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<mods:title id="AE6B6D2B5D91F1D3D1666C20A59795F1">Taxonomic information on some Japanese Chironomidae (Diptera) described by Dr. M. Sasa (†)</mods:title>
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<mods:namePart id="61F8D56A2152D4D6CE4D25FD5E92BDAB">Yamamoto, Nao</mods:namePart>
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<treatment id="03A987B9D529B218FF61FBFCBABA2A95" ID-DOI="http://doi.org/10.5281/zenodo.5973653" ID-GBIF-Taxon="154498462" ID-Zenodo-Dep="5973653" LSID="urn:lsid:plazi:treatment:03A987B9D529B218FF61FBFCBABA2A95" httpUri="http://treatment.plazi.org/id/03A987B9D529B218FF61FBFCBABA2A95" lastPageNumber="517" pageId="1" pageNumber="517">
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<paragraph id="8BBF36AFD529B218FF61FBFCB8A32B59" blockId="1.[151,901,1033,1059]" box="[151,901,1033,1059]" pageId="1" pageNumber="517">
<heading id="D0F781C3D529B218FF61FBFCB8A32B59" bold="true" box="[151,901,1033,1059]" fontSize="11" level="1" pageId="1" pageNumber="517" reason="1">
<emphasis id="B974EABDD529B218FF61FBFCB8A32B59" bold="true" box="[151,901,1033,1059]" pageId="1" pageNumber="517">
<taxonomicName id="4C004D2CD529B218FF61FBFCB8352B59" authority="(Sasa et Okazawa, 1992)" authorityName="Yamamoto &amp; Yamamoto" authorityYear="2018" baseAuthorityName="Sasa et Okazawa" baseAuthorityYear="1992" box="[151,787,1033,1059]" class="Insecta" family="Chironomidae" genus="Chaetocladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="species" species="togatenellus" status="comb. nov.">
<emphasis id="B974EABDD529B218FF61FBFCBAF82B59" bold="true" box="[151,478,1033,1059]" italics="true" pageId="1" pageNumber="517">Chaetocladius togatenellus</emphasis>
(
<bibRefCitation id="EF914B5ED529B218FE1AFBFCB82D2B59" author="Sasa, M. &amp; Okazawa, T." box="[492,779,1033,1059]" pageId="1" pageNumber="517" pagination="92 - 204" refId="ref6048" refString="Sasa, M. &amp; Okazawa, T. (1992) Studies on the chironomid midges (Yusurika) of Toga-Mura, Toyama. Part 2. The subfamily Orthocladiinae. Research Report of Toyama Prefectural Environmental Science Research Center, 1992, 92 - 204." type="journal article" year="1992">
Sasa
<emphasis id="B974EABDD529B218FDD8FBFCB9652B59" bold="true" box="[558,579,1033,1059]" italics="true" pageId="1" pageNumber="517">et</emphasis>
Okazawa, 1992
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A24757C6D529B218FCECFBFCB8A32B59" box="[794,901,1033,1059]" pageId="1" pageNumber="517" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31A6524D529B218FF61FBA5BF382BB5" pageId="1" pageNumber="517" type="reference_group">
<paragraph id="8BBF36AFD529B218FF61FBA5B82F2B1C" blockId="1.[151,777,1104,1126]" box="[151,777,1104,1126]" pageId="1" pageNumber="517">
<treatmentCitationGroup id="AB101181D529B218FF61FBA5B82F2B1C" box="[151,777,1104,1126]" pageId="1" pageNumber="517">
<taxonomicName id="4C004D2CD529B218FF61FBA5B8232B1C" ID-CoL="9HYC8" authority="Sasa et Okazawa, 1992: 135" authorityName="Sasa et Okazawa" authorityPageNumber="135" authorityYear="1992" box="[151,773,1104,1126]" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="species" species="togatenellus">
<emphasis id="B974EABDD529B218FF61FBA5BAF82B1C" box="[151,478,1104,1126]" italics="true" pageId="1" pageNumber="517">Bryophaenocladius togatenellus</emphasis>
<treatmentCitation id="0AA110BED529B218FE12FBA5B8232B1C" author="Sasa, M. &amp; Okazawa, T." box="[484,773,1104,1126]" page="135" pageId="1" pageNumber="517" year="1992">
<bibRefCitation id="EF914B5ED529B218FE12FBA5B9F42B1C" author="Sasa, M. &amp; Okazawa, T." box="[484,722,1104,1126]" pageId="1" pageNumber="517" pagination="92 - 204" refId="ref6048" refString="Sasa, M. &amp; Okazawa, T. (1992) Studies on the chironomid midges (Yusurika) of Toga-Mura, Toyama. Part 2. The subfamily Orthocladiinae. Research Report of Toyama Prefectural Environmental Science Research Center, 1992, 92 - 204." type="journal article" year="1992">
Sasa
<emphasis id="B974EABDD529B218FDEFFBA4B90D2B1C" box="[537,555,1105,1126]" italics="true" pageId="1" pageNumber="517">et</emphasis>
Okazawa, 1992
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: 135
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.
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</paragraph>
<paragraph id="8BBF36AFD529B218FF61FB67BF382BB5" blockId="1.[151,1437,1170,1519]" pageId="1" pageNumber="517">
<emphasis id="B974EABDD529B218FF61FB67BAA82BD1" bold="true" box="[151,398,1170,1195]" pageId="1" pageNumber="517">Specimen examined:</emphasis>
<materialsCitation id="3B683CF2D529B218FE63FB66BF3C2BB5" ID-GBIF-Occurrence="2028438470" collectingDate="1990-05-31" collectorName="T. Okazawa &amp; M. Sasa" country="Japan" location="Momose" municipality="Toga-Mura" pageId="1" pageNumber="517" specimenCount="1" specimenCount-male="1" stateProvince="Toyama" typeStatus="holotype">
<collectingCountry id="F317763FD529B218FE63FB66BAFF2BD0" box="[405,473,1171,1194]" name="Japan" pageId="1" pageNumber="517">Japan</collectingCountry>
:
<collectingRegion id="49C4F84DD529B218FE12FB66B9182BD1" box="[484,574,1171,1195]" country="Japan" name="Toyama" pageId="1" pageNumber="517">Toyama</collectingRegion>
Pref.,
<collectingMunicipality id="6BDBACD5D529B218FD7CFB66B82C2BD1" box="[650,778,1171,1195]" pageId="1" pageNumber="517">Toga-Mura</collectingMunicipality>
,
<location id="8EDF6074D529B218FCE0FB66B85D2BD0" LSID="urn:lsid:plazi:treatment:03A987B9D529B218FF61FBFCBABA2A95:8EDF6074D529B218FCE0FB66B85D2BD0" box="[790,891,1171,1194]" country="Japan" municipality="Toga-Mura" name="Momose" pageId="1" pageNumber="517" stateProvince="Toyama">Momose</location>
,
<typeStatus id="54BB880DD529B218FC70FB67B8CE2BD1" box="[902,1000,1170,1195]" pageId="1" pageNumber="517" type="holotype">holotype</typeStatus>
<specimenCount id="9D06FD26D529B218FC06FB66BF232BD0" box="[1008,1029,1171,1194]" pageId="1" pageNumber="517" type="male"></specimenCount>
of
<taxonomicName id="4C004D2CD529B218FBDDFB66BEB32BD1" ID-CoL="9HYC8" authorityName="Sasa et Okazawa" authorityYear="1992" box="[1067,1429,1171,1195]" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="species" species="togatenellus">
<emphasis id="B974EABDD529B218FBDDFB66BEB32BD1" box="[1067,1429,1171,1195]" italics="true" pageId="1" pageNumber="517">Bryophaenocladius togatenellus</emphasis>
</taxonomicName>
,
<date id="FFBE106FD529B218FF61FB42BA302BB5" box="[151,278,1207,1231]" pageId="1" pageNumber="517" value="1990-05-31">
<collectingDate id="EFFAE987D529B218FF61FB42BA302BB5" box="[151,278,1207,1231]" pageId="1" pageNumber="517" value="1990-05-31">31. v. 1990</collectingDate>
</date>
,
<collectorName id="26F55379D529B218FED7FB42BA802BB5" box="[289,422,1206,1231]" pageId="1" pageNumber="517">T. Okazawa</collectorName>
&amp;
<collectorName id="26F55379D529B218FE3CFB42B9032BB5" box="[458,549,1207,1231]" pageId="1" pageNumber="517">M. Sasa</collectorName>
[specimen No. 182:73 (NSMT-I-Dip 4716)]
</materialsCitation>
.
</paragraph>
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<subSubSection id="C31A6524D529B218FF31FB2FBABA2A95" pageId="1" pageNumber="517" type="description">
<paragraph id="8BBF36AFD529B218FF31FB2FBABA2A95" blockId="1.[151,1437,1170,1519]" pageId="1" pageNumber="517">
The species was treated as a member of
<taxonomicName id="4C004D2CD529B218FD43FB2EB8A92B89" authorityName="Thienemann" authorityYear="1934" box="[693,911,1243,1267]" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974EABDD529B218FD43FB2EB8A92B89" box="[693,911,1243,1267]" italics="true" pageId="1" pageNumber="517">Bryophaenocladius</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF914B5ED529B218FC32FB2EBFB32B89" author="Yamamoto, M." box="[964,1173,1243,1267]" pageId="1" pageNumber="517" pagination="1 - 121" refId="ref6606" refString="Yamamoto, M. (2004) A catalog of Japanese Orthocladiinae (Diptera: Chironomidae). Makunagi / Acta Dipterologica, 21, 1 - 121." type="journal article" year="2004">Yamamoto (2004)</bibRefCitation>
based on the original description. However, it becomes clear that the
<typeStatus id="54BB880DD529B218FD42FB0BB8302A6D" box="[692,790,1278,1303]" pageId="1" pageNumber="517" type="holotype">holotype</typeStatus>
of the species bears such features as follows: tarsomeres 12 of mid and hind legs with pseudospurs, sensilla chaetica present on tarsomere 1 of hind leg; lateral spines of mid and hind tibiae diverge from shaft of spur; small virga preset, anal point hyaline and bare. These features indicate that the species should be transferred to the genus
<taxonomicName id="4C004D2CD529B218FCC5FA9EBE232AF9" authority="Kieffer. Kobayashi (2012)" authorityName="Kieffer. Kobayashi" authorityYear="2012" box="[819,1285,1386,1411]" class="Insecta" family="Chironomidae" genus="Chaetocladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974EABDD529B218FCC5FA9EB8F32AF9" box="[819,981,1387,1411]" italics="true" pageId="1" pageNumber="517">Chaetocladius</emphasis>
Kieffer.
<bibRefCitation id="EF914B5ED529B218FBCDFA9FBE232AF9" author="Kobayashi, T." box="[1083,1285,1386,1411]" pageId="1" pageNumber="517" pagination="13 - 15" refId="ref5239" refString="Kobayashi, T. (2012) Chaetocladius (s. str.) eugenyii sp. n. (Diptera, Chironomidae, Orthocladiinae) from Japan. Euroasian Entomological Journal, 11, 13 - 15." type="journal article" year="2012">Kobayashi (2012)</bibRefCitation>
</taxonomicName>
reported that
<taxonomicName id="4C004D2CD529B218FF61FA7ABA632ADC" authorityName="Sasa et Okazawa" authorityYear="1992" box="[151,325,1422,1446]" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="species" species="togatenellus">
<emphasis id="B974EABDD529B218FF61FA7ABB8E2ADC" box="[151,168,1423,1446]" italics="true" pageId="1" pageNumber="517">B</emphasis>
.
<emphasis id="B974EABDD529B218FF4AFA7BBA632ADC" box="[188,325,1422,1446]" italics="true" pageId="1" pageNumber="517">togatenellus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C004D2CD529B218FE71FA7BB8262ADC" authorityName="Sasa et Tanaka" authorityYear="2001" box="[391,768,1422,1446]" class="Insecta" family="Chironomidae" genus="Paratrissocladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="species" species="sudagaicedeus">
<emphasis id="B974EABDD529B218FE71FA7BB8262ADC" box="[391,768,1422,1446]" italics="true" pageId="1" pageNumber="517">Paratrissocladius sudagaicedeus</emphasis>
</taxonomicName>
each is a member of
<taxonomicName id="4C004D2CD529B218FBE4FA7BBF922ADC" authorityName="Kieffer. Kobayashi" authorityYear="2012" box="[1042,1204,1422,1446]" class="Insecta" family="Chironomidae" genus="Chaetocladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B974EABDD529B218FBE4FA7BBF922ADC" box="[1042,1204,1422,1446]" italics="true" pageId="1" pageNumber="517">Chaetocladius</emphasis>
</taxonomicName>
by citing from the abstract of the meeting for the study of Japanese chironomid workers (
<bibRefCitation id="EF914B5ED529B218FC45FA46BE332AB1" author="Yamamoto, M. &amp; Yamamoto, N." box="[947,1301,1459,1483]" pageId="1" pageNumber="517" pagination="18 - 19" refId="ref6635" refString="Yamamoto, M. &amp; Yamamoto, N. (2007) Japanese Bryophaenocladius Thienemann. Yusurika, No. 32, 18 - 19. [in Japanese]" type="book chapter" year="2007">Yamamoto &amp; Yamamoto, 2007</bibRefCitation>
) which was not published formally.
</paragraph>
</subSubSection>
</treatment>
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