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<mods:title>Taxonomic study on the spider wasp genus Ceropales Latreille, 1796 (Pompilidae: Ceropalinae) with description of two new species from India</mods:title>
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<mods:namePart>Anju, K.</mods:namePart>
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<mods:affiliation>P. G. &amp; Research Department of Zoology, Government College Madappally, Vatakara, Kozhikode, Kerala- 673102, India, Affiliated to University of Calicut. &amp; anjukappu 91 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3312 - 7721</mods:affiliation>
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<mods:namePart>Thejass, P.</mods:namePart>
<mods:affiliation>P. G. &amp; Research Department of Zoology, Government College Madappally, Vatakara, Kozhikode, Kerala- 673102, India, Affiliated to University of Calicut.</mods:affiliation>
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<mods:namePart>Binoy, C.</mods:namePart>
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<mods:affiliation>Department of Zoology, Sree Narayana Guru College, Chelannur, Kozhikode, Kerala- 673616, India, Affiliated to University of Calicut. binoy _ doz @ uoc. ac. in; https: // orcid. org / 0000 - 0003 - 4309 - 5460</mods:affiliation>
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<mods:namePart>Kumar, P. Girish</mods:namePart>
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<mods:affiliation>Western Ghats Regional Centre, Zoological Survey of India, Eranhipalam, Kozhikode, Kerala- 673006, India. kpgiris @ gmail. com; https: // orcid. org / 0000 - 0003 - 2121 - 0165</mods:affiliation>
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Anju, Girish Kumar &amp; Thejass
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The new species runs close to 
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<taxonomicName authority="Lu &amp; Li, 2019" authorityName="Lu &amp; Li" authorityYear="2019" box="[795,1157,151,177]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="longisulcata" subGenus="Ceropales">
(
<emphasis box="[805,824,151,177]" italics="true" pageId="3" pageNumber="122">C</emphasis>
.) 
<emphasis box="[847,985,151,177]" italics="true" pageId="3" pageNumber="122">longisulcata</emphasis>
<bibRefCitation author="Lu, H. &amp; Bashir, N. H. &amp; Li, Q. &amp; Ma, L." box="[993,1157,151,177]" pageId="3" pageNumber="122" pagination="211 - 220" refId="ref7077" refString="Lu, H., Bashir, N. H., Li, Q. &amp; Ma, L. (2019) Two new species of Ceropales (Ceropales) Latreille (Hymenoptera, Pompilidae), with a key to species from China. Zootaxa, 4551 (2), 211 - 220. https: // doi. org / 10.11646 / zootaxa. 4551.2.5." type="journal article" year="2019">
Lu &amp; Li, 
<quantity box="[1101,1157,151,177]" metricMagnitude="1" metricUnit="m" metricValue="5.12826" pageId="3" pageNumber="122" unit="in" value="2019.0">2019</quantity>
</bibRefCitation>
</taxonomicName>
in the key to the species of the subgenus 
<taxonomicName authority="Latreille" authorityName="Latreille" authorityYear="1796" box="[337,554,187,213]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis box="[337,451,187,213]" italics="true" pageId="3" pageNumber="122">Ceropales</emphasis>
Latreille
</taxonomicName>
from 
<collectingCountry box="[625,692,187,213]" name="China" pageId="3" pageNumber="122">China</collectingCountry>
(
<bibRefCitation author="Lu, H. &amp; Bashir, N. H. &amp; Li, Q. &amp; Ma, L." box="[707,871,187,213]" pageId="3" pageNumber="122" pagination="211 - 220" refId="ref7077" refString="Lu, H., Bashir, N. H., Li, Q. &amp; Ma, L. (2019) Two new species of Ceropales (Ceropales) Latreille (Hymenoptera, Pompilidae), with a key to species from China. Zootaxa, 4551 (2), 211 - 220. https: // doi. org / 10.11646 / zootaxa. 4551.2.5." type="journal article" year="2019">
Lu 
<emphasis box="[748,806,187,213]" italics="true" pageId="3" pageNumber="122">et al.</emphasis>
2019
</bibRefCitation>
) in having flat propodeum with lateral side rather shallowly impressed and with deep medial longitudinal groove; frons remarkably flat anterior to antennal sockets and bent in obtuse-angle on 2/3 away from antennal sockets; ocelli forming acute triangle; and scutellum and postscutellum remarkably raised, much higher than mesonotum in lateral view. However, the new species differs from 
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[213,384,331,357]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[213,384,331,357]" italics="true" pageId="3" pageNumber="122">C. longisulcata</emphasis>
</taxonomicName>
in having fore wing with upper margin of SMC2 and SMC3 equal in length (in 
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[1291,1429,331,357]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[1291,1429,331,357]" italics="true" pageId="3" pageNumber="122">longisulcata</emphasis>
</taxonomicName>
, upper margin of SMC3 shorter than SMC2); frons and vertex with combination of small punctures mixed with sparse large punctures (in 
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[446,584,403,429]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[446,584,403,429]" italics="true" pageId="3" pageNumber="122">longisulcata</emphasis>
</taxonomicName>
, deeper and densely set punctures on frons and vertex); labrum pale yellow without any dark patches (in 
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[477,615,439,465]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[477,615,439,465]" italics="true" pageId="3" pageNumber="122">longisulcata</emphasis>
</taxonomicName>
, labrum with a median dark patch); and frons black without any marking in the middle (in 
<taxonomicName authorityName="Lu &amp; Li" authorityYear="2019" box="[344,482,475,501]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="longisulcata">
<emphasis box="[344,482,475,501]" italics="true" pageId="3" pageNumber="122">longisulcata</emphasis>
</taxonomicName>
, frons with a yellow spot medially).
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="124" pageId="3" pageNumber="122" type="description">
<paragraph blockId="3.[151,1437,150,2013]" box="[199,1230,510,537]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,345,510,536]" pageId="3" pageNumber="122">Description.</emphasis>
<typeStatus box="[352,459,511,537]" pageId="3" pageNumber="122">Holotype</typeStatus>
, 
<specimenCount box="[470,491,511,537]" pageId="3" pageNumber="122" type="female">♀</specimenCount>
(
<figureCitation box="[506,622,511,537]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Figs 1–10</figureCitation>
). Body length 
<quantity box="[786,889,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" pageId="3" pageNumber="122" unit="mm" value="6.05">6.05 mm</quantity>
; length of fore wing 
<quantity box="[1123,1226,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="5.27" pageId="3" pageNumber="122" unit="mm" value="5.27">5.27 mm</quantity>
.
</paragraph>
<paragraph blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,287,546,573]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,280,546,573]" italics="true" pageId="3" pageNumber="122">Colour</emphasis>
.
</emphasis>
Overall colour black with antenna brownish black, ventral side of scape pale yellowish, lower frons below antenna and between antennal socket, pale yellow; inner eye margin adjacent to the medial emargination of compound eye, pale yellow; malar space and bands on outer margin of eyes pale yellow; clypeus pale yellowish with black spot medially; labrum pale yellow; basal 2/3 
<superScript attach="left" box="[724,738,654,668]" fontSize="6" pageId="3" pageNumber="122">rd</superScript>
of mandible pale yellow, apex reddish brown; maxillary palp brown; pronotum black with band on posterior margin and posterolateral angle pale yellowish; mesonotum entirely black; metanotum black with medial patch pale yellow; tegula pale yellow; propodeum black with posterolateral corner pale yellow; large spot on outer face of all coxae pale yellow, remainder black; all femora ferruginous; fore tibia and tarsus light brown; mid tibia honey brown with black patches; mid tarsus light brown; hind tibia and tarsus dark brown; mid and hind tibial spur pale yellow; metasoma black; posterior yellow band on T1–T5; T6 largely white; wings hyaline, apical region of fore wing with light brown infumation; veins and pterostigma dark brown.
</paragraph>
<paragraph blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,341,906,933]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,334,906,933]" italics="true" pageId="3" pageNumber="122">Pubescence</emphasis>
.
</emphasis>
Body covered with white pubescence; small impressed white setae on head and mesosoma; metasoma smooth, asetose; frons densely pubescent, concentrated adjacent to compound eyes, remainder of area with scattered white setae; clypeus densely pubescent with appressed short white setae; pubescence on mesosoma concentrated along lateral margins; pubescence on propodeum moderate, confined to lateral sides, midline somewhat bare.
</paragraph>
<paragraph blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,268,1086,1113]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,261,1086,1113]" italics="true" pageId="3" pageNumber="122">Head</emphasis>
.
</emphasis>
Head slightly broader than long (1.1×) in frontal view (
<figureCitation box="[908,979,1087,1113]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 4</figureCitation>
); frons occupying the interantennal area conspicuously elevated; a shallow median line present from anterior ocellus to interantennal space; frons and vertex coarsely pitted with few large punctures in between; compound eyes reniform, inner margins distinctly sinuate, strongly converging below (
<figureCitation box="[475,547,1195,1221]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 4</figureCitation>
); UID as long as MID, LID 0.5× UID and MID; ocellar area slightly raised, anterior ocellus little larger than posterior ocelli; OOD 2.1× POD (
<figureCitation box="[904,973,1230,1257]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 5</figureCitation>
); clypeus slightly convex in lateral view, about twice broad as long, free margin of clypeus slightly emarginated medially (
<figureCitation box="[1077,1148,1267,1293]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 4</figureCitation>
); labrum slightly broader than long, exposed beneath clypeus (
<figureCitation box="[573,644,1303,1329]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 4</figureCitation>
); mandible bidentate; malar space very much reduced; antenna thick, scape 1.4× as long as wide and 2.2× pedicel in profile, flagellomeres almost twice as long as broad, F1–F4 almost of equal length (
<figureCitation box="[334,403,1375,1401]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 3</figureCitation>
).
</paragraph>
<paragraph blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,329,1411,1437]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,322,1411,1437]" italics="true" pageId="3" pageNumber="122">Mesosoma</emphasis>
.
</emphasis>
Pronotum 0.5× as long as wide, densely pitted with large punctures scattered irregularly in between; anterolateral corners of pronotum angulate with posterior margin emarginated inwards (
<figureCitation box="[1134,1202,1447,1473]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 7</figureCitation>
); mesoscutum 1.16× as wide as long, densely pitted with scattered large punctures along the length; parapsidal furrow present through lateral sides of mesoscutum (
<figureCitation box="[473,541,1519,1545]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 6</figureCitation>
); mesoscutellum with conspicuous prescutellar sulcus (
<figureCitation box="[1158,1225,1519,1545]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 6</figureCitation>
); scutellum raised/ bulged in profile, punctuations similar to that on mesoscutum; mesopleuron with scattered punctures (
<figureCitation box="[1348,1421,1555,1581]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 7</figureCitation>
); metanotum rectangular, conspicuously elevated, surface micropunctured; metapostnotum much reduced and seen as small slit with few transverse rugae (
<figureCitation box="[588,655,1627,1653]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 6</figureCitation>
); metapleuron with fine, sparse punctures (
<figureCitation box="[1134,1201,1627,1653]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 7</figureCitation>
); propodeum 1.3× as wide as long along the middle, conspicuously flat, surface with fine, transverse rugulosity; median sulcus reaching nearly half length of propodeum; posterolateral angle rounded, posterior margin nearly straight (
<figureCitation box="[1225,1293,1699,1725]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 6</figureCitation>
); lateral side of propodeum with shallow rugulosity (
<figureCitation box="[596,664,1735,1761]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 7</figureCitation>
).
</paragraph>
<paragraph blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,261,1771,1797]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,253,1771,1797]" italics="true" pageId="3" pageNumber="122">Legs</emphasis>
.
</emphasis>
Tarsal claws normal, bent apically; short, erect and acute subapical tooth present; hind tibial spur long, 0.6× as long as hind basitarsus.
</paragraph>
<paragraph blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,278,1843,1869]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,271,1843,1869]" italics="true" pageId="3" pageNumber="122">Wings</emphasis>
.
</emphasis>
Fore wing with upper margin of SMC2 and SMC3 equal in length, lower margin of SMC3 longer than that of SMC2; SMC2 receiving crossvein 
<emphasis box="[636,707,1879,1905]" italics="true" pageId="3" pageNumber="122">1m-cu</emphasis>
medially, SMC3 receiving crossvein 
<emphasis box="[1139,1209,1879,1905]" italics="true" pageId="3" pageNumber="122">2m-cu</emphasis>
medially; length of SMC3 less than its distance from the wing apex; vein 
<emphasis box="[756,773,1915,1941]" italics="true" pageId="3" pageNumber="122">A</emphasis>
and crossvein 
<emphasis box="[940,990,1915,1940]" italics="true" pageId="3" pageNumber="122">cu-a</emphasis>
of hind wing forming obtuse angle.
</paragraph>
<paragraph blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,326,1951,1977]" pageId="3" pageNumber="122">
<emphasis bold="true" box="[199,319,1951,1977]" italics="true" pageId="3" pageNumber="122">Metasoma</emphasis>
.
</emphasis>
Fusiform; as broad and long as mesosoma; smooth (
<figureCitation box="[911,979,1951,1977]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[215,1372,181,1885]" captionTargetId="figure-19@4.[215,1372,181,1885]" captionTargetPageId="4" captionText="FIGURES 1–10. Ceropales (Ceropales) anaghae Anju, Girish Kumar &amp; Thejass, sp. nov., holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="3" pageNumber="122">Fig. 9</figureCitation>
); T1 sessile, parallel sided; lower margin of S6 flat in lateral view, pointed apically.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836282" ID-Zenodo-Dep="7836282" httpUri="https://zenodo.org/record/7836282/files/figure.png" pageId="4" pageNumber="123" startId="4.[151,264,1909,1933]" targetBox="[215,1372,181,1885]" targetPageId="4">
<paragraph blockId="4.[151,1437,1909,2042]" pageId="4" pageNumber="123">
<emphasis bold="true" box="[151,326,1909,1933]" pageId="4" pageNumber="123">FIGURES 1–10.</emphasis>
<taxonomicName authority="Anju, Girish Kumar &amp; Thejass" authorityName="Anju, Girish Kumar &amp; Thejass" authorityYear="2023" box="[332,970,1909,1934]" class="Insecta" family="Pompilidae" genus="Ceropales" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="anaghae" status="sp. nov." subGenus="Ceropales">
<emphasis box="[332,435,1909,1933]" italics="true" pageId="4" pageNumber="123">Ceropales</emphasis>
(
<emphasis box="[449,552,1909,1933]" italics="true" pageId="4" pageNumber="123">Ceropales</emphasis>
) 
<emphasis box="[566,653,1909,1933]" italics="true" pageId="4" pageNumber="123">anaghae</emphasis>
Anju, Girish Kumar &amp; Thejass
</taxonomicName>
, 
<emphasis bold="true" box="[979,1059,1910,1934]" pageId="4" pageNumber="123">
<taxonomicNameLabel box="[979,1059,1910,1934]" pageId="4" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, ♀. 1, habitus, lateral view; 2, habitus, dorsal view; 3, head and antenna, lateral view; 4, head, frontal view; 5, head, dorsal view; 6, mesosoma, dorsal view; 7, mesosoma, lateral view; 8, wings; 9, metasoma, lateral view; 10, metasoma, dorsal view. Scale bar: 1, 2, 8–2 mm; 3, 6, 9, 10–1 mm; 4, 5, 7–0.5 mm.
</paragraph>
</caption>
<paragraph blockId="5.[151,1436,151,357]" box="[199,392,151,177]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,267,151,177]" pageId="5" pageNumber="124">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="124" type="etymology">
<paragraph blockId="5.[151,1436,151,357]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,334,187,213]" pageId="5" pageNumber="124">Etymology.</emphasis>
The species name in feminine gender is derived from Anagha, after S. Anagha (research scholar, Zoological Survey of 
<collectingCountry box="[396,456,223,249]" name="India" pageId="5" pageNumber="124">India</collectingCountry>
, Western Ghats Regional Centre) who collected the 
<typeStatus box="[1041,1139,223,249]" pageId="5" pageNumber="124">holotype</typeStatus>
specimen.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="124" type="description">
<paragraph blockId="5.[151,1436,151,357]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,319,258,284]" pageId="5" pageNumber="124">Variation.</emphasis>
Relative density of pubescence varies from scattered to moderate setosity in 
<typeStatus box="[1200,1307,259,285]" pageId="5" pageNumber="124">paratypes</typeStatus>
along with variations in intensity of body coloration.
</paragraph>
</subSubSection>
<subSubSection box="[199,823,330,357]" pageId="5" pageNumber="124" type="distribution">
<paragraph blockId="5.[151,1436,151,357]" box="[199,823,330,357]" pageId="5" pageNumber="124">
<emphasis bold="true" box="[199,353,330,356]" pageId="5" pageNumber="124">Distribution.</emphasis>
<collectingCountry box="[359,417,331,357]" name="India" pageId="5" pageNumber="124">India</collectingCountry>
(
<collectingRegion box="[432,511,331,357]" country="India" name="Kerala" pageId="5" pageNumber="124">Kerala</collectingRegion>
, 
<collectingRegion box="[521,656,331,357]" country="India" name="Tamil Nadu" pageId="5" pageNumber="124">Tamil Nadu</collectingRegion>
, 
<collectingRegion box="[668,812,331,357]" country="India" name="Uttarakhand" pageId="5" pageNumber="124">Uttarakhand</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>