treatments-xml/data/03/B0/B2/03B0B202FFF8BF7AC0AAF372FE8AF9F1.xml
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<document id="541CB72C5F64168459C23799994D04BD" ID-CLB-Dataset="121335" ID-DOI="10.11646/zootaxa.5174.4.2" ID-GBIF-Dataset="d6b7a54e-37b1-44c0-963d-c68720fa9e97" ID-ISSN="1175-5326" ID-Zenodo-Dep="6986320" ID-ZooBank="B09A5F4E-31FA-44CE-BF50-007D1DEF43D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660326602367" checkinUser="plazi" docAuthor="Janák, Jiří" docDate="2022" docId="03B0B202FFF8BF7AC0AAF372FE8AF9F1" docLanguage="en" docName="zootaxa.5174.4.2.pdf" docOrigin="Zootaxa 5174 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stenus spiniferoides Janak 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="350" masterDocId="FF89CA7AFFEABF69C03DF62DFF81FFDC" masterDocTitle="Four new species and additional records of the genus Stenus of southern Africa (Coleoptera: Staphylinidae: Steninae)" masterLastPageNumber="356" masterPageNumber="331" pageNumber="349" updateTime="1699366806495" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3337E7ABD1DC47A8F739CF4818CF2D31">Four new species and additional records of the genus Stenus of southern Africa (Coleoptera: Staphylinidae: Steninae)</mods:title>
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<mods:name id="AF9B72645C275C53423FD892EB046BBB" type="personal">
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<mods:namePart id="817E170D2597AD7BB198147133C9DE28">Janák, Jiří</mods:namePart>
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<mods:title id="F7BC2307201AD9BCF4D1F4D8DAD40E6D">Zootaxa</mods:title>
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<mods:part id="DB69CE150BC284EFED4DE109FD6F473F">
<mods:date id="36E468502CAA198005033105AEF9C453">2022</mods:date>
<mods:detail id="796B06F89B1C70003C0E3A305BCC8E82" type="pubDate">
<mods:number id="8C975714FED3FEEB1C56F6ABF2BCD6BA">2022-08-11</mods:number>
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<mods:number id="71DD6BBA25812E837A4331AC99B3021A">5174</mods:number>
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<mods:classification id="73886DD37EAFEC0C4C184FEDF7EDC9B9">journal article</mods:classification>
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<mods:identifier id="0CD8ECA9CD15B6B42D3D8AAD8E83DBC4" type="DOI">10.11646/zootaxa.5174.4.2</mods:identifier>
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<treatment id="03B0B202FFF8BF7AC0AAF372FE8AF9F1" ID-DOI="http://doi.org/10.5281/zenodo.6986799" ID-GBIF-Taxon="199056736" ID-Zenodo-Dep="6986799" LSID="urn:lsid:plazi:treatment:03B0B202FFF8BF7AC0AAF372FE8AF9F1" httpUri="http://treatment.plazi.org/id/03B0B202FFF8BF7AC0AAF372FE8AF9F1" lastPageId="19" lastPageNumber="350" pageId="18" pageNumber="349">
<subSubSection id="C303509FFFF8BF7BC0AAF372FDCAFAA5" box="[151,587,1374,1402]" pageId="18" pageNumber="349" type="nomenclature">
<paragraph id="8BA60314FFF8BF7BC0AAF372FDCAFAA5" blockId="18.[151,587,1374,1438]" box="[151,587,1374,1402]" pageId="18" pageNumber="349">
<heading id="D0EEB478FFF8BF7BC0AAF372FDCAFAA5" bold="true" box="[151,587,1374,1402]" fontSize="11" level="1" pageId="18" pageNumber="349" reason="1">
<emphasis id="B96DDF06FFF8BF7BC0AAF372FDCAFAA5" bold="true" box="[151,587,1374,1402]" pageId="18" pageNumber="349">
<taxonomicName id="4C197897FFF8BF7BC0AAF372FE61FAA5" ID-CoL="9GDLW" authority="Janak" authorityName="Janak" authorityYear="2022" box="[151,480,1374,1402]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="spiniferoides" status="sp. nov.">
<emphasis id="B96DDF06FFF8BF7BC0AAF372FE0AFAA6" bold="true" box="[151,395,1375,1402]" italics="true" pageId="18" pageNumber="349">Stenus spiniferoides</emphasis>
Janák
</taxonomicName>
,
<taxonomicNameLabel id="A25E627DFFF8BF7BC1D0F372FDCAFAA5" box="[493,587,1375,1401]" pageId="18" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C303509FFFF8BF7BC0AAF3AEFED9FA41" box="[151,344,1411,1438]" pageId="18" pageNumber="349" type="description">
<paragraph id="8BA60314FFF8BF7BC0AAF3AEFED9FA41" blockId="18.[151,587,1374,1438]" box="[151,344,1411,1438]" pageId="18" pageNumber="349">
(
<figureCitation id="13221F91FFF8BF7BC0A2F3AEFEA9FA42" box="[159,296,1411,1438]" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="18" pageNumber="349">Figs. 5966</figureCitation>
,
<figureCitation id="13221F91FFF8BF7BC10EF3AEFED1FA41" box="[307,336,1411,1437]" captionStart="FIGURES 6772" captionStartId="21.[151,264,1641,1665]" captionTargetBox="[154,1433,181,1617]" captionTargetId="figure-17@21.[154,1433,181,1617]" captionTargetPageId="21" captionText="FIGURES 6772. Localities where Stenus species were collected. S. hippoamicus Janák, sp. nov. (67, St. Lucia; 68, Ndumo); S. volkerputhzianus Janák, sp. nov. (69, Himeville, 70, Tugella river); S. hessei L. Benick, S. capicola L. Benick, S. depressiusculus L. Benick, S. cf. grandipennis L. Benick, 1933 (71, Olifantsbos); S. spiniferoides Janák, sp. nov. (72, Langvlei)." figureDoi="http://doi.org/10.5281/zenodo.6986334" httpUri="https://zenodo.org/record/6986334/files/figure.png" pageId="18" pageNumber="349">72</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C303509FFFF8BF7BC0AAF3E6FA0DF9A9" pageId="18" pageNumber="349" type="materials_examined">
<paragraph id="8BA60314FFF8BF7BC0AAF3E6FCDBFA39" blockId="18.[151,1437,1483,2014]" box="[151,858,1483,1510]" pageId="18" pageNumber="349">
<materialsCitation id="3B710949FFF8BF7BC0AAF3E6FCD7FA39" ID-GBIF-Occurrence="3888061528" box="[151,854,1483,1510]" country="South Africa" location="Bo-Langvlei Lake" pageId="18" pageNumber="349" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="B96DDF06FFF8BF7BC0AAF3E6FEB8FA39" bold="true" box="[151,313,1483,1509]" pageId="18" pageNumber="349">
<typeStatus id="54A2BDB6FFF8BF7BC0AAF3E6FF53FA39" box="[151,210,1483,1509]" pageId="18" pageNumber="349">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F30E4384FFF8BF7BC17DF3E6FE50FA39" box="[320,465,1483,1509]" name="South Africa" pageId="18" pageNumber="349">South Africa</collectingCountry>
,
<collectingRegion id="49DDCDF6FFF8BF7BC1E1F3E1FDFBFA39" box="[476,634,1483,1510]" country="South Africa" name="Western Cape" pageId="18" pageNumber="349">Western Cape</collectingRegion>
,
<location id="8EC655CFFFF8BF7BC2BBF3E6FCD7FA39" LSID="urn:lsid:plazi:treatment:03B0B202FFF8BF7AC0AAF372FE8AF9F1:8EC655CFFFF8BF7BC2BBF3E6FCD7FA39" box="[646,854,1483,1509]" country="South Africa" name="Bo-Langvlei Lake" pageId="18" pageNumber="349" stateProvince="Western Cape">Bo-Langvlei Lake</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA60314FFF8BF7BC0FAF3C2FA0DF9A9" blockId="18.[151,1437,1483,2014]" pageId="18" pageNumber="349">
<emphasis id="B96DDF06FFF8BF7BC0FAF3C2FEF8F9D5" bold="true" box="[199,377,1519,1545]" pageId="18" pageNumber="349">Type material.</emphasis>
<materialsCitation id="3B710949FFF8BF7BC1BEF3C2FEBAF98D" ID-GBIF-Occurrence="3888061426" collectingDate="2012-02-06" collectionCode="TMSA" collectorName="J. Janak" country="South Africa" county="South Africa" latitude="-33.985" location="Bo-Langvlei lake" longLatPrecision="119" longitude="22.673334" municipality="Wilderness" pageId="18" pageNumber="349" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="B96DDF06FFF8BF7BC1BEF3C2FE71F9D5" bold="true" box="[387,496,1519,1545]" pageId="18" pageNumber="349">
<typeStatus id="54A2BDB6FFF8BF7BC1BEF3C2FE71F9D5" box="[387,496,1519,1545]" pageId="18" pageNumber="349" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D1FC89DFFF8BF7BC1C7F3DDFD93F9D6" box="[506,530,1520,1546]" pageId="18" pageNumber="349" type="male"></specimenCount>
:
<emphasis id="B96DDF06FFF8BF7BC21FF3C2FC82F9D5" bold="true" box="[546,771,1519,1545]" pageId="18" pageNumber="349">
<collectingCountry id="F30E4384FFF8BF7BC21FF3C2FD7FF9D5" box="[546,766,1519,1545]" name="South Africa" pageId="18" pageNumber="349">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DDCDF6FFF8BF7BC333F3DDFC30F9D5" box="[782,945,1519,1546]" country="South Africa" name="Western Cape" pageId="18" pageNumber="349">Western Cape</collectingRegion>
: “
<collectingCounty id="62C77B98FFF8BF7BC3F4F3C2FBE1F9D5" box="[969,1120,1519,1545]" pageId="18" pageNumber="349">South Africa</collectingCounty>
,
<collectingRegion id="49DDCDF6FFF8BF7BC452F3DDFA90F9D5" box="[1135,1297,1519,1546]" country="South Africa" name="Western Cape" pageId="18" pageNumber="349">Western Cape</collectingRegion>
,
<collectingMunicipality id="6BC2996EFFF8BF7BC522F3C2FA1DF9D5" box="[1311,1436,1519,1545]" pageId="18" pageNumber="349">Wilderness</collectingMunicipality>
section of Garden Route NP;
<geoCoordinate id="EE2D65D3FFF8BF7BC1D3F03EFDE3F9F2" box="[494,610,1555,1582]" degrees="33" direction="south" minutes="59.1" orientation="latitude" pageId="18" pageNumber="349" precision="92" value="-33.985">33°59.1´S</geoCoordinate>
,
<geoCoordinate id="EE2D65D3FFF8BF7BC24DF03EFD64F9F1" box="[624,741,1555,1581]" degrees="22" direction="east" minutes="40.4" orientation="longitude" pageId="18" pageNumber="349" precision="92" value="22.673334">22°40.4´E</geoCoordinate>
,
<location id="8EC655CFFFF8BF7BC2CEF03EFC38F9F1" LSID="urn:lsid:plazi:treatment:03B0B202FFF8BF7AC0AAF372FE8AF9F1:8EC655CFFFF8BF7BC2CEF03EFC38F9F1" box="[755,953,1555,1581]" country="South Africa" county="South Africa" latitude="-33.985" longLatPrecision="119" longitude="22.673334" municipality="Wilderness" name="Bo-Langvlei lake" pageId="18" pageNumber="349" stateProvince="Western Cape">Bo-Langvlei lake</location>
nr.
<location id="8EC655CFFFF8BF7BC3D4F03EFBCCF9F1" LSID="urn:lsid:plazi:treatment:03B0B202FFF8BF7AC0AAF372FE8AF9F1:8EC655CFFFF8BF7BC3D4F03EFBCCF9F1" box="[1001,1101,1555,1581]" country="South Africa" county="South Africa" latitude="-33.985" longLatPrecision="119" longitude="22.673334" municipality="Wilderness" name="Malachit" pageId="18" pageNumber="349" stateProvince="Western Cape">Malachit</location>
bird hide,
<date id="FFA725D4FFF8BF7BC4F0F03EFAB2F9F1" box="[1229,1331,1555,1581]" pageId="18" pageNumber="349" value="2012-02-06">
<collectingDate id="EFE3DC3CFFF8BF7BC4F0F03EFAB2F9F1" box="[1229,1331,1555,1581]" pageId="18" pageNumber="349" value="2012-02-06">6.ii.2012</collectingDate>
</date>
,
<collectorName id="26EC66C2FFF8BF7BC57CF039FA1DF9F1" box="[1345,1436,1555,1582]" pageId="18" pageNumber="349">J. Janák</collectorName>
lgt. (
<collectionCode id="ED089BD1FFF8BF7BC0F6F01AFE9FF98D" box="[203,286,1591,1617]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="18" pageNumber="349">TMSA</collectionCode>
), “
</materialsCitation>
<materialsCitation id="3B710949FFF8BF7BC106F01AFBF9F98D" ID-GBIF-Occurrence="3888061436" box="[315,1144,1591,1618]" collectionCode="TMSA" country="South Africa" determinerName="J. Janak" pageId="18" pageNumber="349" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="54A2BDB6FFF8BF7BC106F01AFE65F98D" box="[315,484,1591,1617]" pageId="18" pageNumber="349" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C197897FFF8BF7BC1D6F01AFD4FF98D" authority="Janák, 2022" authorityName="Janák" authorityYear="2022" box="[491,718,1591,1617]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="spiniferoides" status="sp. nov.">Stenus spiniferoides</taxonomicName>
<taxonomicNameLabel id="A25E627DFFF8BF7BC2E9F015FCACF98E" box="[724,813,1592,1618]" pageId="18" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
,
<determinerName id="7E7AEFD0FFF8BF7BC305F015FC10F98D" box="[824,913,1591,1618]" pageId="18" pageNumber="349">J. Janák</determinerName>
det. 2021” (
<collectionCode id="ED089BD1FFF8BF7BC421F01AFBF1F98D" box="[1052,1136,1591,1617]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="18" pageNumber="349">TMSA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B710949FFF8BF7BC4BFF015FED0F9A9" ID-GBIF-Occurrence="3888061325" collectionCode="JJRC" country="South Africa" determinerName="J. Janak" pageId="18" pageNumber="349" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus id="54A2BDB6FFF8BF7BC4BFF015FB65F98E" box="[1154,1252,1592,1618]" pageId="18" pageNumber="349" type="paratype">Paratype</typeStatus>
<specimenCount id="9D1FC89DFFF8BF7BC4D6F015FA83F98E" box="[1259,1282,1592,1618]" pageId="18" pageNumber="349" type="female"></specimenCount>
: same data as holotype (
<collectionCode id="ED089BD1FFF8BF7BC135F076FEC8F9A9" box="[264,329,1627,1653]" pageId="18" pageNumber="349">JJRC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B710949FFF8BF7BC161F071FA09F9A9" ID-GBIF-Occurrence="3888061451" box="[348,1416,1627,1654]" country="South Africa" determinerName="J. Janak" pageId="18" pageNumber="349" specimenCount="2" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus id="54A2BDB6FFF8BF7BC161F071FE3FF9AA" box="[348,446,1628,1654]" pageId="18" pageNumber="349" type="paratype">Paratype</typeStatus>
with additional label “
<typeStatus id="54A2BDB6FFF8BF7BC286F076FCDDF9A9" box="[699,860,1627,1653]" pageId="18" pageNumber="349" type="paratype">PARATYPUS</typeStatus>
<taxonomicName id="4C197897FFF8BF7BC35EF076FBC7F9A9" authority="Janák, 2022" authorityName="Janák" authorityYear="2022" box="[867,1094,1627,1653]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="349" phylum="Arthropoda" rank="species" species="spiniferoides" status="sp. nov.">Stenus spiniferoides</taxonomicName>
<taxonomicNameLabel id="A25E627DFFF8BF7BC471F071FB24F9AA" box="[1100,1189,1628,1654]" pageId="18" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
,
<determinerName id="7E7AEFD0FFF8BF7BC48CF071FA88F9A9" box="[1201,1289,1627,1654]" pageId="18" pageNumber="349">J. Janák</determinerName>
det. 2021”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C303509FFFF8BF7AC0FAF052FBCAFB7D" lastPageId="19" lastPageNumber="350" pageId="18" pageNumber="349" type="description">
<paragraph id="8BA60314FFF8BF7BC0FAF052FC03F8AD" blockId="18.[151,1437,1483,2014]" pageId="18" pageNumber="349">
<emphasis id="B96DDF06FFF8BF7BC0FAF052FED0F945" bold="true" box="[199,337,1663,1689]" pageId="18" pageNumber="349">Description</emphasis>
(n = 2). Body length
<quantity id="4CE1AEF1FFF8BF7BC271F052FD22F946" box="[588,675,1663,1690]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="18" pageNumber="349" unit="mm" value="3.6">3.6 mm</quantity>
(HT) and
<quantity id="4CE1AEF1FFF8BF7BC321F052FCF2F946" box="[796,883,1663,1690]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="18" pageNumber="349" unit="mm" value="3.9">3.9 mm</quantity>
(PT), fore parts 2.0 mm (HT) and
<quantity id="4CE1AEF1FFF8BF7BC53EF052FADBF946" box="[1283,1370,1663,1690]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="18" pageNumber="349" unit="mm" value="2.1">2.1 mm</quantity>
(PT). Macropterous, black, shiny, fore parts moderately coarsely and moderately densely punctate, abdomen moderately coarsely and moderately densely punctate; pubescence distinct, recumbent (
<figureCitation id="13221F91FFF8BF7BC3D0F0EAFBC3F93E" box="[1005,1090,1735,1762]" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="18" pageNumber="349">Fig. 59</figureCitation>
). Two first segments and basal parts of segments 35 of antennae reddish, remaining parts of segments more or less infuscate, club brown. Basal segments of maxillary palpi yellow, apical half of segment 3 slightly infuscate, apical three quarters of segment 4 brownish. Basal third of femora reddish, apical two thirds of femora and tibiae brown to dark brown, tarsi brown. Clypeus black, labrum dark brown, moderately densely pubescent.
</paragraph>
<paragraph id="8BA60314FFF8BF7AC0FAF156FB58FF25" blockId="18.[151,1437,1483,2014]" lastBlockId="19.[151,1437,151,1581]" lastPageId="19" lastPageNumber="350" pageId="18" pageNumber="349">Head moderately narrower than elytra (HT 68.1/85.8 = 0.79, PT 0.74), frons broad (average distance between eyes 3336), with two distinct longitudinal furrows, median part as broad as each of lateral parts, moderately elevated, not extending to level of inner eye margins; punctation moderately coarse and dense, diameters of punctures as large as basal diameter of second antennal segment; antennal tubercles, median part and posteriolateral parts largely impunctate, surface smooth, without any reticulation. Antennae moderately long, when reflexed extending approximately to posterior margin of pronotum, penultimate segments slightly longer than broad.</paragraph>
<paragraph id="8BA60314FFF9BF7AC0FAF72EFE5DFE71" blockId="19.[151,1437,151,1581]" pageId="19" pageNumber="350">Pronotum strongly convex, distinctly longer than broad (HT 61.3/55.9 = 1.10), broadest near middle, distinctly constricted behind; with distinct anterior transverse impression and lateral impressions in posterior half, thus anterior margin seems to be elevated; punctation coarse and irregularly dense, diameter of average punctures as large as basal diameter of second antennal segment or slightly larger, punctation in median part in general less dense and with unpunctured elevations.</paragraph>
<paragraph id="8BA60314FFF9BF7AC0FAF79AFC81FD75" blockId="19.[151,1437,151,1581]" pageId="19" pageNumber="350">Elytra subquadrate, moderately broader than head, about as long as broad or slightly broader than long (R length/width HT = 85.3/85.8 = 0.99, PT 0.95), shoulders slightly oblique, sides slightly rounded and widened behind, distinctly constricted in posterior one-third, posterior margin moderately emarginated (sutural length 74 76); sutural and humeral impressions distinct; two lateral impression present, first in middle and second in posterior quarter; punctation markedly coarser than on pronotum, moderately dense, diameter of punctures slightly larger than maximal diameter of second antennal segment, interstices mostly as large as diameter of punctures, in some areas less dense and with few small unpunctured areas.</paragraph>
<paragraph id="8BA60314FFF9BF7AC0FAF49EFCF1FC79" blockId="19.[151,1437,151,1581]" pageId="19" pageNumber="350">
Abdomen quite narrow, basal constriction of segments deep, tergite 7 with distinct membranous fringe apically; punctation on tergite 3 moderately coarse and dense, interstices about one half or up to twice larger than diameter of punctures, on following tergites gradually finer but not sparser; punctures on tergite 5 as large as one medial eye facet, punctures on tergite 7 markedly finer. Tergite 10 apicomedially spinose (
<figureCitation id="13221F91FFF9BF7AC47CF532FB4DFCE5" box="[1089,1228,798,825]" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="19" pageNumber="350">Figs. 6465</figureCitation>
). Legs moderately slender, metatarsi two thirds as long as metatibiae, segment 1 slightly shorter than segments 2 and 3 combined, markedly shorter than the last, segment 4 deeply bilobed. Fore parts and abdomen not reticulate, only tergite 8 with very faint traces of reticulation (visible at magnification of 70 x).
</paragraph>
<paragraph id="8BA60314FFF9BF7AC0FAF582FAA9FBA1" blockId="19.[151,1437,151,1581]" pageId="19" pageNumber="350">
Male. Apical half of femora moderately enlarged. Sternite 6 apico-medially moderately densely and finely punctured and long setose, sternite 7 apico-medially flattened, very finely and sparsely punctured and long setose with posterior margin markedly emarginated, sternite 8 along middle slightly flattened, with deep, inwards narrowed subtriangular emargination in about posterior one-fifth (
<figureCitation id="13221F91FFF9BF7AC33FF236FCD4FBE9" box="[770,853,1051,1077]" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="19" pageNumber="350">Fig. 62</figureCitation>
). Sternite 9 deeply emarginated apico-medially, with long apicolateral teeth (
<figureCitation id="13221F91FFF9BF7AC19EF212FE79FB85" box="[419,504,1087,1113]" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="19" pageNumber="350">Fig. 63</figureCitation>
). Aedeagus
<quantity id="4CE1AEF1FFF9BF7AC2BEF212FD6BFB85" box="[643,746,1087,1113]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="19" pageNumber="350" unit="mm" value="0.67">0.67 mm</quantity>
, narrowed apically, in apical fourth slightly emarginate (
<figureCitation id="13221F91FFF9BF7AC552F212FF34FBA1" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="19" pageNumber="350">Fig. 61</figureCitation>
), parameres exceed markedly the median lobe, widened apically, with numerous (about 3133) setae.
</paragraph>
<paragraph id="8BA60314FFF9BF7AC0FAF2AAFBCAFB7D" blockId="19.[151,1437,151,1581]" box="[199,1099,1159,1185]" pageId="19" pageNumber="350">
Female. Sternite 8 broadly rounded apically. Valvifer acute posteriorly (
<figureCitation id="13221F91FFF9BF7AC3D7F2AAFBBEFB7D" box="[1002,1087,1159,1185]" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="19" pageNumber="350">Fig. 66</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C303509FFFF9BF7AC0FAF287FC0DFAA5" pageId="19" pageNumber="350" type="diagnosis">
<paragraph id="8BA60314FFF9BF7AC0FAF287FC0DFAA5" blockId="19.[151,1437,151,1581]" pageId="19" pageNumber="350">
<emphasis id="B96DDF06FFF9BF7AC0FAF287FE4FFB18" bold="true" box="[199,462,1194,1220]" pageId="19" pageNumber="350">Differential diagnosis.</emphasis>
<taxonomicName id="4C197897FFF9BF7AC1E9F286FD48FB19" authority="Janak" authorityName="Janak" authorityYear="2022" box="[468,713,1195,1221]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="350" phylum="Arthropoda" rank="species" species="spiniferoides" status="sp. nov.">
<emphasis id="B96DDF06FFF9BF7AC1E9F286FD01FB19" box="[468,640,1195,1221]" italics="true" pageId="19" pageNumber="350">S. spiniferoides</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B96DDF06FFF9BF7AC2E9F286FCACFB19" bold="true" box="[724,813,1195,1221]" pageId="19" pageNumber="350">
<taxonomicNameLabel id="A25E627DFFF9BF7AC2E9F286FCACFB19" box="[724,813,1195,1221]" pageId="19" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is very similar to
<taxonomicName id="4C197897FFF9BF7AC3CAF286FA80FB19" authority="Fauvel, 1907" authorityName="Fauvel" authorityYear="1907" box="[1015,1281,1195,1221]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="350" phylum="Arthropoda" rank="species" species="spinifer">
<emphasis id="B96DDF06FFF9BF7AC3CAF286FBE8FB19" box="[1015,1129,1195,1221]" italics="true" pageId="19" pageNumber="350">S. spinifer</emphasis>
Fauvel, 1907
</taxonomicName>
(the
<typeStatus id="54A2BDB6FFF9BF7AC507F286FA1DFB19" box="[1338,1436,1195,1221]" pageId="19" pageNumber="350" type="holotype">holotype</typeStatus>
was examined,
<figureCitation id="13221F91FFF9BF7AC17BF2E2FE4FFB35" box="[326,462,1230,1257]" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="19" pageNumber="350">Figs. 5358</figureCitation>
), but differs by larger and more markedly unpunctured areas and more coarse and less dense punctation on pronotum and elytra, by more slender aedeagus with more apically enlarged parameres and by much more modified last sternites in male (male sternites 6 and
<quantity id="4CE1AEF1FFF9BF7AC3AEF33AFC3EFAED" box="[915,959,1303,1329]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="19" pageNumber="350" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="4C197897FFF9BF7AC3FAF33AFBBBFAED" authorityName="Fauvel" authorityYear="1907" box="[967,1082,1303,1329]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="350" phylum="Arthropoda" rank="species" species="spinifer">
<emphasis id="B96DDF06FFF9BF7AC3FAF33AFBBBFAED" box="[967,1082,1303,1329]" italics="true" pageId="19" pageNumber="350">S. spinifer</emphasis>
</taxonomicName>
are almost unmodified, shortly setose, posterior margin of sternite 7 only with almost indistinct very narrow and shallow emargination (
<figureCitation id="13221F91FFF9BF7AC52BF316FAE7FA89" box="[1302,1382,1338,1365]" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="19" pageNumber="350">Fig. 54</figureCitation>
) and emargination of male sternite 8 is larger and less deep: see
<figureCitation id="13221F91FFF9BF7AC311F372FCFEFAA5" box="[812,895,1374,1401]" captionStart="FIGURES 5366" captionStartId="20.[151,264,1850,1874]" captionTargetBox="[167,1419,191,1809]" captionTargetId="figure-20@20.[155,1433,181,1827]" captionTargetPageId="20" captionText="FIGURES 5366. 5358: Stenus spinifer Puthz, holotype. 5966: Stenus spiniferoides Janák, sp. nov., 5964, holotype; 6566, paratype. 53, 59, habitus; 54, 60, abdomen ventral; 56, male sternite 8, transmitted light; 62, male sternites 68, reflected light; 57, 63, male sternite 9; 58, 64, male tergite 10; 65, female tergite 10; 66, valves. Scale 1 mm: 5354, 5960; 0,5 mm: 55, 61; 0,5 mm: 5658; 6266." figureDoi="http://doi.org/10.5281/zenodo.6986332" httpUri="https://zenodo.org/record/6986332/files/figure.png" pageId="19" pageNumber="350">Fig. 56</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C303509FFFF9BF7AC0FAF3AFFC68FA41" box="[199,1001,1410,1437]" pageId="19" pageNumber="350" type="etymology">
<paragraph id="8BA60314FFF9BF7AC0FAF3AFFC68FA41" blockId="19.[151,1437,151,1581]" box="[199,1001,1410,1437]" pageId="19" pageNumber="350">
<emphasis id="B96DDF06FFF9BF7AC0FAF3AFFE25FA40" bold="true" box="[199,420,1410,1436]" pageId="19" pageNumber="350">Derivatio nominis.</emphasis>
The name refers to the similarity to
<taxonomicName id="4C197897FFF9BF7AC300F3AEFC62FA41" authorityName="Fauvel" authorityYear="1907" box="[829,995,1411,1437]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="350" phylum="Arthropoda" rank="species" species="spinifer">
<emphasis id="B96DDF06FFF9BF7AC300F3AEFC62FA41" box="[829,995,1411,1437]" italics="true" pageId="19" pageNumber="350">Stenus spinifer</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C303509FFFF9BF7AC0FAF38BFE8AF9F1" pageId="19" pageNumber="350" type="distribution">
<paragraph id="8BA60314FFF9BF7AC0FAF38BFE83FA39" blockId="19.[151,1437,151,1581]" pageId="19" pageNumber="350">
<emphasis id="B96DDF06FFF9BF7AC0FAF38BFEE0FA1C" bold="true" box="[199,353,1446,1472]" pageId="19" pageNumber="350">Distribution.</emphasis>
<taxonomicName id="4C197897FFF9BF7AC151F38AFDEBFA1D" authority="Janak" authorityName="Janak" authorityYear="2022" box="[364,618,1447,1473]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="350" phylum="Arthropoda" rank="species" species="spiniferoides" status="sp. nov.">
<emphasis id="B96DDF06FFF9BF7AC151F38AFD9DFA1D" box="[364,540,1447,1473]" italics="true" pageId="19" pageNumber="350">S. spiniferoides</emphasis>
Janák
</taxonomicName>
,
<emphasis id="B96DDF06FFF9BF7AC244F38AFD56FA1D" bold="true" box="[633,727,1447,1473]" pageId="19" pageNumber="350">
<taxonomicNameLabel id="A25E627DFFF9BF7AC244F38AFD56FA1D" box="[633,727,1447,1473]" pageId="19" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently known from Bo-Langvlei lake in
<collectingRegion id="49DDCDF6FFF9BF7AC4C1F38AFF7FFA39" country="South Africa" name="Western Cape" pageId="19" pageNumber="350">Western Cape Province</collectingRegion>
.
</paragraph>
<paragraph id="8BA60314FFF9BF7AC0FAF3C3FE8AF9F1" blockId="19.[151,1437,151,1581]" pageId="19" pageNumber="350">
<emphasis id="B96DDF06FFF9BF7AC0FAF3C3FECAF9D4" bold="true" box="[199,331,1518,1544]" pageId="19" pageNumber="350">Bionomics.</emphasis>
The
<typeStatus id="54A2BDB6FFF9BF7AC142F3C2FE2EF9D5" box="[383,431,1519,1545]" pageId="19" pageNumber="350">type</typeStatus>
specimens were found by treading of dense reeds on a lake banks together with
<taxonomicName id="4C197897FFF9BF7AC528F3C2FA1DF9D4" authorityName="L. Benick" authorityYear="1933" box="[1301,1436,1519,1545]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="350" phylum="Arthropoda" rank="species" species="separatus">
<emphasis id="B96DDF06FFF9BF7AC528F3C2FA1DF9D4" box="[1301,1436,1519,1545]" italics="true" pageId="19" pageNumber="350">S. separatus</emphasis>
</taxonomicName>
(
<figureCitation id="13221F91FFF9BF7AC0A2F03EFF7EF9F1" box="[159,255,1555,1581]" captionStart="FIGURES 6772" captionStartId="21.[151,264,1641,1665]" captionTargetBox="[154,1433,181,1617]" captionTargetId="figure-17@21.[154,1433,181,1617]" captionTargetPageId="21" captionText="FIGURES 6772. Localities where Stenus species were collected. S. hippoamicus Janák, sp. nov. (67, St. Lucia; 68, Ndumo); S. volkerputhzianus Janák, sp. nov. (69, Himeville, 70, Tugella river); S. hessei L. Benick, S. capicola L. Benick, S. depressiusculus L. Benick, S. cf. grandipennis L. Benick, 1933 (71, Olifantsbos); S. spiniferoides Janák, sp. nov. (72, Langvlei)." figureDoi="http://doi.org/10.5281/zenodo.6986334" httpUri="https://zenodo.org/record/6986334/files/figure.png" pageId="19" pageNumber="350">Figs. 72</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>