522 lines
68 KiB
XML
522 lines
68 KiB
XML
<document id="FBE9A0D4F7DD128041F8D7F418261621" ID="10.11646/zootaxa.4277.4.1SLASH11426" ID-CLB-Dataset="32677" ID-DOI="10.11646/zootaxa.4277.4.1" ID-GBIF-Dataset="bc0690ff-ca74-4ba8-bc70-7376f04cf203" ID-ISSN="1175-5326" ID-Zenodo-Dep="829833" ID-ZooBank="1A18DA8-2AFF-4998-AE87-00D713EC531D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497944538512" checkinUser="plazi" docAuthor="Garg, Sonali & Biju, S. D." docDate="2017" docId="039287B7313FFF82FF5DB280D464FCBB" docLanguage="en" docName="zootaxa.4277.4.1SLASH11426.pdf" docOrigin="Zootaxa 4277 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fejervarya cepfi Garg & Biju 2017, sp. nov." docType="treatment" docUuid="6A7B7C4D-31C3-4C43-B91B-EE78532847DA" docUuidSource="ZooBank" docVersion="8" lastPageNumber="475" masterDocId="FFABFFCF312AFF9AFFCAB132D536FFAF" masterDocTitle="Description of four new species of Burrowing Frogs in the Fejervarya rufescens complex (Dicroglossidae) with notes on morphological affinities of Fejervarya species in the Western Ghats" masterLastPageNumber="490" masterPageNumber="451" pageNumber="472" updateTime="1698447367985" updateUser="ExternalLinkService">
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<mods:title id="F2836C9E8657EA36B24F97580B3BDE75">Description of four new species of Burrowing Frogs in the Fejervarya rufescens complex (Dicroglossidae) with notes on morphological affinities of Fejervarya species in the Western Ghats</mods:title>
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<mods:namePart id="6F9DC099E6F13C945306D2DABD042514">Garg, Sonali</mods:namePart>
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<mods:namePart id="30B96C9F1BCDA65B5E3B605A171F5DEB">Biju, S. D.</mods:namePart>
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<mods:title id="7BB51FB1472E8B382D6F066C3AE55843">Zootaxa</mods:title>
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<mods:date id="082D69311C9ADBC1EE77FF60570945A5">2017</mods:date>
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<mods:number id="0DEC7DBDCDEEA9FCE8F4FAA7D1F810EF">4277</mods:number>
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<treatment id="039287B7313FFF82FF5DB280D464FCBB" ID-DOI="http://doi.org/10.5281/zenodo.6052412" ID-GBIF-Taxon="132549772" ID-Zenodo-Dep="6052412" LSID="urn:lsid:zoobank.org:act:6A7B7C4D-31C3-4C43-B91B-EE78532847DA" httpUri="http://treatment.plazi.org/id/039287B7313FFF82FF5DB280D464FCBB" lastPageId="24" lastPageNumber="475" pageId="21" pageNumber="472">
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<subSubSection id="C321652A313FFF8FFF5DB280D4A0FBBD" pageId="21" pageNumber="472" type="nomenclature">
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<paragraph id="8B8436A1313FFF8FFF5DB280D4F7FC62" blockId="21.[151,1189,946,1078]" box="[151,449,946,973]" pageId="21" pageNumber="472">
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<heading id="D0CC81CD313FFF8FFF5DB280D4F7FC62" bold="true" box="[151,449,946,973]" fontSize="11" level="1" pageId="21" pageNumber="472" reason="1">
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<emphasis id="B94FEAB3313FFF8FFF5DB280D4F7FC62" bold="true" box="[151,449,946,973]" pageId="21" pageNumber="472">
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<taxonomicName id="4C3B4D22313FFF8FFF5DB280D46BFC63" ID-CoL="3DXDP" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[151,349,946,972]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="21" pageNumber="472" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
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<emphasis id="B94FEAB3313FFF8FFF5DB280D46BFC63" bold="true" box="[151,349,946,972]" italics="true" pageId="21" pageNumber="472">Fejervarya cepfi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27C57C8313FFF8FFEAEB281D4F7FC62" box="[356,449,947,973]" pageId="21" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8436A1313FFF8FFF5DB2E7D4A0FBBD" blockId="21.[151,1189,946,1078]" pageId="21" pageNumber="472">http://zoobank.org/urn:lsid:zoobank.org:act:6A7B7C4D-31C3-4C43-B91B-EE78532847DA CEPF Burrowing Frog</paragraph>
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</subSubSection>
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<subSubSection id="C321652A313FFF8FFF5DB52FD480FB99" box="[151,438,1053,1078]" pageId="21" pageNumber="472" type="description">
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<paragraph id="8B8436A1313FFF8FFF5DB52FD480FB99" blockId="21.[151,1189,946,1078]" box="[151,438,1053,1078]" pageId="21" pageNumber="472">
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(
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<tableCitation id="C6B9031A313FFF8FFF54B52FD42BFB99" box="[158,285,1053,1078]" captionStart-0="TABLE 1" captionStart-1="TABLE 4" captionStart-2="TABLE 5" captionStart-3="TABLE 6" captionStart-4="TABLE 7" captionStartId-0="11.[125,217,515,539]" captionStartId-1="14.[151,239,940,963]" captionStartId-2="15.[151,239,581,604]" captionStartId-3="15.[151,239,1081,1104]" captionStartId-4="18.[119,204,562,584]" captionText-0="TABLE 1. List of DNA sequences used in the study. Taxa are arranged in the order of their phylogenetic position in Figure 1 (Top to bottom)." captionText-1="TABLE 4. Factor loadings and percent variance explained by Principal Component (PC) 1 – 5 for 17 size-corrected morphometric variables from males of five species in the Fejervarya rufescens group. Values in bold indicate variables with the highest loadings." captionText-2="TABLE 5. Scores for discriminant function analysis of the principal components (PC 1 – 5). Values in bold indicate variables with the highest loadings for discriminant function roots of eigenvalue> 1.0." captionText-3="TABLE 6. Classification matrix for adult male specimens of the Fejervarya rufescens group from the discriminant function analysis." captionText-4="TABLE 7. Morphometric measurements for five species in the Fejervarya rufescens group. All measurements are in millimeters (mm). The status of each specimen is given the museum number: Holotype (HT), Paratype (PT), Neotype (NT), Referred specimen (RS)." pageId="21" pageNumber="472">Tables 1–7</tableCitation>
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;
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<figureCitation id="13002A24313FFF8FFEE3B52FD4A5FB99" box="[297,403,1053,1078]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="4.[151,250,1885,1907]" captionStartId-1="5.[151,250,1618,1640]" captionStartId-2="6.[151,250,1765,1787]" captionStartId-3="8.[151,250,1820,1842]" captionStartId-4="9.[151,250,1518,1540]" captionStartId-5="10.[151,250,1726,1748]" captionTargetBox-0="[161,1435,204,1854]" captionTargetBox-1="[155,1424,193,1582]" captionTargetBox-2="[151,1436,197,1739]" captionTargetBox-3="[151,1436,180,1813]" captionTargetBox-4="[151,1436,193,1495]" captionTargetBox-5="[158,1434,882,1697]" captionTargetId-0="figure@4.[151,1436,193,1864]" captionTargetId-1="figure@5.[151,1436,193,1597]" captionTargetId-2="figure@6.[151,1436,193,1744]" captionTargetId-3="figure@8.[151,1436,180,1818]" captionTargetId-4="figure@9.[151,1436,193,1497]" captionTargetId-5="figure@10.[151,1437,879,1704]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 1. Maximum Likelihood phylogram based on the 16 S mitochondrial DNA dataset of 545 basepairs. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of> 50 % are indicated above and below the branches, respectively. The uncorrected pairwise distances between species in the Fejervarya rufescens group are shown near the clade labels. Voucher number and locality details of the samples are referenced in Table 1." captionText-1="FIGURE 2. Phylogenetic analyses. A. Maximum Likelihood phylogram based on mitochondrial (16 S, COI and Cytb) and nuclear (Rag 1) DNA of 2314 basepairs, showing phylogenetic relationships among members of the Fejervarya rufescens group. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of> 50 % are indicated above and below the branches, respectively. Fejervarya granosa (2) was used as the outgroup taxon. Voucher number and locality details of the samples are referenced in Table 1; B. Median Joining haplotype network inferred from 554 basepair fragment of the nuclear Rag 1 gene. Circle sizes are proportional to the number of haplotypes, branch lengths are proportional to the number of mutational steps, and black bars represent mutational steps. Number of individuals used for each species: Fejervarya cepfi (3), Fejervarya kadar (1), Fejervarya manoharani (3), Fejervarya neilcoxi (2), and Fejervarya rufescens (5)." captionText-2="FIGURE 3. Geographical distribution of five species in Fejervarya rufescens group of the Western Ghats, Peninsular India. The Western Ghats biodiversity hotspot region is shown in orange colour. Locality details are referenced in Table 3." captionText-3="FIGURE 4. Four morphologically recognized Fejervarya groups in the Western Ghats. A – B. Fejervarya nilagirica group: A. Fejervarya nilagirica in life (male, SDBDU 2014.2772); B. Fejervarya keralensis in life (male, SDBDU 2015.3139); C. Fejervarya rufescens group, Fejervarya rufescens in life (male, SDBDU 2015.3070); D. Fejervarya sahyadris group, Fejervarya sahyadris in life (male, SDBDU 2015.3046); E – F. Fejervarya syhadrensis group: E. Fejervarya syhadrensis in life (male, SDBDU 2015.2934); F. Fejervarya granosa in life (male, SDBDU 2014.2541). Numbers on the panels represents: 1. dorsal view, 2. ventral view of foot showing the inner metatarsal tubercle, 3. schematic illustration of foot webbing, 4. thighs showing the marking and colour pattern, 5. lateral view showing the groin markings." captionText-4="FIGURE 5. Morphological comparison between members of the genus Sphaerotheca and Fejervarya rufescens group of the Western Ghats. A – H. Genus Sphaerotheca: A – E. Sphaerotheca cf. breviceps, in life (SDBDU 2014.2822): A. dorsolateral view, B. ventral view, C. ventral skin texture, D. lateral view showing the groin markings, E. thighs showing the marking and colour pattern; F – H. Inner metatarsal tubercle: F. Sphaerotheca breviceps (syntype ZMB 3351), G. S. dobsonii (Holotype BMNH 1947.2.28.45), H. S. rolandae (Holotype BMNH 1973.3024). I – R. Fejervarya rufescens group: I – K. F. rufescens in life (SDBDU 2015.3070): I. dorsolateral view, J. ventral view, K. ventral skin texture; L – M. F. kadar sp. nov. (Holotype ZSI / WGRC / V / A / 940): L. lateral view showing the groin markings, M. thighs showing the marking and colour pattern; N – R. Inner metatarsal tubercle: N. F. cepfi sp. nov. (Holotype ZSI / WGRC / V / A / 937), O. F. kadar sp. nov. (Holotype ZSI / WGRC / V / A / 940), P. F. manoharani sp. nov. (Holotype ZSI / WGRC / V / A / 945), Q. F. neilcoxi sp. nov. (Holotype ZSI / WGRC / V / A / 951), R. F. rufescens (SDBDU 2015.3070)." captionText-5="FIGURE 6. PCA plot for the first two principal component axes showing morphometric differentiation among adult males of five species in the Fejervarya rufescens groups of the Western Ghats. Factor loadings for each component are provided in Table 4." httpUri-0="https://zenodo.org/record/829835/files/figure.png" httpUri-1="https://zenodo.org/record/829837/files/figure.png" httpUri-2="https://zenodo.org/record/829839/files/figure.png" httpUri-3="https://zenodo.org/record/829841/files/figure.png" httpUri-4="https://zenodo.org/record/829843/files/figure.png" httpUri-5="https://zenodo.org/record/829845/files/figure.png" pageId="21" pageNumber="472">Figs. 1–6</figureCitation>
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,
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<figureCitation id="13002A24313FFF8FFE55B52CD498FB99" box="[415,430,1054,1078]" captionStart="FIGURE 9" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[151,1436,302,1896]" captionTargetId="figure@22.[151,1437,301,1896]" captionTargetPageId="22" captionText="FIGURE 9. Holotype of Fejervarya cepfi sp. nov. (ZSI / WGRC / V / A / 937): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829851/files/figure.png" pageId="21" pageNumber="472">9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C321652A313FFF8FFF5DB557D6FCFAA2" pageId="21" pageNumber="472" type="etymology">
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<paragraph id="8B8436A1313FFF8FFF5DB557D6FCFAA2" blockId="21.[151,1437,1125,2014]" pageId="21" pageNumber="472">
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<emphasis id="B94FEAB3313FFF8FFF5DB557D428FBD1" bold="true" box="[151,286,1125,1150]" pageId="21" pageNumber="472">Etymology.</emphasis>
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The species is named after the Critical Ecosystem Partnership Fund http://www.cepf.net (CEPF) for its effort to protect global biodiversity hotspots by providing grants in general, and specifically for a grant supporting research and conservation planning in the
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<collectingRegion id="49FFF843313FFF8FFDB1B59CD7E0FB6A" box="[635,726,1198,1221]" country="Sri Lanka" name="Western" pageId="21" pageNumber="472">Western</collectingRegion>
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Ghats biodiversity hotspot through the Project
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<collectingRegion id="49FFF843313FFF8FFB3DB59CD064FB6A" box="[1271,1362,1198,1221]" country="Sri Lanka" name="Western" pageId="21" pageNumber="472">Western</collectingRegion>
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Ghats Network of Protected Areas for Threatened Amphibians http://www.wnpata.org (WNPATA) to SDB (University of
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<collectingRegion id="49FFF843313FFF8FFF5DB5C7D5ECFAA1" box="[151,218,1269,1294]" country="India" name="Delhi" pageId="21" pageNumber="472">Delhi</collectingRegion>
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). The specific epithet
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<taxonomicName id="4C3B4D22313FFF8FFE1AB5C4D730FAA1" authorityName="Garg & Biju" authorityYear="2017" box="[464,518,1270,1294]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="21" pageNumber="472" phylum="Chordata" rank="species" species="cepfi">
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<emphasis id="B94FEAB3313FFF8FFE1AB5C4D730FAA1" box="[464,518,1270,1294]" italics="true" pageId="21" pageNumber="472">cepfi</emphasis>
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</taxonomicName>
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is treated as a noun in the genitive case.
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</paragraph>
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<paragraph id="8B8436A1313FFF8FFF0DB42BD14CFAF9" blockId="21.[151,1437,1125,2014]" pageId="21" pageNumber="472">
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<materialsCitation id="3B533CFC313FFF8FFF0DB42BD14CFAF9" ID-GBIF-Occurrence="1572402627" collectingDate="2013-07-22" collectionCode="WGRC" country="India" elevation="750" latitude="15.954614" location="Amboli" longLatPrecision="1" longitude="73.9986" municipality="Sindhudurg district" pageId="21" pageNumber="472" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
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<emphasis id="B94FEAB3313FFF8FFF0DB42BD40DFA9D" bold="true" box="[199,315,1305,1330]" pageId="21" pageNumber="472">
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<typeStatus id="54808803313FFF8FFF0DB42BD400FA9D" box="[199,310,1305,1330]" pageId="21" pageNumber="472" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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ZSI/
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<collectionCode id="ED2AAE64313FFF8FFEBCB428D4FAFA9D" box="[374,460,1306,1330]" pageId="21" pageNumber="472">WGRC</collectionCode>
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/V/A/937, an adult male, from
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(
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,
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asl
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</elevation>
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),
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<collectingMunicipality id="6BE0ACDB313FFF8FFF5DB40FD447FAF9" box="[151,369,1341,1366]" pageId="21" pageNumber="472">Sindhudurg district</collectingMunicipality>
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,
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<collectingRegion id="49FFF843313FFF8FFEB6B40FD73EFAF9" box="[380,520,1341,1366]" country="India" name="Maharashtra" pageId="21" pageNumber="472">Maharashtra</collectingRegion>
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state,
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<collectingCountry id="F32C7631313FFF8FFD9AB40FD7BAFAF9" box="[592,652,1341,1366]" name="India" pageId="21" pageNumber="472">India</collectingCountry>
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, collected by SDB and SG on
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<collectingDate id="EFC1E989313FFF8FFC28B40CD140FAF9" box="[994,1142,1341,1366]" pageId="21" pageNumber="472" value="2013-07-22">
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<date id="FF851061313FFF8FFC28B40CD140FAF9" box="[994,1142,1341,1366]" pageId="21" pageNumber="472" value="2013-07-22">22 July 2013</date>
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</collectingDate>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B8436A1313FFF8FFF0DB453D455FA31" blockId="21.[151,1437,1125,2014]" pageId="21" pageNumber="472">
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<emphasis id="B94FEAB3313FFF8FFF0DB453D470FAD5" bold="true" box="[199,326,1377,1402]" pageId="21" pageNumber="472">Paratypes.</emphasis>
|
||
ZSI/WGRC/V/A/938, an adult male, and ZSI/WGRC/V/A/939, an adult female, collected along with the holotype.
|
||
</paragraph>
|
||
<paragraph id="8B8436A1313FFF8FFF0DB49BD4D8F9FD" blockId="21.[151,1437,1125,2014]" pageId="21" pageNumber="472">
|
||
<materialsCitation id="3B533CFC313FFF8FFF0DB49BD1B2FA49" ID-GBIF-Occurrence="1572402620" collectingDate="2012-12-03" collectionCode="SDBDU" country="India" elevation="221" latitude="18.45514" location="Raigad district" longLatPrecision="1" longitude="72.92619" pageId="21" pageNumber="472" specimenCode="SDBDU 2012.1429" specimenCount="1" stateProvince="Maharashtra">
|
||
<emphasis id="B94FEAB3313FFF8FFF0DB49BD4DDFA6D" bold="true" box="[199,491,1449,1474]" pageId="21" pageNumber="472">Other referred material.</emphasis>
|
||
<specimenCode id="DB9D9EDA313FFF8FFE3BB498D7E7FA6D" box="[497,721,1450,1474]" collectionCode="SDBDU" pageId="21" pageNumber="472">
|
||
<collectionCode id="ED2AAE64313FFF8FFE3BB498D766FA6D" box="[497,592,1450,1474]" pageId="21" pageNumber="472">SDBDU</collectionCode>
|
||
2012.1429
|
||
</specimenCode>
|
||
, from Phansad WLS (
|
||
<geoCoordinate id="EE0F5066313FFF8FFC02B49BD15AFA6D" box="[968,1132,1449,1474]" direction="north" orientation="latitude" pageId="21" pageNumber="472" precision="1" value="18.45514">18°27’18.5’’N</geoCoordinate>
|
||
<geoCoordinate id="EE0F5066313FFF8FFBB8B49BD025FA6D" box="[1138,1299,1449,1474]" direction="east" orientation="longitude" pageId="21" pageNumber="472" precision="1" value="72.92619">72°55’34.3’’E</geoCoordinate>
|
||
,
|
||
<elevation id="0016D192313FFF8FFAD5B498D0B9FA6D" box="[1311,1423,1449,1474]" metricMagnitude="2" metricUnit="m" metricValue="2.21" pageId="21" pageNumber="472" unit="m" value="221.0">
|
||
<quantity id="4CC39B44313FFF8FFAD5B498D050FA6E" box="[1311,1382,1450,1474]" metricMagnitude="2" metricUnit="m" metricValue="2.21" pageId="21" pageNumber="472" unit="m" value="221.0">221 m</quantity>
|
||
asl
|
||
</elevation>
|
||
),
|
||
<location id="8EE4607A313FFF8FFF5DB4FFD408FA49" LSID="urn:lsid:plazi:treatment:039287B7313FFF82FF5DB280D464FCBB:8EE4607A313FFF8FFF5DB4FFD408FA49" box="[151,318,1485,1510]" country="India" latitude="18.45514" longLatPrecision="1" longitude="72.92619" name="Raigad district" pageId="21" pageNumber="472" stateProvince="Maharashtra">Raigad district</location>
|
||
,
|
||
<collectingRegion id="49FFF843313FFF8FFE83B4FFD4E3FA49" box="[329,469,1485,1510]" country="India" name="Maharashtra" pageId="21" pageNumber="472">Maharashtra</collectingRegion>
|
||
state,
|
||
<collectingCountry id="F32C7631313FFF8FFDD7B4FFD76FFA49" box="[541,601,1485,1510]" name="India" pageId="21" pageNumber="472">India</collectingCountry>
|
||
, collected by SDB and SG on 0
|
||
<collectingDate id="EFC1E989313FFF8FFC76B4FCD1B2FA49" box="[956,1156,1485,1510]" pageId="21" pageNumber="472" value="2012-12-03">
|
||
<date id="FF851061313FFF8FFC76B4FCD1B2FA49" box="[956,1156,1485,1510]" pageId="21" pageNumber="472" value="2012-12-03">3 December 2012</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B533CFC313FFF8FFB5AB4FCD4D8F9FD" ID-GBIF-Occurrence="1572402633" collectingDate="2007-08-25" collectionCode="SDBDU" country="India" elevation="820" latitude="17.405066" location="Satara district" longLatPrecision="1" longitude="73.73831" municipality="These" pageId="21" pageNumber="472" specimenCode="SDBDU 2007.1561, SDBDU 2007.1569" specimenCount="1" stateProvince="Maharashtra">
|
||
<specimenCode id="DB9D9EDA313FFF8FFB5AB4FCD05BFA49" box="[1168,1389,1485,1510]" collectionCode="SDBDU" pageId="21" pageNumber="472">
|
||
<collectionCode id="ED2AAE64313FFF8FFB5AB4FCD1D9FA49" box="[1168,1263,1486,1510]" pageId="21" pageNumber="472">SDBDU</collectionCode>
|
||
2007.1561
|
||
</specimenCode>
|
||
and
|
||
<specimenCode id="DB9D9EDA313FFF8FFF5DB4C0D442F9A5" box="[151,372,1521,1546]" collectionCode="SDBDU" pageId="21" pageNumber="472">
|
||
<collectionCode id="ED2AAE64313FFF8FFF5DB4C0D5C0F9A5" box="[151,246,1522,1546]" pageId="21" pageNumber="472">SDBDU</collectionCode>
|
||
2007.1569
|
||
</specimenCode>
|
||
(only tissue sample, voucher not collected), from Koyna (
|
||
<geoCoordinate id="EE0F5066313FFF8FFC35B4C0D186F9A5" box="[1023,1200,1522,1546]" direction="north" orientation="latitude" pageId="21" pageNumber="472" precision="1" value="17.405066">17°24’18.24’’N</geoCoordinate>
|
||
<geoCoordinate id="EE0F5066313FFF8FFB7DB4C0D050F9A5" box="[1207,1382,1522,1546]" direction="east" orientation="longitude" pageId="21" pageNumber="472" precision="1" value="73.73831">73°44’17.93’’E</geoCoordinate>
|
||
,
|
||
<elevation id="0016D192313FFF8FFAB8B4C0D5EEF981" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="21" pageNumber="472" unit="m" value="820.0">
|
||
<quantity id="4CC39B44313FFF8FFAB8B4C0D59BF982" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="21" pageNumber="472" unit="m" value="820.0">820 m</quantity>
|
||
asl
|
||
</elevation>
|
||
),
|
||
<location id="8EE4607A313FFF8FFF24B724D4B8F981" LSID="urn:lsid:plazi:treatment:039287B7313FFF82FF5DB280D464FCBB:8EE4607A313FFF8FFF24B724D4B8F981" box="[238,398,1557,1582]" country="India" latitude="17.405066" longLatPrecision="1" longitude="73.73831" municipality="These" name="Satara district" pageId="21" pageNumber="472" stateProvince="Maharashtra">Satara district</location>
|
||
,
|
||
<collectingRegion id="49FFF843313FFF8FFE51B727D711F981" box="[411,551,1557,1582]" country="India" name="Maharashtra" pageId="21" pageNumber="472">Maharashtra</collectingRegion>
|
||
state,
|
||
<collectingCountry id="F32C7631313FFF8FFDB9B727D799F981" box="[627,687,1557,1582]" name="India" pageId="21" pageNumber="472">India</collectingCountry>
|
||
, collected by SDB on
|
||
<collectingDate id="EFC1E989313FFF8FFC7FB727D146F981" box="[949,1136,1557,1582]" pageId="21" pageNumber="472" value="2007-08-25">
|
||
<date id="FF851061313FFF8FFC7FB727D146F981" box="[949,1136,1557,1582]" pageId="21" pageNumber="472" value="2007-08-25">25 August 2007</date>
|
||
</collectingDate>
|
||
.
|
||
<collectingMunicipality id="6BE0ACDB313FFF8FFBB6B727D189F981" box="[1148,1215,1557,1582]" pageId="21" pageNumber="472">These</collectingMunicipality>
|
||
samples were only used for genetic identification.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B8436A1313FFF8FFF0DB76FD6A2F8A9" blockId="21.[151,1437,1125,2014]" pageId="21" pageNumber="472">
|
||
<emphasis id="B94FEAB3313FFF8FFF0DB76FD4F6F9D9" bold="true" box="[199,448,1629,1654]" pageId="21" pageNumber="472">Genetic relationship.</emphasis>
|
||
Phylogenetically,
|
||
<taxonomicName id="4C3B4D22313FFF8FFD5AB76CD67CF9D9" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[656,842,1630,1654]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="21" pageNumber="472" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313FFF8FFD5AB76CD67CF9D9" box="[656,842,1630,1654]" italics="true" pageId="21" pageNumber="472">Fejervarya cepfi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313FFF8FFC99B76FD69BF9D9" box="[851,941,1629,1654]" pageId="21" pageNumber="472" rank="species">
|
||
<emphasis id="B94FEAB3313FFF8FFC99B76FD69BF9D9" bold="true" box="[851,941,1629,1654]" pageId="21" pageNumber="472">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is nested in the
|
||
<taxonomicName id="4C3B4D22313FFF8FFBAFB76CD065F9D9" authorityName="Jerdon" authorityYear="1853" box="[1125,1363,1630,1654]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="21" pageNumber="472" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis id="B94FEAB3313FFF8FFBAFB76CD065F9D9" box="[1125,1363,1630,1654]" italics="true" pageId="21" pageNumber="472">Fejervarya rufescens</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<figureCitation id="13002A24313FFF8FFF55B7B3D43AF935" box="[159,268,1665,1690]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1885,1907]" captionStartId-1="5.[151,250,1618,1640]" captionTargetBox-0="[161,1435,204,1854]" captionTargetBox-1="[155,1424,193,1582]" captionTargetId-0="figure@4.[151,1436,193,1864]" captionTargetId-1="figure@5.[151,1436,193,1597]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Maximum Likelihood phylogram based on the 16 S mitochondrial DNA dataset of 545 basepairs. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of> 50 % are indicated above and below the branches, respectively. The uncorrected pairwise distances between species in the Fejervarya rufescens group are shown near the clade labels. Voucher number and locality details of the samples are referenced in Table 1." captionText-1="FIGURE 2. Phylogenetic analyses. A. Maximum Likelihood phylogram based on mitochondrial (16 S, COI and Cytb) and nuclear (Rag 1) DNA of 2314 basepairs, showing phylogenetic relationships among members of the Fejervarya rufescens group. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of> 50 % are indicated above and below the branches, respectively. Fejervarya granosa (2) was used as the outgroup taxon. Voucher number and locality details of the samples are referenced in Table 1; B. Median Joining haplotype network inferred from 554 basepair fragment of the nuclear Rag 1 gene. Circle sizes are proportional to the number of haplotypes, branch lengths are proportional to the number of mutational steps, and black bars represent mutational steps. Number of individuals used for each species: Fejervarya cepfi (3), Fejervarya kadar (1), Fejervarya manoharani (3), Fejervarya neilcoxi (2), and Fejervarya rufescens (5)." httpUri-0="https://zenodo.org/record/829835/files/figure.png" httpUri-1="https://zenodo.org/record/829837/files/figure.png" pageId="21" pageNumber="472">Figs. 1–2</figureCitation>
|
||
) of the
|
||
<collectingRegion id="49FFF843313FFF8FFEA4B7B0D4FFF936" box="[366,457,1666,1689]" country="Sri Lanka" name="Western" pageId="21" pageNumber="472">Western</collectingRegion>
|
||
Ghats. The average uncorrected pairwise genetic divergences with
|
||
<taxonomicName id="4C3B4D22313FFF8FFB14B7B0D051F935" authorityName="Jerdon" authorityYear="1853" box="[1246,1383,1666,1690]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="21" pageNumber="472" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis id="B94FEAB3313FFF8FFB14B7B0D051F935" box="[1246,1383,1666,1690]" italics="true" pageId="21" pageNumber="472">F. rufescens</emphasis>
|
||
</taxonomicName>
|
||
are: 4.5% (range 3.9–5.2%,
|
||
<emphasis id="B94FEAB3313FFF8FFE7CB795D4FFF911" box="[438,457,1703,1726]" italics="true" pageId="21" pageNumber="472">N</emphasis>
|
||
= 28) for 16S, 11.3% (range 10.7–11.9%,
|
||
<emphasis id="B94FEAB3313FFF8FFC2AB795D6C5F911" box="[992,1011,1703,1726]" italics="true" pageId="21" pageNumber="472">N</emphasis>
|
||
= 15) for
|
||
<collectionCode id="ED2AAE64313FFF8FFB4BB794D182F911" box="[1153,1204,1702,1726]" country="Portugal" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="21" pageNumber="472">COI</collectionCode>
|
||
, and 13.1% (range 12.7–13.9%,
|
||
<emphasis id="B94FEAB3313FFF8FFEE4B7F8D477F94E" box="[302,321,1738,1761]" italics="true" pageId="21" pageNumber="472">N</emphasis>
|
||
= 15) for Cytb (Table 2). For comparison with
|
||
<taxonomicName id="4C3B4D22313FFF8FFCA2B7F8D6F1F94D" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[872,967,1737,1762]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="21" pageNumber="472" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313FFF8FFCA2B7F8D6F1F94D" box="[872,967,1737,1762]" italics="true" pageId="21" pageNumber="472">F. kadar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313FFF8FFC1AB7FBD11DF94D" box="[976,1067,1737,1762]" pageId="21" pageNumber="472" rank="species">
|
||
<emphasis id="B94FEAB3313FFF8FFC1AB7FBD11DF94D" bold="true" box="[976,1067,1737,1762]" pageId="21" pageNumber="472">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C3B4D22313FFF8FFBF1B7F8D1D4F94D" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[1083,1250,1737,1762]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="21" pageNumber="472" phylum="Chordata" rank="species" species="manoharani" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313FFF8FFBF1B7F8D1D4F94D" box="[1083,1250,1737,1762]" italics="true" pageId="21" pageNumber="472">F. manoharani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313FFF8FFB21B7FBD070F94D" box="[1259,1350,1737,1762]" pageId="21" pageNumber="472" rank="species">
|
||
<emphasis id="B94FEAB3313FFF8FFB21B7FBD070F94D" bold="true" box="[1259,1350,1737,1762]" pageId="21" pageNumber="472">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C3B4D22313FFF8FFA4DB7F8D5C6F8A9" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="21" pageNumber="472" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313FFF8FFA4DB7F8D5C6F8A9" italics="true" pageId="21" pageNumber="472">F. neilcoxi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313FFF8FFF3CB7DFD466F8A9" box="[246,336,1773,1798]" pageId="21" pageNumber="472" rank="species">
|
||
<emphasis id="B94FEAB3313FFF8FFF3CB7DFD466F8A9" bold="true" box="[246,336,1773,1798]" pageId="21" pageNumber="472">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, see ‘Genetic relationship’ section of those species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321652A313FFF8DFF0DB623D466FC29" lastPageId="23" lastPageNumber="474" pageId="21" pageNumber="472" type="diagnosis">
|
||
<paragraph id="8B8436A1313FFF8CFF0DB623D691FF57" blockId="21.[151,1437,1125,2014]" lastBlockId="22.[151,1436,151,248]" lastPageId="22" lastPageNumber="473" pageId="21" pageNumber="472">
|
||
<emphasis id="B94FEAB3313FFF8FFF0DB623D477F885" bold="true" box="[199,321,1809,1834]" pageId="21" pageNumber="472">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C3B4D22313FFF8FFE98B620D723F885" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[338,533,1810,1834]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="21" pageNumber="472" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313FFF8FFE98B620D723F885" box="[338,533,1810,1834]" italics="true" pageId="21" pageNumber="472">Fejervarya cepfi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313FFF8FFDEDB623D7BDF885" box="[551,651,1809,1834]" pageId="21" pageNumber="472" rank="species">
|
||
<emphasis id="B94FEAB3313FFF8FFDEDB623D7BDF885" bold="true" box="[551,651,1809,1834]" pageId="21" pageNumber="472">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from known congeners by the following combination of morphological characters: (1) medium male adult size (SVL
|
||
<quantity id="4CC39B44313FFF8FFC3BB604D1A0F8E2" box="[1009,1174,1846,1870]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.31" metricValueMin="2.9899999999999998" pageId="21" pageNumber="472" unit="mm" value="31.5" valueMax="33.1" valueMin="29.9">29.9–33.1 mm</quantity>
|
||
,
|
||
<emphasis id="B94FEAB3313FFF8FFB6BB605D182F8E1" box="[1185,1204,1847,1870]" italics="true" pageId="21" pageNumber="472">N</emphasis>
|
||
= 2); (2) stout body; (3) snout subovoid in dorsal view and obtuse in lateral view; (4) presence of rictal gland at labial commissure of the mouth; (5) eye length shorter than snout length (male EL/SL ratio 72.9–73.9%,
|
||
<emphasis id="B94FEAB3313FFF8FFBFBB64DD172F839" box="[1073,1092,1919,1942]" italics="true" pageId="21" pageNumber="472">N</emphasis>
|
||
= 2); (6) tympanum diameter nearly half of eye length, (male TYD/EL ratio 51.4–52.9%,
|
||
<emphasis id="B94FEAB3313FFF8FFCF3B690D67AF816" box="[825,844,1954,1977]" italics="true" pageId="21" pageNumber="472">N</emphasis>
|
||
= 2); (7) inter upper eyelid width nearly equal to the upper eyelid width (male IUE/UEW ratio 96.0–96.2%,
|
||
<emphasis id="B94FEAB3313FFF8FFCFEB6F5D671F871" box="[820,839,1991,2014]" italics="true" pageId="21" pageNumber="472">N</emphasis>
|
||
= 2) and internarial distance (male IUE/IN ratio 96–100%,
|
||
<emphasis id="B94FEAB3313CFF8CFED9B1ABD410FF1F" box="[275,294,153,176]" italics="true" pageId="22" pageNumber="473">N</emphasis>
|
||
= 2); (8) thigh length shorter than shank length (male TL/SHL ratio 93.9–94.3%,
|
||
<emphasis id="B94FEAB3313CFF8CFB2FB1ABD1CEFF1F" box="[1253,1272,153,176]" italics="true" pageId="22" pageNumber="473">N</emphasis>
|
||
= 2) and foot length (male TL/FOL ratio 89.7–90.1%,
|
||
<emphasis id="B94FEAB3313CFF8CFDAFB18FD74EFF7B" box="[613,632,189,212]" italics="true" pageId="22" pageNumber="473">N</emphasis>
|
||
= 2); (9) prominent shovel-shaped inner metatarsal tubercle prominent and small outer metatarsal tubercle; (10) webbing between toes small.
|
||
</paragraph>
|
||
<caption id="DF446629313CFF8CFF5DB64FD6E6F87F" httpUri="https://zenodo.org/record/829851/files/figure.png" pageId="22" pageNumber="473" targetBox="[151,1436,302,1896]" targetPageId="22">
|
||
<paragraph id="8B8436A1313CFF8CFF5DB64FD6E6F87F" blockId="22.[151,1436,1917,2000]" pageId="22" pageNumber="473">
|
||
<emphasis id="B94FEAB3313CFF8CFF5DB64FD422F83D" bold="true" box="[151,276,1917,1939]" pageId="22" pageNumber="473">FIGURE 9.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C3B4D22313CFF8CFE73B64CD755F83C" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[441,611,1917,1939]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="22" pageNumber="473" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313CFF8CFE73B64CD755F83C" box="[441,611,1917,1939]" italics="true" pageId="22" pageNumber="473">Fejervarya cepfi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313CFF8CFDA0B64CD78AF83C" box="[618,700,1918,1939]" pageId="22" pageNumber="473" rank="species">
|
||
<emphasis id="B94FEAB3313CFF8CFDA0B64CD78AF83C" bold="true" box="[618,700,1918,1939]" pageId="22" pageNumber="473">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(ZSI/WGRC/V/A/937):
|
||
<emphasis id="B94FEAB3313CFF8CFC77B64FD6E0F83D" bold="true" box="[957,982,1917,1938]" pageId="22" pageNumber="473">A.</emphasis>
|
||
dorsolateral view, in life;
|
||
<emphasis id="B94FEAB3313CFF8CFB2CB64CD1CBF83C" bold="true" box="[1254,1277,1918,1939]" pageId="22" pageNumber="473">B.</emphasis>
|
||
dorsal view, in preservation;
|
||
<emphasis id="B94FEAB3313CFF8CFEEBB6AED40CF81D" bold="true" box="[289,314,1948,1970]" pageId="22" pageNumber="473">C.</emphasis>
|
||
ventral view, in preservation;
|
||
<emphasis id="B94FEAB3313CFF8CFDA6B6AED7B3F81E" bold="true" box="[620,645,1948,1969]" pageId="22" pageNumber="473">D.</emphasis>
|
||
lateral view of head, in preservation;
|
||
<emphasis id="B94FEAB3313CFF8CFBCEB6AED12DF81E" bold="true" box="[1028,1051,1948,1969]" pageId="22" pageNumber="473">E.</emphasis>
|
||
ventral view of hand, in preservation;
|
||
<emphasis id="B94FEAB3313CFF8CFF5DB689D59EF87F" bold="true" box="[151,168,1979,2000]" pageId="22" pageNumber="473">F.</emphasis>
|
||
ventral view of foot, in preservation;
|
||
<emphasis id="B94FEAB3313CFF8CFDE0B688D708F87F" bold="true" box="[554,574,1978,2000]" pageId="22" pageNumber="473">G.</emphasis>
|
||
schematic illustration of foot webbing.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8436A1313DFF8DFF0DB1A5D7A4FF57" blockId="23.[151,1437,151,2028]" pageId="23" pageNumber="474">
|
||
<emphasis id="B94FEAB3313DFF8DFF0DB1A5D725FF1F" bold="true" box="[199,531,151,176]" pageId="23" pageNumber="474">Morphological comparison.</emphasis>
|
||
Based on the overall morphology and comparable body size,
|
||
<taxonomicName id="4C3B4D22313DFF8DFB2AB1ABD0ADFF1F" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[1248,1435,152,176]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313DFF8DFB2AB1ABD0ADFF1F" box="[1248,1435,152,176]" italics="true" pageId="23" pageNumber="474">Fejervarya cepfi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313DFF8DFF5DB18ED5C3FF7A" box="[151,245,188,213]" pageId="23" pageNumber="474" rank="species">
|
||
<emphasis id="B94FEAB3313DFF8DFF5DB18ED5C3FF7A" bold="true" box="[151,245,188,213]" pageId="23" pageNumber="474">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
could be confused with the known species
|
||
<taxonomicName id="4C3B4D22313DFF8DFCC9B18FD6BBFF7B" authorityName="Jerdon" authorityYear="1853" box="[771,909,188,212]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis id="B94FEAB3313DFF8DFCC9B18FD6BBFF7B" box="[771,909,188,212]" italics="true" pageId="23" pageNumber="474">F. rufescens</emphasis>
|
||
</taxonomicName>
|
||
and three new species
|
||
<taxonomicName id="4C3B4D22313DFF8DFB62B18FD03CFF7A" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[1192,1290,188,213]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313DFF8DFB62B18FD03CFF7A" box="[1192,1290,188,213]" italics="true" pageId="23" pageNumber="474">F. kadar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313DFF8DFADCB18ED043FF7A" box="[1302,1397,188,213]" pageId="23" pageNumber="474" rank="species">
|
||
<emphasis id="B94FEAB3313DFF8DFADCB18ED043FF7A" bold="true" box="[1302,1397,188,213]" pageId="23" pageNumber="474">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C3B4D22313DFF8DFA4DB18FD416FF57" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="manoharani" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313DFF8DFA4DB18FD416FF57" italics="true" pageId="23" pageNumber="474">F. manoharani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313DFF8DFEEDB1EDD4B6FF57" box="[295,384,223,248]" pageId="23" pageNumber="474" rank="species">
|
||
<emphasis id="B94FEAB3313DFF8DFEEDB1EDD4B6FF57" bold="true" box="[295,384,223,248]" pageId="23" pageNumber="474">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C3B4D22313DFF8DFE77B1D3D704FF57" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[445,562,223,248]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
|
||
<emphasis id="B94FEAB3313DFF8DFE77B1D3D704FF57" box="[445,562,223,248]" italics="true" pageId="23" pageNumber="474">F. neilcoxi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C8313DFF8DFDF3B1EDD7A4FF57" box="[569,658,223,248]" pageId="23" pageNumber="474" rank="species">
|
||
<emphasis id="B94FEAB3313DFF8DFDF3B1EDD7A4FF57" bold="true" box="[569,658,223,248]" pageId="23" pageNumber="474">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8B8436A1313DFF8DFF0DB037D07CFC94" blockId="23.[151,1437,151,2028]" pageId="23" pageNumber="474">
|
||
However,
|
||
<taxonomicName id="4C3B4D22313DFF8DFEF4B037D4CDFEB3" authorityName="Garg & Biju" authorityYear="2017" box="[318,507,260,284]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="cepfi">
|
||
<emphasis id="B94FEAB3313DFF8DFEF4B037D4CDFEB3" box="[318,507,260,284]" italics="true" pageId="23" pageNumber="474">Fejervarya cepfi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3B4D22313DFF8DFD51B037D613FEB3" authorityName="Jerdon" authorityYear="1853" box="[667,805,260,284]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis id="B94FEAB3313DFF8DFD51B037D613FEB3" box="[667,805,260,284]" italics="true" pageId="23" pageNumber="474">F. rufescens</emphasis>
|
||
</taxonomicName>
|
||
by its snout obtuse in lateral view (vs. rounded); eye length relatively shorter compared to snout length, male EL
|
||
<quantity id="4CC39B44313DFF8DFCFCB015D689FE90" box="[822,959,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="23" pageNumber="474" unit="mm" value="3.45" valueMax="3.5" valueMin="3.4">3.4–3.5 mm</quantity>
|
||
, SL 4.6–4.8, EL/SL ratio 72.9–73.9%,
|
||
<emphasis id="B94FEAB3313DFF8DFAB9B01BD0B0FEEF" box="[1395,1414,297,320]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 2 (vs. relatively longer, male EL
|
||
<quantity id="4CC39B44313DFF8DFDECB07ED785FECB" box="[550,691,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.9" metricValueMin="3.9" pageId="23" pageNumber="474" unit="mm" value="4.4" valueMax="4.9" valueMin="3.9">3.9–4.9 mm</quantity>
|
||
, SL 4.2–5.0 mm, EL/SL ratio 88.6–98.0%,
|
||
<emphasis id="B94FEAB3313DFF8DFB75B07FD1E4FECB" box="[1215,1234,333,356]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 6); tympanum diameter relatively larger compared to eye length, male TYD
|
||
<quantity id="4CC39B44313DFF8DFC81B042D695FE28" box="[843,931,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="474" unit="mm" value="1.8">1.8 mm</quantity>
|
||
, SL 4.6–4.8, TYD/EL ratio 51.4–52.9%,
|
||
<emphasis id="B94FEAB3313DFF8DFAB8B043D0B3FE27" box="[1394,1413,369,392]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 2 (vs. relatively smaller, male TYD
|
||
<quantity id="4CC39B44313DFF8DFDE3B0A6D784FE03" box="[553,690,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="23" pageNumber="474" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.6–2.2 mm</quantity>
|
||
, SL 4.2–5.0 mm, TYD/EL ratio 39.5–46.2%,
|
||
<emphasis id="B94FEAB3313DFF8DFB78B0A7D1F3FE03" box="[1202,1221,405,428]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 6); tympanum to eye distance relatively larger compared to tympanum diameter, male TYE 1.5–1.6, TYD
|
||
<quantity id="4CC39B44313DFF8DFB4FB08AD1EBFE60" box="[1157,1245,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="474" unit="mm" value="1.8">1.8 mm</quantity>
|
||
, TYE/TYD ratio 83.3–88.9%,
|
||
<emphasis id="B94FEAB3313DFF8DFEE6B0EFD409FE5B" box="[300,319,477,500]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 2 (vs. relatively smaller, male TYE
|
||
<quantity id="4CC39B44313DFF8DFD26B0EED643FE5B" box="[748,885,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="23" pageNumber="474" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 mm</quantity>
|
||
, TYD
|
||
<quantity id="4CC39B44313DFF8DFC0BB0EED17FFE5B" box="[961,1097,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="23" pageNumber="474" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.6–2.2 mm</quantity>
|
||
, TYE/TYD ratio 50.0–76.5%,
|
||
<emphasis id="B94FEAB3313DFF8DFF5DB333D59CFDB7" box="[151,170,513,536]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 6); inter upper eyelid width nearly equal to the upper eyelid width, male IUE
|
||
<quantity id="4CC39B44313DFF8DFBF8B0CDD18DFDB8" box="[1074,1211,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="23" pageNumber="474" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
|
||
, UEW
|
||
<quantity id="4CC39B44313DFF8DFAC4B0CDD0AEFDB8" box="[1294,1432,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="23" pageNumber="474" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 mm</quantity>
|
||
, IUE/UEW ratio 96.0–96.2%,
|
||
<emphasis id="B94FEAB3313DFF8DFE25B317D734FD93" box="[495,514,549,572]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 2 (vs. narrower, male IUE
|
||
<quantity id="4CC39B44313DFF8DFCAFB316D6C4FD93" box="[869,1010,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="23" pageNumber="474" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.4–1.8 mm</quantity>
|
||
, UEW
|
||
<quantity id="4CC39B44313DFF8DFB81B316D1EEFD93" box="[1099,1240,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="23" pageNumber="474" unit="mm" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 mm</quantity>
|
||
, IUE/UEW ratio 42.9–58.6%,
|
||
<emphasis id="B94FEAB3313DFF8DFEE4B37BD477FDCF" box="[302,321,585,608]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 6); inter upper eyelid width nearly equal to internarial distance, male IUE
|
||
<quantity id="4CC39B44313DFF8DFB70B375D073FDF0" box="[1210,1349,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="23" pageNumber="474" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
|
||
, IN
|
||
<quantity id="4CC39B44313DFF8DFAB3B375D5F3FD2B" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="474" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, IUE/IN ratio 96–100%,
|
||
<emphasis id="B94FEAB3313DFF8DFE34B35FD727FD2B" box="[510,529,621,644]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 2 (vs. narrower, male IUE
|
||
<quantity id="4CC39B44313DFF8DFC5BB35ED114FD2B" box="[913,1058,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="23" pageNumber="474" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.4–1.8 mm</quantity>
|
||
, IN
|
||
<quantity id="4CC39B44313DFF8DFBABB35ED1C4FD2B" box="[1121,1266,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.2" metricValueMin="2.4" pageId="23" pageNumber="474" unit="mm" value="2.8" valueMax="3.2" valueMin="2.4">2.4–3.2 mm</quantity>
|
||
, IUE/IN ratio 46.9–72.0%,
|
||
<emphasis id="B94FEAB3313DFF8DFEE6B3A3D409FD07" box="[300,319,657,680]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 6); thigh length shorter than shank length, male TL
|
||
<quantity id="4CC39B44313DFF8DFC6AB3BDD173FD08" box="[928,1093,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" metricValueMax="1.54" metricValueMin="1.48" pageId="23" pageNumber="474" unit="mm" value="15.1" valueMax="15.4" valueMin="14.8">14.8–15.4 mm</quantity>
|
||
, SHL
|
||
<quantity id="4CC39B44313DFF8DFB47B3BDD004FD08" box="[1165,1330,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="1.6049999999999998" metricValueMax="1.64" metricValueMin="1.5699999999999998" pageId="23" pageNumber="474" unit="mm" value="16.049999999999997" valueMax="16.4" valueMin="15.7">15.7–16.4 mm</quantity>
|
||
, TL/SHL ratio 93.9–94.3%,
|
||
<emphasis id="B94FEAB3313DFF8DFEAFB387D44EFD63" box="[357,376,693,716]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 2 (vs. nearly equal, male TL
|
||
<quantity id="4CC39B44313DFF8DFD1CB386D64DFD63" box="[726,891,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.525" metricValueMax="1.58" metricValueMin="1.47" pageId="23" pageNumber="474" unit="mm" value="15.25" valueMax="15.8" valueMin="14.7">14.7–15.8 mm</quantity>
|
||
, SHL
|
||
<quantity id="4CC39B44313DFF8DFC0BB386D150FD63" box="[961,1126,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.515" metricValueMax="1.58" metricValueMin="1.45" pageId="23" pageNumber="474" unit="mm" value="15.15" valueMax="15.8" valueMin="14.5">14.5–15.8 mm</quantity>
|
||
, TL/SHL 100–102.1%,
|
||
<emphasis id="B94FEAB3313DFF8DFAB9B387D0B0FD63" box="[1395,1414,693,716]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 6); thigh length shorter than foot length, male TL
|
||
<quantity id="4CC39B44313DFF8DFD06B3E5D647FD40" box="[716,881,727,752]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" metricValueMax="1.54" metricValueMin="1.48" pageId="23" pageNumber="474" unit="mm" value="15.1" valueMax="15.4" valueMin="14.8">14.8–15.4 mm</quantity>
|
||
, FOL
|
||
<quantity id="4CC39B44313DFF8DFC70B3E5D156FD40" box="[954,1120,727,752]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" metricValueMax="1.71" metricValueMin="1.65" pageId="23" pageNumber="474" unit="mm" value="16.8" valueMax="17.1" valueMin="16.5">16.5–17.1 mm</quantity>
|
||
, TL/FOL ratio 89.7–90.1%,
|
||
<emphasis id="B94FEAB3313DFF8DFF5DB3CFD59CFCBB" box="[151,170,765,788]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 2 (vs. nearly equal, male TL
|
||
<quantity id="4CC39B44313DFF8DFDC5B3CED782FCBB" box="[527,692,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.525" metricValueMax="1.58" metricValueMin="1.47" pageId="23" pageNumber="474" unit="mm" value="15.25" valueMax="15.8" valueMin="14.7">14.7–15.8 mm</quantity>
|
||
, FOL
|
||
<quantity id="4CC39B44313DFF8DFD37B3CED694FCBB" box="[765,930,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.59" metricValueMin="1.48" pageId="23" pageNumber="474" unit="mm" value="15.35" valueMax="15.9" valueMin="14.8">14.8–15.9 mm</quantity>
|
||
, TL/FOL 98.0–99.4%,
|
||
<emphasis id="B94FEAB3313DFF8DFB60B3CFD18BFCBB" box="[1194,1213,765,788]" italics="true" pageId="23" pageNumber="474">N</emphasis>
|
||
= 6); and relatively more webbing between toes, male
|
||
<date id="FF851061313DFF8DFDD6B211D705FC94" box="[540,563,803,827]" pageId="23" pageNumber="474">I1</date>
|
||
+–2–
|
||
<date id="FF851061313DFF8DFD95B211D749FC94" box="[607,639,803,827]" pageId="23" pageNumber="474">II1</date>
|
||
+–3–III2–
|
||
<date id="FF851061313DFF8DFD22B211D622FC94" box="[744,788,803,827]" pageId="23" pageNumber="474">3IV</date>
|
||
3–1 1/
|
||
<date id="FF851061313DFF8DFC83B203D652FC95" box="[841,868,803,831]" pageId="23" pageNumber="474">2V</date>
|
||
(vs. less, male
|
||
<date id="FF851061313DFF8DFBDBB211D11FFC94" box="[1041,1065,803,827]" pageId="23" pageNumber="474">I2</date>
|
||
– –
|
||
<date id="FF851061313DFF8DFBF5B211D156FC94" box="[1087,1120,803,827]" pageId="23" pageNumber="474">2II</date>
|
||
2– –3–III2–
|
||
<date id="FF851061313DFF8DFB1BB211D1CDFC94" box="[1233,1275,803,827]" pageId="23" pageNumber="474">3IV</date>
|
||
3–
|
||
<date id="FF851061313DFF8DFAD3B211D008FC94" box="[1305,1342,803,827]" pageId="23" pageNumber="474">2V</date>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8436A1313DFF8DFF0DB278D466FC29" blockId="23.[151,1437,151,2028]" pageId="23" pageNumber="474">
|
||
For differences with
|
||
<taxonomicName id="4C3B4D22313DFF8DFE65B278D744FCCD" authorityName="Garg & Biju" authorityYear="2017" box="[431,626,841,866]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="kadar">
|
||
<emphasis id="B94FEAB3313DFF8DFE65B278D744FCCD" box="[431,626,841,866]" italics="true" pageId="23" pageNumber="474">Fejervarya kadar</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D22313DFF8DFDB5B278D615FCCD" authorityName="Garg & Biju" authorityYear="2017" box="[639,803,841,866]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="manoharani">
|
||
<emphasis id="B94FEAB3313DFF8DFDB5B278D615FCCD" box="[639,803,841,866]" italics="true" pageId="23" pageNumber="474">F. manoharani</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3B4D22313DFF8DFC93B278D6FBFCCD" authorityName="Garg & Biju" authorityYear="2017" box="[857,973,841,866]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="neilcoxi">
|
||
<emphasis id="B94FEAB3313DFF8DFC93B278D6FBFCCD" box="[857,973,841,866]" italics="true" pageId="23" pageNumber="474">F. neilcoxi</emphasis>
|
||
</taxonomicName>
|
||
, see ‘Morphological comparison’ section of those species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321652A313DFF82FF0DB2A3D1C8FE7F" lastPageId="24" lastPageNumber="475" pageId="23" pageNumber="474" type="description">
|
||
<paragraph id="8B8436A1313DFF8DFF0DB2A3D49AF9B8" blockId="23.[151,1437,151,2028]" pageId="23" pageNumber="474">
|
||
<emphasis id="B94FEAB3313DFF8DFF0DB2A3D7CEFC05" bold="true" box="[199,760,913,938]" pageId="23" pageNumber="474">
|
||
Description of
|
||
<typeStatus id="54808803313DFF8DFEB2B2A3D4E9FC05" box="[376,479,913,938]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
|
||
<emphasis id="B94FEAB3313DFF8DFE2CB2A3D7C6FC05" bold="true" box="[486,752,913,938]" italics="true" pageId="23" pageNumber="474">(measurements in mm)</emphasis>
|
||
.
|
||
</emphasis>
|
||
Adult male (SVL 29.9), rather stout; head nearly as long as wide (HW 11.2, HL 11.1); snout subovoid in dorsal view, obtuse in lateral view, snout length (SL 4.6) longer than horizontal diameter of eye (EL 3.4); loreal region acute, rounded canthus rostralis; interorbital space flat, nearly as wide (IUE 2.4) as upper eyelid (UEW 2.5) and internarial distance (IN 2.5); nostril oval, as close to snout (NS 1.6) as to eye (EN 1.6); tympanum (TYD 1.8) 52.9% of eye diameter (EL 3.4); tympanum-eye distance (TYE 1.5), 83.3% of the tympanum diameter (TYD 1.8); supratympanic fold well developed, extends from posterior corner of eye to near the shoulder; vomerine ridge present, bearing small teeth, at an angle of 45° to the body axis, as close to choanae as to each other; tongue moderately large, emarginated, bearing no median lingual process; rictal gland present at labial commissure of the mouth. Arms short, forearm length (FAL 6.2) shorter then the hand (HAL 7.2); relative length of fingers IV<II<I<III (FL I 3.6, FL II 2.6, FL III 4.3, FL IV 2.2); finger tips rounded, slightly enlarged without discs, fingers without fringes, webbing between fingers absent; subarticular tubercles prominent, circular; one distinct palmar tubercle, oval, bifid; supernumerary tubercles absent. Hind limbs short, thigh (TL 14.8) shorter than shank (SHL 15.7) and foot (FOL 16.5); distance from the base of tarsus to the tip of toe IV (TFOL 23.5); toes long, relative length of toes I<II<III<V<IV; toe tips rounded, slightly enlarged without discs, toes without fringes, webbing between toes small:
|
||
<date id="FF851061313DFF8DFE2EB4A2D4CCFA07" box="[484,506,1424,1448]" pageId="23" pageNumber="474">I1</date>
|
||
+–2–
|
||
<date id="FF851061313DFF8DFDEDB4A2D771FA07" box="[551,583,1424,1448]" pageId="23" pageNumber="474">II1</date>
|
||
+–3–III2–
|
||
<date id="FF851061313DFF8DFD7AB4A2D7EAFA07" box="[688,732,1424,1448]" pageId="23" pageNumber="474">3IV</date>
|
||
3–1 1/
|
||
<date id="FF851061313DFF8DFCDAB4ADD618FA07" box="[784,814,1425,1453]" pageId="23" pageNumber="474">2V</date>
|
||
; inner toe length (ITL 2.9); minute spinular projections on the outer margins of toe V; subarticular tubercles prominent, all present, circular; inner metatarsal tubercle prominent (IMT 2.0), shovel-shaped; outer metatarsal tubercle, small (OMT 0.5), rounded; supernumerary tubercles absent (
|
||
<figureCitation id="13002A24313DFF8DFE90B4CDD4A8F9B8" box="[346,414,1535,1560]" captionStart="FIGURE 9" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[151,1436,302,1896]" captionTargetId="figure@22.[151,1437,301,1896]" captionTargetPageId="22" captionText="FIGURE 9. Holotype of Fejervarya cepfi sp. nov. (ZSI / WGRC / V / A / 937): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829851/files/figure.png" pageId="23" pageNumber="474">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8436A1313DFF8DFF0DB711D407F964" blockId="23.[151,1437,151,2028]" pageId="23" pageNumber="474">
|
||
Skin of snout shagreened with scattered granular projections, upper eyelids prominently tuberculate, anterior and posterior parts of back, and upper and lower parts of flank shagreened with prominent granular projections and scattered larger glandular warts; dorsal surfaces of forelimb, thigh and shank shagreened with prominent granular projections. Ventral surface of throat, chest, belly and limbs smooth; fejervaryan line present on both sides of the belly (
|
||
<figureCitation id="13002A24313DFF8DFF14B781D415F964" box="[222,291,1715,1740]" captionStart="FIGURE 9" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[151,1436,302,1896]" captionTargetId="figure@22.[151,1437,301,1896]" captionTargetPageId="22" captionText="FIGURE 9. Holotype of Fejervarya cepfi sp. nov. (ZSI / WGRC / V / A / 937): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829851/files/figure.png" pageId="23" pageNumber="474">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8436A1313DFF82FF0DB7E5D461FEB3" blockId="23.[151,1437,151,2028]" lastBlockId="24.[151,1437,151,789]" lastPageId="24" lastPageNumber="475" pageId="23" pageNumber="474">
|
||
<emphasis id="B94FEAB3313DFF8DFF0DB7E5D49BF95F" bold="true" box="[199,429,1751,1776]" pageId="23" pageNumber="474">
|
||
Colour of
|
||
<typeStatus id="54808803313DFF8DFEF5B7E5D49EF95F" box="[319,424,1751,1776]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B94FEAB3313DFF8DFE7EB7EAD4CBF95F" box="[436,509,1751,1776]" italics="true" pageId="23" pageNumber="474">In life.</emphasis>
|
||
Dorsum, upper eyelids and lateral side of snout brick red with prominent greenishbrown markings (
|
||
<figureCitation id="13002A24313DFF8DFEAEB7C9D49CF8BC" box="[356,426,1787,1812]" captionStart="FIGURE 9" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[151,1436,302,1896]" captionTargetId="figure@22.[151,1437,301,1896]" captionTargetPageId="22" captionText="FIGURE 9. Holotype of Fejervarya cepfi sp. nov. (ZSI / WGRC / V / A / 937): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829851/files/figure.png" pageId="23" pageNumber="474">Fig. 9</figureCitation>
|
||
A); lateral surfaces of head light yellowish-brown; upper and lower lip with prominent greenish-brown cross bands; tympanum light yellowish-brown with a greenish-brown patch; forelimbs and hind limbs (including toes) light brick red with greenish-brown transverse bands; anterior parts of flank light greenishbrown and posterior parts light greenish-brown to yellow; groin light yellow; webbing light brown; anterior parts of thigh light yellowish with faint grey reticulation. Ventral surface of throat light flesh colour with minute dark brown speckles and two lateral black calling patches; belly white; forearm and foreleg light flesh red in colour with dark brown mottling on the margins.
|
||
<emphasis id="B94FEAB3313DFF8DFDA9B6E6D623F844" box="[611,789,2004,2027]" italics="true" pageId="23" pageNumber="474">In preservation</emphasis>
|
||
. Dorsum greyish-brown with blackish-brown patches, forelimbs and hind limbs light brown with dark greyish-brown transverse bands, fingers yellowish-white with minute dark brown speckles on the third and fourth fingers. Ventral surface of throat light grey with dark grey speckles and two lateral black calling patches; belly off-white; margins of the limbs light grey with greyish-brown mottling (
|
||
<figureCitation id="13002A243132FF82FECFB036D47FFEB3" box="[261,329,260,285]" captionStart="FIGURE 9" captionStartId="22.[151,250,1917,1939]" captionTargetBox="[151,1436,302,1896]" captionTargetId="figure@22.[151,1437,301,1896]" captionTargetPageId="22" captionText="FIGURE 9. Holotype of Fejervarya cepfi sp. nov. (ZSI / WGRC / V / A / 937): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829851/files/figure.png" pageId="24" pageNumber="475">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8436A13132FF82FF0DB015D794FE27" blockId="24.[151,1437,151,789]" pageId="24" pageNumber="475">
|
||
<emphasis id="B94FEAB33132FF82FF0DB015D47CFEEF" bold="true" box="[199,330,295,320]" pageId="24" pageNumber="475">Variations.</emphasis>
|
||
Morphometric data from two adult males and an adult female, including the holotype, is given in
|
||
<tableCitation id="C6B9031A3132FF82FF5DB07ED5DAFECB" box="[151,236,332,357]" captionStart="TABLE 7" captionStartId="18.[119,204,562,584]" captionText="TABLE 7. Morphometric measurements for five species in the Fejervarya rufescens group. All measurements are in millimeters (mm). The status of each specimen is given the museum number: Holotype (HT), Paratype (PT), Neotype (NT), Referred specimen (RS)." pageId="24" pageNumber="475">Table 7</tableCitation>
|
||
.
|
||
<emphasis id="B94FEAB33132FF82FF3DB07ED4C3FECB" box="[247,501,332,357]" italics="true" pageId="24" pageNumber="475">Colour in preservation</emphasis>
|
||
. ZSI/WGRC/V/A/938 and ZSI/WGRC/V/A/939: dorsum greyish-brown with more prominent and larger blackish-brown blotches.
|
||
</paragraph>
|
||
<paragraph id="8B8436A13132FF82FF0DB0A6D1C8FE7F" blockId="24.[151,1437,151,789]" pageId="24" pageNumber="475">
|
||
<emphasis id="B94FEAB33132FF82FF0DB0A6D712FE02" bold="true" box="[199,548,404,429]" pageId="24" pageNumber="475">Secondary sexual characters.</emphasis>
|
||
<emphasis id="B94FEAB33132FF82FDE6B0A6D753FE02" box="[556,613,404,429]" italics="true" pageId="24" pageNumber="475">Male</emphasis>
|
||
: nuptial pad on finger I present; distinct calling patches on either side of the throat.
|
||
<emphasis id="B94FEAB33132FF82FEC5B085D455FE7F" box="[271,355,439,464]" italics="true" pageId="24" pageNumber="475">Female</emphasis>
|
||
(ZSI/WGRC/V/A/939): pigmented eggs present (diameter 1.9 ±
|
||
<quantity id="4CC39B443132FF82FBF7B08AD1A3FE60" box="[1085,1173,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="475" unit="mm" value="0.4">0.4 mm</quantity>
|
||
,
|
||
<emphasis id="B94FEAB33132FF82FB6BB08BD182FE7F" box="[1185,1204,441,464]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 15).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321652A3132FF82FF0DB0EED464FCBB" pageId="24" pageNumber="475" type="distribution">
|
||
<paragraph id="8B8436A13132FF82FF0DB0EED464FCBB" blockId="24.[151,1437,151,789]" pageId="24" pageNumber="475">
|
||
<emphasis id="B94FEAB33132FF82FF0DB0EED768FE5A" bold="true" box="[199,606,476,501]" pageId="24" pageNumber="475">Distribution and natural history.</emphasis>
|
||
<taxonomicName id="4C3B4D223132FF82FDA1B0EFD61CFE5B" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[619,810,476,500]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FDA1B0EFD61CFE5B" box="[619,810,476,500]" italics="true" pageId="24" pageNumber="475">Fejervarya cepfi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FCFDB0EED6A0FE5A" box="[823,918,476,501]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FCFDB0EED6A0FE5A" bold="true" box="[823,918,476,501]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is currently known only from the northern
|
||
<collectingRegion id="49FFF8433132FF82FF5DB332D5C4FDB8" box="[151,242,512,535]" country="Sri Lanka" name="Western" pageId="24" pageNumber="475">Western</collectingRegion>
|
||
Ghats state of
|
||
<collectingRegion id="49FFF8433132FF82FE6AB0CDD71AFDB7" box="[416,556,511,536]" country="India" name="Maharashtra" pageId="24" pageNumber="475">Maharashtra</collectingRegion>
|
||
(
|
||
<figureCitation id="13002A243132FF82FDF7B0CDD7B5FDB7" box="[573,643,511,536]" captionStart="FIGURE 3" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[151,1436,197,1739]" captionTargetId="figure@6.[151,1436,193,1744]" captionTargetPageId="6" captionText="FIGURE 3. Geographical distribution of five species in Fejervarya rufescens group of the Western Ghats, Peninsular India. The Western Ghats biodiversity hotspot region is shown in orange colour. Locality details are referenced in Table 3." httpUri="https://zenodo.org/record/829839/files/figure.png" pageId="24" pageNumber="475">Fig. 3</figureCitation>
|
||
). Apart from the
|
||
<typeStatus id="548088033132FF82FC86B332D64AFDB7" box="[844,892,512,536]" pageId="24" pageNumber="475">type</typeStatus>
|
||
locality Amboli, we also genetically confirmed the presence of this species in Phansad WLS (Raigad district) and Koyna (Satara district). During the monsoon season (June–July), large congregations of males and females were observed on emerging laterite rock surfaces inside open grassland areas at Amboli. These sites were usually adjacent to shallow streams or temporary water collections. Calling males were extremely sensitive to nearby movements, and would immediately stop calling and try to hide upon being approached. This species was observed to be locally abundant during the breeding season but only for a very short duration lasting a couple of weeks. It was difficult to locate individuals outside the breeding season.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |