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<document id="BF2280427A9609BB0DE5D4862CC44347" ID-CLB-Dataset="297560" ID-DOI="10.5252/zoosystema2024v46a13" ID-GBIF-Dataset="9d97ce13-f0b1-4c8e-a281-034d4df4fe41" ID-ISSN="1638-9387" ID-Zenodo-Dep="11391415" ID-ZooBank="7547F492-D2B2-4881-B3D9-12F730B4DA65" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1716971933471" checkinUser="plazi" docAuthor="Langlois, Dominique &amp; Daugeron, Christophe" docDate="2024" docId="03A987F75414FFAEC7902CD8A86BF844" docLanguage="en" docName="zoosystema.2024.46.13.pdf" docOrigin="Zoosystema 46 (13)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a13.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Empis (Xanthempis) montsegurensis Langlois &amp; Daugeron 2024, n. sp." docType="treatment" docUuid="AEE7D705-F046-4877-B160-E215A6FA0835" docUuidSource="ZooBank" docVersion="2" lastPageNumber="323" masterDocId="FF90FF8F5411FFA8C773296BAD09FFA1" masterDocTitle="Two new species of the genus Empis Linnaeus, 1758 (Diptera, Empididae, Empidinae) from French nature reserves" masterLastPageNumber="325" masterPageNumber="319" pageNumber="322" updateTime="1717154445900" updateUser="julia" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="06AE87AAA4DB1974981A25F179C45A99">Two new species of the genus Empis Linnaeus, 1758 (Diptera, Empididae, Empidinae) from French nature reserves</mods:title>
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<mods:namePart id="D47FD7D7C8551C94E7F0B4DDC9E686C2">Langlois, Dominique</mods:namePart>
<mods:affiliation id="592084FBA8069828584DA73A22FC6359">Foucherans, Bonnevaux-le-Prieuré, F- 25620 Ornans (France) dl. entomo @ gmx. com</mods:affiliation>
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<mods:namePart id="881F12381CD7F59EE64C34833B870016">Daugeron, Christophe</mods:namePart>
<mods:affiliation id="2BA5B76A937ECB0879DB7460126E59B5">Muséum national dHistoire naturelle, Centre national de la Recherche scientifique, Mécanismes adaptatifs et évolution, UMR 7179 MNHN-CNRS, case postale 50, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) christophe. daugeron @ mnhn. fr (corresponding author)</mods:affiliation>
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<mods:date id="D7329E4A19A7F6E0B875CC5706DC103A">2024</mods:date>
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<taxonomicName id="4C004D625414FFADC7902CD8AF6EFA6C" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" box="[227,615,1458,1485]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="montsegurensis" status="sp. nov." subGenus="Xanthempis">
<emphasis id="B974EAF35414FFADC7902CD8AC2FFA6C" bold="true" box="[227,294,1459,1485]" italics="true" pageId="5" pageNumber="322">Empis</emphasis>
(
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)
<emphasis id="B974EAF35414FFADC6B52CD8AF6EFA6C" bold="true" box="[454,615,1459,1485]" italics="true" pageId="5" pageNumber="322">montsegurensis</emphasis>
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(
<figureCitation id="133B2A645414FFADC6D62CB8ACECFA4C" box="[421,485,1491,1517]" captionStart="FIG" captionStartId="6.[132,143,938,956]" captionTargetBox="[132,1405,219,892]" captionTargetId="figure-431@6.[132,888,219,894]" captionTargetPageId="6" captionText="FIG. 2. — Empis (Xanthempis) montsegurensis n. sp.: A, male habitus;B, male hypopygium.Abbreviations:see Material and methods.Scale bars:A,1 mm;B, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11391419" httpUri="https://zenodo.org/record/11391419/files/figure.png" pageId="5" pageNumber="322">Fig. 2</figureCitation>
)
</paragraph>
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urn:lsid:zoobank.org:act:
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<materialsCitation id="3B683CBC5414FFADC7F72F22AF52F968" ID-GBIF-Occurrence="4869303301" collectingDate="2019-06-26" collectionCode="TYPE, MATERIAL" collectorName="de Saint-Barthelemy &amp; L. Serviere" country="France" county="Caramille" elevation="1745" latitude="42.828888" location="France" longLatPrecision="19" longitude="1.7947222" municipality="Reserve" pageId="5" pageNumber="322" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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. —
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<typeStatus id="54BB88435414FFADC6222F23ACBBF9C1" box="[337,434,1608,1632]" pageId="5" pageNumber="322" type="holotype">Holotype</typeStatus>
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.
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<collectingCountry id="F31776715414FFADC6B12F23AF0FF9C1" box="[450,518,1608,1632]" name="France" pageId="5" pageNumber="322">France</collectingCountry>
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<specimenCount id="9D06FD685414FFADC5522F20AF3CF9FE" box="[545,565,1611,1631]" count="1" pageId="5" pageNumber="322" type="male"></specimenCount>
;
<location id="8EDF603A5414FFADC5342F22AF84F9C1" LSID="urn:lsid:plazi:treatment:03A987F75414FFAEC7902CD8A86BF844:8EDF603A5414FFADC5342F22AF84F9C1" box="[583,653,1609,1632]" country="France" county="Caramille" latitude="42.828888" longLatPrecision="19" longitude="1.7947222" municipality="Reserve" name="France" pageId="5" pageNumber="322">France</location>
,
<location id="8EDF603A5414FFADC5EA2F23AE0EF9C1" LSID="urn:lsid:plazi:treatment:03A987F75414FFAEC7902CD8A86BF844:8EDF603A5414FFADC5EA2F23AE0EF9C1" box="[665,775,1608,1632]" country="France" county="Caramille" latitude="42.828888" longLatPrecision="19" longitude="1.7947222" municipality="Reserve" name="Montsegur" pageId="5" pageNumber="322">Montségur</location>
(09211),
<collectingMunicipality id="6BDBAC9B5414FFADC7932F08AC20F9DA" box="[224,297,1635,1659]" pageId="5" pageNumber="322">Réserve</collectingMunicipality>
naturelle régionale du massif
<collectorName id="26F553375414FFADC5322F08AE0CF9DA" box="[577,773,1635,1659]" pageId="5" pageNumber="322">de Saint-Barthélémy</collectorName>
,
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;
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,
<geoCoordinate id="EE3450265414FFADC60C2F16ACEEF934" box="[383,487,1661,1685]" degrees="1" direction="east" minutes="47" orientation="longitude" pageId="5" pageNumber="322" precision="15" seconds="41" value="1.7947222">1°4741”E</geoCoordinate>
;
<elevation id="002DD1D25414FFADC6852F16AF74F934" box="[502,637,1661,1685]" metricMagnitude="3" metricUnit="m" metricValue="1.7449999999999999" pageId="5" pageNumber="322" unit="m" value="1745.0">
<quantity id="4CF89B045414FFADC6852F16AF4FF934" box="[502,582,1661,1685]" metricMagnitude="3" metricUnit="m" metricValue="1.7449999999999999" pageId="5" pageNumber="322" unit="m" value="1745.0">1745 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBE10215414FFADC5F82F16AE0BF934" box="[651,770,1661,1685]" pageId="5" pageNumber="322" value="2019-06-26">
<collectingDate id="EFFAE9C95414FFADC5F82F16AE0BF934" box="[651,770,1661,1685]" pageId="5" pageNumber="322" value="2019-06-26">26.VI.2019</collectingDate>
</date>
;
<collectorName id="26F553375414FFADC7F52FF3ADFBF90E" box="[134,242,1687,1711]" pageId="5" pageNumber="322">L. Servière</collectorName>
;
<collectingMethod id="52414EF65414FFADC78F2FFCAC7CF90E" box="[252,373,1687,1711]" pageId="5" pageNumber="322">Malaise trap</collectingMethod>
no. 4, wet meadow, near a pine and acidophilous beech forest; MNHN-ED-ED10936.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A656A5414FFADC7F72F8DAC5CF8B9" pageId="5" pageNumber="322" type="etymology">
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. — Named after Montségur, the locality where the species was collected.
</paragraph>
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<smallCapsWord id="8D59A03D5414FFADC7F72E5FADE6F8EB" baselines="1861,1862" box="[132,239,1844,1867]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="5" pageNumber="322">DIAGNOSIS</smallCapsWord>
. — A species of medium size belonging to the subgenus
<taxonomicName id="4C004D625414FFADC7F02E25ADFAF8C7" authorityName="Bezzi" authorityYear="1909" box="[131,243,1870,1894]" pageId="5" pageNumber="322" rank="subGenus" subGenus="Xanthempis">
<emphasis id="B974EAF35414FFADC7F02E25ADFAF8C7" box="[131,243,1870,1894]" italics="true" pageId="5" pageNumber="322">Xanthempis</emphasis>
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with the occiput dusted blackish dorsally to yellowish ventrally, a dusted greyish to blackish scutum except around postpronotal lobe and postalar callus. Hypandrium bare, phallus characteriscally sinuous at middle.
</paragraph>
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<subSubSection id="C31A656A5414FFADC7F72EBAAF3CF848" box="[132,565,2000,2025]" pageId="5" pageNumber="322" type="distribution">
<paragraph id="8BBF36E15414FFADC7F72EBAAF3CF848" blockId="5.[132,565,2000,2025]" box="[132,565,2000,2025]" pageId="5" pageNumber="322">
<smallCapsWord id="8D59A03D5414FFADC7F72EBAAC1BF846" baselines="2018,2019" box="[132,274,2001,2024]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="5" pageNumber="322">DISTRIBUTION</smallCapsWord>
. —
<collectingCountry id="F31776715414FFADC64D2EBAAC8BF849" box="[318,386,2001,2024]" name="France" pageId="5" pageNumber="322">France</collectingCountry>
: Pyrenees (
<figureCitation id="133B2A645414FFADC69E2EBBAF21F848" box="[493,552,2000,2025]" captionStart="FIG" captionStartId="7.[133,144,948,966]" captionTargetBox="[132,1456,94,878]" captionTargetId="figure-528@7.[130,1453,-323,1260]" captionTargetPageId="7" captionText="FIG. 3. — Map of France showing the distribution of Empis (Empis) cartusiana n. sp. (●) and Empis (Xanthempis) montsegurensis n. sp. (Ω). Background map: Sémhur, Wikimedia Commons." figureDoi="http://doi.org/10.5281/zenodo.11391423" httpUri="https://zenodo.org/record/11391423/files/figure.png" pageId="5" pageNumber="322">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A656A5414FFADC45F29B3AEA8F8AD" pageId="5" pageNumber="322" type="description">
<paragraph id="8BBF36E15414FFADC45F29B3AEB4FF51" blockId="5.[810,1457,216,560]" box="[812,957,216,242]" pageId="5" pageNumber="322">
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<emphasis id="B974EAF35414FFADC45F299CAE68FEB0" box="[812,865,247,273]" italics="true" pageId="5" pageNumber="322">Male</emphasis>
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<paragraph id="8BBF36E15414FFADC45F287CA95DFD91" blockId="5.[810,1457,216,560]" pageId="5" pageNumber="322">
<emphasis id="B974EAF35414FFADC45F287CAE6FFE90" bold="true" box="[812,870,279,305]" pageId="5" pageNumber="322">Head</emphasis>
. Ocellar triangle dusted blackish with pair of rather strong setae, occiput dusted blackish dorsally, yellowish ventrally, with strong, short setae. Frons brownish with setulae along the margin of eyes. Face, gena yellowish. Labrum yellowish, twice head height, labium brownish. Palpus yellowish with a few black hairs. Scape and pedicel brownish, postpedicel blackish; scape and postpedicel long, three and six times pedicel length respectively, style very short. Eyes dichoptic, all ommatidia of equal size.
</paragraph>
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<emphasis id="B974EAF35414FFADC45F2B3DAE8FFDD1" bold="true" box="[812,902,598,624]" pageId="5" pageNumber="322">Thorax.</emphasis>
Antepronotum yellowish to blackish at middle, bilobed with 5-6 short spine-like setae. Postpronotum yellow with one anterior and one apical distinct setae, the latter stronger and longer. Prosternum, proepisternum yellow with 1-2 minute black hairs. Scutum dusted greyish to blackish, but yellowish around postpronotal lobe, postalar callus, notopleural area. Acrostichals absent, dorsocentrals uniserial, distinct, becoming strong, long in presculettar depression. Strong, long following setae: one postpronotal, one presutural supraalar, one notopleural, one postalar. Scutellum dusted greyish, with one pair of strong, long apical setae. Lateral parts of thorax yellow, laterotergite with a few strong, long setae. Postscutellum blackish to yellowish laterally. Anterior and posterior spiracles yellow.
</paragraph>
<paragraph id="8BBF36E15414FFADC45F2D5FA9C9FB0F" blockId="5.[810,1455,1076,1198]" pageId="5" pageNumber="322">
<emphasis id="B974EAF35414FFADC45F2D5FAE7BFBEF" bold="true" box="[812,882,1076,1102]" pageId="5" pageNumber="322">Wing.</emphasis>
Length:
<quantity id="4CF89B045414FFADC4A12D5FA922FBEE" box="[978,1067,1076,1103]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="322" unit="mm" value="6.5">6.5 mm</quantity>
. Rather clear feebly tinted of brown, veins brown, well sclerotized except anal vein feebly sclerotized basally but complete; anal lobe obtuse. Haltere slightly brownish. Stigma brownish not well visible.
</paragraph>
<paragraph id="8BBF36E15414FFADC45F2DB8A903FA6C" blockId="5.[810,1456,1235,1485]" pageId="5" pageNumber="322">
<emphasis id="B974EAF35414FFADC45F2DB8AE95FB4C" bold="true" box="[812,924,1235,1261]" pageId="5" pageNumber="322">Abdomen.</emphasis>
Yellow with dorsal stripe formed by brown T-shaped spots on each tergite in dorsal view, another brownish stripe on each side along margin of tergites. Thin, short yellowish hairs or black thin, short setae at margin of tergite and sternite. Posterior margins of tergites 6 and 7 with a few strong, long and black dorsal setae, posterior margins of tergite and sternite 8 with many strong, long and black setae. Tergite and sternite 8 fused.
</paragraph>
<paragraph id="8BBF36E15414FFADC45F2C99AEFDF90D" blockId="5.[811,1457,1522,1709]" pageId="5" pageNumber="322">
<emphasis id="B974EAF35414FFADC45F2C99AECDF9AD" bold="true" box="[812,964,1522,1548]" pageId="5" pageNumber="322">Hypopygium</emphasis>
. Entirely yellow. Cercus higher than wide, lengthened ventrally, with dense rather thin, short and black dorsal setae, epandrium subrectangular, pointed apically, with strong and long dorsal setae subapically and apically, hypandrium reduced, bare, phallus long, curved, characteristically sinuous at middle.
</paragraph>
<paragraph id="8BBF36E15414FFADC45F2FB9AE7CF94D" blockId="5.[812,929,1746,1804]" box="[812,885,1746,1772]" pageId="5" pageNumber="322">
<emphasis id="B974EAF35414FFADC45F2FB9AE7CF94D" box="[812,885,1746,1772]" italics="true" pageId="5" pageNumber="322">Female</emphasis>
</paragraph>
<paragraph id="8BBF36E15414FFADC45F2F99AEA8F8AD" blockId="5.[812,929,1746,1804]" box="[812,929,1778,1804]" pageId="5" pageNumber="322">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31A656A5414FFADC45F2E59AE99F8EB" box="[812,912,1842,1868]" pageId="5" pageNumber="322" type="discussion">
<paragraph id="8BBF36E15414FFADC45F2E59AE99F8EB" blockId="5.[812,1457,1842,2028]" box="[812,912,1842,1868]" pageId="5" pageNumber="322">
<smallCapsWord id="8D59A03D5414FFADC45F2E59AE99F8EB" baselines="1861,1861" box="[812,912,1842,1868]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="5" pageNumber="322">REMARKS</smallCapsWord>
</paragraph>
</subSubSection>
<subSubSection id="C31A656A5414FFAEC45F2E39AE8DFBCF" lastPageId="6" lastPageNumber="323" pageId="5" pageNumber="322" type="reference_group">
<paragraph id="8BBF36E15414FFAEC45F2E39AE8DFBCF" blockId="5.[812,1457,1842,2028]" lastBlockId="6.[811,1455,1044,1134]" lastPageId="6" lastPageNumber="323" pageId="5" pageNumber="322">
<taxonomicName id="4C004D625414FFADC45F2E39A92FF8CA" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" box="[812,1062,1874,1900]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="montsegurensis" status="n. sp." subGenus="Xanthempis">
<emphasis id="B974EAF35414FFADC45F2E39AE4CF8CA" box="[812,837,1874,1899]" italics="true" pageId="5" pageNumber="322">E.</emphasis>
(
<emphasis id="B974EAF35414FFADC42B2E39AE78F8CA" box="[856,881,1874,1899]" italics="true" pageId="5" pageNumber="322">X.</emphasis>
)
<emphasis id="B974EAF35414FFADC4F02E39A92FF8CA" box="[899,1062,1874,1899]" italics="true" pageId="5" pageNumber="322">montsegurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757885414FFADC35D2E39A966F8CD" box="[1070,1135,1874,1900]" pageId="5" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
can be easily distinguished from all other species of
<taxonomicName id="4C004D625414FFADC3242E1AA9D2F82A" ID-CoL="8YQX2" authorityName="Bezzi" authorityYear="1909" box="[1111,1243,1905,1931]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="324" phylum="Arthropoda" rank="subGenus" subGenus="Xanthempis">
<emphasis id="B974EAF35414FFADC3242E1AA9D2F82A" box="[1111,1243,1905,1931]" italics="true" pageId="5" pageNumber="322">Xanthempis</emphasis>
</taxonomicName>
. For the moment it is known from a single specimen collected in the Pyrenees, and in addition to
<taxonomicName id="4C004D625414FFADC3232ED9AE62F84A" ID-CoL="8BZRG" authority="Daugeron, 2000" authorityName="Daugeron" authorityYear="2000" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="hypandrialis" subGenus="Xanthempis">
<emphasis id="B974EAF35414FFADC3232ED9A960F86A" box="[1104,1129,1970,1995]" italics="true" pageId="5" pageNumber="322">E.</emphasis>
(
<emphasis id="B974EAF35414FFADC30A2ED9A99BF86A" box="[1145,1170,1970,1995]" italics="true" pageId="5" pageNumber="322">X.</emphasis>
)
<emphasis id="B974EAF35414FFADC3D12EDAA825F86A" box="[1186,1324,1969,1995]" italics="true" pageId="5" pageNumber="322">hypandrialis</emphasis>
<bibRefCitation id="EF914B105414FFADC2402ED9AE62F84A" author="DAUGERON C." pageId="5" pageNumber="322" pagination="371 - 388" refId="ref3963" refString="DAUGERON C. 2000. - The subgenus Xanthempis: new species and taxonomical data (Diptera: Empididae). Annales de la Societe Entomologique de France (N. S) 36: 371 - 388." type="journal article" year="2000">Daugeron, 2000</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D625414FFADC40B2EB9A803F84A" ID-CoL="8BZRY" authority="Daugeron, 2000" authorityName="Daugeron" authorityYear="2000" box="[888,1290,2001,2028]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="montivaga" subGenus="Xanthempis">
<emphasis id="B974EAF35414FFADC40B2EB9AE98F84A" box="[888,913,2002,2027]" italics="true" pageId="5" pageNumber="322">E.</emphasis>
(
<emphasis id="B974EAF35414FFADC4D22EB9AEB3F84A" box="[929,954,2002,2027]" italics="true" pageId="5" pageNumber="322">X.</emphasis>
)
<emphasis id="B974EAF35414FFADC4B82EB9A948F84A" box="[971,1089,2002,2027]" italics="true" pageId="5" pageNumber="322">montivaga</emphasis>
<bibRefCitation id="EF914B105414FFADC3342EB9A803F84A" author="DAUGERON C." box="[1095,1290,2001,2028]" pageId="5" pageNumber="322" pagination="371 - 388" refId="ref3963" refString="DAUGERON C. 2000. - The subgenus Xanthempis: new species and taxonomical data (Diptera: Empididae). Annales de la Societe Entomologique de France (N. S) 36: 371 - 388." type="journal article" year="2000">Daugeron, 2000</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D625414FFAEC2652EB9AC4FFB8F" ID-CoL="9HKL8" authority="Daugeron, 2009" authorityName="Daugeron" authorityYear="2009" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" lastPageId="6" lastPageNumber="323" order="Diptera" pageId="5" pageNumber="323" phylum="Arthropoda" rank="species" species="fagina" subGenus="Xanthempis">
<emphasis id="B974EAF35414FFADC2652EB9A826F84A" box="[1302,1327,2002,2027]" italics="true" pageId="5" pageNumber="322">E.</emphasis>
(
<emphasis id="B974EAF35414FFADC2332EB9A858F84A" box="[1344,1361,2002,2027]" italics="true" pageId="5" pageNumber="322">X</emphasis>
.)
<emphasis id="B974EAF35414FFADC21A2EBAA8A6F84A" box="[1385,1455,2001,2027]" italics="true" pageId="5" pageNumber="322">fagina</emphasis>
<bibRefCitation id="EF914B105417FFAEC7F72D7EAC4FFB8F" author="DAUGERON C." box="[132,326,1044,1071]" pageId="6" pageNumber="323" pagination="59 - 64" refId="ref4002" refString="DAUGERON C. 2009. - Two new species of Xanthempis Bezzi (Diptera, Empididae, Empidinae) endemic to the Pyrenees. Zootaxa 2087 (1): 59 - 64. https: // doi. org / 10.11646 / zootaxa. 2087.1.4" type="journal article" year="2009">Daugeron, 2009</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C004D625417FFAEC6F02D7EAE0BFB8F" ID-CoL="9HKLD" authority="Daugeron, 2009" authorityName="Daugeron" authorityYear="2009" box="[387,770,1044,1071]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="323" phylum="Arthropoda" rank="species" species="virgulata" subGenus="Xanthempis">
<emphasis id="B974EAF35417FFAEC6F02D7EAC95FB8F" box="[387,412,1045,1070]" italics="true" pageId="6" pageNumber="323">E.</emphasis>
(
<emphasis id="B974EAF35417FFAEC6DF2D7EACCCFB8F" box="[428,453,1045,1070]" italics="true" pageId="6" pageNumber="323">X.</emphasis>
)
<emphasis id="B974EAF35417FFAEC6A62D7FAF33FB8F" box="[469,570,1044,1070]" italics="true" pageId="6" pageNumber="323">virgulata</emphasis>
<bibRefCitation id="EF914B105417FFAEC5332D7EAE0BFB8F" author="DAUGERON C." box="[576,770,1044,1071]" pageId="6" pageNumber="323" pagination="59 - 64" refId="ref4002" refString="DAUGERON C. 2009. - Two new species of Xanthempis Bezzi (Diptera, Empididae, Empidinae) endemic to the Pyrenees. Zootaxa 2087 (1): 59 - 64. https: // doi. org / 10.11646 / zootaxa. 2087.1.4" type="journal article" year="2009">Daugeron, 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C004D625417FFAEC7F72D5EAC76FBEF" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" box="[132,383,1076,1102]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="323" phylum="Arthropoda" rank="species" species="montsegurensis" status="n. sp." subGenus="Xanthempis">
<emphasis id="B974EAF35417FFAEC7F72D5EAD94FBEF" box="[132,157,1077,1102]" italics="true" pageId="6" pageNumber="323">E.</emphasis>
(
<emphasis id="B974EAF35417FFAEC7C32D5EADC3FBEF" box="[176,202,1077,1102]" italics="true" pageId="6" pageNumber="323">X.</emphasis>
)
<emphasis id="B974EAF35417FFAEC7AF2D5EAC76FBEF" box="[220,383,1077,1102]" italics="true" pageId="6" pageNumber="323">montsegurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757885417FFAEC6FB2D5EACC0FBEE" box="[392,457,1077,1103]" pageId="6" pageNumber="323" rank="species">n. sp.</taxonomicNameLabel>
is the fifth species of
<taxonomicName id="4C004D625417FFAEC5BC2D5EADD1FBCF" ID-CoL="8YQX2" authorityName="Bezzi" authorityYear="1909" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="324" phylum="Arthropoda" rank="subGenus" subGenus="Xanthempis">
<emphasis id="B974EAF35417FFAEC5BC2D5EADD1FBCF" italics="true" pageId="6" pageNumber="323">Xanthempis</emphasis>
</taxonomicName>
only known from the Pyrenees. The key proposed previously by
<bibRefCitation id="EF914B105417FFAEC4B82D7EA99EFB8F" author="DAUGERON C." box="[971,1175,1044,1071]" pageId="6" pageNumber="323" pagination="59 - 64" refId="ref4002" refString="DAUGERON C. 2009. - Two new species of Xanthempis Bezzi (Diptera, Empididae, Empidinae) endemic to the Pyrenees. Zootaxa 2087 (1): 59 - 64. https: // doi. org / 10.11646 / zootaxa. 2087.1.4" type="journal article" year="2009">Daugeron (2009)</bibRefCitation>
for species known from the mountain area is here modified to include the new species.
</paragraph>
</subSubSection>
<caption id="DF7F66695417FFAEC7F72AC1A8B8FC1D" ID-DOI="http://doi.org/10.5281/zenodo.11391419" ID-Zenodo-Dep="11391419" box="[132,1457,938,956]" httpUri="https://zenodo.org/record/11391419/files/figure.png" pageId="6" pageNumber="323" startId="6.[132,143,938,956]" targetBox="[132,1405,219,892]" targetPageId="6" targetType="figure">
<paragraph id="8BBF36E15417FFAEC7F72AC1A8B8FC1D" blockId="6.[132,1457,938,956]" box="[132,1457,938,956]" pageId="6" pageNumber="323">
<smallCapsWord id="8D59A03D5417FFAEC7F72AC1AD94FC1A" baselines="951,952" box="[132,157,938,956]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="323">FIG</smallCapsWord>
. 2. —
<taxonomicName id="4C004D625417FFAEC7A22AC1ACFCFC1D" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" box="[209,501,938,956]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="323" phylum="Arthropoda" rank="species" species="montsegurensis" status="sp. nov." subGenus="Xanthempis">
<emphasis id="B974EAF35417FFAEC7A22AC1AC0DFC1D" box="[209,260,938,956]" italics="true" pageId="6" pageNumber="323">Empis</emphasis>
(
<emphasis id="B974EAF35417FFAEC67F2AC1AC65FC1D" box="[268,364,938,956]" italics="true" pageId="6" pageNumber="323">Xanthempis</emphasis>
)
<emphasis id="B974EAF35417FFAEC6072AC1ACFCFC1D" box="[372,501,938,956]" italics="true" pageId="6" pageNumber="323">montsegurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757885417FFAEC68B2AC1AF21FC1D" box="[504,552,938,956]" pageId="6" pageNumber="323" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B974EAF35417FFAEC55E2AC1AF33FC1D" bold="true" box="[557,570,938,956]" pageId="6" pageNumber="323">A</emphasis>
, male habitus;
<emphasis id="B974EAF35417FFAEC5C22AC1AFB7FC1D" bold="true" box="[689,702,938,956]" pageId="6" pageNumber="323">B</emphasis>
, male hypopygium.Abbreviations:see Material and methods.Scale bars:A,1 mm;B, 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C31A656A5417FFAEC5372DA1A86BF844" pageId="6" pageNumber="323" type="key">
<paragraph id="8BBF36E15417FFAEC5372DA1AEBFFAA0" blockId="6.[208,1377,1225,1363]" pageId="6" pageNumber="323">
<smallCapsWord id="8D59A03D5417FFAEC5372DA1AF66FB43" baselines="1245,1245" box="[580,623,1226,1252]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Key" pageId="6" pageNumber="323">KEY</smallCapsWord>
<smallCapsWord id="8D59A03D5417FFAEC5052DA4AF9CFB43" baselines="1245" box="[630,661,1231,1250]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="to" pageId="6" pageNumber="323">TO</smallCapsWord>
<smallCapsWord id="8D59A03D5417FFAEC5E82DA4AFE0FB43" baselines="1245" box="[667,745,1231,1250]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="species" pageId="6" pageNumber="323">SPECIES</smallCapsWord>
<smallCapsWord id="8D59A03D5417FFAEC5822DA4AE04FB43" baselines="1245" box="[753,781,1231,1250]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="6" pageNumber="323">OF</smallCapsWord>
<taxonomicName id="4C004D625417FFAEC4672DA1AEF9FB42" ID-CoL="8YQX2" authorityName="Bezzi" authorityYear="1909" box="[788,1008,1225,1251]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="324" phylum="Arthropoda" rank="subGenus" subGenus="Xanthempis">
<smallCapsWord id="8D59A03D5417FFAEC4672DA1AE5AFB43" baselines="1245,1245" box="[788,851,1226,1251]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Empis" pageId="6" pageNumber="323">
<emphasis id="B974EAF35417FFAEC4672DA1AE5AFB43" box="[788,851,1226,1251]" italics="true" pageId="6" pageNumber="323">EMPIS</emphasis>
</smallCapsWord>
(
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<emphasis id="B974EAF35417FFAEC4112DA1AEEEFB43" box="[866,999,1226,1251]" italics="true" pageId="6" pageNumber="323">XANTHEMPIS</emphasis>
</smallCapsWord>
)
</taxonomicName>
<smallCapsWord id="8D59A03D5417FFAEC50E2D85AFD8FAA0" baselines="1276" box="[637,721,1262,1281]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="known" pageId="6" pageNumber="323">KNOWN</smallCapsWord>
<smallCapsWord id="8D59A03D5417FFAEC5AB2D85AE1CFAA0" baselines="1276" box="[728,789,1262,1281]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="from" pageId="6" pageNumber="323">FROM</smallCapsWord>
<smallCapsWord id="8D59A03D5417FFAEC46F2D84AE4EFAA3" baselines="1277" box="[796,839,1263,1282]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="the" pageId="6" pageNumber="323">THE</smallCapsWord>
<smallCapsWord id="8D59A03D5417FFAEC43D2D82AEBFFAA0" baselines="1276,1276" box="[846,950,1257,1283]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pyrenees" pageId="6" pageNumber="323">PYRENEES</smallCapsWord>
</paragraph>
<key id="FF9161D05417FFAEC7A32C75A86BF844" pageId="6" pageNumber="323">
<keyStep id="30F42E445417FFAEC7A32C75A868FAF2" pageId="6" pageNumber="323">
<paragraph id="8BBF36E15417FFAEC7A32C75A868FA97" blockId="6.[208,1377,1225,1363]" box="[208,1377,1310,1334]" pageId="6" pageNumber="323">
<keyLead id="30F195D45417FFAEC7A32C75A868FA97" box="[208,1377,1310,1334]" pageId="6" pageNumber="323">1. Scutum entirely yellow ................................................................................................................................ 2</keyLead>
</paragraph>
<paragraph id="8BBF36E15417FFAEC7A32C50A868FAF2" blockId="6.[208,1377,1225,1363]" box="[208,1377,1339,1363]" pageId="6" pageNumber="323">
<keyLead id="30F195D45417FFAEC7A32C50A868FAF2" box="[208,1377,1339,1363]" pageId="6" pageNumber="323">— Scutum with a blackish pattern ................................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F42E445417FFAEC7A32C05A868FA58" pageId="6" pageNumber="323">
<paragraph id="8BBF36E15417FFAEC7A32C05A868FA02" blockId="6.[208,1380,1390,1529]" pageId="6" pageNumber="323">
<keyLead id="30F195D45417FFAEC7A32C05A868FA02" pageId="6" pageNumber="323">
2. Scape and pedicel brownish, occiput with a blackish-brown spot, hypandrium with pair of short apical spines, phallus pointed at tip (
<bibRefCitation id="EF914B105417FFAEC69A2CE0AF85FA02" author="DAUGERON C." box="[489,652,1419,1443]" pageId="6" pageNumber="323" pagination="371 - 388" refId="ref3963" refString="DAUGERON C. 2000. - The subgenus Xanthempis: new species and taxonomical data (Diptera: Empididae). Annales de la Societe Entomologique de France (N. S) 36: 371 - 388." type="journal article" year="2000">Daugeron 2000</bibRefCitation>
: fig. 6) .........................................
<taxonomicName id="4C004D625417FFAEC48C2CE7A868FA02" authority="Daugeron, 2000" authorityName="Daugeron" authorityYear="2000" box="[1023,1377,1419,1443]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="323" phylum="Arthropoda" rank="species" species="montivaga" subGenus="Xanthempis">
<emphasis id="B974EAF35417FFAEC48C2CE7A91FFA02" box="[1023,1046,1420,1443]" italics="true" pageId="6" pageNumber="323">E.</emphasis>
(
<emphasis id="B974EAF35417FFAEC3572CE7A932FA02" box="[1060,1083,1420,1443]" italics="true" pageId="6" pageNumber="323">X.</emphasis>
)
<emphasis id="B974EAF35417FFAEC3392CE0A9BBFA03" box="[1098,1202,1419,1442]" italics="true" pageId="6" pageNumber="323">montivaga</emphasis>
<bibRefCitation id="EF914B105417FFAEC3CB2CE0A868FA02" author="DAUGERON C." box="[1208,1377,1419,1443]" pageId="6" pageNumber="323" pagination="371 - 388" refId="ref3963" refString="DAUGERON C. 2000. - The subgenus Xanthempis: new species and taxonomical data (Diptera: Empididae). Annales de la Societe Entomologique de France (N. S) 36: 371 - 388." type="journal article" year="2000">Daugeron, 2000</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBF36E15417FFAEC7A32CC3A868FA58" blockId="6.[208,1380,1390,1529]" pageId="6" pageNumber="323">
<keyLead id="30F195D45417FFAEC7A32CC3A868FA58" pageId="6" pageNumber="323">
— Scape and pedicel yellowish, occiput with pair of subrectangular blackish spots (
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: fig. 3A), hypandrium with two pairs of short apical spines, phallus truncate at tip (
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: fig. 1) ..................... ........................................................................................................................
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<emphasis id="B974EAF35417FFAEC35A2C89A949FA58" box="[1065,1088,1506,1529]" italics="true" pageId="6" pageNumber="323">E.</emphasis>
(
<emphasis id="B974EAF35417FFAEC33D2C89A96CFA58" box="[1102,1125,1506,1529]" italics="true" pageId="6" pageNumber="323">X.</emphasis>
)
<emphasis id="B974EAF35417FFAEC3072C8AA9BBFA58" box="[1140,1202,1505,1529]" italics="true" pageId="6" pageNumber="323">fagina</emphasis>
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<keyLead id="30F195D45417FFAEC7A32F7FA868F9E8" pageId="6" pageNumber="323">3. Scutum with at least one more or less broad central black stripe between dorsocentrals, epandrial lamella sub-triangular .................................................................................................................................................... 4</keyLead>
</paragraph>
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<keyLead id="30F195D45417FFAEC7A32F25A868F9C7" box="[208,1377,1614,1638]" pageId="6" pageNumber="323">
— Scutum entirely blackish, epandrium subrectangular (
<bibRefCitation id="EF914B105417FFAEC4562F25AEA0F9C7" author="CHVALA M." box="[805,937,1614,1638]" pageId="6" pageNumber="323" refId="ref3771" refString="CHVALA M. 1994. - The Empidoidea (Diptera) of Fennoscandia and Denmark. III - Genus Empis. Fauna entomologica scandinavica 29. E. J. Brill, Leiden, New York, Koln, 192 p." type="book" year="1994">Chvála 1994</bibRefCitation>
: fig. 301) .............................................. 6
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<keyLead id="30F195D45417FFAEC7A32FEAA868F938" box="[208,1377,1665,1689]" pageId="6" pageNumber="323">4. Scutum with a broad central black stripe ..................................................................................................... 5</keyLead>
</paragraph>
<paragraph id="8BBF36E15417FFAEC7A32FF5A868F917" blockId="6.[208,1377,1665,1718]" box="[208,1377,1694,1718]" pageId="6" pageNumber="323">
<keyLead id="30F195D45417FFAEC7A32FF5A868F917" box="[208,1377,1694,1718]" pageId="6" pageNumber="323">
— Scutum with a single narrow central black stripe (
<bibRefCitation id="EF914B105417FFAEC58F2FF5AE76F917" author="CHVALA M." box="[764,895,1694,1718]" pageId="6" pageNumber="323" refId="ref3771" refString="CHVALA M. 1994. - The Empidoidea (Diptera) of Fennoscandia and Denmark. III - Genus Empis. Fauna entomologica scandinavica 29. E. J. Brill, Leiden, New York, Koln, 192 p." type="book" year="1994">Chvála 1994</bibRefCitation>
: fig. 284) ...........
<taxonomicName id="4C004D625417FFAEC3332FF4A868F917" authority="Linne, 1761" authorityName="Linnaeus" authorityYear="1761" box="[1088,1377,1694,1718]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="323" phylum="Arthropoda" rank="species" species="stercorea" subGenus="Xanthempis">
<emphasis id="B974EAF35417FFAEC3332FF4A95EF917" box="[1088,1111,1695,1718]" italics="true" pageId="6" pageNumber="323">E.</emphasis>
(
<emphasis id="B974EAF35417FFAEC3162FF4A975F917" box="[1125,1148,1695,1718]" italics="true" pageId="6" pageNumber="323">X.</emphasis>
)
<emphasis id="B974EAF35417FFAEC3F92FF4A9D5F917" box="[1162,1244,1695,1718]" italics="true" pageId="6" pageNumber="323">stercorea</emphasis>
Linné, 1761
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5. Scutum yellow between dorsocentrals and notopleuron, hypandrium lengthened with many apical and subapical spines (
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: figs 2; 3) ......................................................
<taxonomicName id="4C004D625417FFAEC49D2F84A868F8A7" authority="Daugeron, 2000" authorityName="Daugeron" authorityYear="2000" box="[1006,1377,1774,1799]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="323" phylum="Arthropoda" rank="species" species="hypandrialis" subGenus="Xanthempis">
<emphasis id="B974EAF35417FFAEC49D2F84A90DF8A7" box="[1006,1028,1775,1798]" italics="true" pageId="6" pageNumber="323">E.</emphasis>
(
<emphasis id="B974EAF35417FFAEC3612F84A920F8A7" box="[1042,1065,1775,1798]" italics="true" pageId="6" pageNumber="323">X.</emphasis>
)
<emphasis id="B974EAF35417FFAEC34B2F85A9BBF8A7" box="[1080,1202,1774,1798]" italics="true" pageId="6" pageNumber="323">hypandrialis</emphasis>
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<keyLead id="30F195D45417FFAEC7A32E60A868F8FD" pageId="6" pageNumber="323">
— Scutum with pair of brownish to black spots, sometimes entirely fused in a single stripe, between dorsocentrals and notopleuron (
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: fig. 3B), hypandrium not lengthened, bearing a single pair of subapical spines (
<bibRefCitation id="EF914B105417FFAEC63F2E2EACE6F8FD" author="DAUGERON C." box="[332,495,1860,1885]" pageId="6" pageNumber="323" pagination="59 - 64" refId="ref4002" refString="DAUGERON C. 2009. - Two new species of Xanthempis Bezzi (Diptera, Empididae, Empidinae) endemic to the Pyrenees. Zootaxa 2087 (1): 59 - 64. https: // doi. org / 10.11646 / zootaxa. 2087.1.4" type="journal article" year="2009">Daugeron 2009</bibRefCitation>
: fig. 2) ...................................................................
<taxonomicName id="4C004D625417FFAEC37D2E2EA868F8FD" authority="Daugeron, 2009" authorityName="Daugeron" authorityYear="2009" box="[1038,1377,1860,1885]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="323" phylum="Arthropoda" rank="species" species="virgulata" subGenus="Xanthempis">
<emphasis id="B974EAF35417FFAEC37D2E2EA92DF8FD" box="[1038,1060,1861,1884]" italics="true" pageId="6" pageNumber="323">E.</emphasis>
(
<emphasis id="B974EAF35417FFAEC3402E2EA943F8FD" box="[1075,1098,1861,1884]" italics="true" pageId="6" pageNumber="323">X.</emphasis>
)
<emphasis id="B974EAF35417FFAEC32B2E2FA9B8F8FD" box="[1112,1201,1860,1884]" italics="true" pageId="6" pageNumber="323">virgulata</emphasis>
<bibRefCitation id="EF914B105417FFAEC3CB2E2EA868F8FD" author="DAUGERON C." box="[1208,1377,1860,1885]" pageId="6" pageNumber="323" pagination="59 - 64" refId="ref4002" refString="DAUGERON C. 2009. - Two new species of Xanthempis Bezzi (Diptera, Empididae, Empidinae) endemic to the Pyrenees. Zootaxa 2087 (1): 59 - 64. https: // doi. org / 10.11646 / zootaxa. 2087.1.4" type="journal article" year="2009">Daugeron, 2009</bibRefCitation>
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<keyLead id="30F195D45417FFAEC7A32E13A868F80D" pageId="6" pageNumber="323">
6. Large species (wing length&gt;
<quantity id="4CF89B045417FFAEC5512E13AF6BF831" box="[546,610,1912,1936]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="323" unit="mm" value="7.0">7 mm</quantity>
), abdomen usually entirely yellow, without stripe, two notopleurals, phallus regularly curved ..............................................................................................
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<emphasis id="B974EAF35417FFAEC35B2EFEA936F80D" box="[1064,1087,1941,1964]" italics="true" pageId="6" pageNumber="323">E.</emphasis>
(
<emphasis id="B974EAF35417FFAEC33E2EFEA96DF80D" box="[1101,1124,1941,1964]" italics="true" pageId="6" pageNumber="323">X.</emphasis>
)
<emphasis id="B974EAF35417FFAEC3012EFEA9B2F80D" box="[1138,1211,1941,1964]" italics="true" pageId="6" pageNumber="323">testacea</emphasis>
Fabricius, 1805
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<keyLead id="30F195D45417FFAEC7A32ED9A86BF844" pageId="6" pageNumber="323">
— Smaller species (wing =
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), abdomen yellow with a well visible dorsal stripe, one notopleural, phallus somewhat sinuous at middle ...............................................................................
<taxonomicName id="4C004D625417FFAEC3392EA4A82AF844" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" box="[1098,1315,1998,2022]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="323" phylum="Arthropoda" rank="species" species="montsegurensis" status="n. sp." subGenus="Xanthempis">
<emphasis id="B974EAF35417FFAEC3392EA4A969F847" box="[1098,1120,1999,2022]" italics="true" pageId="6" pageNumber="323">E.</emphasis>
(
<emphasis id="B974EAF35417FFAEC31C2EA4A98FF847" box="[1135,1158,1999,2022]" italics="true" pageId="6" pageNumber="323">X.</emphasis>
)
<emphasis id="B974EAF35417FFAEC3E72EA5A82AF844" box="[1172,1315,1998,2021]" italics="true" pageId="6" pageNumber="323">montsegurensis</emphasis>
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<taxonomicNameLabel id="A24757885417FFAEC2592EA5A86BF844" box="[1322,1378,1998,2021]" pageId="6" pageNumber="323" rank="species">n. sp.</taxonomicNameLabel>
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