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<document id="47628F328AB9F5FAF69FE5BFADE2BD05" ID-CLB-Dataset="4293" ID-DOI="10.18590/euscorpius.2010.vol2010.iss92.1" ID-GBIF-Dataset="b4c2af58-5d10-4fef-8f6a-302d7ce970d5" ID-ISSN="1536-9307" ID-Zenodo-Dep="4649069" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617137184749" checkinUser="carolina" docAuthor="Teruel, Rolando &amp; Chazal, Léonard" docDate="2010" docId="039287DE4034267ECB3AFA4FFEB5F8B0" docLanguage="en" docName="Euscorpius.92.1-9.pdf" docOrigin="Euscorpius 92 (92)" docSource="https://mds.marshall.edu/euscorpius/vol2010/iss92/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Oiclus nanus Teruel et Chazal 2010, sp. nov." docType="treatment" docUuid="E4154796-2759-4DCB-BCE1-0D857FDDC8B7" docUuidSource="ZooBank" docVersion="9" lastPageNumber="9" masterDocId="FFABFFA640362674C846FF9BFF8CFF93" masterDocTitle="A new species of the genus Oiclus Simon, 1880 (Scorpiones: Scorpionidae: Diplocentrinae) from Guadeloupe, Lesser Antilles" masterLastPageNumber="9" masterPageNumber="1" pageNumber="1" updateTime="1698935551947" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="A5831D2A14593496176F68151DBC822D">A new species of the genus Oiclus Simon, 1880 (Scorpiones: Scorpionidae: Diplocentrinae) from Guadeloupe, Lesser Antilles</mods:title>
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<mods:namePart id="98836951020220A398A175FB615C2272">Teruel, Rolando</mods:namePart>
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<treatment id="039287DE4034267ECB3AFA4FFEB5F8B0" ID-DOI="http://doi.org/10.5281/zenodo.5507481" ID-GBIF-Taxon="187974479" ID-Zenodo-Dep="5507481" LSID="urn:lsid:zoobank.org:act:E4154796-2759-4DCB-BCE1-0D857FDDC8B7" httpUri="http://treatment.plazi.org/id/039287DE4034267ECB3AFA4FFEB5F8B0" lastPageId="10" lastPageNumber="9" pageId="2" pageNumber="1">
<subSubSection id="C321654340342676CB3AFA4FFAEEFA63" box="[892,1378,1492,1520]" pageId="2" pageNumber="1" type="nomenclature">
<paragraph id="8B8436C840342676CB3AFA4FFAEEFA63" blockId="2.[892,1378,1492,1520]" box="[892,1378,1492,1520]" pageId="2" pageNumber="1">
<heading id="D0CC81A440342676CB3AFA4FFAEEFA63" box="[892,1378,1492,1520]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1" reason="3">
<emphasis id="B94FEADA40342676CB3AFA4FFAEEFA63" bold="true" box="[892,1378,1492,1520]" pageId="2" pageNumber="1">
<taxonomicName id="4C3B4D4B40342676CB3AFA4FFB7BFA63" ID-CoL="48ZG7" authority="Teruel et Chazal" authorityName="Teruel et Chazal" authorityYear="2010" box="[892,1271,1492,1520]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA40342676CB3AFA4FFB90FA7C" bold="true" box="[892,1052,1492,1519]" italics="true" pageId="2" pageNumber="1">Oiclus nanus</emphasis>
Teruel et Chazal
</taxonomicName>
,
<taxonomicNameLabel id="A27C57A140342676CD45FA4DFAEEFA63" box="[1283,1378,1494,1520]" pageId="2" pageNumber="1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C321654340342676CBBEFA6DFB6BF91B" pageId="2" pageNumber="1" type="description">
<paragraph id="8B8436C840342676CBBEFA6DFB6BF99E" blockId="2.[1016,1255,1526,1549]" box="[1016,1255,1526,1549]" pageId="2" pageNumber="1">
(
<figureCitation id="13002A4D40342676CBB9FA6DFB0DF99E" box="[1023,1153,1526,1549]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="3.[192,268,1894,1917]" captionStartId-1="5.[192,268,1821,1844]" captionStartId-2="6.[192,268,1468,1491]" captionStartId-3="8.[192,268,1875,1898]" captionStartId-4="9.[192,268,1159,1182]" captionTargetBox-0="[191,1458,180,1873]" captionTargetBox-1="[192,1439,167,1800]" captionTargetBox-2="[192,1439,192,1439]" captionTargetBox-3="[191,1440,167,1846]" captionTargetBox-4="[191,1442,191,1130]" captionTargetId-0="figure-6@3.[192,1457,181,1872]" captionTargetId-1="figure-5@5.[192,1439,167,1800]" captionTargetId-2="figure-50@6.[192,1439,192,1439]" captionTargetId-3="figure-39@8.[192,1439,168,1845]" captionTargetId-4="figure-861@9.[192,1441,192,1129]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Figure 1: Adult male holotype of Oiclus nanus sp. n.: a) entire dorsal view; b) entire ventral view." captionText-1="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." captionText-2="Figure 3: Type specimens of Oiclus nanus sp. n., still alive when photographed: a) adult male holotype, in its natural habitat; bc) adult male paratypes, in their natural habitat; d) juvenile male paratype, in captivity." captionText-3="Figure 4: Two views of localities where Oiclus nanus sp. n. occurs at Pointe des Châteaux: a) habitat; b) microhabitat." captionText-4="Figure 5: Known distribution of the genus Oiclus: O. nanus sp. n. (white square with black cross, including the potential records from Bouillantes and Les Saintes), Oiclus questeli (black square with white cross), Oiclus purvesii purvesii (black square), Oiclus purvesii sabae (white square), and Oiclus sp. (white square with black dot)." figureDoi-0="http://doi.org/10.5281/zenodo.4649071" figureDoi-1="http://doi.org/10.5281/zenodo.4649073" figureDoi-2="http://doi.org/10.5281/zenodo.4649075" figureDoi-3="http://doi.org/10.5281/zenodo.4649077" figureDoi-4="http://doi.org/10.5281/zenodo.4649079" httpUri-0="https://zenodo.org/record/4649071/files/figure.png" httpUri-1="https://zenodo.org/record/4649073/files/figure.png" httpUri-2="https://zenodo.org/record/4649075/files/figure.png" httpUri-3="https://zenodo.org/record/4649077/files/figure.png" httpUri-4="https://zenodo.org/record/4649079/files/figure.png" pageId="2" pageNumber="1">Figures 15</figureCitation>
,
<tableCitation id="C6B9037340342676CCCBFA6DFB50F99E" box="[1165,1244,1526,1549]" captionStart="Table 1" captionStartId="4.[192,257,1091,1114]" captionTargetPageId="4" captionText="Table 1: Measurements of the three adult types of Oiclus nanus, sp. n. Abbreviations: length (L), width (W), posterior width (Wp), depth (H)." httpUri="http://table.plazi.org/id/DF44664040322670C886FBD8FEEEFBE0" pageId="2" pageNumber="1" tableUuid="DF44664040322670C886FBD8FEEEFBE0">Table 1</tableCitation>
)
</paragraph>
<paragraph id="8B8436C840342676CB78F9AFFAE0F9D8" blockId="2.[830,1441,1588,1672]" box="[830,1388,1588,1611]" pageId="2" pageNumber="1">
(?)
<taxonomicName id="4C3B4D4B40342676CB25F9AFFAB1F9D8" authority="purvesii: Francke, 1978: 33" authorityName="Francke" authorityPageNumber="33" authorityYear="1978" box="[867,1341,1588,1611]" class="Arachnida" family="Diplocentridae" genus="Oieclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="purvesii" subSpecies="purvesii">
<emphasis id="B94FEADA40342676CB25F9AFFBE1F9D8" box="[867,1133,1588,1611]" italics="true" pageId="2" pageNumber="1">Oieclus purvesii purvesii</emphasis>
: Francke, 1978: 33
</taxonomicName>
, 35.
</paragraph>
<paragraph id="8B8436C840342676CB78F9C9FB6BF91B" blockId="2.[830,1441,1588,1672]" pageId="2" pageNumber="1">
(?)
<taxonomicName id="4C3B4D4B40342676CB2EF9C9FC15F91B" authority="purvesii: Teruel &amp; Francke, 2006: 286" authorityName="purvesii: Teruel &amp; Francke" authorityPageNumber="286" authorityYear="2006" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="purvesii" subSpecies="purvesii">
<emphasis id="B94FEADA40342676CB2EF9C9FBFFF9FA" box="[872,1139,1618,1641]" italics="true" pageId="2" pageNumber="1">Oiclus purvesii purvesii</emphasis>
: Teruel &amp; Francke, 2006: 286
</taxonomicName>
, fig. 2; Teruel, 2008: 95, fig. 5.
</paragraph>
</subSubSection>
<subSubSection id="C321654340342670CB78F935FEDBFA48" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1" type="diagnosis">
<paragraph id="8B8436C840342670CB78F935FEDBFA48" blockId="2.[830,1441,1710,1917]" lastBlockId="4.[192,802,1192,1499]" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1">
<emphasis id="B94FEADA40342676CB78F935FC20F955" bold="true" box="[830,940,1710,1734]" italics="true" pageId="2" pageNumber="1">Diagnosis</emphasis>
(males only, females unknown): species of small size (
<quantity id="4CC39B2D40342676CBFFF956FBA2F977" box="[953,1070,1741,1764]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="2" pageNumber="1" unit="mm" value="21.5" valueMax="23.0" valueMin="20.0">2023 mm</quantity>
) for the genus. Body light brown, diffusely patterned with dark brown; legs yellowish; pedipalps and metasoma with carinae and fingers faintly infuscate. Entire body conspicuously hirsute, especially on pedipalps, mesosoma and metasoma. Carapace polished, with very finely and densely granulose areas symmetrically scattered; tergites finely and densely granulose. Punctate tegument restricted to pedipalp patella and chela in adults, absent in juveniles. Pedipalp chela robust, with dorsoexternal surfaces granulose and strongly reticulate. Metasoma moderately to weakly carinated in dorsal and lateral surfaces, intercarinal tegument smooth and polished. Pectines with fulcra variable from moderately well developed to essentially absent; tooth count 7/7. Modal tarsal spine formula 3/3: 4/4: 5/5: 5/5.
</paragraph>
</subSubSection>
<caption id="DF44664040352677C886F8FDFB08F8EE" ID-DOI="http://doi.org/10.5281/zenodo.4649071" ID-Zenodo-Dep="4649071" box="[192,1156,1894,1917]" httpUri="https://zenodo.org/record/4649071/files/figure.png" pageId="3" pageNumber="2" startId="3.[192,268,1894,1917]" targetBox="[191,1458,180,1873]" targetPageId="3">
<paragraph id="8B8436C840352677C886F8FDFB08F8EE" blockId="3.[192,1156,1894,1917]" box="[192,1156,1894,1917]" pageId="3" pageNumber="2">
<emphasis id="B94FEADA40352677C886F8FDFEA5F8EE" bold="true" box="[192,297,1894,1917]" pageId="3" pageNumber="2">Figure 1:</emphasis>
Adult male holotype of
<taxonomicName id="4C3B4D4B40352677CA53F8F3FD1EF8EE" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[533,658,1896,1917]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA40352677CA53F8F3FD1EF8EE" box="[533,658,1896,1917]" italics="true" pageId="3" pageNumber="2">Oiclus nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA40352677CADEF8F3FD43F8EE" bold="true" box="[664,719,1896,1917]" pageId="3" pageNumber="2">
<taxonomicNameLabel id="A27C57A140352677CADEF8F3FD43F8EE" box="[664,719,1896,1917]" pageId="3" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="B94FEADA40352677CA9DF8F3FD63F8EE" bold="true" box="[731,751,1896,1917]" pageId="3" pageNumber="2">a)</emphasis>
entire dorsal view;
<emphasis id="B94FEADA40352677CBE8F8F3FC4FF8EE" bold="true" box="[942,963,1896,1917]" pageId="3" pageNumber="2">b)</emphasis>
entire ventral view.
</paragraph>
</caption>
<paragraph id="8B8436C840322670C91DFF7FFB50FB81" pageId="4" pageNumber="3">
<table id="F93BC4684032D98BC89EFF7FFA90FB81" box="[216,1308,228,1042]" gridcols="4" gridrows="20" pageId="4" pageNumber="3">
<tr id="350B348A4032D98BC89EFF7FFA90FE93" box="[216,1308,228,256]" gridrow="0" pageId="4" pageNumber="3" rowspan-1="1">
<th id="76DA5DF64032D98BC89EFF7FFE75FE93" box="[216,505,228,256]" gridcol="0" gridrow="0" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C91DFF7FFE75FE93" bold="true" box="[347,505,228,256]" pageId="4" pageNumber="3">Dimensions</emphasis>
</th>
<th id="76DA5DF64032D98BCAB1FF7FFC33FE93" box="[759,959,228,256]" gridcol="2" gridrow="0" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670CB76FF7FFC29FE93" bold="true" box="[816,933,228,256]" pageId="4" pageNumber="3">holotype</emphasis>
</th>
<th id="76DA5DF64032D98BCC13FF7FFA90FE93" box="[1109,1308,228,256]" gridcol="3" gridrow="0" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670CCCDFF7FFA8AFE93" bold="true" box="[1163,1286,228,256]" pageId="4" pageNumber="3">paratype</emphasis>
</th>
</tr>
<tr id="350B348A4032D98BC89EFE89FA90FEBD" box="[216,1308,274,302]" gridrow="1" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFE89FE75FEBD" box="[216,505,274,302]" gridcol="0" gridrow="1" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C89EFE89FED0FEBD" bold="true" box="[216,348,274,302]" pageId="4" pageNumber="3">Carapace</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FE89FD13FEBD" box="[548,671,274,302]" gridcol="1" gridrow="1" pageId="4" pageNumber="3">L / Wp</td>
<td id="76DA5DF64032D98BCAB1FE89FC33FEBD" box="[759,959,274,302]" gridcol="2" gridrow="1" pageId="4" pageNumber="3">2.70 / 2.70</td>
<td id="76DA5DF64032D98BCC13FE89FA90FEBD" box="[1109,1308,274,302]" gridcol="3" gridrow="1" pageId="4" pageNumber="3">3.20 / 3.10</td>
</tr>
<tr id="350B348A4032D98BC89EFEA0FA90FEC4" box="[216,1308,315,343]" gridrow="2" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFEA0FE75FEC4" box="[216,505,315,343]" gridcol="0" gridrow="2" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C89EFEA0FEE4FEC4" bold="true" box="[216,360,315,343]" pageId="4" pageNumber="3">Mesosoma</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FEA0FD13FEC4" box="[548,671,315,343]" gridcol="1" gridrow="2" pageId="4" pageNumber="3">L</td>
<td id="76DA5DF64032D98BCAB1FEA0FC33FEC4" box="[759,959,315,343]" gridcol="2" gridrow="2" pageId="4" pageNumber="3">5.80</td>
<td id="76DA5DF64032D98BCC13FEA0FA90FEC4" box="[1109,1308,315,343]" gridcol="3" gridrow="2" pageId="4" pageNumber="3">6.50</td>
</tr>
<tr id="350B348A4032D98BC89EFEF8FA90FEEC" box="[216,1308,355,383]" gridrow="3" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFEF8FE75FEEC" box="[216,505,355,383]" gridcol="0" gridrow="3" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8ADFEF8FE0BFEEC" bold="true" box="[235,391,355,383]" pageId="4" pageNumber="3">Tergite VII</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FEF8FD13FEEC" box="[548,671,355,383]" gridcol="1" gridrow="3" pageId="4" pageNumber="3">L</td>
<td id="76DA5DF64032D98BCAB1FEF8FC33FEEC" box="[759,959,355,383]" gridcol="2" gridrow="3" pageId="4" pageNumber="3">1.20 / 2.30</td>
<td id="76DA5DF64032D98BCC13FEF8FA90FEEC" box="[1109,1308,355,383]" gridcol="3" gridrow="3" pageId="4" pageNumber="3">1.50 / 2.80</td>
</tr>
<tr id="350B348A4032D98BC89EFE17FA90FE3B" box="[216,1308,396,424]" gridrow="4" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFE17FE75FE3B" box="[216,505,396,424]" gridcol="0" gridrow="4" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C89EFE17FEEAFE3B" bold="true" box="[216,358,396,424]" pageId="4" pageNumber="3">Metasoma</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FE17FD13FE3B" box="[548,671,396,424]" gridcol="1" gridrow="4" pageId="4" pageNumber="3">L</td>
<td id="76DA5DF64032D98BCAB1FE17FC33FE3B" box="[759,959,396,424]" gridcol="2" gridrow="4" pageId="4" pageNumber="3">11.60</td>
<td id="76DA5DF64032D98BCC13FE17FA90FE3B" box="[1109,1308,396,424]" gridcol="3" gridrow="4" pageId="4" pageNumber="3">13.50</td>
</tr>
<tr id="350B348A4032D98BC89EFE2FFA90FE43" box="[216,1308,436,464]" gridrow="5" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFE2FFE75FE43" box="[216,505,436,464]" gridcol="0" gridrow="5" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8AFFE2FFEFFFE43" bold="true" box="[233,371,436,464]" pageId="4" pageNumber="3">Segment I</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FE2FFD13FE43" box="[548,671,436,464]" gridcol="1" gridrow="5" pageId="4" pageNumber="3">L / W</td>
<td id="76DA5DF64032D98BCAB1FE2FFC33FE43" box="[759,959,436,464]" gridcol="2" gridrow="5" pageId="4" pageNumber="3">1.40 / 1.70</td>
<td id="76DA5DF64032D98BCC13FE2FFA90FE43" box="[1109,1308,436,464]" gridcol="3" gridrow="5" pageId="4" pageNumber="3">1.60 / 1.90</td>
</tr>
<tr id="350B348A4032D98BC89EFE46FA90FE6A" box="[216,1308,477,505]" gridrow="6" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFE46FE75FE6A" box="[216,505,477,505]" gridcol="0" gridrow="6" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8ADFE46FE0EFE6A" bold="true" box="[235,386,477,505]" pageId="4" pageNumber="3">Segment II</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FE46FD13FE6A" box="[548,671,477,505]" gridcol="1" gridrow="6" pageId="4" pageNumber="3">L /W</td>
<td id="76DA5DF64032D98BCAB1FE46FC33FE6A" box="[759,959,477,505]" gridcol="2" gridrow="6" pageId="4" pageNumber="3">1.60 / 1.55</td>
<td id="76DA5DF64032D98BCC13FE46FA90FE6A" box="[1109,1308,477,505]" gridcol="3" gridrow="6" pageId="4" pageNumber="3">1.80 / 1.75</td>
</tr>
<tr id="350B348A4032D98BC89EFD9EFA90FDB2" box="[216,1308,517,545]" gridrow="7" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFD9EFE75FDB2" box="[216,505,517,545]" gridcol="0" gridrow="7" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8ADFD9EFE02FDB2" bold="true" box="[235,398,517,545]" pageId="4" pageNumber="3">Segment III</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FD9EFD13FDB2" box="[548,671,517,545]" gridcol="1" gridrow="7" pageId="4" pageNumber="3">L / W</td>
<td id="76DA5DF64032D98BCAB1FD9EFC33FDB2" box="[759,959,517,545]" gridcol="2" gridrow="7" pageId="4" pageNumber="3">1.70 / 1.50</td>
<td id="76DA5DF64032D98BCC13FD9EFA90FDB2" box="[1109,1308,517,545]" gridcol="3" gridrow="7" pageId="4" pageNumber="3">2.00 / 1.70</td>
</tr>
<tr id="350B348A4032D98BC89EFDB5FA90FDD9" box="[216,1308,558,586]" gridrow="8" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFDB5FE75FDD9" box="[216,505,558,586]" gridcol="0" gridrow="8" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8ADFDB5FE00FDD9" bold="true" box="[235,396,558,586]" pageId="4" pageNumber="3">Segment IV</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FDB5FD13FDD9" box="[548,671,558,586]" gridcol="1" gridrow="8" pageId="4" pageNumber="3">L / W</td>
<td id="76DA5DF64032D98BCAB1FDB5FC33FDD9" box="[759,959,558,586]" gridcol="2" gridrow="8" pageId="4" pageNumber="3">2.00 / 1.40</td>
<td id="76DA5DF64032D98BCC13FDB5FA90FDD9" box="[1109,1308,558,586]" gridcol="3" gridrow="8" pageId="4" pageNumber="3">2.30 / 1.60</td>
</tr>
<tr id="350B348A4032D98BC89EFDCDFA90FDE1" box="[216,1308,598,626]" gridrow="9" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFDCDFE75FDE1" box="[216,505,598,626]" gridcol="0" gridrow="9" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8ADFDCDFE0CFDE1" bold="true" box="[235,384,598,626]" pageId="4" pageNumber="3">Segment V</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FDCDFD13FDE1" box="[548,671,598,626]" gridcol="1" gridrow="9" pageId="4" pageNumber="3">L / W</td>
<td id="76DA5DF64032D98BCAB1FDCDFC33FDE1" box="[759,959,598,626]" gridcol="2" gridrow="9" pageId="4" pageNumber="3">2.50 / 1.35</td>
<td id="76DA5DF64032D98BCC13FDCDFA90FDE1" box="[1109,1308,598,626]" gridcol="3" gridrow="9" pageId="4" pageNumber="3">2.90 / 1.55</td>
</tr>
<tr id="350B348A4032D98BC89EFDE4FA90FD08" box="[216,1308,639,667]" gridrow="10" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFDE4FE75FD08" box="[216,505,639,667]" gridcol="0" gridrow="10" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C89EFDE4FEBFFD08" bold="true" box="[216,307,639,667]" pageId="4" pageNumber="3">Telson</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FDE4FD13FD08" box="[548,671,639,667]" gridcol="1" gridrow="10" pageId="4" pageNumber="3">L</td>
<td id="76DA5DF64032D98BCAB1FDE4FC33FD08" box="[759,959,639,667]" gridcol="2" gridrow="10" pageId="4" pageNumber="3">2.40</td>
<td id="76DA5DF64032D98BCC13FDE4FA90FD08" box="[1109,1308,639,667]" gridcol="3" gridrow="10" pageId="4" pageNumber="3">2.90</td>
</tr>
<tr id="350B348A4032D98BC89EFD3CFA90FD50" box="[216,1308,679,707]" gridrow="11" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFD3CFE75FD50" box="[216,505,679,707]" gridcol="0" gridrow="11" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8AFFD3CFEC5FD50" bold="true" box="[233,329,679,707]" pageId="4" pageNumber="3">Vesicle</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FD3CFD13FD50" box="[548,671,679,707]" gridcol="1" gridrow="11" pageId="4" pageNumber="3">L / W / H</td>
<td id="76DA5DF64032D98BCAB1FD3CFC33FD50" box="[759,959,679,707]" gridcol="2" gridrow="11" pageId="4" pageNumber="3">1.70 / 1.20 / 1.00</td>
<td id="76DA5DF64032D98BCC13FD3CFA90FD50" box="[1109,1308,679,707]" gridcol="3" gridrow="11" pageId="4" pageNumber="3">2.10 / 1.40 / 1.10</td>
</tr>
<tr id="350B348A4032D98BC89EFD54FA90FD7F" box="[216,1308,719,748]" gridrow="12" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFD54FE75FD7F" box="[216,505,719,748]" gridcol="0" gridrow="12" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8ABFD4BFED5FD7F" bold="true" box="[237,345,720,748]" pageId="4" pageNumber="3">Aculeus</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FD54FD13FD7F" box="[548,671,719,748]" gridcol="1" gridrow="12" pageId="4" pageNumber="3">L</td>
<td id="76DA5DF64032D98BCAB1FD54FC33FD7F" box="[759,959,719,748]" gridcol="2" gridrow="12" pageId="4" pageNumber="3">0.70</td>
<td id="76DA5DF64032D98BCC13FD54FA90FD7F" box="[1109,1308,719,748]" gridcol="3" gridrow="12" pageId="4" pageNumber="3">0.80</td>
</tr>
<tr id="350B348A4032D98BC89EFD63FA90FC87" box="[216,1308,760,788]" gridrow="13" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFD63FE75FC87" box="[216,505,760,788]" gridcol="0" gridrow="13" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C89EFD63FEDDFC87" bold="true" box="[216,337,760,788]" pageId="4" pageNumber="3">Pedipalp</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FD63FD13FC87" box="[548,671,760,788]" gridcol="1" gridrow="13" pageId="4" pageNumber="3">L</td>
<td id="76DA5DF64032D98BCAB1FD63FC33FC87" box="[759,959,760,788]" gridcol="2" gridrow="13" pageId="4" pageNumber="3">7.30</td>
<td id="76DA5DF64032D98BCC13FD63FA90FC87" box="[1109,1308,760,788]" gridcol="3" gridrow="13" pageId="4" pageNumber="3">8.90</td>
</tr>
<tr id="350B348A4032D98BC89EFCBBFA90FCAE" box="[216,1308,800,829]" gridrow="14" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFCBBFE75FCAE" box="[216,505,800,829]" gridcol="0" gridrow="14" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8AFFCBAFEC9FCAE" bold="true" box="[233,325,801,829]" pageId="4" pageNumber="3">Femur</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FCBBFD13FCAE" box="[548,671,800,829]" gridcol="1" gridrow="14" pageId="4" pageNumber="3">L / W</td>
<td id="76DA5DF64032D98BCAB1FCBBFC33FCAE" box="[759,959,800,829]" gridcol="2" gridrow="14" pageId="4" pageNumber="3">1.80 / 0.90</td>
<td id="76DA5DF64032D98BCC13FCBBFA90FCAE" box="[1109,1308,800,829]" gridcol="3" gridrow="14" pageId="4" pageNumber="3">2.10 / 1.00</td>
</tr>
<tr id="350B348A4032D98BC89EFCD2FA90FCF6" box="[216,1308,841,869]" gridrow="15" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFCD2FE75FCF6" box="[216,505,841,869]" gridcol="0" gridrow="15" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8AFFCD2FECBFCF6" bold="true" box="[233,327,841,869]" pageId="4" pageNumber="3">Patella</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FCD2FD13FCF6" box="[548,671,841,869]" gridcol="1" gridrow="15" pageId="4" pageNumber="3">L / W</td>
<td id="76DA5DF64032D98BCAB1FCD2FC33FCF6" box="[759,959,841,869]" gridcol="2" gridrow="15" pageId="4" pageNumber="3">2.00 / 0.95</td>
<td id="76DA5DF64032D98BCC13FCD2FA90FCF6" box="[1109,1308,841,869]" gridcol="3" gridrow="15" pageId="4" pageNumber="3">2.50 / 1.10</td>
</tr>
<tr id="350B348A4032D98BC89EFCEAFA90FC1D" box="[216,1308,881,910]" gridrow="16" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFCEAFE75FC1D" box="[216,505,881,910]" gridcol="0" gridrow="16" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8AFFCE9FEB5FC1D" bold="true" box="[233,313,882,910]" pageId="4" pageNumber="3">Chela</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FCEAFD13FC1D" box="[548,671,881,910]" gridcol="1" gridrow="16" pageId="4" pageNumber="3">L</td>
<td id="76DA5DF64032D98BCAB1FCEAFC33FC1D" box="[759,959,881,910]" gridcol="2" gridrow="16" pageId="4" pageNumber="3">3.50</td>
<td id="76DA5DF64032D98BCC13FCEAFA90FC1D" box="[1109,1308,881,910]" gridcol="3" gridrow="16" pageId="4" pageNumber="3">4.30</td>
</tr>
<tr id="350B348A4032D98BC89EFC01FA90FC25" box="[216,1308,922,950]" gridrow="17" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFC01FE75FC25" box="[216,505,922,950]" gridcol="0" gridrow="17" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8BAFC01FEC4FC25" bold="true" box="[252,328,922,950]" pageId="4" pageNumber="3">Hand</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FC01FD13FC25" box="[548,671,922,950]" gridcol="1" gridrow="17" pageId="4" pageNumber="3">L / W / H</td>
<td id="76DA5DF64032D98BCAB1FC01FC33FC25" box="[759,959,922,950]" gridcol="2" gridrow="17" pageId="4" pageNumber="3">1.50 / 1.70 / 1.60</td>
<td id="76DA5DF64032D98BCC13FC01FA90FC25" box="[1109,1308,922,950]" gridcol="3" gridrow="17" pageId="4" pageNumber="3">1.80 / 2.00 / 2.30</td>
</tr>
<tr id="350B348A4032D98BC89EFC59FA90FC4C" box="[216,1308,962,991]" gridrow="18" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFC59FE75FC4C" box="[216,505,962,991]" gridcol="0" gridrow="18" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C8B9FC58FE5CFC4C" bold="true" box="[255,464,963,991]" pageId="4" pageNumber="3">Movable finger</emphasis>
</th>
<td id="76DA5DF64032D98BCA62FC59FD13FC4C" box="[548,671,962,991]" gridcol="1" gridrow="18" pageId="4" pageNumber="3">L</td>
<td id="76DA5DF64032D98BCAB1FC59FC33FC4C" box="[759,959,962,991]" gridcol="2" gridrow="18" pageId="4" pageNumber="3">2.00</td>
<td id="76DA5DF64032D98BCC13FC59FA90FC4C" box="[1109,1308,962,991]" gridcol="3" gridrow="18" pageId="4" pageNumber="3">2.50</td>
</tr>
<tr id="350B348A4032D98BC89EFC6DFA90FB81" box="[216,1308,1014,1042]" gridrow="19" pageId="4" pageNumber="3">
<th id="76DA5DF64032D98BC89EFC6DFE75FB81" box="[216,505,1014,1042]" gridcol="0" gridrow="19" pageId="4" pageNumber="3">Total</th>
<td id="76DA5DF64032D98BCA62FC6DFD13FB81" box="[548,671,1014,1042]" gridcol="1" gridrow="19" pageId="4" pageNumber="3">L</td>
<td id="76DA5DF64032D98BCAB1FC6DFC33FB81" box="[759,959,1014,1042]" gridcol="2" gridrow="19" pageId="4" pageNumber="3">20.10</td>
<td id="76DA5DF64032D98BCC13FC6DFA90FB81" box="[1109,1308,1014,1042]" gridcol="3" gridrow="19" pageId="4" pageNumber="3">23.20</td>
</tr>
</table>
</paragraph>
<caption id="DF44664040322670C886FBD8FEEEFBE0" ID-Table-UUID="DF44664040322670C886FBD8FEEEFBE0" httpUri="http://table.plazi.org/id/DF44664040322670C886FBD8FEEEFBE0" pageId="4" pageNumber="3" startId="4.[192,257,1091,1114]" targetBox="[216,1308,228,1042]" targetIsTable="true" targetPageId="4">
<paragraph id="8B8436C840322670C886FBD8FEEEFBE0" blockId="4.[192,1440,1091,1139]" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C886FBD8FEACFBC9" bold="true" box="[192,288,1091,1114]" pageId="4" pageNumber="3">Table 1:</emphasis>
Measurements of the three adult types of
<taxonomicName id="4C3B4D4B40322670CA83FBDEFCC9FBC9" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[709,837,1093,1114]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA40322670CA83FBDEFCC9FBC9" box="[709,837,1093,1114]" italics="true" pageId="4" pageNumber="3">Oiclus nanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27C57A140322670CB15FBDEFC05FBC9" box="[851,905,1093,1114]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
Abbreviations: length (L), width (W), posterior width (Wp), depth (H).
</paragraph>
</caption>
<subSubSection id="C321654340322670C886F999FEA9F90B" pageId="4" pageNumber="3" type="materials_examined">
<paragraph id="8B8436C840322670C886F999FEA9F90B" blockId="4.[192,802,1538,1688]" pageId="4" pageNumber="3">
<materialsCitation id="3B533C9540322670C886F999FE50F9CA" ID-GBIF-Occurrence="3352114304" collectingDate="2009-07-09" collectionCode="RTO" collectorName="L. Chazal." country="Guadeloupe" county="Pointe des Chateaux" location="Grande-Terre" municipality="Saint Francois" pageId="4" pageNumber="3" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B94FEADA40322670C886F999FEAFF989" bold="true" box="[192,291,1538,1562]" italics="true" pageId="4" pageNumber="3">
<typeStatus id="5480886A40322670C886F999FEAFF989" box="[192,291,1538,1562]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
</emphasis>
: adult
<specimenCount id="9D3DFD4140322670C9CDF998FE13F989" box="[395,415,1539,1562]" pageId="4" pageNumber="3" type="male"></specimenCount>
(
<collectionCode id="ED2AAE0D40322670C9FAF998FE7BF989" box="[444,503,1539,1562]" pageId="4" pageNumber="3">RTO</collectionCode>
: Sco.0409):
<emphasis id="B94FEADA40322670CADFF998FC97F989" box="[665,795,1539,1562]" italics="true" pageId="4" pageNumber="3">
<collectingCountry id="F32C765840322670CADFF998FC97F989" box="[665,795,1539,1562]" name="Guadeloupe" pageId="4" pageNumber="3">Guadeloupe</collectingCountry>
</emphasis>
,
<location id="8EE4601340322670C886F9B9FED8F9AA" LSID="urn:lsid:plazi:treatment:039287DE4034267ECB3AFA4FFEB5F8B0:8EE4601340322670C886F9B9FED8F9AA" box="[192,340,1570,1593]" country="Guadeloupe" county="Pointe des Chateaux" municipality="Saint Francois" name="Grande-Terre" pageId="4" pageNumber="3">Grande-Terre</location>
,
<collectingMunicipality id="6BE0ACB240322670C923F9B9FD84F9AA" box="[357,520,1570,1593]" pageId="4" pageNumber="3">Saint François</collectingMunicipality>
,
<collectingCounty id="62E54E4440322670CA5FF9B9FC8FF9AA" box="[537,771,1570,1593]" pageId="4" pageNumber="3">Pointe des Châteaux</collectingCounty>
,
<date id="FF85100840322670CB52F9B9FEA1F9CA" pageId="4" pageNumber="3" value="2009-07-09">
<collectingDate id="EFC1E9E040322670CB52F9B9FEA1F9CA" pageId="4" pageNumber="3" value="2009-07-09">9 July 2009</collectingDate>
</date>
, leg.
<collectorName id="26CE531E40322670C92CF9D9FE50F9CA" box="[362,476,1602,1625]" pageId="4" pageNumber="3">L. Chazal.</collectorName>
</materialsCitation>
<materialsCitation id="3B533C9540322670C9A2F9D9FEADF90B" ID-GBIF-Occurrence="3352114301" collectingDate="2009-07-09" collectionCode="RTO" collectorName="L. Chazal." country="Guadeloupe" county="Pointe des Chateaux" location="Grande-Terre" municipality="Saint Francois" pageId="4" pageNumber="3" specimenCount="5" specimenCount-adult="2" specimenCount-juvenile="1" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B94FEADA40322670C9A2F9D9FDDCF9CA" bold="true" box="[484,592,1602,1625]" italics="true" pageId="4" pageNumber="3">
<typeStatus id="5480886A40322670C9A2F9D9FDDCF9CA" box="[484,592,1602,1625]" pageId="4" pageNumber="3" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D3DFD4140322670CA26F9D9FD27F9CA" box="[608,683,1602,1625]" pageId="4" pageNumber="3" type="adult">2 adult</specimenCount>
<specimenCount id="9D3DFD4140322670CAF2F9D9FD50F9CA" box="[692,732,1602,1625]" pageId="4" pageNumber="3" type="male">♂♂</specimenCount>
and
<specimenCount id="9D3DFD4140322670CB52F9D9FE9AF9EB" pageId="4" pageNumber="3" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="9D3DFD4140322670C96FF9FAFEB1F9EB" box="[297,317,1633,1656]" pageId="4" pageNumber="3" type="male"></specimenCount>
(
<collectionCode id="ED2AAE0D40322670C911F9FAFE1EF9EB" box="[343,402,1633,1656]" pageId="4" pageNumber="3">RTO</collectionCode>
: Sco. 0410), with same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C321654340322670C886F924FC68FB4D" pageId="4" pageNumber="3" type="etymology">
<paragraph id="8B8436C840322670C886F924FC68FB4D" blockId="4.[192,802,1727,1846]" lastBlockId="4.[830,1440,1192,1246]" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670C886F924FEB2F944" bold="true" box="[192,318,1727,1751]" italics="true" pageId="4" pageNumber="3">Etymology:</emphasis>
the selected name is a Latin adjective that means “dwarf” and literally reflects the tiny size of this scorpion, which represents together with
<taxonomicName id="4C3B4D4B40322670CA38F964FD7FF885" authorityName="Teruel" authorityYear="2008" box="[638,755,1791,1814]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="questeli">
<emphasis id="B94FEADA40322670CA38F964FD7FF885" box="[638,755,1791,1814]" italics="true" pageId="4" pageNumber="3">O. questeli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3B4D4B40322670C886F884FDCCF8A5" authority="Armas 1981" authorityName="Armas" authorityYear="1981" box="[192,576,1823,1846]" class="Arachnida" family="Diplocentridae" genus="Heteronebo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B94FEADA40322670C886F884FE2EF8A5" box="[192,418,1823,1846]" italics="true" pageId="4" pageNumber="3">Heteronebo pumilus</emphasis>
Armas 1981
</taxonomicName>
(from southeastern
<collectingCountry id="F32C765840322670CB78FB33FCFBFB2C" box="[830,887,1192,1215]" name="Haiti" pageId="4" pageNumber="3">Haiti</collectingCountry>
), the smallest adult size known so far amongst all Diplocentrinae.
</paragraph>
</subSubSection>
<subSubSection id="C321654340322670CB78FA9EFA0AFA28" pageId="4" pageNumber="3" type="distribution">
<paragraph id="8B8436C840322670CB78FA9EFA0AFA28" blockId="4.[830,1441,1285,1467]" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670CB78FA9EFC48FA8E" bold="true" box="[830,964,1285,1309]" italics="true" pageId="4" pageNumber="3">Distribution</emphasis>
(
<figureCitation id="13002A4D40322670CB99FA9DFBA2FA8E" box="[991,1070,1286,1309]" captionStart="Figure 5" captionStartId="9.[192,268,1159,1182]" captionTargetBox="[191,1442,191,1130]" captionTargetId="figure-861@9.[192,1441,192,1129]" captionTargetPageId="9" captionText="Figure 5: Known distribution of the genus Oiclus: O. nanus sp. n. (white square with black cross, including the potential records from Bouillantes and Les Saintes), Oiclus questeli (black square with white cross), Oiclus purvesii purvesii (black square), Oiclus purvesii sabae (white square), and Oiclus sp. (white square with black dot)." figureDoi="http://doi.org/10.5281/zenodo.4649079" httpUri="https://zenodo.org/record/4649079/files/figure.png" pageId="4" pageNumber="3">Fig. 5</figureCitation>
): this scorpion appears to be endemic from the
<collectingCountry id="F32C765840322670CC55FABDFB18FAAE" box="[1043,1172,1318,1341]" name="Guadeloupe" pageId="4" pageNumber="3">Guadeloupe</collectingCountry>
insular bank, as it has been collected from a single locality in the eastern tip of Grande-Terre, but it seems to occur also in western Basse-Terre (Bouillante) and Les Saintes (Terre-de- Haut); see more details below, in the Remarks section.
</paragraph>
</subSubSection>
<subSubSection id="C32165434032267DCB78FA78FC00FA05" lastPageId="9" lastPageNumber="8" pageId="4" pageNumber="3" type="description">
<paragraph id="8B8436C840322673CB78FA78FB11FE40" blockId="4.[830,1441,1507,1846]" lastBlockId="7.[830,1441,199,467]" lastPageId="7" lastPageNumber="6" pageId="4" pageNumber="3">
<emphasis id="B94FEADA40322670CB78FA78FC32FA68" bold="true" box="[830,958,1507,1531]" italics="true" pageId="4" pageNumber="3">Description</emphasis>
(adult male
<typeStatus id="5480886A40322670CC16FA78FB3EFA69" box="[1104,1202,1507,1530]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
).
<emphasis id="B94FEADA40322670CC8EFA78FACFFA69" bold="true" box="[1224,1347,1507,1530]" pageId="4" pageNumber="3">Coloration</emphasis>
(
<figureCitation id="13002A4D40322670CD10FA78FA15FA69" box="[1366,1433,1507,1530]" captionStart="Figure 1" captionStartId="3.[192,268,1894,1917]" captionTargetBox="[191,1458,180,1873]" captionTargetId="figure-6@3.[192,1457,181,1872]" captionTargetPageId="3" captionText="Figure 1: Adult male holotype of Oiclus nanus sp. n.: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649071" httpUri="https://zenodo.org/record/4649071/files/figure.png" pageId="4" pageNumber="3">Fig. 1</figureCitation>
,
<figureCitation id="13002A4D40322670CB78F998FCC1F989" box="[830,845,1539,1562]" captionStart="Figure 2" captionStartId="5.[192,268,1821,1844]" captionTargetBox="[192,1439,167,1800]" captionTargetId="figure-5@5.[192,1439,167,1800]" captionTargetPageId="5" captionText="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="4" pageNumber="3">2</figureCitation>
,
<figureCitation id="13002A4D40322670CB19F998FCF4F989" box="[863,888,1539,1562]" captionStart="Figure 3" captionStartId="6.[192,268,1468,1491]" captionTargetBox="[192,1439,192,1439]" captionTargetId="figure-50@6.[192,1439,192,1439]" captionTargetPageId="6" captionText="Figure 3: Type specimens of Oiclus nanus sp. n., still alive when photographed: a) adult male holotype, in its natural habitat; bc) adult male paratypes, in their natural habitat; d) juvenile male paratype, in captivity." figureDoi="http://doi.org/10.5281/zenodo.4649075" httpUri="https://zenodo.org/record/4649075/files/figure.png" pageId="4" pageNumber="3">3a</figureCitation>
) basically light brown, with a dense but diffuse pattern of dark brown reticulate spots over carapace and tergites; pedipalps and metasoma with all carinae and fingers faintly infuscate; carapace with anterior margin, ocular tubercle and eyes blackish; mesosoma venter and pectines yellowish immaculate; legs yellowish, immaculate to very faintly spotted.
<emphasis id="B94FEADA40322670CCEDF95BFA95F944" bold="true" box="[1195,1305,1728,1751]" pageId="4" pageNumber="3">Carapace</emphasis>
(
<figureCitation id="13002A4D40322670CD6CF95BFAF4F944" box="[1322,1400,1728,1751]" captionStart="Figure 2" captionStartId="5.[192,268,1821,1844]" captionTargetBox="[192,1439,167,1800]" captionTargetId="figure-5@5.[192,1439,167,1800]" captionTargetPageId="5" captionText="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="4" pageNumber="3">Fig. 2a</figureCitation>
) as long as wide, anterior margin with 23 pairs of macrosetae alternated with abundant shorter setae of different size, frontal lobes very wide and rounded, frontal notch very wide and shallow. Tegument basically smooth and polished, with circum-ocular, mediolateral and posterolateral areas very finely and densely granulose. All furrows obsolete except for the lateral ocular, posterior median, posterior lateral and posterior marginal, which are relatively narrow and deep. Median eyes relatively small but larger than the lateral eyes, and separated by clearly less than one ocular diameter, median tubercle subtly raised; two pairs of lateral eyes.
<emphasis id="B94FEADA40302672C886F8AEFE91F8DF" bold="true" box="[192,285,1845,1868]" pageId="6" pageNumber="5">Tergites</emphasis>
(
<figureCitation id="13002A4D40302672C968F8AEFEF1F8DF" box="[302,381,1845,1868]" captionStart="Figure 2" captionStartId="5.[192,268,1821,1844]" captionTargetBox="[192,1439,167,1800]" captionTargetId="figure-5@5.[192,1439,167,1800]" captionTargetPageId="5" captionText="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="6" pageNumber="5">Fig. 2b</figureCitation>
) with median carina vestigial to weak on IVI, flanked on each side by narrowly depressed furrows; tegument very finely and densely granulose, with irregular and polished lateral areas increasing in size from IVI. Tergite VII with moderately bilobed lateroposterior region and with two pairs of lateral carinae which are all similar and composed by several large, rounded granules.
<emphasis id="B94FEADA40302672CC2CF921FB68F942" bold="true" box="[1130,1252,1722,1745]" pageId="6" pageNumber="5">Chelicerae</emphasis>
(
<figureCitation id="13002A4D40302672CCB8F921FADAF942" box="[1278,1366,1722,1745]" captionStart="Figure 2" captionStartId="5.[192,268,1821,1844]" captionTargetBox="[192,1439,167,1800]" captionTargetId="figure-5@5.[192,1439,167,1800]" captionTargetPageId="5" captionText="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="6" pageNumber="5">Fig. 2a</figureCitation>
) with dentition typical for the family, tegument smooth and polished.
<emphasis id="B94FEADA40302672CBEBF963FB90F89C" bold="true" box="[941,1052,1784,1807]" pageId="6" pageNumber="5">Pedipalps</emphasis>
(
<figureCitation id="13002A4D40302672CC74F963FB26F89C" box="[1074,1194,1784,1807]" captionStart="Figure 2" captionStartId="5.[192,268,1821,1844]" captionTargetBox="[192,1439,167,1800]" captionTargetId="figure-5@5.[192,1439,167,1800]" captionTargetPageId="5" captionText="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="6" pageNumber="5">Figs. 2cd</figureCitation>
) orthobothriotaxic C. Femur deeper than wide, with dorsal surface markedly convex; dorsointernal and ventrointernal carinae very poorly defined, irregularly granulose, ventroexternal carina absent; tegument smooth and polished, dorsal surface with some coarse granules scattered. Patella with all carinae obsolete except for the dorsointernal (moderate, costate to subgranulose) and the dorsoexternal (weak, costate to subgranulose); tegument smooth and polished except on the internal surface, which is very finely and densely granulose; dorsal and external surfaces sparsely punctate. Chela very short and robust, moderately depressed laterally in cross-section and much deeper than wide; hand with all carinae obsolete to absent, digital carina vestigial, subgranulose, ventrointernal carinae absent, ventroexternal carinae strong, costate to subgranulose and directed essentially towards its articulation condyle, dorsal marginal and dorsointernal carinae strong, irregularly granulose; tegument densely punctate and covered with granulose reticulations which cross even over some of the carinae, dorsointernal surface densely granulose. Fingers very short, densely punctate, acarinate and densely setose, without lobe/notch combination; opposable edges with irregular granulation not arranged in rows.
<emphasis id="B94FEADA40312673CAEBFCB7FD6EFCD0" bold="true" box="[685,738,812,835]" pageId="7" pageNumber="6">Legs</emphasis>
(
<figureCitation id="13002A4D40312673CAB0FCB7FF78FCF1" captionStart="Figure 1" captionStartId="3.[192,268,1894,1917]" captionTargetBox="[191,1458,180,1873]" captionTargetId="figure-6@3.[192,1457,181,1872]" captionTargetPageId="3" captionText="Figure 1: Adult male holotype of Oiclus nanus sp. n.: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649071" httpUri="https://zenodo.org/record/4649071/files/figure.png" pageId="7" pageNumber="6">Fig. 1ab</figureCitation>
) with tegument smooth and polished; pedal spurs absent; tarsomere II without laterodistal lobes; tarsal spine formula 3/3: 4/4: 5/5: 5/5.
<emphasis id="B94FEADA40312673CA12FC13FD3BFC0C" bold="true" box="[596,695,904,927]" pageId="7" pageNumber="6">Sternum</emphasis>
(
<figureCitation id="13002A4D40312673CA8FFC13FC96FC0C" box="[713,794,904,927]" captionStart="Figure 2" captionStartId="5.[192,268,1821,1844]" captionTargetBox="[192,1439,167,1800]" captionTargetId="figure-5@5.[192,1439,167,1800]" captionTargetPageId="5" captionText="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="7" pageNumber="6">Fig. 2e</figureCitation>
) type 2, strongly pentagonal, with parallel sides.
<emphasis id="B94FEADA40312673CA89FC3CFE91FC4E" bold="true" pageId="7" pageNumber="6">Genital operculi</emphasis>
(
<figureCitation id="13002A4D40312673C970FC5DFE07FC4E" box="[310,395,966,989]" captionStart="Figure 2" captionStartId="5.[192,268,1821,1844]" captionTargetBox="[192,1439,167,1800]" captionTargetId="figure-5@5.[192,1439,167,1800]" captionTargetPageId="5" captionText="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="7" pageNumber="6">Fig. 2e</figureCitation>
) ellipsoidal; genital papillae moderately developed and subtly exposed.
<emphasis id="B94FEADA40312673CA27FC7FFD32FC68" bold="true" box="[609,702,996,1019]" pageId="7" pageNumber="6">Pectines</emphasis>
(
<figureCitation id="13002A4D40312673CA8AFC7FFC95FC68" box="[716,793,996,1019]" captionStart="Figure 2" captionStartId="5.[192,268,1821,1844]" captionTargetBox="[192,1439,167,1800]" captionTargetId="figure-5@5.[192,1439,167,1800]" captionTargetPageId="5" captionText="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="7" pageNumber="6">Fig. 2e</figureCitation>
) hirsute, with 7/7 teeth; fulcra well developed basally but becoming reduced distally in each pecten; basal plate much wider than long; anterior margin weakly notched, posterior margin straight.
<emphasis id="B94FEADA40312673C9AFFBC4FDC2FBE5" bold="true" box="[489,590,1119,1142]" pageId="7" pageNumber="6">Sternites</emphasis>
(
<figureCitation id="13002A4D40312673CA24FBC4FD3AFBE5" box="[610,694,1119,1142]" captionStart="Figure 1" captionStartId="3.[192,268,1894,1917]" captionTargetBox="[191,1458,180,1873]" captionTargetId="figure-6@3.[192,1457,181,1872]" captionTargetPageId="3" captionText="Figure 1: Adult male holotype of Oiclus nanus sp. n.: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649071" httpUri="https://zenodo.org/record/4649071/files/figure.png" pageId="7" pageNumber="6">Fig. 1b</figureCitation>
) smooth, polished and moderately hirsute, especially on lateral and posterior margins; VII with lateral carinae moderately granulose and ventrosubmedian carinae weakly granulose; spiracles narrow, oval-elongate.
<emphasis id="B94FEADA40312673CAEDFB41FCAEFB62" bold="true" box="[683,802,1242,1265]" pageId="7" pageNumber="6">Metasoma</emphasis>
(
<figureCitation id="13002A4D40312673C881FB63FEB4FA9C" box="[199,312,1272,1295]" captionStart="Figure 1" captionStartId="3.[192,268,1894,1917]" captionTargetBox="[191,1458,180,1873]" captionTargetId="figure-6@3.[192,1457,181,1872]" captionTargetPageId="3" captionText="Figure 1: Adult male holotype of Oiclus nanus sp. n.: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649071" httpUri="https://zenodo.org/record/4649071/files/figure.png" pageId="7" pageNumber="6">Figs. 1ab</figureCitation>
,
<figureCitation id="13002A4D40312673C900FB63FEFAFA9C" box="[326,374,1272,1295]" captionStart="Figure 2" captionStartId="5.[192,268,1821,1844]" captionTargetBox="[192,1439,167,1800]" captionTargetId="figure-5@5.[192,1439,167,1800]" captionTargetPageId="5" captionText="Figure 2: Adult male holotype of Oiclus nanus sp. n.: a) carapace; b) tergites; c) pedipalp, dorsal view; d) pedipalp, ventral view; e) sternopectinal region; f) metasomal segments IIII, dorsal view; g) metasomal segments IIII, lateral view; h) metasomal segments IIII, ventral view; i) metasomal segments IVV and telson, dorsal view; j) metasomal segments IVV and telson, lateral view; k) metasomal segment V and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="7" pageNumber="6">2fk</figureCitation>
) with segment I wider than long and II- V longer than wide; intercarinal tegument smooth and polished, with only a few small and medium-sized granules scattered on dorsal and lateral surfaces, especially on V; segments IIII, with ten carinae, IV with eight, V with five, all densely covered by rigid macrosetae; dorsolateral carinae moderate and irregularly granulose on IIV, absent on V; lateral supramedian carinae moderately granulose on all segments; lateral inframedian carinae very weak and subcostate on I, vestigial and smooth on IIIII, absent on IVV; ventrolateral carinae moderately crenulate to serratocrenulate on III, weakly crenulate on III, very weakly subcrenulate on IV, composed by irregularly arranged conical granules on basal two-thirds of V but replaced on distal third by the ventral transverse carina; ventral submedian carinae strongly crenulate to serratocrenulate on III, moderately crenulate on III, very weakly subcostate on IV, absent in V; ventromedian carinae on V strong and composed by irregularly arranged conical granules; ventral transverse carina strong, dentate and evenly arched; segment V slightly longer than telson, with anal arc denticulate, laterodistal lobes bluntly triangular and not projected. Telson ovaldepressed and moderately slender; vesicle polished, irregularly granulose and densely covered by rigid macrosetae, ventrobasal area coarsely granulose, subaculear tubercle large, laterally compressed and covered by many rigid setae and a few coarse granules; aculeus short, sharp and strongly curved.
</paragraph>
<caption id="DF44664040332671C886F886FCEAF8EC" ID-DOI="http://doi.org/10.5281/zenodo.4649073" ID-Zenodo-Dep="4649073" httpUri="https://zenodo.org/record/4649073/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,268,1821,1844]" targetBox="[192,1439,167,1800]" targetPageId="5">
<paragraph id="8B8436C840332671C886F886FCEAF8EC" blockId="5.[192,1441,1821,1919]" pageId="5" pageNumber="4">
<emphasis id="B94FEADA40332671C886F886FEA7F8A7" bold="true" box="[192,299,1821,1844]" pageId="5" pageNumber="4">Figure 2:</emphasis>
Adult male holotype of
<taxonomicName id="4C3B4D4B40332671CA67F884FD2DF8A7" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[545,673,1823,1844]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA40332671CA67F884FD2DF8A7" box="[545,673,1823,1844]" italics="true" pageId="5" pageNumber="4">Oiclus nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA40332671CAEFF884FD6EF8A7" bold="true" box="[681,738,1823,1844]" pageId="5" pageNumber="4">
<taxonomicNameLabel id="A27C57A140332671CAEFF884FD6EF8A7" box="[681,738,1823,1844]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="B94FEADA40332671CAB7F884FC89F8A7" bold="true" box="[753,773,1823,1844]" pageId="5" pageNumber="4">a)</emphasis>
carapace;
<emphasis id="B94FEADA40332671CB36F884FC09F8A7" bold="true" box="[880,901,1823,1844]" pageId="5" pageNumber="4">b)</emphasis>
tergites;
<emphasis id="B94FEADA40332671CBA5F884FC79F8A7" bold="true" box="[995,1013,1823,1844]" pageId="5" pageNumber="4">c)</emphasis>
pedipalp, dorsal view;
<emphasis id="B94FEADA40332671CC99F884FB78F8A7" bold="true" box="[1247,1268,1823,1844]" pageId="5" pageNumber="4">d)</emphasis>
pedipalp, ventral view;
<emphasis id="B94FEADA40332671C944F8A3FE99F8DE" bold="true" box="[258,277,1848,1869]" pageId="5" pageNumber="4">e)</emphasis>
sternopectinal region;
<emphasis id="B94FEADA40332671CA40F8A3FD9AF8DE" bold="true" box="[518,534,1848,1869]" pageId="5" pageNumber="4">f)</emphasis>
metasomal segments IIII, dorsal view;
<emphasis id="B94FEADA40332671CB83F8A3FC55F8DE" bold="true" box="[965,985,1848,1869]" pageId="5" pageNumber="4">g)</emphasis>
metasomal segments IIII, lateral view;
<emphasis id="B94FEADA40332671CDCCF8A3FA2CF8DE" bold="true" box="[1418,1440,1848,1869]" pageId="5" pageNumber="4">h)</emphasis>
metasomal segments IIII, ventral view;
<emphasis id="B94FEADA40332671CA0BF8CAFDD0F8F5" bold="true" box="[589,604,1873,1894]" pageId="5" pageNumber="4">i)</emphasis>
metasomal segments IVV and telson, dorsal view;
<emphasis id="B94FEADA40332671CC1AF8CAFBE7F8F5" bold="true" box="[1116,1131,1873,1894]" pageId="5" pageNumber="4">j)</emphasis>
metasomal segments IVV and telson, lateral view;
<emphasis id="B94FEADA40332671C9C5F8F1FE14F8EC" bold="true" box="[387,408,1898,1919]" pageId="5" pageNumber="4">k)</emphasis>
metasomal segment V and telson, ventral view.
</paragraph>
</caption>
<caption id="DF44664040302672C886FA27FB8DFA7F" ID-DOI="http://doi.org/10.5281/zenodo.4649075" ID-Zenodo-Dep="4649075" httpUri="https://zenodo.org/record/4649075/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,268,1468,1491]" targetBox="[192,1439,192,1439]" targetPageId="6">
<paragraph id="8B8436C840302672C886FA27FB8DFA7F" blockId="6.[192,1440,1468,1516]" pageId="6" pageNumber="5">
<emphasis id="B94FEADA40302672C886FA27FEA4FA40" bold="true" box="[192,296,1468,1491]" pageId="6" pageNumber="5">Figure 3:</emphasis>
Type specimens of
<taxonomicName id="4C3B4D4B40302672C9ACFA25FDE4FA40" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[490,616,1470,1491]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA40302672C9ACFA25FDE4FA40" box="[490,616,1470,1491]" italics="true" pageId="6" pageNumber="5">Oiclus nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA40302672CA28FA25FD28FA40" bold="true" box="[622,676,1470,1491]" pageId="6" pageNumber="5">
<taxonomicNameLabel id="A27C57A140302672CA28FA25FD28FA40" box="[622,676,1470,1491]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, still alive when photographed:
<emphasis id="B94FEADA40302672CB9CFA25FC62FA40" bold="true" box="[986,1006,1470,1491]" pageId="6" pageNumber="5">a)</emphasis>
adult male holotype, in its natural habitat;
<emphasis id="B94FEADA40302672CDCDFA25FF5EFA7F" bold="true" pageId="6" pageNumber="5">bc)</emphasis>
adult male paratypes, in their natural habitat;
<emphasis id="B94FEADA40302672CACBFA4CFD2EFA7F" bold="true" box="[653,674,1495,1516]" pageId="6" pageNumber="5">d)</emphasis>
juvenile male paratype, in captivity.
</paragraph>
</caption>
<paragraph id="8B8436C840312673CB28FD9FFAF4FC85" blockId="7.[830,1441,516,1563]" pageId="7" pageNumber="6">
<emphasis id="B94FEADA40312673CB28FD9FFC6DFD8F" bold="true" box="[878,993,516,540]" italics="true" pageId="7" pageNumber="6">Variation:</emphasis>
one adult male
<typeStatus id="5480886A40312673CCE8FD9FFA86FD88" box="[1198,1290,516,539]" pageId="7" pageNumber="6" type="paratype">paratype</typeStatus>
(
<figureCitation id="13002A4D40312673CD67FD9FFAFAFD88" box="[1313,1398,516,539]" captionStart="Figure 3" captionStartId="6.[192,268,1468,1491]" captionTargetBox="[192,1439,192,1439]" captionTargetId="figure-50@6.[192,1439,192,1439]" captionTargetPageId="6" captionText="Figure 3: Type specimens of Oiclus nanus sp. n., still alive when photographed: a) adult male holotype, in its natural habitat; bc) adult male paratypes, in their natural habitat; d) juvenile male paratype, in captivity." figureDoi="http://doi.org/10.5281/zenodo.4649075" httpUri="https://zenodo.org/record/4649075/files/figure.png" pageId="7" pageNumber="6">Fig. 3b</figureCitation>
) is identical to the
<typeStatus id="5480886A40312673CBBFFDBFFBDBFDA8" box="[1017,1111,548,571]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
in size, coloration, morphometrics, sculpture of the tegument, pectinal tooth count and tarsal spine formula, but the other adult male
<typeStatus id="5480886A40312673CB78FD19FC16FD0A" box="[830,922,642,665]" pageId="7" pageNumber="6" type="paratype">paratype</typeStatus>
(
<figureCitation id="13002A4D40312673CBF5FD19FB80FD0A" box="[947,1036,642,665]" captionStart="Figure 3" captionStartId="6.[192,268,1468,1491]" captionTargetBox="[192,1439,192,1439]" captionTargetId="figure-50@6.[192,1439,192,1439]" captionTargetPageId="6" captionText="Figure 3: Type specimens of Oiclus nanus sp. n., still alive when photographed: a) adult male holotype, in its natural habitat; bc) adult male paratypes, in their natural habitat; d) juvenile male paratype, in captivity." figureDoi="http://doi.org/10.5281/zenodo.4649075" httpUri="https://zenodo.org/record/4649075/files/figure.png" pageId="7" pageNumber="6">Fig. 3c</figureCitation>
;
<tableCitation id="C6B9037340312673CC64FD19FBFCFD0A" box="[1058,1136,642,665]" captionStart="Table 1" captionStartId="4.[192,257,1091,1114]" captionTargetPageId="4" captionText="Table 1: Measurements of the three adult types of Oiclus nanus, sp. n. Abbreviations: length (L), width (W), posterior width (Wp), depth (H)." httpUri="http://table.plazi.org/id/DF44664040322670C886FBD8FEEEFBE0" pageId="7" pageNumber="6" tableUuid="DF44664040322670C886FBD8FEEEFBE0">Tab. 1</tableCitation>
) shows some subtle differences: size slightly larger (supposed to represent another size class), coloration somewhat lighter and less conspicuously spotted, prolateral row of leg I with 34 spines, and a few trivial morphometric discrepancies.
</paragraph>
<paragraph id="8B8436C840312673CB28FC85FC69FBBC" blockId="7.[830,1441,516,1563]" pageId="7" pageNumber="6">
A very interesting and strong variation was observed in the development of the pectinal fulcra, which is an unusual trend among scorpions in general: all type specimens show individual variations of this structure both within each pecten and between both pectines. The largest male has fulcra weakly developed to absent, but the other two males (including the
<typeStatus id="5480886A40312673CB78FC62FC11FB83" box="[830,925,1017,1040]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
), and the juvenile have well developed to vestigial fulcra.
</paragraph>
<paragraph id="8B8436C840312673CB28FBA3FBF5F988" blockId="7.[830,1441,516,1563]" pageId="7" pageNumber="6">
The single juvenile specimen available (
<figureCitation id="13002A4D40312673CD04FBA3FA14FBDC" box="[1346,1432,1080,1103]" captionStart="Figure 3" captionStartId="6.[192,268,1468,1491]" captionTargetBox="[192,1439,192,1439]" captionTargetId="figure-50@6.[192,1439,192,1439]" captionTargetPageId="6" captionText="Figure 3: Type specimens of Oiclus nanus sp. n., still alive when photographed: a) adult male holotype, in its natural habitat; bc) adult male paratypes, in their natural habitat; d) juvenile male paratype, in captivity." figureDoi="http://doi.org/10.5281/zenodo.4649075" httpUri="https://zenodo.org/record/4649075/files/figure.png" pageId="7" pageNumber="6">Fig. 3d</figureCitation>
) differs from the adults by the same basic features as in all other diplocentrine scorpions:
<emphasis id="B94FEADA40312673CCF7FBEEFB4BFB1F" bold="true" box="[1201,1223,1141,1164]" pageId="7" pageNumber="6">1)</emphasis>
coloration with the base conspicuously paler, and the spotted pattern darker and more contrasting;
<emphasis id="B94FEADA40312673CC71FB29FBC2FB5A" bold="true" box="[1079,1102,1202,1225]" pageId="7" pageNumber="6">2)</emphasis>
body overall more slender;
<emphasis id="B94FEADA40312673CDCCFB29FA2CFB5A" bold="true" box="[1418,1440,1202,1225]" pageId="7" pageNumber="6">3)</emphasis>
pedipalps and metasoma with weaker carinae;
<emphasis id="B94FEADA40312673CDCCFB4AFA2CFB7B" bold="true" box="[1418,1440,1233,1256]" pageId="7" pageNumber="6">4)</emphasis>
carapace and tergites smooth and polished;
<emphasis id="B94FEADA40312673CD5BFB6BFABFFA94" bold="true" box="[1309,1331,1264,1287]" pageId="7" pageNumber="6">5)</emphasis>
pedipalps not punctate. The last feature is very important from a taxonomic point of view, as this is the first time that such variation is observed in Diplocentrinae: in all other species of this subfamily with punctate tegument, this character is invariably present in all juvenile and adult instars (R. Teruel, personal observation); this fact must be taken into account when identifying
<taxonomicName id="4C3B4D4B40312673CD1CFA5DFA2CFA4E" authorityName="Simon" authorityYear="1880" box="[1370,1440,1478,1501]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA40312673CD1CFA5DFA2CFA4E" box="[1370,1440,1478,1501]" italics="true" pageId="7" pageNumber="6">Oiclus</emphasis>
</taxonomicName>
populations from small samples to avoid taxonomic errors and misinterpretations.
</paragraph>
<paragraph id="8B8436C840312673CB78F9D2FCF7F8E6" blockId="7.[830,1440,1609,1909]" pageId="7" pageNumber="6">
<emphasis id="B94FEADA40312673CB78F9D2FC72F9F2" bold="true" box="[830,1022,1609,1633]" italics="true" pageId="7" pageNumber="6">Ecological notes:</emphasis>
all specimens of
<taxonomicName id="4C3B4D4B40312673CCF9F9D1FAACF9F2" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[1215,1312,1610,1633]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA40312673CCF9F9D1FAACF9F2" box="[1215,1312,1610,1633]" italics="true" pageId="7" pageNumber="6">O. nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA40312673CD61F9D1FAE8F9F2" bold="true" box="[1319,1380,1610,1633]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A27C57A140312673CD61F9D1FAE8F9F2" box="[1319,1380,1610,1633]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
were collected under small rocks in dry coastal forest, at an altitude of
<quantity id="4CC39B2D40312673CBF2F913FC60F90C" box="[948,1004,1672,1695]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="6" unit="m" value="12.0">12 m</quantity>
a.s.l.; every scorpion burrows a shallow horizontal gallery which follows the lower contour of the rock, and has an enlarged bottom where each individual rests and retreats for shelter (
<figureCitation id="13002A4D40312673CD4FF97FFAF8F968" box="[1289,1396,1764,1787]" captionStart="Figure 4" captionStartId="8.[192,268,1875,1898]" captionTargetBox="[191,1440,167,1846]" captionTargetId="figure-39@8.[192,1439,168,1845]" captionTargetPageId="8" captionText="Figure 4: Two views of localities where Oiclus nanus sp. n. occurs at Pointe des Châteaux: a) habitat; b) microhabitat." figureDoi="http://doi.org/10.5281/zenodo.4649077" httpUri="https://zenodo.org/record/4649077/files/figure.png" pageId="7" pageNumber="6">Fig. 4ab</figureCitation>
). It occurs both sympatrically and syntopically with two buthid scorpions:
<taxonomicName id="4C3B4D4B40312673CC54F8BAFCF1F8C4" authority="(Pocock, 1898)" baseAuthorityName="Pocock" baseAuthorityYear="1898" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="barbudensis">
<emphasis id="B94FEADA40312673CC54F8BAFABDF8AB" box="[1042,1329,1825,1848]" italics="true" pageId="7" pageNumber="6">Centruroides barbudensis</emphasis>
(Pocock, 1898)
</taxonomicName>
and
<taxonomicName id="4C3B4D4B40312673CB85F8DBFCFFF8E6" authority="Sissom et Francke, 1983" authorityName="Sissom et Francke" authorityYear="1983" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="pococki">
<emphasis id="B94FEADA40312673CB85F8DBFB3FF8C4" box="[963,1203,1856,1879]" italics="true" pageId="7" pageNumber="6">Centruroides pococki</emphasis>
Sissom et Francke, 1983
</taxonomicName>
.
</paragraph>
<caption id="DF446640403E267CC886F8C8FACAF8F9" ID-DOI="http://doi.org/10.5281/zenodo.4649077" ID-Zenodo-Dep="4649077" box="[192,1350,1875,1898]" httpUri="https://zenodo.org/record/4649077/files/figure.png" pageId="8" pageNumber="7" startId="8.[192,268,1875,1898]" targetBox="[191,1440,167,1846]" targetPageId="8">
<paragraph id="8B8436C8403E267CC886F8C8FACAF8F9" blockId="8.[192,1350,1875,1898]" box="[192,1350,1875,1898]" pageId="8" pageNumber="7">
<emphasis id="B94FEADA403E267CC886F8C8FEA5F8F9" bold="true" box="[192,297,1875,1898]" pageId="8" pageNumber="7">Figure 4:</emphasis>
Two views of localities where
<taxonomicName id="4C3B4D4B403E267CCA10F8CEFD5FF8F9" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[598,723,1877,1898]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA403E267CCA10F8CEFD5FF8F9" box="[598,723,1877,1898]" italics="true" pageId="8" pageNumber="7">Oiclus nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA403E267CCA9FF8CEFC9CF8F9" bold="true" box="[729,784,1877,1898]" pageId="8" pageNumber="7">
<taxonomicNameLabel id="A27C57A1403E267CCA9FF8CEFC9CF8F9" box="[729,784,1877,1898]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
occurs at Pointe des Châteaux:
<emphasis id="B94FEADA403E267CCC04F8CEFBDAF8F9" bold="true" box="[1090,1110,1877,1898]" pageId="8" pageNumber="7">a)</emphasis>
habitat;
<emphasis id="B94FEADA403E267CCCECF8CEFB33F8F9" bold="true" box="[1194,1215,1877,1898]" pageId="8" pageNumber="7">b)</emphasis>
microhabitat.
</paragraph>
</caption>
<caption id="DF446640403F267DC886FB1CFBA1FB43" ID-DOI="http://doi.org/10.5281/zenodo.4649079" ID-Zenodo-Dep="4649079" httpUri="https://zenodo.org/record/4649079/files/figure.png" pageId="9" pageNumber="8" startId="9.[192,268,1159,1182]" targetBox="[191,1442,191,1130]" targetPageId="9">
<paragraph id="8B8436C8403F267DC886FB1CFBA1FB43" blockId="9.[192,1440,1159,1232]" pageId="9" pageNumber="8">
<emphasis id="B94FEADA403F267DC886FB1CFEA0FB0D" bold="true" box="[192,300,1159,1182]" pageId="9" pageNumber="8">Figure 5:</emphasis>
Known distribution of the genus
<taxonomicName id="4C3B4D4B403F267DCAC0FB12FD49FB0D" authorityName="Simon" authorityYear="1880" box="[646,709,1161,1182]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403F267DCAC0FB12FD49FB0D" box="[646,709,1161,1182]" italics="true" pageId="9" pageNumber="8">Oiclus</emphasis>
</taxonomicName>
:
<taxonomicName id="4C3B4D4B403F267DCA93FB12FCA3FB0D" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[725,815,1161,1182]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA403F267DCA93FB12FCA3FB0D" box="[725,815,1161,1182]" italics="true" pageId="9" pageNumber="8">O. nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA403F267DCB7FFB12FCF8FB0D" bold="true" box="[825,884,1161,1182]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="A27C57A1403F267DCB7FFB12FCF8FB0D" box="[825,884,1161,1182]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(white square with black cross, including the potential records from Bouillantes and Les Saintes),
<taxonomicName id="4C3B4D4B403F267DCA3AFB39FC9CFB24" authorityName="Teruel" authorityYear="2008" box="[636,784,1186,1207]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="questeli">
<emphasis id="B94FEADA403F267DCA3AFB39FC9CFB24" box="[636,784,1186,1207]" italics="true" pageId="9" pageNumber="8">Oiclus questeli</emphasis>
</taxonomicName>
(black square with white cross),
<taxonomicName id="4C3B4D4B403F267DCC2CFB39FAD5FB24" authority="purvesii" authorityName="purvesii: Teruel &amp; Francke" authorityYear="2006" box="[1130,1369,1186,1207]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="purvesii" subSpecies="purvesii">
<emphasis id="B94FEADA403F267DCC2CFB39FAD5FB24" box="[1130,1369,1186,1207]" italics="true" pageId="9" pageNumber="8">Oiclus purvesii purvesii</emphasis>
</taxonomicName>
(black square),
<taxonomicName id="4C3B4D4B403F267DC955FB20FE6FFB43" authorityName="Francke" authorityYear="1978" box="[275,483,1211,1232]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="purvesii" subSpecies="sabae">
<emphasis id="B94FEADA403F267DC955FB20FE6FFB43" box="[275,483,1211,1232]" italics="true" pageId="9" pageNumber="8">Oiclus purvesii sabae</emphasis>
</taxonomicName>
(white square), and
<taxonomicName id="4C3B4D4B403F267DCAE1FB20FD6AFB43" authorityName="Simon" authorityYear="1880" box="[679,742,1211,1232]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403F267DCAE1FB20FD6AFB43" box="[679,742,1211,1232]" italics="true" pageId="9" pageNumber="8">Oiclus</emphasis>
</taxonomicName>
sp. (white square with black dot).
</paragraph>
</caption>
<paragraph id="8B8436C8403F267DC886FA9FFE45F939" blockId="9.[192,804,1284,1890]" pageId="9" pageNumber="8">
<emphasis id="B94FEADA403F267DC886FA9FFEDDFA8F" bold="true" box="[192,337,1284,1308]" italics="true" pageId="9" pageNumber="8">Comparisons</emphasis>
(adult males only):
<taxonomicName id="4C3B4D4B403F267DCA6CFA9EFD06FA8F" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[554,650,1285,1308]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA403F267DCA6CFA9EFD06FA8F" box="[554,650,1285,1308]" italics="true" pageId="9" pageNumber="8">O. nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA403F267DCAD7FA9EFD42FA8F" bold="true" box="[657,718,1285,1308]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="A27C57A1403F267DCAD7FA9EFD42FA8F" box="[657,718,1285,1308]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is most closely related to
<taxonomicName id="4C3B4D4B403F267DC938FAB8FE79FAA9" baseAuthorityName="Becker" baseAuthorityYear="1880" box="[382,501,1315,1338]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" sensu="stricto" species="purvesii">
<emphasis id="B94FEADA403F267DC938FAB8FE79FAA9" box="[382,501,1315,1338]" italics="true" pageId="9" pageNumber="8">O. purvesii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57A1403F267DC9BAFAB8FDAAFAA9" box="[508,550,1315,1338]" pageId="9" pageNumber="8" sensu="stricto">s. s.</taxonomicNameLabel>
in overall morphology, but the latter can be unequivocally distinguished by having:
<emphasis id="B94FEADA403F267DC95FFAFAFEA3FAEB" bold="true" box="[281,303,1377,1400]" pageId="9" pageNumber="8">1)</emphasis>
sternites and metasomal segment V punctate;
<emphasis id="B94FEADA403F267DC884FAE4FF54FA05" bold="true" box="[194,216,1407,1430]" pageId="9" pageNumber="8">2)</emphasis>
size considerably larger (
<quantity id="4CC39B2D403F267DC9BFFAE4FDFDFA05" box="[505,625,1407,1430]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="9" pageNumber="8" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">2832 mm</quantity>
);
<emphasis id="B94FEADA403F267DCACBFAE4FD2FFA05" bold="true" box="[653,675,1407,1430]" pageId="9" pageNumber="8">3)</emphasis>
metasomal segment II wider than long. The two populations assigned to
<taxonomicName id="4C3B4D4B403F267DC905FA26FE30FA47" baseAuthorityName="Becker" baseAuthorityYear="1880" box="[323,444,1469,1492]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" sensu="lato" species="purvesii">
<emphasis id="B94FEADA403F267DC905FA26FE30FA47" box="[323,444,1469,1492]" italics="true" pageId="9" pageNumber="8">O. purvesii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57A1403F267DC980FA26FE62FA47" box="[454,494,1469,1492]" pageId="9" pageNumber="8" sensu="lato">s. l.</taxonomicNameLabel>
by Francke (1978) as
<emphasis id="B94FEADA403F267DCAACFA26FF72FA61" italics="true" pageId="9" pageNumber="8">O. p. sabae</emphasis>
(from
<collectingCountry id="F32C7658403F267DC90DFA40FE0DFA61" box="[331,385,1499,1522]" name="Bonaire, Sint Eustatius and Saba" pageId="9" pageNumber="8">Saba</collectingCountry>
) and a supposed hybrid between
<emphasis id="B94FEADA403F267DCAABFA40FE9AF982" italics="true" pageId="9" pageNumber="8">
O. p.
<taxonomicName id="4C3B4D4B403F267DC887FA61FE9AF982" baseAuthorityName="Becker" baseAuthorityYear="1880" box="[193,278,1530,1553]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="purvesii">purvesii</taxonomicName>
</emphasis>
and
<emphasis id="B94FEADA403F267DC908FA61FE44F982" box="[334,456,1530,1553]" italics="true" pageId="9" pageNumber="8">O. p. sabae</emphasis>
(from
<collectingCountry id="F32C7658403F267DCA53FA61FD05F982" box="[533,649,1530,1553]" name="Saint Kitts and Nevis" pageId="9" pageNumber="8">Saint Kitts</collectingCountry>
) are in urgent need of a thorough revision, as both will possibly prove to be distinct at species level from
<taxonomicName id="4C3B4D4B403F267DCA7AF9ACFD3FF9DD" baseAuthorityName="Becker" baseAuthorityYear="1880" box="[572,691,1591,1614]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" sensu="stricto" species="purvesii">
<emphasis id="B94FEADA403F267DCA7AF9ACFD3FF9DD" box="[572,691,1591,1614]" italics="true" pageId="9" pageNumber="8">O. purvesii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57A1403F267DCAFCF9ACFD6AF9DD" box="[698,742,1591,1614]" pageId="9" pageNumber="8" sensu="stricto">s. s.</taxonomicNameLabel>
; they are not referable to
<taxonomicName id="4C3B4D4B403F267DC9D4F9CDFE7FF9FE" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[402,499,1622,1645]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA403F267DC9D4F9CDFE7FF9FE" box="[402,499,1622,1645]" italics="true" pageId="9" pageNumber="8">O. nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA403F267DC9BFF9CDFDBAF9FE" bold="true" box="[505,566,1622,1645]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="A27C57A1403F267DC9BFF9CDFDBAF9FE" box="[505,566,1622,1645]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
either, on the basis of the strong differences in size, metasomal proportions and tarsal spine formula.
</paragraph>
<paragraph id="8B8436C8403F267DC8B6F929FC00FA05" blockId="9.[192,804,1284,1890]" lastBlockId="9.[830,1441,1285,1430]" pageId="9" pageNumber="8">
On the other hand,
<taxonomicName id="4C3B4D4B403F267DC9F8F929FDBDF95A" authorityName="Teruel" authorityYear="2008" box="[446,561,1714,1737]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="questeli">
<emphasis id="B94FEADA403F267DC9F8F929FDBDF95A" box="[446,561,1714,1737]" italics="true" pageId="9" pageNumber="8">O. questeli</emphasis>
</taxonomicName>
is about the same size and base coloration of
<taxonomicName id="4C3B4D4B403F267DC9FBF94AFDADF97B" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[445,545,1745,1768]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA403F267DC9FBF94AFDADF97B" box="[445,545,1745,1768]" italics="true" pageId="9" pageNumber="8">O. nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA403F267DCA6FF94AFDE4F97B" bold="true" box="[553,616,1745,1768]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="A27C57A1403F267DCA6FF94AFDE4F97B" box="[553,616,1745,1768]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but differs conspicuously by:
<emphasis id="B94FEADA403F267DC935F974FE05F895" bold="true" box="[371,393,1775,1798]" pageId="9" pageNumber="8">1)</emphasis>
entire body much less hirsute;
<emphasis id="B94FEADA403F267DCB4AF974FCAEF895" bold="true" box="[780,802,1775,1798]" pageId="9" pageNumber="8">2)</emphasis>
pedipalp chela more robust, with fingers shorter and hand globose (not flattened);
<emphasis id="B94FEADA403F267DCA48F8B6FDA8F8D7" bold="true" box="[526,548,1837,1860]" pageId="9" pageNumber="8">3)</emphasis>
metasoma remarkably less slender, with segments I-III each wider than long;
<emphasis id="B94FEADA403F267DCB4AF8D0FCAEF8F1" bold="true" box="[780,802,1867,1890]" pageId="9" pageNumber="8">4)</emphasis>
sternite VII without ventrosubmedian carinae;
<emphasis id="B94FEADA403F267DCDCCFA9EFA2CFA8F" bold="true" box="[1418,1440,1285,1308]" pageId="9" pageNumber="8">5)</emphasis>
pedipalp patella not punctate;
<emphasis id="B94FEADA403F267DCCD6FAB8FB2BFAA9" bold="true" box="[1168,1191,1315,1338]" pageId="9" pageNumber="8">6)</emphasis>
carapace, tergites and pedipalp chela more densely and strongly granulose;
<emphasis id="B94FEADA403F267DCDCCFAD9FA2CFACA" bold="true" box="[1418,1440,1346,1369]" pageId="9" pageNumber="8">7)</emphasis>
coloration with the spotted pattern much darker and denser.
</paragraph>
</subSubSection>
<subSubSection id="C3216543403F267ECB78FA20FEB5F8B0" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="discussion">
<paragraph id="8B8436C8403F267DCB78FA20FC37FA44" blockId="9.[830,955,1467,1495]" box="[830,955,1467,1495]" pageId="9" pageNumber="8">
<heading id="D0CC81A4403F267DCB78FA20FC37FA44" box="[830,955,1467,1495]" pageId="9" pageNumber="8">
<emphasis id="B94FEADA403F267DCB78FA20FC37FA44" bold="true" box="[830,955,1467,1495]" pageId="9" pageNumber="8">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8436C8403F267ECB28FA61FEF2FECA" blockId="9.[830,1442,1530,1890]" lastBlockId="10.[192,802,199,1197]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
Francke (1978) recorded
<emphasis id="B94FEADA403F267DCCC3FA61FA96F982" box="[1157,1306,1530,1553]" italics="true" pageId="9" pageNumber="8">
O. p.
<taxonomicName id="4C3B4D4B403F267DCC82FA61FA96F982" baseAuthorityName="Becker" baseAuthorityYear="1880" box="[1220,1306,1530,1553]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="purvesii">purvesii</taxonomicName>
</emphasis>
from Terrede-Haut, a small islet of Les Saintes just off the south shore of Basse-Terre, but this record appears to be based upon a misidentification. Very recently, one of us (RT) had the chance to examine a series of high-quality color pictures of one adult male, one adult female and one juvenile from this population (taken by François Meurgey and kindly shared by our friend and arachnid enthusiast Karl Questel), which closely match the
<typeStatus id="5480886A403F267DCD2EF974FA2CF895" box="[1384,1440,1775,1798]" pageId="9" pageNumber="8">types</typeStatus>
of
<taxonomicName id="4C3B4D4B403F267DCB18F895FC4DF8B6" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[862,961,1806,1829]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA403F267DCB18F895FC4DF8B6" box="[862,961,1806,1829]" italics="true" pageId="9" pageNumber="8">O. nanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57A1403F267DCB8CF895FB85F8B6" box="[970,1033,1806,1829]" pageId="9" pageNumber="8" rank="species">
<emphasis id="B94FEADA403F267DCB8CF895FB8EF8B6" bold="true" box="[970,1026,1806,1829]" pageId="9" pageNumber="8">sp. n</emphasis>
.
</taxonomicNameLabel>
Also, together with these images we received a second series of high-quality color pictures of five juvenile specimens of
<taxonomicName id="4C3B4D4B403F267DCC27F8D0FB2AF8F1" authorityName="Simon" authorityYear="1880" box="[1121,1190,1867,1890]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403F267DCC27F8D0FB2AF8F1" box="[1121,1190,1867,1890]" italics="true" pageId="9" pageNumber="8">Oiclus</emphasis>
</taxonomicName>
collected in Bouillante (western Basse-Terre), which also look referable to
<taxonomicName id="4C3B4D4B403C267ECB4EFF5CFE8CFF6E" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA403C267ECB4EFF5CFE8CFF6E" italics="true" pageId="10" pageNumber="9">O. nanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57A1403C267EC941FF7DFEC8FF6E" box="[263,324,230,253]" pageId="10" pageNumber="9" rank="species">
<emphasis id="B94FEADA403C267EC941FF7DFEB1FF6E" bold="true" box="[263,317,230,253]" pageId="10" pageNumber="9">sp. n</emphasis>
.
</taxonomicNameLabel>
Both records are here tentatively referred to this species and appear to indicate that it could be widely distributed in mainland
<collectingCountry id="F32C7658403C267EC987FEB8FDCEFEA9" box="[449,578,291,314]" name="Guadeloupe" pageId="10" pageNumber="9">Guadeloupe</collectingCountry>
and at least some of its offshore islets.
</paragraph>
<paragraph id="8B8436C8403C267EC8B6FEC4FEFBFB89" blockId="10.[192,802,199,1197]" pageId="10" pageNumber="9">
Since the classic diplocentrine revision of Francke (1978),
<taxonomicName id="4C3B4D4B403C267EC95EFEE6FED2FE07" authorityName="Simon" authorityYear="1880" box="[280,350,381,404]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403C267EC95EFEE6FED2FE07" box="[280,350,381,404]" italics="true" pageId="10" pageNumber="9">Oiclus</emphasis>
</taxonomicName>
appeared to be the only Antillean genus of this subfamily represented by a single and widespread species, a biogeographical pattern totally different from the one typical of the remaining genera of Diplocentrinae occurring throughout the insular Caribbean:
<taxonomicName id="4C3B4D4B403C267EC886FD8BFE47FDB4" authority="Francke, 1978" authorityName="Francke" authorityYear="1978" box="[192,459,528,551]" class="Arachnida" family="Diplocentridae" genus="Cazierius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403C267EC886FD8BFEAAFDB4" box="[192,294,528,551]" italics="true" pageId="10" pageNumber="9">Cazierius</emphasis>
Francke, 1978
</taxonomicName>
and
<taxonomicName id="4C3B4D4B403C267ECA40FD8BFCADFDB4" authority="Pocock, 1899" authorityName="Pocock" authorityYear="1899" box="[518,801,528,551]" class="Arachnida" family="Diplocentridae" genus="Heteronebo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403C267ECA40FD8BFD08FDB4" box="[518,644,528,551]" italics="true" pageId="10" pageNumber="9">Heteronebo</emphasis>
Pocock,
<quantity id="4CC39B2D403C267ECAAAFD8BFCADFDB4" box="[748,801,528,551]" metricMagnitude="1" metricUnit="m" metricValue="4.82346" pageId="10" pageNumber="9" unit="in" value="1899.0">1899</quantity>
</taxonomicName>
in the Greater Antilles,
<taxonomicName id="4C3B4D4B403C267EC980FDB6FCADFDD7" authority="Kraepelin, 1905" authorityName="Kraepelin" authorityYear="1905" box="[454,801,557,580]" class="Arachnida" family="Diplocentridae" genus="Didymocentrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403C267EC980FDB6FDE5FDD7" box="[454,617,557,580]" italics="true" pageId="10" pageNumber="9">Didymocentrus</emphasis>
Kraepelin,
<quantity id="4CC39B2D403C267ECAAAFDB6FCADFDD7" box="[748,801,557,580]" metricMagnitude="1" metricUnit="m" metricValue="4.8387" pageId="10" pageNumber="9" unit="in" value="1905.0">1905</quantity>
</taxonomicName>
in the Lesser Antilles (Windward Islands) and southern
<collectingCountry id="F32C7658403C267EC886FDF3FE2EFDEC" box="[192,418,616,639]" name="Curaçao" pageId="10" pageNumber="9">Netherlands Antilles</collectingCountry>
; each of these genera is composed by not less than 10 species, none of which is present in more than one independent island. When
<taxonomicName id="4C3B4D4B403C267ECA3BFD38FD7DFD29" authorityName="Teruel" authorityYear="2008" box="[637,753,675,698]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="questeli">
<emphasis id="B94FEADA403C267ECA3BFD38FD7DFD29" box="[637,753,675,698]" italics="true" pageId="10" pageNumber="9">O. questeli</emphasis>
</taxonomicName>
was recently described, Teruel (2008: 98) hypothesized that
<taxonomicName id="4C3B4D4B403C267EC886FD46FE8AFD67" authorityName="Simon" authorityYear="1880" box="[192,262,733,756]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403C267EC886FD46FE8AFD67" box="[192,262,733,756]" italics="true" pageId="10" pageNumber="9">Oiclus</emphasis>
</taxonomicName>
was possibly more diverse than suspected. This was quickly confirmed with the discovery of
<taxonomicName id="4C3B4D4B403C267ECAF8FD60FCAEFC81" authority="Teruel &amp; Chazal, 2010" authorityName="Teruel &amp; Chazal" authorityYear="2010" box="[702,802,763,786]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="B94FEADA403C267ECAF8FD60FCAEFC81" box="[702,802,763,786]" italics="true" pageId="10" pageNumber="9">O. nanus</emphasis>
</taxonomicName>
<emphasis id="B94FEADA403C267EC886FC83FE89FCBC" bold="true" box="[192,261,792,815]" pageId="10" pageNumber="9">
<taxonomicNameLabel id="A27C57A1403C267EC886FC83FE89FCBC" box="[192,261,792,815]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, which represents both the third species-level taxon added to this genus and also its southernmost known geographical occurrence. Based on the general correlation of this confirmed species diversity to their distribution, now it seems very likely that
<taxonomicName id="4C3B4D4B403C267ECAE3FC15FD67FC36" authorityName="Simon" authorityYear="1880" box="[677,747,910,933]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403C267ECAE3FC15FD67FC36" box="[677,747,910,933]" italics="true" pageId="10" pageNumber="9">Oiclus</emphasis>
</taxonomicName>
also shares the same biogeographical pattern as the other members of the subfamily: multiple species endemic from each independent insular bank, in this case in the Leeward Islands.
</paragraph>
<paragraph id="8B8436C8403C267EC8B6FBBAFE99FB3E" blockId="10.[192,802,199,1197]" pageId="10" pageNumber="9">
Last, it is not yet possible to prepare a key to the species of
<taxonomicName id="4C3B4D4B403C267EC97BFBA5FE0FFBC6" authorityName="Simon" authorityYear="1880" box="[317,387,1086,1109]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403C267EC97BFBA5FE0FFBC6" box="[317,387,1086,1109]" italics="true" pageId="10" pageNumber="9">Oiclus</emphasis>
</taxonomicName>
until its
<typeStatus id="5480886A403C267EC9B4FBA5FDACFBC6" box="[498,544,1086,1109]" pageId="10" pageNumber="9">type</typeStatus>
species
<taxonomicName id="4C3B4D4B403C267ECAC1FBA5FC8FFBC6" baseAuthorityName="Becker" baseAuthorityYear="1880" box="[647,771,1086,1109]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="purvesii">
<emphasis id="B94FEADA403C267ECAC1FBA5FC8FFBC6" box="[647,771,1086,1109]" italics="true" pageId="10" pageNumber="9">O. purvesii</emphasis>
</taxonomicName>
is redescribed according to current taxonomy, and the true identity of all populations previously assigned to it is revised.
</paragraph>
<paragraph id="8B8436C8403C267EC886FB57FD0AFB7B" blockId="10.[192,646,1228,1256]" box="[192,646,1228,1256]" pageId="10" pageNumber="9">
<heading id="D0CC81A4403C267EC886FB57FD0AFB7B" box="[192,646,1228,1256]" pageId="10" pageNumber="9">
<emphasis id="B94FEADA403C267EC886FB57FD0AFB7B" bold="true" box="[192,646,1228,1256]" pageId="10" pageNumber="9">Comparative Material Examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8436C8403C267EC886FA9DFDDCFA05" blockId="10.[192,802,1286,1827]" pageId="10" pageNumber="9">
<emphasis id="B94FEADA403C267EC886FA9DFD35FAB2" bold="true" box="[192,697,1286,1314]" pageId="10" pageNumber="9">
<taxonomicName id="4C3B4D4B403C267EC8A0FA92FD38FAB2" authority="purvesii (Becker, 1880)" authorityName="purvesii (Becker" authorityYear="1880" baseAuthorityName="Becker" baseAuthorityYear="1880" box="[230,692,1289,1313]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="purvesii" subSpecies="purvesii">
<emphasis id="B94FEADA403C267EC8A0FA92FE7BFAB2" bold="true" box="[230,503,1289,1313]" italics="true" pageId="10" pageNumber="9">Oiclus purvesii purvesii</emphasis>
(Becker, 1880)
</taxonomicName>
:
</emphasis>
<materialsCitation id="3B533C95403C267ECA81FA91FEFCFAC8" ID-GBIF-Occurrence="3352114306" collectingDate="1918-01" collectionCode="R" country="Antigua and Barbuda" location="English Harbour" pageId="10" pageNumber="9" specimenCount="1">
<emphasis id="B94FEADA403C267ECA81FA91FC97FAB2" box="[711,795,1290,1313]" italics="true" pageId="10" pageNumber="9">
<collectingCountry id="F32C7658403C267ECA81FA91FC97FAB2" box="[711,795,1290,1313]" name="Antigua and Barbuda" pageId="10" pageNumber="9">Antigua</collectingCountry>
</emphasis>
,
<location id="8EE46013403C267EC8A0FABCFE2DFAAD" LSID="urn:lsid:plazi:treatment:039287DE4034267ECB3AFA4FFEB5F8B0:8EE46013403C267EC8A0FABCFE2DFAAD" box="[230,417,1319,1342]" country="Antigua and Barbuda" name="English Harbour" pageId="10" pageNumber="9">English Harbour</location>
, south end of the island;
<date id="FF851008403C267ECA89FABCFE92FAC8" pageId="10" pageNumber="9" value="1918-01">
<collectingDate id="EFC1E9E0403C267ECA89FABCFE92FAC8" pageId="10" pageNumber="9" value="1918-01">January 1918</collectingDate>
</date>
, leg.
<collectionCode id="ED2AAE0D403C267EC91DFADFFEFCFAC8" box="[347,368,1348,1371]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="9">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B533C95403C267EC93AFADFFDC0FA05" ID-GBIF-Occurrence="3352114302" accessionNumber="MCZ 12422" collectingDate="2002-05-31" collectionCode="MCZ, MSU" collectorName="K. Marske" country="Antigua" county="Cassara Ghant" location="Forrest" municipality="Montserrat" pageId="10" pageNumber="9" specimenCount="6" specimenCount-adult="2" specimenCount-female="2" specimenCount-juvenile="1" specimenCount-male="1">
<location id="8EE46013403C267EC93AFADFFE46FAC8" LSID="urn:lsid:plazi:treatment:039287DE4034267ECB3AFA4FFEB5F8B0:8EE46013403C267EC93AFADFFE46FAC8" box="[380,458,1348,1371]" country="Antigua" county="Cassara Ghant" municipality="Montserrat" name="Forrest" pageId="10" pageNumber="9">Forrest</location>
,
<specimenCount id="9D3DFD41403C267EC991FADFFDAEFAC8" box="[471,546,1348,1371]" pageId="10" pageNumber="9" type="adult">1 adult</specimenCount>
<specimenCount id="9D3DFD41403C267ECA6DFADFFDB3FAC8" box="[555,575,1348,1371]" pageId="10" pageNumber="9" type="female"></specimenCount>
,
<specimenCount id="9D3DFD41403C267ECA08FADFFD36FAC8" box="[590,698,1348,1371]" pageId="10" pageNumber="9" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="9D3DFD41403C267ECA84FADFFD5AFAC8" box="[706,726,1348,1371]" pageId="10" pageNumber="9" type="male"></specimenCount>
(
<collectionCode id="ED2AAE0D403C267ECAA0FADFFCADFAC8" box="[742,801,1348,1371]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="10" pageNumber="9" type="Museum">MCZ</collectionCode>
12422).
<emphasis id="B94FEADA403C267EC903FAF9FE31FAEA" box="[325,445,1378,1401]" italics="true" pageId="10" pageNumber="9">
<collectingMunicipality id="6BE0ACB2403C267EC903FAF9FE31FAEA" box="[325,445,1378,1401]" pageId="10" pageNumber="9">Montserrat</collectingMunicipality>
</emphasis>
,
<collectingCounty id="62E54E44403C267EC989FAF9FDFEFAEA" box="[463,626,1378,1401]" pageId="10" pageNumber="9">Cassara Ghant</collectingCounty>
,
<date id="FF851008403C267ECAC4FAF9FC91FAEA" box="[642,797,1378,1401]" pageId="10" pageNumber="9" value="2002-05-31">
<collectingDate id="EFC1E9E0403C267ECAC4FAF9FC91FAEA" box="[642,797,1378,1401]" pageId="10" pageNumber="9" value="2002-05-31">31 May 2002</collectingDate>
</date>
, leg.
<collectorName id="26CE531E403C267EC952FAE4FE0BFA05" box="[276,391,1407,1430]" pageId="10" pageNumber="9">K. Marske</collectorName>
,
<specimenCount id="9D3DFD41403C267EC9D4FAE4FE57FA05" box="[402,475,1407,1430]" pageId="10" pageNumber="9" type="adult">1 adult</specimenCount>
<specimenCount id="9D3DFD41403C267EC9A4FAE4FE7AFA05" box="[482,502,1407,1430]" pageId="10" pageNumber="9" type="female"></specimenCount>
(
<collectionCode id="ED2AAE0D403C267ECA43FAE4FDC8FA05" box="[517,580,1407,1430]" country="USA" httpUri="http://grbio.org/cool/53w1-8r3k" name="Michigan State University Museum" pageId="10" pageNumber="9" type="University or college">MSU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8436C8403C267EC886FA3BFCADF91A" blockId="10.[192,802,1286,1827]" pageId="10" pageNumber="9">
<emphasis id="B94FEADA403C267EC886FA3BFDB1FA28" bold="true" box="[192,573,1440,1468]" pageId="10" pageNumber="9">
<taxonomicName id="4C3B4D4B403C267EC8A0FA38FDBBFA28" authority="Teruel, 2008" authorityName="Teruel" authorityYear="2008" box="[230,567,1443,1467]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="questeli">
<emphasis id="B94FEADA403C267EC8A0FA38FE03FA28" bold="true" box="[230,399,1443,1467]" italics="true" pageId="10" pageNumber="9">Oiclus questeli</emphasis>
Teruel, 2008
</taxonomicName>
:
</emphasis>
<materialsCitation id="3B533C95403C267ECA0DFA3FFC92F91A" ID-GBIF-Occurrence="3352114305" collectingDate="2005-10-16" collectingDateMax="2008-02-14" collectingDateMin="2005-10-16" collectionCode="RTO" collectorName="K. Questel" country="Guadeloupe" location="Barthelemy" municipality="Petit Anse" pageId="10" pageNumber="9" specimenCount="10" specimenCount-adult="3" specimenCount-female="3" specimenCount-juvenile="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B94FEADA403C267ECA0DFA3FFD41FA28" box="[587,717,1444,1467]" italics="true" pageId="10" pageNumber="9">
<collectingCountry id="F32C7658403C267ECA0DFA3FFD41FA28" box="[587,717,1444,1467]" name="Guadeloupe" pageId="10" pageNumber="9">Guadeloupe</collectingCountry>
</emphasis>
, Saint-
<location id="8EE46013403C267EC8A0FA5AFEE9FA4B" LSID="urn:lsid:plazi:treatment:039287DE4034267ECB3AFA4FFEB5F8B0:8EE46013403C267EC8A0FA5AFEE9FA4B" box="[230,357,1473,1496]" country="Guadeloupe" municipality="Petit Anse" name="Barthelemy" pageId="10" pageNumber="9">Barthélemy</location>
,
<collectingMunicipality id="6BE0ACB2403C267EC933FA5AFE60FA4B" box="[373,492,1473,1496]" pageId="10" pageNumber="9">Petit Anse</collectingMunicipality>
,
<date id="FF851008403C267EC9BDFA5AFD49FA4B" box="[507,709,1473,1496]" pageId="10" pageNumber="9" value="2008-02-14">
<collectingDate id="EFC1E9E0403C267EC9BDFA5AFD49FA4B" box="[507,709,1473,1496]" pageId="10" pageNumber="9" value="2008-02-14">14 February 2008</collectingDate>
</date>
, leg.
<collectorName id="26CE531E403C267ECB4EFA5AFEB6FA65" pageId="10" pageNumber="9">K. Questel</collectorName>
,
<specimenCount id="9D3DFD41403C267EC901FA44FE1FFA65" box="[327,403,1503,1526]" pageId="10" pageNumber="9" type="adult">1 adult</specimenCount>
<specimenCount id="9D3DFD41403C267EC9DAFA44FE3CFA65" box="[412,432,1503,1526]" pageId="10" pageNumber="9" type="male"></specimenCount>
<typeStatus id="5480886A403C267EC9FFFA44FD9AFA65" box="[441,534,1503,1526]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED2AAE0D403C267ECA61FA44FDEEFA65" box="[551,610,1503,1526]" pageId="10" pageNumber="9">RTO</collectionCode>
: Sco.0379); same locality,
<date id="FF851008403C267EC903FA67FE76F980" box="[325,506,1532,1555]" pageId="10" pageNumber="9" value="2008-02-07">
<collectingDate id="EFC1E9E0403C267EC903FA67FE76F980" box="[325,506,1532,1555]" pageId="10" pageNumber="9" value="2008-02-07">7 February 2008</collectingDate>
</date>
, leg.
<collectorName id="26CE531E403C267ECA73FA67FD27F980" box="[565,683,1532,1555]" pageId="10" pageNumber="9">K. Questel</collectorName>
,
<specimenCount id="9D3DFD41403C267ECAF0FA67FCADF980" box="[694,801,1532,1555]" pageId="10" pageNumber="9" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="9D3DFD41403C267EC8A0F982FF76F9A3" box="[230,250,1561,1584]" pageId="10" pageNumber="9" type="male"></specimenCount>
and
<specimenCount id="9D3DFD41403C267EC979F982FE3EF9A3" box="[319,434,1561,1584]" pageId="10" pageNumber="9" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="9D3DFD41403C267EC986F982FE58F9A3" box="[448,468,1561,1584]" pageId="10" pageNumber="9" type="female"></specimenCount>
<typeStatus id="5480886A403C267EC9A5F982FDC5F9A3" box="[483,585,1561,1584]" pageId="10" pageNumber="9" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED2AAE0D403C267ECA26F982FD16F9A3" box="[608,666,1561,1584]" pageId="10" pageNumber="9">RTO</collectionCode>
: Sco.0378); Saline,
<date id="FF851008403C267EC97FF9ACFE79F9DD" box="[313,501,1591,1614]" pageId="10" pageNumber="9" value="2005-10-16">
<collectingDate id="EFC1E9E0403C267EC97FF9ACFE79F9DD" box="[313,501,1591,1614]" pageId="10" pageNumber="9" value="2005-10-16">16 October 2005</collectingDate>
</date>
, leg.
<collectorName id="26CE531E403C267ECA75F9ACFD26F9DD" box="[563,682,1591,1614]" pageId="10" pageNumber="9">K. Questel</collectorName>
,
<specimenCount id="9D3DFD41403C267ECAFEF9ACFC89F9DD" box="[696,773,1591,1614]" pageId="10" pageNumber="9" type="adult">1 adult</specimenCount>
<specimenCount id="9D3DFD41403C267ECB48F9ACFCAEF9DD" box="[782,802,1591,1614]" pageId="10" pageNumber="9" type="female"></specimenCount>
<typeStatus id="5480886A403C267EC8A0F9CFFECEF9F8" box="[230,322,1620,1643]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED2AAE0D403C267EC912F9CFFE03F9F8" box="[340,399,1620,1643]" pageId="10" pageNumber="9">RTO</collectionCode>
: Sco.0314); Flamand,
<date id="FF851008403C267ECACFF9CFFC92F9F8" box="[649,798,1620,1643]" pageId="10" pageNumber="9" value="2007-07-30">
<collectingDate id="EFC1E9E0403C267ECACFF9CFFC92F9F8" box="[649,798,1620,1643]" pageId="10" pageNumber="9" value="2007-07-30">30 July 2007</collectingDate>
</date>
; leg.
<collectorName id="26CE531E403C267EC952F9E9FE05F91A" box="[276,393,1650,1673]" pageId="10" pageNumber="9">K. Questel</collectorName>
,
<specimenCount id="9D3DFD41403C267EC9D2F9E9FE51F91A" box="[404,477,1650,1673]" pageId="10" pageNumber="9" type="adult">1 adult</specimenCount>
<specimenCount id="9D3DFD41403C267EC9A2F9E9FE74F91A" box="[484,504,1650,1673]" pageId="10" pageNumber="9" type="female"></specimenCount>
<typeStatus id="5480886A403C267EC9B8F9E9FDD6F91A" box="[510,602,1650,1673]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED2AAE0D403C267ECA2FF9E9FD28F91A" box="[617,676,1650,1673]" pageId="10" pageNumber="9">RTO</collectionCode>
: Sco.0365)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8436C8403C267EC886F909FEB5F8B0" blockId="10.[192,802,1286,1827]" pageId="10" pageNumber="9">
<emphasis id="B94FEADA403C267EC886F909FEDBF93E" bold="true" box="[192,343,1682,1710]" pageId="10" pageNumber="9">
<taxonomicName id="4C3B4D4B403C267EC8A0F90EFEA1F93E" authorityName="Simon" authorityYear="1880" box="[230,301,1685,1709]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEADA403C267EC8A0F90EFEA1F93E" bold="true" box="[230,301,1685,1709]" italics="true" pageId="10" pageNumber="9">Oiclus</emphasis>
</taxonomicName>
sp.
</emphasis>
(supposed hybrid between
<emphasis id="B94FEADA403C267ECACCF90DFCAEF93E" box="[650,802,1686,1709]" italics="true" pageId="10" pageNumber="9">
O. p.
<taxonomicName id="4C3B4D4B403C267ECA8AF90DFCAEF93E" baseAuthorityName="Becker" baseAuthorityYear="1880" box="[716,802,1686,1709]" class="Arachnida" family="Diplocentridae" genus="Oiclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="purvesii">purvesii</taxonomicName>
</emphasis>
and
<emphasis id="B94FEADA403C267EC952F92FFE03F958" box="[276,399,1716,1739]" italics="true" pageId="10" pageNumber="9">O. p. sabae</emphasis>
):
<materialsCitation id="3B533C95403C267EC9E1F92FFEBAF8B0" ID-GBIF-Occurrence="3352114303" collectingDate="2003-07-04" collectionCode="MSU" collectorName="M. A. Ivie" country="Saint Kitts and Nevis" county="Peter Manning Trail" location="St. Thomas Middle Island Parish" municipality="Wingfield National Park" pageId="10" pageNumber="9" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1">
<emphasis id="B94FEADA403C267EC9E1F92FFE70F958" box="[423,508,1716,1739]" italics="true" pageId="10" pageNumber="9">
<collectingCountry id="F32C7658403C267EC9E1F92FFE70F958" box="[423,508,1716,1739]" name="Saint Kitts and Nevis" pageId="10" pageNumber="9">St. Kitts</collectingCountry>
</emphasis>
,
<location id="8EE46013403C267ECA4CF92FFEA0F97B" LSID="urn:lsid:plazi:treatment:039287DE4034267ECB3AFA4FFEB5F8B0:8EE46013403C267ECA4CF92FFEA0F97B" country="Saint Kitts and Nevis" county="Peter Manning Trail" municipality="Wingfield National Park" name="St. Thomas Middle Island Parish" pageId="10" pageNumber="9">St. Thomas Middle Island Parish</location>
,
<collectingMunicipality id="6BE0ACB2403C267EC904F94AFDEEF97B" box="[322,610,1745,1768]" pageId="10" pageNumber="9">Wingfield National Park</collectingMunicipality>
,
<collectingCounty id="62E54E44403C267ECA3EF94AFE90F896" pageId="10" pageNumber="9">Peter Manning Trail</collectingCounty>
,
<date id="FF851008403C267EC968F975FE36F896" box="[302,442,1774,1797]" pageId="10" pageNumber="9" value="2003-07-04">
<collectingDate id="EFC1E9E0403C267EC968F975FE36F896" box="[302,442,1774,1797]" pageId="10" pageNumber="9" value="2003-07-04">4 July 2003</collectingDate>
</date>
, leg.
<collectorName id="26CE531E403C267ECA46F975FDF2F896" box="[512,638,1774,1797]" pageId="10" pageNumber="9">M. A. Ivie</collectorName>
,
<specimenCount id="9D3DFD41403C267ECAD6F975FC8CF896" box="[656,768,1774,1797]" pageId="10" pageNumber="9" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="9D3DFD41403C267ECB48F975FCAEF896" box="[782,802,1774,1797]" pageId="10" pageNumber="9" type="female"></specimenCount>
(
<collectionCode id="ED2AAE0D403C267EC8A8F897FEA1F8B0" box="[238,301,1804,1827]" country="USA" httpUri="http://grbio.org/cool/53w1-8r3k" name="Michigan State University Museum" pageId="10" pageNumber="9" type="University or college">MSU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>