageId="25" pageNumber="554">
<emphasis id="B96D8534FF9AFFC756BCF892B98A23AD" box="[179,399,404,428]" italics="true" pageId="25" pageNumber="554">Material examined.</emphasis>
<materialsCitation id="3B71537BFF9AFFC757A5F892B8BB23ED" collectingDate="1905-01-21" collectionCode="USNM" collectorName="Wilson &amp; Note" country="Fiji" location="Fiji" pageId="25" pageNumber="554" specimenCount="1">
<collectingCountry id="F30E19B6FF9AFFC757A5F892B9D723AD" box="[426,466,404,428]" name="Fiji" pageId="25" pageNumber="554">Fiji</collectingCountry>
Is. (16
<emphasis id="B96D8534FF9AFFC7542BF892BA2B23AD" bold="true" box="[548,558,404,428]" italics="true" pageId="25" pageNumber="554">ss</emphasis>
32
<emphasis id="B96D8534FF9AFFC75446F892BA4A23AD" bold="true" box="[585,591,404,428]" italics="true" pageId="25" pageNumber="554">¾</emphasis>
S, 119
<emphasis id="B96D8534FF9AFFC75499F892BAA523AD" bold="true" box="[662,672,404,428]" italics="true" pageId="25" pageNumber="554">ss</emphasis>
59
<emphasis id="B96D8534FF9AFFC754B4F892BAC423AD" bold="true" box="[699,705,404,428]" italics="true" pageId="25" pageNumber="554">¾</emphasis>
W), 40 fathoms,
<date id="FFA77FE6FF9AFFC75585F892BC4323AD" box="[906,1094,404,429]" pageId="25" pageNumber="554" value="1905-01-21">
<collectingDate id="EFE3860EFF9AFFC75585F892BC4323AD" box="[906,1094,404,429]" pageId="25" pageNumber="554" value="1905-01-21">21 January 1905</collectingDate>
</date>
(
<collectorName id="26EC3CF0FF9AFFC75697F8B2B8ED23CD" box="[152,232,436,460]" pageId="25" pageNumber="554">Wilson</collectorName>
leg.): 7 and 6.
<collectorName id="26EC3CF0FF9AFFC757ECF8B3BA1E23CD" box="[483,539,437,460]" pageId="25" pageNumber="554">Note</collectorName>
: the ledger book for the
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numbers says only
</materialsCitation>
<materialsCitation id="3B71537BFF9AFFC756C9F8D2BAB7200D" collectorName="Tuamotu Is." country="Fiji" location="South Paci" pageId="25" pageNumber="554" specimenCount="1">
<collectingCountry id="F30E19B6FF9AFFC756C9F8D2B8E823ED" box="[198,237,468,492]" name="Fiji" pageId="25" pageNumber="554">Fiji</collectingCountry>
, and the latitude/longitude of the locality described in the vial plots out to the
<location id="8EC60FFDFF9AFFC756B5F8F2B93C200D" LSID="urn:lsid:plazi:treatment:03B0E830FF9AFFC0579BF823B8F423DD:8EC60FFDFF9AFFC756B5F8F2B93C200D" box="[186,313,500,524]" country="Fiji" name="South Paci" pageId="25" pageNumber="554">South Paci</location>
®c near
<collectorName id="26EC3CF0FF9AFFC75792F8F3BA28200D" box="[413,557,501,524]" pageId="25" pageNumber="554">Tuamotu Is.</collectorName>
but not in
</materialsCitation>
<materialsCitation id="3B71537BFF9AFFC754B2F8F2BA78202D" country="Fiji" location="Hence" pageId="25" pageNumber="554" specimenCount="1">
<collectingCountry id="F30E19B6FF9AFFC754B2F8F2BAE1200D" box="[701,740,500,524]" name="Fiji" pageId="25" pageNumber="554">Fiji</collectingCountry>
.
<location id="8EC60FFDFF9AFFC754F7F8F3BB3A200D" LSID="urn:lsid:plazi:treatment:03B0E830FF9AFFC0579BF823B8F423DD:8EC60FFDFF9AFFC754F7F8F3BB3A200D" box="[760,831,501,524]" country="Fiji" name="Hence" pageId="25" pageNumber="554">Hence</location>
the
<typeStatus id="54A2E784FF9AFFC75576F8F3BBAF200C" box="[889,938,501,525]" pageId="25" pageNumber="554">type</typeStatus>
locality is in question (Walter, personal communication)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA65926FF9AFFC756BCFB32BC43208A" blockId="25.[140,1095,372,939]" pageId="25" pageNumber="554">
<emphasis id="B96D8534FF9AFFC756BCFB32B8F9204A" box="[179,252,564,587]" italics="true" pageId="25" pageNumber="554">Types.</emphasis>
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<typeStatus id="54A2E784FF9AFFC75718FB32B98F204A" box="[279,394,563,587]" pageId="25" pageNumber="554" type="holotype">HOLOTYPE</typeStatus>
: 1, appendages dissected and mounted on glass slides, body in vial,
<specimenCode id="DBBFF15DFF9AFFC7573EFB55B9D1206A" box="[305,468,595,619]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="554" type="Museum">USNM 67242</specimenCode>
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.
<materialsCitation id="3B71537BFF9AFFC757E6FB52BC3B208A" collectionCode="USNM" pageId="25" pageNumber="554" specimenCode="USNM 232823" specimenCount="1" typeStatus="paratype">
<typeStatus id="54A2E784FF9AFFC757E6FB52BA65206A" box="[489,608,595,619]" pageId="25" pageNumber="554" type="paratype">PARATYPES</typeStatus>
: 1 and 3, appendages dissected and mounted on glass slides, body in vial; 5 and 3, whole specimens,
<specimenCode id="DBBFF15DFF9AFFC75599FB75BC3B208A" box="[918,1086,627,651]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="554" type="Museum">USNM 232823</specimenCode>
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.
</paragraph>
<paragraph id="8BA65926FF9AFFC756BCFB95B91620CA" blockId="25.[140,1095,372,939]" pageId="25" pageNumber="554">
<emphasis id="B96D8534FF9AFFC756BCFB95B93A20AA" box="[179,319,659,683]" italics="true" pageId="25" pageNumber="554">Body length.</emphasis>
: 2.91±3.23 mm (
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3.04
<emphasis id="B96D8534FF9AFFC7545AFB95BA6A20AA" bold="true" box="[597,623,659,683]" italics="true" pageId="25" pageNumber="554">Ô</emphasis>
</geoCoordinate>
0.11 mm,
<emphasis id="B96D8534FF9AFFC754D2FB92BAEF20AA" box="[733,746,660,683]" italics="true" pageId="25" pageNumber="554">n</emphasis>
57).: 3.16±3.74 mm (
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3.45
<emphasis id="B96D8534FF9AFFC75223FB95BC4320AA" bold="true" box="[1068,1094,659,683]" italics="true" pageId="25" pageNumber="554">Ô</emphasis>
</geoCoordinate>
0.23,
<emphasis id="B96D8534FF9AFFC756C4FBB2B8DD20CA" box="[203,216,692,715]" italics="true" pageId="25" pageNumber="554">n</emphasis>
56).
</paragraph>
<paragraph id="8BA65926FF9AFFC756BCFBD2BAEE212B" blockId="25.[140,1095,372,939]" pageId="25" pageNumber="554">
<emphasis id="B96D8534FF9AFFC756BCFBD2B93C20EA" box="[179,313,724,747]" italics="true" pageId="25" pageNumber="554">Description.</emphasis>
The new species was brie¯y described as
<taxonomicName id="4C1922A5FF9AFFC75533FBD5BC4220EA" authorityName="sensu Wilson" authorityYear="1950" box="[828,1095,723,747]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="25" pageNumber="554" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9AFFC75533FBD5BC4220EA" box="[828,1095,723,747]" italics="true" pageId="25" pageNumber="554">Macandrewella chelipes</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8824D7FF9AFFC756BFFBF5B952210A" author="WILSON, C. B." box="[176,343,755,779]" pageId="25" pageNumber="554" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilson (1950)</bibRefCitation>
, but several characters were overlooked or misunderstood. These are newly or precisely described in the present study.
</paragraph>
<paragraph id="8BA65926FF9AFFC256BCFA35BB9A22BC" blockId="25.[140,1095,372,939]" lastBlockId="28.[140,1095,133,1687]" lastPageId="28" lastPageNumber="557" pageId="25" pageNumber="554">
<emphasis id="B96D8534FF9AFFC756BCFA35B904214A" box="[179,257,819,843]" italics="true" pageId="25" pageNumber="554">Female</emphasis>
: Prosome (®gure 19A) with fourth and ®fth pedigers almost completely coalescent; prosomal ends (®gure 19A±D) slightly asymmetrical, with dorsolateral processes reaching posterior end of genital double-somite. Urosome (®gure 19A, D) less than one-third length of prosome; genital double-somite asymmetrical, with two anterior round knobs and posterior swollen corner on right side; genital operculum (®gure 19E) wider than long; seminal receptacles extending laterally (®gure 19A); second and third urosomites with striated frill along posterior margin; anal somite small, almost telescoped into preceding somite; anal operculum rounded posteriorly; caudal rami symmetrical in original description, but not con®rmed in present description, as in all specimens examined caudal seta V was missing.
</paragraph>
<caption id="DF6609AEFF9AFFC75683FF6EBBA82499" ID-DOI="http://doi.org/10.5281/zenodo.5299895" ID-Zenodo-Dep="5299895" httpUri="https://zenodo.org/record/5299895/files/figure.png" pageId="25" pageNumber="554" startId="25.[140,156,1640,1661]" targetBox="[141,1094,990,1613]" targetPageId="25">
<paragraph id="8BA65926FF9AFFC75683FF6EBBA82499" blockId="25.[140,1094,1640,1688]" pageId="25" pageNumber="554">
FIG. 18. Gut contents of
<taxonomicName id="4C1922A5FF9AFFC75797FF6EBA72247F" authority="Ohtsuka &amp; Nishida &amp; Nakaguchi, 2002" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[408,631,1640,1662]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="25" pageNumber="554" phylum="Arthropoda" rank="species" species="omorii" status="sp. nov.">
<emphasis id="B96D8534FF9AFFC75797FF6EBA72247F" box="[408,631,1640,1662]" italics="true" pageId="25" pageNumber="554">Macandrewella omorii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E384FFF9AFFC7548CFF6EBABC247C" box="[643,697,1640,1661]" pageId="25" pageNumber="554" rank="species">n. sp.</taxonomicNameLabel>
, female. (A) Fragments of radiolarians and crustaceans. (B) Radiolarian fragments. Scales5 0.01 mm.
</paragraph>
</caption>
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<paragraph id="8BA65926FF99FFC45683FCBEBB6B24A3" blockId="26.[140,1095,1464,1698]" pageId="26" pageNumber="555">
FIG. 19.
<taxonomicName id="4C1922A5FF99FFC456FCFCBEB9F027CF" authority="Ohtsuka &amp; Nishida &amp; Nakaguchi, 2002" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[243,501,1464,1486]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="26" pageNumber="555" phylum="Arthropoda" rank="species" species="serratipes" status="sp. nov.">
<emphasis id="B96D8534FF99FFC456FCFCBEB9F027CF" box="[243,501,1464,1486]" italics="true" pageId="26" pageNumber="555">Macandrewella serratipes</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E384FFF99FFC45408FCBEBA4127CC" box="[519,580,1464,1485]" pageId="26" pageNumber="555" rank="species">n. sp.</taxonomicNameLabel>
, female (holotype: A±C, F, G, H±J; paratype: D, E), male (paratypes: K±O). (A) Habitus, lateral view. (B) Prosomal end, right lateral view. (C) Prosomal end, left lateral view. (D) Prosomal ends and urosome, dorsal view. (E) Genital double-somite, ventral view. (F, K) Mandibular basis and ®rst endopodal segment, setae showing sexual dimorphism indicated by arrows. (G) Maxillipedal syncoxa. (H) Leg 2, anterior surface, coxa omitted. (I) Leg 3, anterior surface, coxa omitted. (J) Leg 4, anterior surface, coxa omitted. (L) Tuft of short setules on maxillipedal syncoxa. (M) Endopod of leg 2, anterior surface. (N) Endopod of leg 3, anterior surface. (O) Endopod of leg 4, anterior surface.
</paragraph>
</caption>
<caption id="DF6609AEFF98FFC55683FC0BBB6B27DC" ID-DOI="http://doi.org/10.5281/zenodo.5299899" ID-Zenodo-Dep="5299899" httpUri="https://zenodo.org/record/5299899/files/figure.png" pageId="27" pageNumber="556" startId="27.[140,156,1293,1314]" targetBox="[174,1062,133,1264]" targetPageId="27">
<paragraph id="8BA65926FF98FFC55683FC0BBB6B27DC" blockId="27.[140,1095,1293,1501]" pageId="27" pageNumber="556">
FIG. 20.
<taxonomicName id="4C1922A5FF98FFC556FCFC0BB9FD2722" authority="Ohtsuka &amp; Nishida &amp; Nakaguchi, 2002" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[243,504,1293,1315]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="27" pageNumber="556" phylum="Arthropoda" rank="species" species="serratipes" status="sp. nov.">
<emphasis id="B96D8534FF98FFC556FCFC0BB9FD2722" box="[243,504,1293,1315]" italics="true" pageId="27" pageNumber="556">Macandrewella serratipes</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E384FFF98FFC55403FC0BBA4F2723" box="[524,586,1293,1314]" pageId="27" pageNumber="556" rank="species">n. sp.</taxonomicNameLabel>
, male (paratypes). (A) Rostrum, lateral view. (B) Prosomal ends and urosome, dorsal view. (C) Prosomal end, right lateral view. (D) Prosomal end, left lateral view. (E) Right leg 5, anterior surface. (F) Base of right endopod of leg 5, posterior surface. (G) Second and third exopodal segments of right leg 5, anterior surface. (H) Left leg 5, anterior surface. (I) Inner serration along distal half of left endopod of leg 5. (J) Terminal portion of left exopod of leg 5. (K) Right antennulary segments I to XVI±XVII. (L) Right antennulary segments XVIII to XXVII±XXVIII. (M) Left antennulary segments XXII to XXIV.
</paragraph>
</caption>
<paragraph id="8BA65926FF9FFFC256BCF9C3BB4D233D" blockId="28.[140,1095,133,1687]" pageId="28" pageNumber="557">
Antennule exceeding beyond prosomal ends by last two segments (®gure 19A); armature and fusion pattern of segments as in
<taxonomicName id="4C1922A5FF9FFFC254C7F9E0BB5422FD" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[712,849,229,253]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="stygiana">
<emphasis id="B96D8534FF9FFFC254C7F9E0BB5422FD" box="[712,849,229,253]" italics="true" pageId="28" pageNumber="557">M. stygiana</emphasis>
</taxonomicName>
. Antenna, mandible (®gure 19F), maxillule and maxilla similar to those of
<taxonomicName id="4C1922A5FF9FFFC25524F800BBB0231D" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[811,949,261,285]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="stygiana">
<emphasis id="B96D8534FF9FFFC25524F800BBB0231D" box="[811,949,261,285]" italics="true" pageId="28" pageNumber="557">M. stygiana</emphasis>
</taxonomicName>
. Maxilliped (®gure 19G) with proximal patch of long setules on syncoxa.
</paragraph>
<paragraph id="8BA65926FF9FFFC256BCF843BAEF239D" blockId="28.[140,1095,133,1687]" pageId="28" pageNumber="557">
Legs 1±4 (®gure 19H±J) basically resembling those of
<taxonomicName id="4C1922A5FF9FFFC25510F840BBA6235D" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[799,931,325,349]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="stygiana">
<emphasis id="B96D8534FF9FFFC25510F840BBA6235D" box="[799,931,325,349]" italics="true" pageId="28" pageNumber="557">M. stygiana</emphasis>
</taxonomicName>
(see ®gure 6), but diOEering in number of minute prominences. Leg 4 (®gure 19J) lacking large prominences on both surfaces of rami. Leg 5 absent.
</paragraph>
<paragraph id="8BA65926FF9FFFC256BCF8A2BC0A203D" blockId="28.[140,1095,133,1687]" pageId="28" pageNumber="557">
<emphasis id="B96D8534FF9FFFC256BCF8A2B8E823BD" box="[179,237,420,444]" italics="true" pageId="28" pageNumber="557">Male</emphasis>
: Habitus similar to two new species described above; rostrum (®gure 20A) with pair of sharply pointed prominences beneath bases of ®laments; prosomal ends (®gure 20B±D) slightly asymmetrical, with right dorsolateral prominence larger than left. Urosome (®gure 20B) with ®rst two somites asymmetrical; genital somite with small round dorsal knob on both sides; second urosomite swollen on left side.
</paragraph>
<paragraph id="8BA65926FF9FFFC256BCFB45BBDE211B" blockId="28.[140,1095,133,1687]" pageId="28" pageNumber="557">
Antennules (®gure 20K±M) reaching posterior margin of third urosomite; armature and fusion pattern of segments of both antennules similar to two new species described above. Antenna with proximal-most seta on endopod longer than in female, as in
<emphasis id="B96D8534FF9FFFC25722FBA2B9BB20BA" box="[301,446,676,699]" italics="true" pageId="28" pageNumber="557">
<taxonomicName id="4C1922A5FF9FFFC25722FBA2B9B020BA" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[301,437,676,699]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="stygiana">M. stygiana</taxonomicName>
.
</emphasis>
Mandible with two basal setae shorter than in female (®gure 19K, arrowed); second seta on ®rst endopodal segment diOEering from female in being plumose, instead of spinulose. Maxillule and maxilla similar to female. Maxilliped (®gure 19L) bearing proximal tuft of short setules on syncoxa.
</paragraph>
<paragraph id="8BA65926FF9FFFC256BCFA25BB72217A" blockId="28.[140,1095,133,1687]" pageId="28" pageNumber="557">
Legs 1±4 similar to female. Number of large processes on posterior surface of endopods of legs 2 and 3 (®gure 19M, N) as in female. Leg 4 (®gure 19O) furnished only with minute prominences on both surfaces, as in
<emphasis id="B96D8534FF9FFFC254F1FA62BB72217A" box="[766,887,868,891]" italics="true" pageId="28" pageNumber="557">
<taxonomicName id="4C1922A5FF9FFFC254F1FA62BB76217A" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[766,883,868,891]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="omorii">M. omorii</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BA65926FF9FFFC256BCFA85BA9326B8" blockId="28.[140,1095,133,1687]" pageId="28" pageNumber="557">
Leg 5 (®gure 20E±J) similar to that of
<taxonomicName id="4C1922A5FF9FFFC254BAFA82BB35219A" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[693,816,900,923]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="omorii">
<emphasis id="B96D8534FF9FFFC254BAFA82BB35219A" box="[693,816,900,923]" italics="true" pageId="28" pageNumber="557">M. omorii</emphasis>
</taxonomicName>
, except for: right leg (®gure 20E±G) with relatively long protopod; endopod abruptly curved outward at distal quarter; ®rst exopodal segment bearing minute outer prominence midway, without inner irregular knobs except proximal swelling; second exopodal segment with two inner proximal processes, ®rst process longer and thicker, second process with truncate tip; third exopodal segment evenly curved inward, with outer middle process. Left leg (®gure 20H±J) with endopod bearing serration along posterior half of inner margin; ®rst exopodal segment with two low round protuberances at midlength; second exopodal segment slightly produced into round projection at outer distal corner; distal processes as in ®gure 20J.
</paragraph>
<paragraph id="8BA65926FF9FFFC256BCFDC4BA202759" blockId="28.[140,1095,133,1687]" pageId="28" pageNumber="557">
<emphasis id="B96D8534FF9FFFC256BCFDC4B92726D8" box="[179,290,1218,1241]" italics="true" pageId="28" pageNumber="557">Variation.</emphasis>
The number and position of minute prominences on the anterior and posterior surfaces of rami of legs 2±4 vary among individuals, and even between the right and left legs of the same individual. The outer middle process on the third exopodal segment of the male right leg 5 is round at the tip in
<specimenCount id="9D1F92AFFF9FFFC25570FC27BC132738" box="[895,1046,1313,1337]" pageId="28" pageNumber="557" type="generic" typeStatus="paratype">one paratype</specimenCount>
but curved distally in another
<typeStatus id="54A2E784FF9FFFC257B4FC47BA192758" box="[443,540,1345,1369]" pageId="28" pageNumber="557" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="8BA65926FF9FFFC356BCFC67BB6D231D" blockId="28.[140,1095,133,1687]" lastBlockId="29.[140,1095,133,763]" lastPageId="29" lastPageNumber="558" pageId="28" pageNumber="557">
<emphasis id="B96D8534FF9FFFC256BCFC67B91B2778" box="[179,286,1377,1401]" italics="true" pageId="28" pageNumber="557">Remarks.</emphasis>
<bibRefCitation id="EF8824D7FF9FFFC25737FC67BA022778" author="GIESBRECHT, W." box="[312,519,1377,1401]" pageId="28" pageNumber="557" pagination="328" refId="ref12632" refString="GIESBRECHT, W., 1896, U E ber pelagische Copepoden des Rothen Meeres, gesammelt vom Marinestabsarrzt Dr. Augustin Kramer, Zoologische Jahrbucher. Abteilung fuEr Systematik, Geographie und Biologie, 9, 315 ± 328." type="journal article" year="1896">Giesbrecht (1896)</bibRefCitation>
described
<taxonomicName id="4C1922A5FF9FFFC2548BFC67BB6C2778" authorityName="Giesbrecht" authorityYear="1896" box="[644,873,1377,1401]" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9FFFC2548BFC67BB6C2778" box="[644,873,1377,1401]" italics="true" pageId="28" pageNumber="557">Scolecithrix chelipes</emphasis>
</taxonomicName>
from the Red Sea, on the basis of a single male. Later,
<bibRefCitation id="EF8824D7FF9FFFC25443FC87BAD82798" author="SCOTT, A." box="[588,733,1409,1433]" pageId="28" pageNumber="557" pagination="408" refId="ref14030" refString="SCOTT, A., 1909, The Copepoda of the Siboga Expedition. Part 1. Free-swimming, littoral and semi-parasitic Copepoda, Siboga-Expeditie, Monograph, 29 a, 1 ± 408." type="journal article" year="1909">Scott (1909)</bibRefCitation>
transferred this species to the genus
<taxonomicName id="4C1922A5FF9FFFC256D9FCA7B97B27B8" authorityName="Scott" authorityYear="1909" box="[214,382,1441,1465]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="genus">
<emphasis id="B96D8534FF9FFFC256D9FCA7B97B27B8" box="[214,382,1441,1465]" italics="true" pageId="28" pageNumber="557">Macandrewella</emphasis>
</taxonomicName>
when he established it with the description of a new species,
<taxonomicName id="4C1922A5FF9FFFC25683FCC7B8FA27D9" authorityName="Scott" authorityYear="1909" box="[140,255,1473,1496]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="joanae">
<emphasis id="B96D8534FF9FFFC25683FCC7B8FA27D9" box="[140,255,1473,1496]" italics="true" pageId="28" pageNumber="557">M. joanae</emphasis>
</taxonomicName>
. The male of
<taxonomicName id="4C1922A5FF9FFFC257A3FCC7BA2827D9" authorityName="Giesbrecht" authorityYear="1896" box="[428,557,1472,1496]" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9FFFC257A3FCC7BA2827D9" box="[428,557,1472,1496]" italics="true" pageId="28" pageNumber="557">M. chelipes</emphasis>
</taxonomicName>
was subsequently recorded from the
<collectingCountry id="F30E19B6FF9FFFC255E6FCC6B8D827F9" name="India" pageId="28" pageNumber="557">Nicobar Islands</collectingCountry>
in the Indian Ocean (
<bibRefCitation id="EF8824D7FF9FFFC257D6FCE6BA6227F9" author="SEWELL, R. B. S." box="[473,615,1504,1528]" pageId="28" pageNumber="557" pagination="221" refId="ref14103" refString="SEWELL, R. B. S., 1929, The Copepoda of Indian Seas, Memoirs of the Indian Museum, 10, 1 ± 221." type="journal article" year="1929">Sewell, 1929</bibRefCitation>
). The female was ®rst described from the southern Paci®c by
<bibRefCitation id="EF8824D7FF9FFFC25769FF06BA0C2419" author="WILSON, C. B." box="[358,521,1536,1560]" pageId="28" pageNumber="557" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilson (1950)</bibRefCitation>
; however the corresponding male is rather diOEerent from those previously recorded from the Indian Ocean. Recently, both sexes of the species have been redescribed from the Gulf of Elat, Red Sea, by
<bibRefCitation id="EF8824D7FF9FFFC2557AFF46BC472459" author="CAMPANER, A. F." box="[885,1090,1600,1624]" pageId="28" pageNumber="557" pagination="235" refId="ref12162" refString="CAMPANER, A. F., 1989, Supplementary description of Macandrewella chelipes (Giesbrecht, 1896) from the Gulf of Elat (Copepoda: Calanoida: Scolecithricidae), and comments on its relationships with Scottocalanus Sars and Scolecocalanus Farran, Israel Journal of Zoology, 35, 229 ± 235." type="journal article" year="1989">Campaner (1989)</bibRefCitation>
. Unfortunately
<bibRefCitation id="EF8824D7FF9FFFC25733FF66BA092479" author="CAMPANER, A. F." box="[316,524,1632,1656]" pageId="28" pageNumber="557" pagination="235" refId="ref12162" refString="CAMPANER, A. F., 1989, Supplementary description of Macandrewella chelipes (Giesbrecht, 1896) from the Gulf of Elat (Copepoda: Calanoida: Scolecithricidae), and comments on its relationships with Scottocalanus Sars and Scolecocalanus Farran, Israel Journal of Zoology, 35, 229 ± 235." type="journal article" year="1989">Campaner (1989)</bibRefCitation>
described only the lens, rostrum, parts of the mouthparts, and legs 1±3 of the female of
<taxonomicName id="4C1922A5FF9FFFC2548FFF86BB042496" authorityName="Giesbrecht" authorityYear="1896" box="[640,769,1663,1687]" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="28" pageNumber="557" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9FFFC2548FFF86BB042496" box="[640,769,1663,1687]" italics="true" pageId="28" pageNumber="557">M. chelipes</emphasis>
</taxonomicName>
, but not the habitus, which makes detailed comparison between the females of
<taxonomicName id="4C1922A5FF9EFFC354E3F980BB6B229C" authorityName="Giesbrecht" authorityYear="1896" box="[748,878,133,157]" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9EFFC354E3F980BB6B229C" box="[748,878,133,157]" italics="true" pageId="29" pageNumber="558">M. chelipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1922A5FF9EFFC355A0F980BC43229C" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[943,1094,134,157]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="serratipes">
<emphasis id="B96D8534FF9EFFC355A0F980BC43229C" box="[943,1094,134,157]" italics="true" pageId="29" pageNumber="558">M. serratipes</emphasis>
</taxonomicName>
di cult. Although other congeners have a restricted distribution (see ®gure 21),
<taxonomicName id="4C1922A5FF9EFFC35683F9C0B90E22DC" authorityName="Giesbrecht" authorityYear="1896" box="[140,267,197,221]" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9EFFC35683F9C0B90E22DC" box="[140,267,197,221]" italics="true" pageId="29" pageNumber="558">M. chelipes</emphasis>
</taxonomicName>
apparently had a broad distribution in the Red Sea, the Indian Ocean, and the southern Paci®c. Hence we suspected that
<bibRefCitation id="EF8824D7FF9EFFC354C1F9E3BB8622FC" author="WILSON, C. B." box="[718,899,229,253]" pageId="29" pageNumber="558" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilsons (1950)</bibRefCitation>
specimens of the southern Paci®c
<taxonomicName id="4C1922A5FF9EFFC35744F800B9CE231C" authorityName="Giesbrecht" authorityYear="1896" box="[331,459,261,285]" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9EFFC35744F800B9CE231C" box="[331,459,261,285]" italics="true" pageId="29" pageNumber="558">M. chelipes</emphasis>
</taxonomicName>
might correspond to a new species.
</paragraph>
<paragraph id="8BA65926FF9EFFC356BCF823B998207A" blockId="29.[140,1095,133,763]" pageId="29" pageNumber="558">
The male of the new species is similar to those of
<taxonomicName id="4C1922A5FF9EFFC3552CF820BC3B233C" authority="Farran, 1936" authorityName="Farran" authorityYear="1936" box="[803,1086,293,317]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="sewelli">
<emphasis id="B96D8534FF9EFFC3552CF820BB9E233C" box="[803,923,293,317]" italics="true" pageId="29" pageNumber="558">M. sewelli</emphasis>
<bibRefCitation id="EF8824D7FF9EFFC355A8F823BC3B233C" author="FARRAN, G. P." box="[935,1086,293,317]" pageId="29" pageNumber="558" pagination="142" refId="ref12335" refString="FARRAN, G. P., 1936, Copepoda, Great Barrier Reef Expedition, 1928 ± 1929, Scienti W c Reports, 5, 73 ± 142." type="journal article" year="1936">Farran, 1936</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1922A5FF9EFFC35683F840B8FA235D" authorityName="Scott" authorityYear="1909" box="[140,255,325,349]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="joanae">
<emphasis id="B96D8534FF9EFFC35683F840B8FA235D" box="[140,255,325,349]" italics="true" pageId="29" pageNumber="558">M. joanae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1922A5FF9EFFC3571CF840B982235D" authority="Ohtsuka &amp; Nishida &amp; Nakaguchi, 2002" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[275,391,325,349]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="omorii" status="sp. nov.">
<emphasis id="B96D8534FF9EFFC3571CF840B982235D" box="[275,391,325,349]" italics="true" pageId="29" pageNumber="558">M. omorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E384FFF9EFFC3579BF843B9D0235D" box="[404,469,325,348]" pageId="29" pageNumber="558" rank="species">n. sp.</taxonomicNameLabel>
and, possibly,
<taxonomicName id="4C1922A5FF9EFFC35482F840BB43235D" authorityName="Farran" authorityYear="1936" box="[653,838,325,349]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="asymmetrica">
<emphasis id="B96D8534FF9EFFC35482F840BB43235D" box="[653,838,325,349]" italics="true" pageId="29" pageNumber="558">M. asymmetrica</emphasis>
</taxonomicName>
, all of which share a serration along the distal half of the inner margin of the left endopod of leg 5. However, the new species is distinguishable from the latter three species by the following characters: (1) the second exopodal segment of left leg 5 without a large triangular process at the outer distal corner; (2) the second exopodal segment of right leg 5 has a proximal inner process much shorter than the segment proper. Although
<taxonomicName id="4C1922A5FF9EFFC3570BFB03B978201D" authorityName="Farran" authorityYear="1936" box="[260,381,516,540]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="sewelli">
<emphasis id="B96D8534FF9EFFC3570BFB03B978201D" box="[260,381,516,540]" italics="true" pageId="29" pageNumber="558">M. sewelli</emphasis>
</taxonomicName>
bears also a relatively short inner proximal process on the second exopodal segment of the right leg, it has a large outer digitiform process at the outer distal corner of the ®rst exopodal segment of the right leg, and a truncate tip on the left endopod.
</paragraph>
<paragraph id="8BA65926FF9EFFC056BCFB85B93422DC" blockId="29.[140,1095,133,763]" lastBlockId="30.[140,1095,133,476]" lastPageId="30" lastPageNumber="559" pageId="29" pageNumber="558">
<bibRefCitation id="EF8824D7FF9EFFC356BCFB85B95D209A" author="WILSON, C. B." box="[179,344,643,667]" pageId="29" pageNumber="558" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilson (1950)</bibRefCitation>
erroneously assigned the male of the new species to
<taxonomicName id="4C1922A5FF9EFFC355CAFB82B94B20BA" authority="Giesbrecht, 1896" authorityName="Giesbrecht" authorityYear="1896" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9EFFC355CAFB82BC43209A" box="[965,1094,643,667]" italics="true" pageId="29" pageNumber="558">M. chelipes</emphasis>
<bibRefCitation id="EF8824D7FF9EFFC35683FBA5B94B20BA" author="GIESBRECHT, W." box="[140,334,675,699]" pageId="29" pageNumber="558" pagination="328" refId="ref12632" refString="GIESBRECHT, W., 1896, U E ber pelagische Copepoden des Rothen Meeres, gesammelt vom Marinestabsarrzt Dr. Augustin Kramer, Zoologische Jahrbucher. Abteilung fuEr Systematik, Geographie und Biologie, 9, 315 ± 328." type="journal article" year="1896">Giesbrecht, 1896</bibRefCitation>
</taxonomicName>
, but the latter is distinctly diOEerent as follows: (1) the second exopodal segment of right leg 5 has a single proximal inner process, which is directed proximally and tapers distally (two processes, the tips of which are truncate, are displayed in
<taxonomicName id="4C1922A5FF9DFFC0572EF980B9B2229C" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[289,439,134,157]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="30" pageNumber="559" phylum="Arthropoda" rank="species" species="serratipes">
<emphasis id="B96D8534FF9DFFC0572EF980B9B2229C" box="[289,439,134,157]" italics="true" pageId="30" pageNumber="559">M. serratipes</emphasis>
</taxonomicName>
); (2) the third exopodal segment of right leg 5 with an outer process half as long as the segment (one-tenth as long as the segment in
<taxonomicName id="4C1922A5FF9DFFC05683F9C0B92522DD" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[140,288,197,221]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="30" pageNumber="559" phylum="Arthropoda" rank="species" species="serratipes">
<emphasis id="B96D8534FF9DFFC05683F9C0B92522DD" box="[140,288,197,221]" italics="true" pageId="30" pageNumber="559">M. serratipes</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF6609AEFF9EFFC35683FCE5BC372499" ID-DOI="http://doi.org/10.5281/zenodo.5299901" ID-Zenodo-Dep="5299901" httpUri="https://zenodo.org/record/5299901/files/figure.png" pageId="29" pageNumber="558" startId="29.[140,156,1507,1528]" targetBox="[144,1093,815,1477]" targetPageId="29">
<paragraph id="8BA65926FF9EFFC35683FCE5BC372499" blockId="29.[140,1095,1507,1688]" pageId="29" pageNumber="558">
FIG. 21. Distribution of
<taxonomicName id="4C1922A5FF9EFFC35795FCE5BA3727F8" authorityName="Scott" authorityYear="1909" box="[410,562,1507,1529]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="genus">
<emphasis id="B96D8534FF9EFFC35795FCE5BA3727F8" box="[410,562,1507,1529]" italics="true" pageId="29" pageNumber="558">Macandrewella</emphasis>
</taxonomicName>
. Note that their distribution is restricted between 30
<emphasis id="B96D8534FF9EFFC356ECFCFBB8E92412" bold="true" box="[227,236,1533,1555]" italics="true" pageId="29" pageNumber="558">ss</emphasis>
N and 20
<emphasis id="B96D8534FF9EFFC35740FCFBB95D2412" bold="true" box="[335,344,1533,1555]" italics="true" pageId="29" pageNumber="558">ss</emphasis>
S in the Indo-Paci®c. (1)
<taxonomicName id="4C1922A5FF9EFFC3546BFCF8BB9B2412" authority="(Campaner, 1989)" baseAuthorityName="Campaner" baseAuthorityYear="1989" box="[612,926,1533,1555]" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9EFFC3546BFCF8BADD2412" box="[612,728,1533,1555]" italics="true" pageId="29" pageNumber="558">M. chelipes</emphasis>
(
<bibRefCitation id="EF8824D7FF9EFFC354E8FCFBBB942412" author="CAMPANER, A. F." box="[743,913,1533,1555]" pageId="29" pageNumber="558" pagination="235" refId="ref12162" refString="CAMPANER, A. F., 1989, Supplementary description of Macandrewella chelipes (Giesbrecht, 1896) from the Gulf of Elat (Copepoda: Calanoida: Scolecithricidae), and comments on its relationships with Scottocalanus Sars and Scolecocalanus Farran, Israel Journal of Zoology, 35, 229 ± 235." type="journal article" year="1989">Campaner, 1989</bibRefCitation>
)
</taxonomicName>
. (2)
<taxonomicName id="4C1922A5FF9EFFC355DCFCF8B997242C" authority="(Giesbrecht, 1896)" baseAuthorityName="Giesbrecht" baseAuthorityYear="1896" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9EFFC355DCFCF8BC422412" box="[979,1095,1533,1555]" italics="true" pageId="29" pageNumber="558">M. chelipes</emphasis>
(
<bibRefCitation id="EF8824D7FF9EFFC356DDFF1EB980242F" author="GIESBRECHT, W." box="[210,389,1560,1582]" pageId="29" pageNumber="558" pagination="328" refId="ref12632" refString="GIESBRECHT, W., 1896, U E ber pelagische Copepoden des Rothen Meeres, gesammelt vom Marinestabsarrzt Dr. Augustin Kramer, Zoologische Jahrbucher. Abteilung fuEr Systematik, Geographie und Biologie, 9, 315 ± 328." type="journal article" year="1896">Giesbrecht, 1896</bibRefCitation>
)
</taxonomicName>
. (3)
<taxonomicName id="4C1922A5FF9EFFC357DFFF1FBB70242C" authority="(Gopalakrishnan, 1973)" baseAuthorityName="Gopalakrishnan" baseAuthorityYear="1973" box="[464,885,1560,1582]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="cochinensis">
<emphasis id="B96D8534FF9EFFC357DFFF1FBA6C242F" box="[464,617,1560,1582]" italics="true" pageId="29" pageNumber="558">M. cochinensis</emphasis>
(
<bibRefCitation id="EF8824D7FF9EFFC35472FF1EBB6D242F" author="GOPALAKRISHNAN, T. C." box="[637,872,1560,1582]" pageId="29" pageNumber="558" pagination="189" refId="ref12793" refString="GOPALAKRISHNAN, T. C., 1973, A new species of Macandrewella (Copepoda: Calanoida) from oOE Cochin, south west coast of India, Handbook to International Zooplankton Collections, Indian Ocean Biological Centre, 5, 180 ± 189." type="journal article" year="1973">Gopalakrishnan, 1973</bibRefCitation>
)
</taxonomicName>
. (4)
<taxonomicName id="4C1922A5FF9EFFC355BCFF1FBC16242F" authorityName="Sewell" authorityYear="1929" box="[947,1043,1561,1582]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="scotti">
<emphasis id="B96D8534FF9EFFC355BCFF1FBC16242F" box="[947,1043,1561,1582]" italics="true" pageId="29" pageNumber="558">M. scotti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1922A5FF9EFFC356C3FF35B9D52448" authority="(Sewell, 1929)" baseAuthorityName="Sewell" baseAuthorityYear="1929" box="[204,464,1587,1609]" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9EFFC356C3FF35B9392448" box="[204,316,1587,1609]" italics="true" pageId="29" pageNumber="558">M. chelipes</emphasis>
(
<bibRefCitation id="EF8824D7FF9EFFC35743FF35B9CC2448" author="SEWELL, R. B. S." box="[332,457,1587,1609]" pageId="29" pageNumber="558" pagination="221" refId="ref14103" refString="SEWELL, R. B. S., 1929, The Copepoda of Indian Seas, Memoirs of the Indian Museum, 10, 1 ± 221." type="journal article" year="1929">Sewell, 1929</bibRefCitation>
)
</taxonomicName>
. (5)
<taxonomicName id="4C1922A5FF9EFFC3540BFF35BB032448" authority="(Wilson, 1950)" baseAuthorityName="Wilson" baseAuthorityYear="1950" box="[516,774,1587,1609]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="sewelli">
<emphasis id="B96D8534FF9EFFC3540BFF35BA6F2448" box="[516,618,1587,1609]" italics="true" pageId="29" pageNumber="558">M. sewelli</emphasis>
(
<bibRefCitation id="EF8824D7FF9EFFC35475FF35BAFA2448" author="WILSON, C. B." box="[634,767,1587,1609]" pageId="29" pageNumber="558" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilson, 1950</bibRefCitation>
)
</taxonomicName>
. (6)
<taxonomicName id="4C1922A5FF9EFFC35535FF35BBAA2449" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[826,943,1587,1608]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="stygiana">
<emphasis id="B96D8534FF9EFFC35535FF35BBAA2449" box="[826,943,1587,1608]" italics="true" pageId="29" pageNumber="558">M. stygiana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1922A5FF9EFFC355ECFF35BC402449" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[995,1093,1587,1608]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="omorii">
<emphasis id="B96D8534FF9EFFC355ECFF35BC402449" box="[995,1093,1587,1608]" italics="true" pageId="29" pageNumber="558">M. omorii</emphasis>
</taxonomicName>
(present study). (7)
<taxonomicName id="4C1922A5FF9EFFC3579BFF48BA862463" authority="(Scott, 1909)" baseAuthorityName="Scott" baseAuthorityYear="1909" box="[404,643,1613,1635]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="joanae">
<emphasis id="B96D8534FF9EFFC3579BFF48B9F32462" box="[404,502,1614,1635]" italics="true" pageId="29" pageNumber="558">M. joanae</emphasis>
(
<bibRefCitation id="EF8824D7FF9EFFC3540BFF4BBA732462" author="SCOTT, A." box="[516,630,1613,1635]" pageId="29" pageNumber="558" pagination="408" refId="ref14030" refString="SCOTT, A., 1909, The Copepoda of the Siboga Expedition. Part 1. Free-swimming, littoral and semi-parasitic Copepoda, Siboga-Expeditie, Monograph, 29 a, 1 ± 408." type="journal article" year="1909">Scott, 1909</bibRefCitation>
)
</taxonomicName>
. (8)
<taxonomicName id="4C1922A5FF9EFFC354BEFF48BB572462" authorityName="Farran" authorityYear="1936" box="[689,850,1614,1635]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="asymmetrica">
<emphasis id="B96D8534FF9EFFC354BEFF48BB572462" box="[689,850,1614,1635]" italics="true" pageId="29" pageNumber="558">M. asymmetrica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1922A5FF9EFFC35552FF48BBB72462" authorityName="Farran" authorityYear="1936" box="[861,946,1614,1635]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="mera">
<emphasis id="B96D8534FF9EFFC35552FF48BBB72462" box="[861,946,1614,1635]" italics="true" pageId="29" pageNumber="558">M. mera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1922A5FF9EFFC355EDFF48B96D247C" authority="(Farran, 1936)" baseAuthorityName="Farran" baseAuthorityYear="1936" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="sewelli">
<emphasis id="B96D8534FF9EFFC355EDFF48BC422462" box="[994,1095,1613,1635]" italics="true" pageId="29" pageNumber="558">M. sewelli</emphasis>
(
<bibRefCitation id="EF8824D7FF9EFFC356D9FF6EB959247F" author="FARRAN, G. P." box="[214,348,1640,1662]" pageId="29" pageNumber="558" pagination="142" refId="ref12335" refString="FARRAN, G. P., 1936, Copepoda, Great Barrier Reef Expedition, 1928 ± 1929, Scienti W c Reports, 5, 73 ± 142." type="journal article" year="1936">Farran, 1936</bibRefCitation>
)
</taxonomicName>
. (9)
<taxonomicName id="4C1922A5FF9EFFC35791FF6FBAB2247F" authority="(Wilson, 1950)" baseAuthorityName="Wilson" baseAuthorityYear="1950" box="[414,695,1640,1662]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="agassizi">
<emphasis id="B96D8534FF9EFFC35791FF6FBA16247F" box="[414,531,1641,1662]" italics="true" pageId="29" pageNumber="558">M. agassizi</emphasis>
(
<bibRefCitation id="EF8824D7FF9EFFC35428FF6EBAB5247F" author="WILSON, C. B." box="[551,688,1640,1662]" pageId="29" pageNumber="558" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilson, 1950</bibRefCitation>
)
</taxonomicName>
. (10)
<taxonomicName id="4C1922A5FF9EFFC354F2FF6FBC0F247F" authority="(Wilson, 1950)" baseAuthorityName="Wilson" baseAuthorityYear="1950" box="[765,1034,1640,1662]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="sewelli">
<emphasis id="B96D8534FF9EFFC354F2FF6FBB63247F" box="[765,870,1640,1662]" italics="true" pageId="29" pageNumber="558">M. sewelli</emphasis>
(
<bibRefCitation id="EF8824D7FF9EFFC35575FF6EBC07247F" author="WILSON, C. B." box="[890,1026,1640,1662]" pageId="29" pageNumber="558" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilson, 1950</bibRefCitation>
)
</taxonomicName>
. (11)
<taxonomicName id="4C1922A5FF9EFFC356C3FF85B9542499" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[204,337,1667,1688]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="serratipes">
<emphasis id="B96D8534FF9EFFC356C3FF85B9542499" box="[204,337,1667,1688]" italics="true" pageId="29" pageNumber="558">M. serratipes</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C1922A5FF9EFFC35771FF85BA802499" authority=", Wilson, 1950" authorityName="Wilson" authorityYear="1950" box="[382,645,1666,1688]" class="Hexanauplia" family="Scolecitrichidae" genus="Scolecithrix" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="chelipes">
<emphasis id="B96D8534FF9EFFC35771FF85B9F42499" box="[382,497,1666,1688]" italics="true" pageId="29" pageNumber="558">M. chelipes</emphasis>
,
<bibRefCitation id="EF8824D7FF9EFFC357F0FF84BA802499" author="WILSON, C. B." box="[511,645,1666,1688]" pageId="29" pageNumber="558" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilson, 1950</bibRefCitation>
</taxonomicName>
; see material examined of
<taxonomicName id="4C1922A5FF9EFFC35592FF85BC262499" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[925,1059,1667,1688]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="29" pageNumber="558" phylum="Arthropoda" rank="species" species="serratipes">
<emphasis id="B96D8534FF9EFFC35592FF85BC262499" box="[925,1059,1667,1688]" italics="true" pageId="29" pageNumber="558">M. serratipes</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="8BA65926FF9DFFC056BCF9E3BAF7233D" blockId="30.[140,1095,133,476]" pageId="30" pageNumber="559">
The new species exhibits sexual dimorphism in the antenna and mandible like
<taxonomicName id="4C1922A5FF9DFFC05683F800B915231D" authorityName="Ohtsuka &amp; Nishida &amp; Nakaguchi" authorityYear="2002" box="[140,272,261,285]" class="Hexanauplia" family="Scolecitrichidae" genus="Macandrewella" kingdom="Animalia" order="Calanoida" pageId="30" pageNumber="559" phylum="Arthropoda" rank="species" species="stygiana">
<emphasis id="B96D8534FF9DFFC05683F800B915231D" box="[140,272,261,285]" italics="true" pageId="30" pageNumber="559">M. stygiana</emphasis>
</taxonomicName>
. In addition, the maxillipedal syncoxa bears proximally a patch of long setules in the female, but of short setules in the male.
</paragraph>
<paragraph id="8BA65926FF9DFFC056BCF843BB1B237C" blockId="30.[140,1095,133,476]" pageId="30" pageNumber="559">
<emphasis id="B96D8534FF9DFFC056BCF843B93A235C" box="[179,319,325,349]" italics="true" pageId="30" pageNumber="559">Distribution.</emphasis>
According to
<bibRefCitation id="EF8824D7FF9DFFC057F0F843BA9F235C" author="WILSON, C. B." box="[511,666,325,349]" pageId="30" pageNumber="559" pagination="141" refId="ref14223" refString="WILSON, C. B., 1950, Copepods gathered by the United States Fisheries Steamer`Albatross' from 1887 to 1909, chie ¯ y in the Paci ® c Ocean. Bulletin of the United States National Museum, 100 (14), 141 ± 441." type="journal article" year="1950">Wilson (1950</bibRefCitation>
, note in vials), the new species was collected at a depth of 40 fathoms oOE
<collectingCountry id="F30E19B6FF9DFFC05449F863BA68237C" box="[582,621,357,381]" name="Fiji" pageId="30" pageNumber="559">Fiji</collectingCountry>
(see ®gure 21).
</paragraph>
<paragraph id="8BA65926FF9DFFC056BCF882B8F423DD" blockId="30.[140,1095,133,476]" pageId="30" pageNumber="559">
<emphasis id="B96D8534FF9DFFC056BCF882B937239D" box="[179,306,388,412]" italics="true" pageId="30" pageNumber="559">Etymology.</emphasis>
The speci®c name,
<emphasis id="B96D8534FF9DFFC05420F882BA9B239D" box="[559,670,388,412]" italics="true" pageId="30" pageNumber="559">`serratipes</emphasis>
is derived from the inner serration along the distal half of the left endopod of male leg 5 (Latin
<emphasis id="B96D8534FF9DFFC05544F8A3BBA523BD" box="[843,928,421,444]" italics="true" pageId="30" pageNumber="559">serratus</emphasis>
, serrated, and
<emphasis id="B96D8534FF9DFFC05683F8C3B8AA23DD" box="[140,175,453,476]" italics="true" pageId="30" pageNumber="559">pes</emphasis>
, leg).
</paragraph>
</subSubSection>
</treatment>
</document>