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<mods:title id="B182B174FC55716F2C85D6F3CEA3D0A1">On the taxonomic status of Chactas camposi Mello-Leitão, 1939 (Scorpiones, Chactidae)</mods:title>
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<mods:namePart id="AE389390B240D26C2619688A3D212AB0">Ochoa, José A.</mods:namePart>
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<treatment id="03928791122C393CFF2AFEF93493B05D" ID-DOI="http://doi.org/10.5281/zenodo.6166620" ID-GBIF-Taxon="119509640" ID-Zenodo-Dep="6166620" LSID="urn:lsid:plazi:treatment:03928791122C393CFF2AFEF93493B05D" httpUri="http://treatment.plazi.org/id/03928791122C393CFF2AFEF93493B05D" lastPageId="7" lastPageNumber="68" pageId="2" pageNumber="63">
<subSubSection id="C321650C122C3939FF2AFEF93523B2EB" pageId="2" pageNumber="63" type="nomenclature">
<paragraph id="8B843687122C3939FF2AFEF93434B2C8" blockId="2.[151,788,359,418]" box="[151,788,359,385]" pageId="2" pageNumber="63">
<heading id="D0CC81EB122C3939FF2AFEF93434B2C8" bold="true" box="[151,788,359,385]" fontSize="11" level="1" pageId="2" pageNumber="63" reason="1">
<emphasis id="B94FEA95122C3939FF2AFEF93434B2C8" bold="true" box="[151,788,359,385]" pageId="2" pageNumber="63">
<taxonomicName id="4C3B4D04122C3939FF2AFEF93582B2C8" ID-CoL="55Y48" authority="Mello-Leitao, 1939" authorityName="Mello-Leitao" authorityYear="1939" box="[151,674,359,385]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122C3939FF2AFEF936BCB2C8" bold="true" box="[151,412,359,385]" italics="true" pageId="2" pageNumber="63">Teuthraustes camposi</emphasis>
(
<bibRefCitation id="EFAA4B76122C3939FE11FEF935B9B2C8" author="Mello-Leitao" box="[428,665,359,385]" pageId="2" pageNumber="63" refString="Mello-Leitao, C. de. (1939) Una nueva especie de alacran del genero Chactas. Physis, 17, 147 - 148." type="journal article" year="1939">Mello-Leitão, 1939</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122C3939FD14FEF93434B2C8" box="[681,788,359,385]" pageId="2" pageNumber="63">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B843687122C3939FF2AFE143523B2EB" blockId="2.[151,788,359,418]" box="[151,515,394,418]" pageId="2" pageNumber="63">
<figureCitation id="13002A02122C3939FF2AFE143619B2EB" box="[151,313,394,418]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="2" pageNumber="63">Figures 1AD</figureCitation>
,
<figureCitation id="13002A02122C3939FEF9FE1436A7B2EB" box="[324,391,394,418]" captionStart="FIGURE 2 A E" captionStartId="5.[151,250,1935,1958]" captionTargetBox="[151,1436,697,1913]" captionTargetId="figure@5.[151,1436,697,1914]" captionTargetPageId="5" captionText="FIGURE 2 A E. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n. Dextral pedipalp: A, femur and patella, dorsal view; B, femur, patella and chela, ventral view; C, patella, external view; D, chela, external view; E, chela, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215452/files/figure.png" pageId="2" pageNumber="63">2AE</figureCitation>
,
<figureCitation id="13002A02122C3939FE29FE143684B2EB" box="[404,420,394,418]" captionStart="FIGURE 3" captionStartId="6.[151,249,1159,1182]" captionTargetBox="[245,1341,193,1138]" captionTargetId="figure@6.[245,1341,193,1138]" captionTargetPageId="6" captionText="FIGURE 3. Map of Ecuador plotting the type locality of Teuthraustes camposi (Mello-Leitão, 1939) comb. n." httpUri="https://zenodo.org/record/215453/files/figure.png" pageId="2" pageNumber="63">3</figureCitation>
;
<tableCitation id="C6B9033C122C3939FE0DFE143523B2EB" box="[432,515,394,418]" captionStart="TABLE 2" captionStartId="3.[151,235,152,173]" captionText="TABLE 2. Selected measurements (mm) of adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n. Carapace:" pageId="2" pageNumber="63">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C321650C122C3939FF2AFE4C3408B16F" pageId="2" pageNumber="63" type="reference_group">
<paragraph id="8B843687122C3939FF2AFE4C3408B16F" blockId="2.[151,1277,466,550]" pageId="2" pageNumber="63">
<treatmentCitationGroup id="AB2B11A9122C3939FF2AFE4C3408B16F" pageId="2" pageNumber="63">
<taxonomicName id="4C3B4D04122C3939FF2AFE4C3563B2A1" authority="Mello-Leitao, 1939: 147" authorityName="Mello-Leitao" authorityPageNumber="147" authorityYear="1939" box="[151,579,466,488]" class="Arachnida" family="Chactidae" genus="Chactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi">
<emphasis id="B94FEA95122C3939FF2AFE4C3664B2A1" box="[151,324,466,488]" italics="true" pageId="2" pageNumber="63">Chactas camposi</emphasis>
<treatmentCitation id="0A9A1096122C3939FEF6FE4C3563B2A1" author="Mello-Leitao" box="[331,579,466,488]" page="147" pageId="2" pageNumber="63" year="1939">
<bibRefCitation id="EFAA4B76122C3939FEF6FE4C3533B2A1" author="Mello-Leitao" box="[331,531,466,488]" pageId="2" pageNumber="63" refString="Mello-Leitao, C. de. (1939) Una nueva especie de alacran del genero Chactas. Physis, 17, 147 - 148." type="journal article" year="1939">Mello-Leitão, 1939</bibRefCitation>
: 147
</treatmentCitation>
</taxonomicName>
148, figs. 16;
<treatmentCitation id="0A9A1096122C3939FD55FE4D349FB2A1" author="Lourenco" box="[744,959,466,488]" page="157" pageId="2" pageNumber="63" year="1994">
<bibRefCitation id="EFAA4B76122C3939FD55FE4D34ACB2A1" author="Lourenco" box="[744,908,467,488]" pageId="2" pageNumber="63" refString="Lourenco, W. R. (1994) Diversity and endemism in tropical versus temperate scorpion communities. Biogeographica, 70 (3), 155 - 160." type="journal article" year="1994">Lourenço, 1994</bibRefCitation>
: 157
</treatmentCitation>
; 1995: 68;
<treatmentCitation id="0A9A1096122C3939FB8DFE4C33D8B2A1" author="Kovarik" box="[1072,1272,466,488]" page="126" pageId="2" pageNumber="63" year="1998">
<bibRefCitation id="EFAA4B76122C3939FB8DFE4C33E5B2A1" author="Kovarik" box="[1072,1221,466,488]" pageId="2" pageNumber="63" refString="Kovarik, F. (1998) Sti r i (Scorpions). Madagaskar, Jihlava, 175 pp. [Czech]." type="book" year="1998">Kovařik, 1998</bibRefCitation>
: 126
</treatmentCitation>
.
<taxonomicName id="4C3B4D04122C3939FF2AFE6F359EB14E" authority="Mello-Leitao, 1945: 48" authorityName="Mello-Leitao" authorityPageNumber="48" authorityYear="1945" box="[151,702,497,519]" class="Arachnida" family="Chactidae" genus="Chactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" subGenus="Euchactas">
<emphasis id="B94FEA95122C3939FF2AFE6F37CAB14E" box="[151,234,497,519]" italics="true" pageId="2" pageNumber="63">Chactas</emphasis>
(
<emphasis id="B94FEA95122C3939FF45FE6F3641B14E" box="[248,353,497,519]" italics="true" pageId="2" pageNumber="63">Euchactas</emphasis>
)
<emphasis id="B94FEA95122C3939FECCFE6C36E5B14E" box="[369,453,498,519]" italics="true" pageId="2" pageNumber="63">camposi</emphasis>
:
<treatmentCitation id="0A9A1096122C3939FE6EFE6F359EB14E" author="Mello-Leitao" box="[467,702,497,519]" page="48" pageId="2" pageNumber="63" year="1945">
<bibRefCitation id="EFAA4B76122C3939FE6EFE6F35BBB14E" author="Mello-Leitao" box="[467,667,497,519]" pageId="2" pageNumber="63" refString="Mello-Leitao, C. de. (1945) Escorpioes sul-americanos. Arquivos do Museu Nacional, Rio de Janeiro, 40, 7 - 468." type="journal article" year="1945">Mello-Leitão, 1945</bibRefCitation>
: 48
</treatmentCitation>
</taxonomicName>
50, figs. 1819;
<treatmentCitation id="0A9A1096122C3939FCCDFE6F330FB14E" author="Sissom" box="[880,1071,497,519]" page="307" pageId="2" pageNumber="63" year="2000">
<bibRefCitation id="EFAA4B76122C3939FCCDFE6F34DCB14E" author="Sissom" box="[880,1020,497,519]" pageId="2" pageNumber="63" refString="Sissom, W. D. (2000) Family Chactidae Pocock, 1893. In V. Fet, W. D. Sissom, G. Lowe, and M. E. Braunwalder, Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 287 - 322." type="book chapter" year="2000">Sissom, 2000</bibRefCitation>
: 307
</treatmentCitation>
.
<taxonomicName id="4C3B4D04122C3939FF2AFD8E36CEB16F" authority="Mello-Leitao" authorityName="Mello-Leitao" box="[151,494,528,550]" class="Arachnida" family="Chactidae" genus="Chactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="rosenbergi">
<emphasis id="B94FEA95122C3939FF2AFD8E367CB16F" box="[151,348,528,550]" italics="true" pageId="2" pageNumber="63">Chactas rosenbergi</emphasis>
: Mello-Leitão
</taxonomicName>
<emphasis id="B94FEA95122C3939FE48FD8F3549B16F" box="[501,617,528,550]" italics="true" pageId="2" pageNumber="63">in schedula</emphasis>
, misidentification.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C321650C122C3939FF2AFDCD3338B1C7" pageId="2" pageNumber="63" type="materials_examined">
<paragraph id="8B843687122C3939FF2AFDCD3338B1C7" blockId="2.[151,1437,594,1986]" pageId="2" pageNumber="63">
<emphasis id="B94FEA95122C3939FF2AFDCD3664B122" bold="true" box="[151,324,594,619]" pageId="2" pageNumber="63">
<typeStatus id="54808825122C3939FF2AFDCD37F2B122" box="[151,210,595,619]" pageId="2" pageNumber="63">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54808825122C3939FEF7FDCC3692B123" box="[330,434,594,618]" pageId="2" pageNumber="63" type="holotype">Holotype</typeStatus>
♀ (
<collectionCode id="ED2AAE42122C3939FE61FDCC3509B123" box="[476,553,594,618]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="63">MNRJ</collectionCode>
58732):
<collectingCountry id="F32C7617122C3939FD31FDCC3439B122" box="[652,793,594,619]" name="Ecuador" pageId="2" pageNumber="63">
<emphasis id="B94FEA95122C3939FD31FDCC3439B122" bold="true" box="[652,793,594,619]" pageId="2" pageNumber="63">ECUADOR</emphasis>
</collectingCountry>
:
<emphasis id="B94FEA95122C3939FC95FDCD34FAB122" box="[808,986,595,619]" italics="true" pageId="2" pageNumber="63">Cañar Province</emphasis>
, Montañas de Cañar, Francisco Campos col. [Labeled as “
<taxonomicName id="4C3B4D04122C3939FEE1FDE835B5B1C7" authority="Pocock" authorityName="Pocock" box="[348,661,630,654]" class="Arachnida" family="Chactidae" genus="Chactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="rosenbergi">
<emphasis id="B94FEA95122C3939FEE1FDE83516B1C7" box="[348,566,630,654]" italics="true" pageId="2" pageNumber="63">Chactas rosenbergi</emphasis>
Pocock
</taxonomicName>
”, most certainly by Mello-Leitão].
</paragraph>
</subSubSection>
<subSubSection id="C321650C122C3939FF74FD043459B6CB" pageId="2" pageNumber="63" type="diagnosis">
<paragraph id="8B843687122C3939FF74FD043459B6CB" blockId="2.[151,1437,594,1986]" pageId="2" pageNumber="63">
<emphasis id="B94FEA95122C3939FF74FD043663B1FA" bold="true" box="[201,323,666,691]" pageId="2" pageNumber="63">Diagnosis.</emphasis>
This species appears to be most closely related with
<taxonomicName id="4C3B4D04122C3939FC23FD053279B1FB" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" box="[926,1369,666,691]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="whymperi">
<emphasis id="B94FEA95122C3939FC23FD053382B1FA" box="[926,1186,667,691]" italics="true" pageId="2" pageNumber="63">Teuthraustes whymperi</emphasis>
(Pocock, 1893)
</taxonomicName>
. Both species are similar and may be distinguished from the other species of the genus based on the shape of the anterior margin of carapace, which presents a well developed median notch (strongly excavated) (
<figureCitation id="13002A02122C3939FB3EFD7C33FCB1B3" box="[1155,1244,738,762]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="2" pageNumber="63">Fig. 1A</figureCitation>
); the ventral carination absent or obsolete on metasomal segments IIII (
<figureCitation id="13002A02122C3939FC9FFC98345DB057" box="[802,893,774,798]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="2" pageNumber="63">Fig. 1D</figureCitation>
); the telson globose with the aculeus elongated (
<figureCitation id="13002A02122C3939FF22FCB43605B00B" box="[159,293,810,834]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="2" pageNumber="63">Fig. 1C, 1D</figureCitation>
); the pedipalp chela with well developed costate granular VE carinae (
<figureCitation id="13002A02122C3939FB8AFCB433AEB00B" box="[1079,1166,810,834]" captionStart="FIGURE 2 A E" captionStartId="5.[151,250,1935,1958]" captionTargetBox="[151,1436,697,1913]" captionTargetId="figure@5.[151,1436,697,1914]" captionTargetPageId="5" captionText="FIGURE 2 A E. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n. Dextral pedipalp: A, femur and patella, dorsal view; B, femur, patella and chela, ventral view; C, patella, external view; D, chela, external view; E, chela, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215452/files/figure.png" pageId="2" pageNumber="63">Fig. 2B</figureCitation>
); and the chela movable finger with evident subbasal notch on dorsal margin (
<figureCitation id="13002A02122C3939FCBDFCD0347DB02F" box="[768,861,846,870]" captionStart="FIGURE 2 A E" captionStartId="5.[151,250,1935,1958]" captionTargetBox="[151,1436,697,1913]" captionTargetId="figure@5.[151,1436,697,1914]" captionTargetPageId="5" captionText="FIGURE 2 A E. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n. Dextral pedipalp: A, femur and patella, dorsal view; B, femur, patella and chela, ventral view; C, patella, external view; D, chela, external view; E, chela, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215452/files/figure.png" pageId="2" pageNumber="63">Fig. 2D</figureCitation>
).
<taxonomicName id="4C3B4D04122C3939FCCFFCD03347B02F" box="[882,1127,846,870]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122C3939FCCFFCD03347B02F" box="[882,1127,846,870]" italics="true" pageId="2" pageNumber="63">Teuthraustes camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122C3939FBCCFCD033F4B02F" box="[1137,1236,846,870]" pageId="2" pageNumber="63">comb. n.</taxonomicNameLabel>
and
<taxonomicName id="4C3B4D04122C3939FAB2FCCE32B9B02F" box="[1295,1433,846,871]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="whymperi">
<emphasis id="B94FEA95122C3939FAB2FCCE32B9B02F" box="[1295,1433,846,871]" italics="true" pageId="2" pageNumber="63">T. whymperi</emphasis>
</taxonomicName>
may be separated by the following respects: the anterior median notch of the carapace is slightly more pronounced in
<taxonomicName id="4C3B4D04122C3939FF09FC06361BB0E7" box="[180,315,918,943]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="whymperi">
<emphasis id="B94FEA95122C3939FF09FC06361BB0E7" box="[180,315,918,943]" italics="true" pageId="2" pageNumber="63">T. whymperi</emphasis>
</taxonomicName>
; the telson is weakly granular on ventral and lateral surfaces in
<taxonomicName id="4C3B4D04122C3939FBB6FC0633A3B0E7" box="[1035,1155,919,943]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi">
<emphasis id="B94FEA95122C3939FBB6FC0633A3B0E7" box="[1035,1155,919,943]" italics="true" pageId="2" pageNumber="63">T. camposi</emphasis>
</taxonomicName>
, whereas in
<taxonomicName id="4C3B4D04122C3939FAAFFC0632B9B0E7" box="[1298,1433,918,943]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="whymperi">
<emphasis id="B94FEA95122C3939FAAFFC0632B9B0E7" box="[1298,1433,918,943]" italics="true" pageId="2" pageNumber="63">T. whymperi</emphasis>
</taxonomicName>
is completely smooth (
<figureCitation id="13002A02122C3939FE27FC243504B09B" box="[410,548,954,978]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="2" pageNumber="63">Fig. 1C, 1D</figureCitation>
); the pectinal tooth count is
<quantity id="4CC39B62122C3939FCD7FC2434E1B09B" box="[874,961,954,978]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="2" pageNumber="63" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">910 in</quantity>
<taxonomicName id="4C3B4D04122C3939FC74FC223363B09A" box="[969,1091,956,979]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122C3939FC74FC223363B09A" box="[969,1091,956,979]" italics="true" pageId="2" pageNumber="63">T. camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122C3939FBF6FC24338EB09B" box="[1099,1198,954,978]" pageId="2" pageNumber="63">comb. n.</taxonomicNameLabel>
(
<figureCitation id="13002A02122C3939FB03FC243235B09B" box="[1214,1301,954,978]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="2" pageNumber="63">Fig. 1B</figureCitation>
), compared with
<quantity id="4CC39B62122C3939FF6FFC40363BB0BF" box="[210,283,990,1014]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="2" pageNumber="63" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">67 in</quantity>
<taxonomicName id="4C3B4D04122C3939FE99FC7E368DB0BF" box="[292,429,990,1015]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="whymperi">
<emphasis id="B94FEA95122C3939FE99FC7E368DB0BF" box="[292,429,990,1015]" italics="true" pageId="2" pageNumber="63">T. whymperi</emphasis>
</taxonomicName>
(females). These two species may be further distinguished by means of the granulation and carination of pedipalps; the dorsal intercarinal surfaces of femur and patella are scattered granular in
<taxonomicName id="4C3B4D04122C3939FAFAFB9A37E6B777" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122C3939FAFAFB9A37E6B777" italics="true" pageId="2" pageNumber="63">T. camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122C3939FF70FBB83610B777" box="[205,304,1062,1086]" pageId="2" pageNumber="63">comb. n.</taxonomicNameLabel>
(
<figureCitation id="13002A02122C3939FEFDFBB836B9B777" box="[320,409,1062,1086]" captionStart="FIGURE 2 A E" captionStartId="5.[151,250,1935,1958]" captionTargetBox="[151,1436,697,1913]" captionTargetId="figure@5.[151,1436,697,1914]" captionTargetPageId="5" captionText="FIGURE 2 A E. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n. Dextral pedipalp: A, femur and patella, dorsal view; B, femur, patella and chela, ventral view; C, patella, external view; D, chela, external view; E, chela, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215452/files/figure.png" pageId="2" pageNumber="63">Fig. 2A</figureCitation>
) whereas in
<taxonomicName id="4C3B4D04122C3939FD96FBB63593B777" box="[555,691,1062,1087]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="whymperi">
<emphasis id="B94FEA95122C3939FD96FBB63593B777" box="[555,691,1062,1087]" italics="true" pageId="2" pageNumber="63">T. whymperi</emphasis>
</taxonomicName>
these are densely granular; the external carinae of pedipalp chela are slightly more developed in
<taxonomicName id="4C3B4D04122C3939FE52FBD23556B72A" box="[495,630,1099,1123]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="whymperi">
<emphasis id="B94FEA95122C3939FE52FBD23556B72A" box="[495,630,1099,1123]" italics="true" pageId="2" pageNumber="63">T. whymperi</emphasis>
</taxonomicName>
. Finally, the number of median spinules of telotarsi IIV is 6-6-8-
<quantity id="4CC39B62122C3939FAEBFBD432A1B72B" box="[1366,1409,1098,1122]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="2" pageNumber="63" unit="in" value="9.0">9 in</quantity>
<taxonomicName id="4C3B4D04122C3939FA3AFBD237D4B7CF" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122C3939FA3AFBD237D4B7CF" italics="true" pageId="2" pageNumber="63">T. camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122C3939FF46FBF0367DB7CF" box="[251,349,1134,1158]" pageId="2" pageNumber="63">comb. n.</taxonomicNameLabel>
compared with 5-6-6-6/
<quantity id="4CC39B62122C3939FDD3FBF035B9B7CF" box="[622,665,1134,1158]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="2" pageNumber="63" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="4C3B4D04122C3939FD1CFBEE3408B7CF" box="[673,808,1134,1159]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="whymperi">
<emphasis id="B94FEA95122C3939FD1CFBEE3408B7CF" box="[673,808,1134,1159]" italics="true" pageId="2" pageNumber="63">T. whymperi</emphasis>
</taxonomicName>
.
<taxonomicName id="4C3B4D04122C3939FC84FBF0330CB7CF" box="[825,1068,1134,1158]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122C3939FC84FBF0330CB7CF" box="[825,1068,1134,1158]" italics="true" pageId="2" pageNumber="63">Teuthraustes camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122C3939FB8EFBF033B4B7CF" box="[1075,1172,1134,1158]" pageId="2" pageNumber="63">comb. n.</taxonomicNameLabel>
may be separated from
<taxonomicName id="4C3B4D04122C3939FF2AFB0A36EFB7E3" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" box="[151,463,1170,1195]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="gervaisii">
<emphasis id="B94FEA95122C3939FF2AFB0A3636B7E2" box="[151,278,1172,1195]" italics="true" pageId="2" pageNumber="63">T. gervaisii</emphasis>
(Pocock, 1893)
</taxonomicName>
, which occurs in close geographical proximity (Cuenca province in central Andes of
<collectingCountry id="F32C7617122C3939FF2AFB2837D9B787" box="[151,249,1206,1230]" name="Ecuador" pageId="2" pageNumber="63">Ecuador</collectingCountry>
), by means of granulation of carapace and pedipalps; the anterior third of carapace is coarsely granular (
<figureCitation id="13002A02122C3939FF22FB4437D6B7BB" box="[159,246,1242,1266]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="2" pageNumber="63">Fig. 1A</figureCitation>
), the dorsal and external surface of pedipalp patella, and the external surface of the chela are granular in
<taxonomicName id="4C3B4D04122C3939FA3AFB4237D4B65F" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122C3939FA3AFB4237D4B65F" italics="true" pageId="2" pageNumber="63">T. camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122C3939FF41FB60367EB65F" box="[252,350,1278,1302]" pageId="2" pageNumber="63">comb. n.</taxonomicNameLabel>
(
<figureCitation id="13002A02122C3939FED3FB60352DB65F" box="[366,525,1278,1302]" captionStart="FIGURE 2 A E" captionStartId="5.[151,250,1935,1958]" captionTargetBox="[151,1436,697,1913]" captionTargetId="figure@5.[151,1436,697,1914]" captionTargetPageId="5" captionText="FIGURE 2 A E. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n. Dextral pedipalp: A, femur and patella, dorsal view; B, femur, patella and chela, ventral view; C, patella, external view; D, chela, external view; E, chela, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215452/files/figure.png" pageId="2" pageNumber="63">Fig. 2A, C, D</figureCitation>
), whereas in
<taxonomicName id="4C3B4D04122C3939FD19FA9E3402B65F" box="[676,802,1279,1303]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="gervaisii">
<emphasis id="B94FEA95122C3939FD19FA9E3402B65F" box="[676,802,1279,1303]" italics="true" pageId="2" pageNumber="63">T. gervaisii</emphasis>
</taxonomicName>
these surfaces are completely smooth. Furthermore, the anterior margin of the carapace possesses a weakly developed median notch in
<taxonomicName id="4C3B4D04122C3939FBA8FABA33B3B672" box="[1045,1171,1316,1339]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="gervaisii">
<emphasis id="B94FEA95122C3939FBA8FABA33B3B672" box="[1045,1171,1316,1339]" italics="true" pageId="2" pageNumber="63">T. gervaisii</emphasis>
</taxonomicName>
, whereas in
<taxonomicName id="4C3B4D04122C3939FA9EFABA32BCB672" box="[1315,1436,1316,1339]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122C3939FA9EFABA32BCB672" box="[1315,1436,1316,1339]" italics="true" pageId="2" pageNumber="63">T. camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122C3939FF2AFAD837DAB617" box="[151,250,1350,1374]" pageId="2" pageNumber="63">comb. n.</taxonomicNameLabel>
the median notch is well developed (
<figureCitation id="13002A02122C3939FD15FAD83423B617" box="[680,771,1350,1374]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="2" pageNumber="63">Fig. 1A</figureCitation>
); the VE carinae of pedipalp chela is costate granular and well developed in
<taxonomicName id="4C3B4D04122C3939FED8FAF236FBB6CA" box="[357,475,1388,1411]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122C3939FED8FAF236FBB6CA" box="[357,475,1388,1411]" italics="true" pageId="2" pageNumber="63">T. camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122C3939FE59FAF43567B6CB" box="[484,583,1386,1410]" pageId="2" pageNumber="63">comb. n.</taxonomicNameLabel>
, but smooth in
<taxonomicName id="4C3B4D04122C3939FD48FAF23452B6CA" box="[757,882,1388,1411]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="gervaisii">
<emphasis id="B94FEA95122C3939FD48FAF23452B6CA" box="[757,882,1388,1411]" italics="true" pageId="2" pageNumber="63">T. gervaisii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C321650C122C393CFF74FA103548B286" lastPageId="7" lastPageNumber="68" pageId="2" pageNumber="63" type="description">
<paragraph id="8B843687122C3939FF74FA103338B6EF" blockId="2.[151,1437,594,1986]" box="[201,1048,1422,1447]" pageId="2" pageNumber="63">
<emphasis id="B94FEA95122C3939FF74FA10367AB6EE" bold="true" box="[201,346,1422,1447]" pageId="2" pageNumber="63">Description.</emphasis>
Measurements of the
<typeStatus id="54808825122C3939FDEEFA103595B6EF" box="[595,693,1422,1446]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
female are recorded in
<tableCitation id="C6B9033C122C3939FC03FA103334B6EF" box="[958,1044,1422,1446]" captionStart="TABLE 2" captionStartId="3.[151,235,152,173]" captionText="TABLE 2. Selected measurements (mm) of adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n. Carapace:" pageId="2" pageNumber="63">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8B843687122C3939FF74FA2D324CB55B" blockId="2.[151,1437,594,1986]" pageId="2" pageNumber="63">
<emphasis id="B94FEA95122C3939FF74FA2D362AB682" box="[201,266,1459,1483]" italics="true" pageId="2" pageNumber="63">Color</emphasis>
: Base color, carapace, tergites and metasoma dark brown, becoming slightly darker on anterior half of carapace; chelicerae chestnut, becoming darker on fingers; pedipalps blackish; legs brownish; coxosternal region, pectines, and sternites brownish. Pigmentation pattern is not evident due the long time preservation in ethanol.
</paragraph>
<paragraph id="8B843687122C3939FF74F98034EAB58F" blockId="2.[151,1437,594,1986]" pageId="2" pageNumber="63">
<emphasis id="B94FEA95122C3939FF74F9803664B57F" box="[201,324,1566,1590]" italics="true" pageId="2" pageNumber="63">Chelicerae</emphasis>
: Manus dorsoexternal surfaces smooth, sparsely setose; ventral and internal surfaces densely setose; with four macrosetae near the base of the fingers. Fixed finger, dorsal margin with four teeth (distal, subdistal, median and basal); median and basal teeth fused into bicuspid. Movable finger, internal distal tooth at most partially overlapping external distal tooth in dorsal view; dorsal margin with five teeth (internal distal, two subdistal, median and basal); ventral surface with small but well developed serrula.
</paragraph>
<paragraph id="8B843687122C3939FF74F94D3410B48B" blockId="2.[151,1437,594,1986]" pageId="2" pageNumber="63">
<emphasis id="B94FEA95122C3939FF74F94D3618B5A3" box="[201,312,1747,1770]" italics="true" pageId="2" pageNumber="63">Carapace</emphasis>
(
<figureCitation id="13002A02122C3939FEF5F94C3682B5A3" box="[328,418,1746,1770]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="2" pageNumber="63">Fig. 1A</figureCitation>
): Anterior margin granular, with a well developed median notch; posterior margin sublinear, slightly granular. Anterior third surface coarsely granular, central lateral and postero-lateral surfaces finely granular. Ocular tubercle well developed, slightly granular, placed slightly ahead the middle of the carapace with two postocular setae. Median eyes well developed, one ocular diameter apart; three pairs of lateral ocelli present, anterior and median pairs largest, posterior pair greatly reduced. Anteromedian longitudinal sulcus broad, well developed, finely granular; posteromedian longitudinal sulcus well developed, finely granular on anterior half; posterolateral and posterior transverse sulci weakly developed.
</paragraph>
<caption id="DF44660F122D3938FF2AFF063625B396" pageId="3" pageNumber="64">
<paragraph id="8B843687122D3938FF2AFF063625B396" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">
<emphasis id="B94FEA95122D3938FF2AFF063624B3E4" bold="true" box="[151,260,152,173]" pageId="3" pageNumber="64">TABLE 2.</emphasis>
Selected measurements (mm) of adult female holotype of
<taxonomicName id="4C3B4D04122D3938FCEBFF06322DB3E7" authority="Mello-Leitao, 1939" authorityName="Mello-Leitao" authorityYear="1939" box="[854,1293,152,174]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122D3938FCEBFF063312B3E7" box="[854,1074,152,174]" italics="true" pageId="3" pageNumber="64">Teuthraustes camposi</emphasis>
(Mello-Leitão, 1939)
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122D3938FAA9FF06324CB3E7" box="[1300,1388,152,174]" pageId="3" pageNumber="64">comb. n.</taxonomicNameLabel>
Carapace:
</paragraph>
</caption>
<paragraph id="8B843687122D3938FF22FF6C37C2B2CD" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">length 9.0 anterior width 5.1 posterior width 8.5 Chela:</paragraph>
<paragraph id="8B843687122D3938FF22FE0637CCB11A" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">length 14.2 width 4.9 height 6.4 movable finger length 8.7 Patella:</paragraph>
<paragraph id="8B843687122D3938FF22FDF837C9B186" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">length 6.5 width 3.1 Femur:</paragraph>
<paragraph id="8B843687122D3938FF22FD7C3635B002" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">length 6.3 width 2.7 Mesosoma:</paragraph>
<paragraph id="8B843687122D3938FF22FCC0363EB0D7" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">total length 14.8 Metasoma I:</paragraph>
<paragraph id="8B843687122D3938FF22FC2F3607B753" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">length 2.8 width 3.9 Metasoma II:</paragraph>
<paragraph id="8B843687122D3938FF22FBB33610B7DF" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">length 3.6 width 3.6 Metasoma III:</paragraph>
<paragraph id="8B843687122D3938FF22FB37360FB65B" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">length 4.0 width 3.4 Metasoma IV:</paragraph>
<paragraph id="8B843687122D3938FF22FABB3607B6C7" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">length 4.4 width 3.2 Metasoma V:</paragraph>
<paragraph id="8B843687122D3938FF22FA3F37C7B57A" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">length 7.3 width 3.3 height 3.0 Telson:</paragraph>
<paragraph id="8B843687122D3938FF22F9D83631B462" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">total length 8.3 vesicle length 5.5 vesicle width 3.7 vesicle height 3.1 aculeus length 2.8 Metasoma:</paragraph>
<paragraph id="8B843687122D3938FF22F8A0349CB437" blockId="3.[151,1388,152,1918]" pageId="3" pageNumber="64">total length 31.5 Total body length: 55.3</paragraph>
<caption id="DF44660F122A393FFF2AF93D3506B5BE" httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="4" pageNumber="65" targetBox="[163,1422,185,1675]" targetPageId="4">
<paragraph id="8B843687122A393FFF2AF93D3506B5BE" blockId="4.[151,1436,1699,1783]" pageId="4" pageNumber="65">
<emphasis id="B94FEA95122A393FFF2AF93D3664B5F0" bold="true" box="[151,324,1699,1722]" pageId="4" pageNumber="65">FIGURE 1AD.</emphasis>
Adult female holotype of
<taxonomicName id="4C3B4D04122A393FFDE9F93A332EB5F3" authority="Mello-Leitao, 1939" authorityName="Mello-Leitao" authorityYear="1939" box="[596,1038,1699,1722]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122A393FFDE9F93A3410B5F3" box="[596,816,1700,1722]" italics="true" pageId="4" pageNumber="65">Teuthraustes camposi</emphasis>
(Mello-Leitão, 1939)
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122A393FFBA8F93D334EB5F3" box="[1045,1134,1699,1722]" pageId="4" pageNumber="65">comb. n.</taxonomicNameLabel>
.
<emphasis id="B94FEA95122A393FFB3FF93A33BBB5F0" bold="true" box="[1154,1179,1700,1721]" pageId="4" pageNumber="65">A,</emphasis>
carapace;
<emphasis id="B94FEA95122A393FFAB4F93A323FB5F0" bold="true" box="[1289,1311,1700,1721]" pageId="4" pageNumber="65">B,</emphasis>
coxosternal region, genital operculum and pectines;
<emphasis id="B94FEA95122A393FFD89F95C356CB590" bold="true" box="[564,588,1730,1753]" pageId="4" pageNumber="65">C,</emphasis>
metasomal segment V and telson, ventral view;
<emphasis id="B94FEA95122A393FFBF9F95D337CB591" bold="true" box="[1092,1116,1731,1752]" pageId="4" pageNumber="65">D,</emphasis>
metasomal segments IIV and telson, lateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8B843687122A393EFF74F88035A1B1CD" blockId="4.[151,1436,1822,2026]" lastBlockId="5.[151,1436,151,644]" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65">
<emphasis id="B94FEA95122A393FFF74F8803619B47F" box="[201,313,1822,1846]" italics="true" pageId="4" pageNumber="65">Pedipalps</emphasis>
(
<figureCitation id="13002A02122A393FFEF5F88036E3B47F" box="[328,451,1822,1846]" captionStart="FIGURE 2 A E" captionStartId="5.[151,250,1935,1958]" captionTargetBox="[151,1436,697,1913]" captionTargetId="figure@5.[151,1436,697,1914]" captionTargetPageId="5" captionText="FIGURE 2 A E. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n. Dextral pedipalp: A, femur and patella, dorsal view; B, femur, patella and chela, ventral view; C, patella, external view; D, chela, external view; E, chela, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215452/files/figure.png" pageId="4" pageNumber="65">Fig. 2AE</figureCitation>
): Femur,
<collectingCountry id="F32C7617122A393FFD97F8813568B47F" box="[554,584,1823,1846]" name="Iceland" pageId="4" pageNumber="65">DI</collectingCountry>
<collectingCountry id="F32C7617122A393FFDF3F8813556B47F" box="[590,630,1823,1846]" name="South Georgia and the South Sandwich Islands" pageId="4" pageNumber="65">DE</collectingCountry>
, VI and IM carinae complete, granular,
<collectingCountry id="F32C7617122A393FFB8FF8813377B47F" box="[1074,1111,1823,1846]" name="South Georgia and the South Sandwich Islands" pageId="4" pageNumber="65">DE</collectingCountry>
and IM more developed; VE carinae restricted to proximal third of segment, granular; EM restricted to distal third, vestigial, comprising fine granulation; dorsal intercarinal surfaces with few scattered granules; internal surface with fine granulation, ventral surface smooth. Patella,
<collectingCountry id="F32C7617122A393FFE15F81536E8B4EB" box="[424,456,1931,1954]" name="Iceland" pageId="4" pageNumber="65">DI</collectingCountry>
, VI, and VE carinae complete, granular;
<collectingCountry id="F32C7617122A393FFC32F8153494B4EB" box="[911,948,1931,1954]" name="South Georgia and the South Sandwich Islands" pageId="4" pageNumber="65">DE</collectingCountry>
and EM carinae obsolete, slightly granular; DPP and VPP each with one prominent granule proximally, VPP slightly bigger; dorsal and external intercarinal surfaces with scattered granulation; internal intercarinal surfaces finely granular; ventral intercarinal surfaces smooth. Chela manus swollen, fingers shorter than manus length. External and dorsal intercarinal surfaces between DS-SMA-DI and D-E-VE carinae coarsely and densely granular more so on distal part of manus; surfaces between D and DS carinae smooth on proximal two-thirds, distal third coarsely granular; ventral intercarinal surface smooth; internal intercarinal surfaces mostly smooth with few scattered granules; D and DS carinae complete, identified by weak granulation and subtle differences in angles between surfaces; SD vestigial, comprising three granules proximally; DMA carina complete, coarsely granular;
<collectingCountry id="F32C7617122B393EFCCAFED334B5B22D" box="[887,917,333,356]" name="Iceland" pageId="5" pageNumber="66">DI</collectingCountry>
carina evident on anterior half, obscured by adjacent granulation on distal half; VI carina vestigial, comprising four granules basally; VE carinae well developed, costate granular; IM carina scarcely granular on proximal three-quarters of manus, obscured by fine scattered granulation on distal quart of manus; E carinae complete, granular, evident by subtle differences in angles between adjacent surfaces on proximal two-thirds of manus. Fingers almost acarinate, only DS and DMA carinae slightly evident; fixed finger, median denticle row comprising six primary subrows of denticles, flanked by six external and seven internal denticles; the basal external denticle greatly enlarged, which corresponds with a well developed subbasal notch in movable finger. Movable finger, median denticle row comprising six subrows of denticles, flanked by seven external and six internal denticles.
</paragraph>
<caption id="DF44660F122B393EFF2AF81136FBB4AB" httpUri="https://zenodo.org/record/215452/files/figure.png" pageId="5" pageNumber="66" targetBox="[151,1436,697,1913]" targetPageId="5">
<paragraph id="8B843687122B393EFF2AF81136FBB4AB" blockId="5.[151,1436,1935,2018]" pageId="5" pageNumber="66">
<emphasis id="B94FEA95122B393EFF2AF8113665B4EC" bold="true" box="[151,325,1935,1958]" pageId="5" pageNumber="66">FIGURE 2AE.</emphasis>
Adult female holotype of
<taxonomicName id="4C3B4D04122B393EFDE6F8113338B4EF" authority="Mello-Leitao, 1939" authorityName="Mello-Leitao" authorityYear="1939" box="[603,1048,1935,1958]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA95122B393EFDE6F8113418B4EC" box="[603,824,1935,1957]" italics="true" pageId="5" pageNumber="66">Teuthraustes camposi</emphasis>
(Mello-Leitão, 1939)
</taxonomicName>
<taxonomicNameLabel id="A27C57EE122B393EFB9CF811335BB4EC" box="[1057,1147,1935,1957]" pageId="5" pageNumber="66">comb. n.</taxonomicNameLabel>
Dextral pedipalp:
<emphasis id="B94FEA95122B393EFA83F8113277B4ED" bold="true" box="[1342,1367,1935,1956]" pageId="5" pageNumber="66">A,</emphasis>
femur and patella, dorsal view;
<emphasis id="B94FEA95122B393EFE2BF830368DB48A" bold="true" box="[406,429,1966,1987]" pageId="5" pageNumber="66">B,</emphasis>
femur, patella and chela, ventral view;
<emphasis id="B94FEA95122B393EFCFCF8333479B48D" bold="true" box="[833,857,1965,1988]" pageId="5" pageNumber="66">C,</emphasis>
patella, external view;
<emphasis id="B94FEA95122B393EFBF4F8303341B48A" bold="true" box="[1097,1121,1966,1987]" pageId="5" pageNumber="66">D,</emphasis>
chela, external view;
<emphasis id="B94FEA95122B393EFAFFF8303278B48A" bold="true" box="[1346,1368,1966,1987]" pageId="5" pageNumber="66">E,</emphasis>
chela, dorsal view. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF44660F1228393DFF2AFB1933D3B7D7" box="[151,1267,1159,1182]" httpUri="https://zenodo.org/record/215453/files/figure.png" pageId="6" pageNumber="67" targetBox="[245,1341,193,1138]" targetPageId="6">
<paragraph id="8B8436871228393DFF2AFB1933D3B7D7" blockId="6.[151,1267,1159,1182]" box="[151,1267,1159,1182]" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFF2AFB193632B7D4" bold="true" box="[151,274,1159,1182]" pageId="6" pageNumber="67">FIGURE 3.</emphasis>
Map of Ecuador plotting the type locality of
<taxonomicName id="4C3B4D041228393DFD60FB1633B5B7D7" authority="Mello-Leitao, 1939" authorityName="Mello-Leitao" authorityYear="1939" box="[733,1173,1160,1182]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA951228393DFD60FB163498B7D7" box="[733,952,1160,1182]" italics="true" pageId="6" pageNumber="67">Teuthraustes camposi</emphasis>
(Mello-Leitão, 1939)
</taxonomicName>
<taxonomicNameLabel id="A27C57EE1228393DFB26FB1633D3B7D7" box="[1179,1267,1160,1182]" pageId="6" pageNumber="67">comb. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B8436871228393DFF74FB5B33C1B606" blockId="6.[151,1436,1221,2008]" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFF74FB5B3645B794" box="[201,357,1221,1245]" italics="true" pageId="6" pageNumber="67">Trichobothria</emphasis>
(
<figureCitation id="13002A021228393DFEC9FB5B36D8B794" box="[372,504,1221,1245]" captionStart="FIGURE 2 A E" captionStartId="5.[151,250,1935,1958]" captionTargetBox="[151,1436,697,1913]" captionTargetId="figure@5.[151,1436,697,1914]" captionTargetPageId="5" captionText="FIGURE 2 A E. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n. Dextral pedipalp: A, femur and patella, dorsal view; B, femur, patella and chela, ventral view; C, patella, external view; D, chela, external view; E, chela, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215452/files/figure.png" pageId="6" pageNumber="67">Figs. 2AE</figureCitation>
): Femur with three trichobothria (
<emphasis id="B94FEA951228393DFCCBFB5834A2B794" box="[886,898,1222,1245]" italics="true" pageId="6" pageNumber="67">e</emphasis>
,
<emphasis id="B94FEA951228393DFC2CFB5B34BFB794" box="[913,927,1221,1245]" italics="true" pageId="6" pageNumber="67">d</emphasis>
,
<emphasis id="B94FEA951228393DFC10FB583495B794" box="[941,949,1222,1245]" italics="true" pageId="6" pageNumber="67">i</emphasis>
). Patella with 25 trichobothria: five ventral (
<date id="FF8510471228393DFF1DFB7437F7B64F" box="[160,215,1258,1286]" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFF1DFB74378CB648" box="[160,172,1258,1281]" italics="true" pageId="6" pageNumber="67">v</emphasis>
1
<emphasis id="B94FEA951228393DFF7FFB7437EEB648" box="[194,206,1258,1281]" italics="true" pageId="6" pageNumber="67">v</emphasis>
5
</date>
); 17 external (
<emphasis id="B94FEA951228393DFEC2FB7436B3B648" box="[383,403,1258,1281]" italics="true" pageId="6" pageNumber="67">et</emphasis>
1
<emphasis id="B94FEA951228393DFE26FB74369DB648" box="[411,445,1258,1281]" italics="true" pageId="6" pageNumber="67">et</emphasis>
5,
<emphasis id="B94FEA951228393DFE69FB7436D3B648" box="[468,499,1258,1281]" italics="true" pageId="6" pageNumber="67">est</emphasis>
1
<emphasis id="B94FEA951228393DFDB4FB743508B648" box="[521,552,1258,1281]" italics="true" pageId="6" pageNumber="67">est</emphasis>
3,
<emphasis id="B94FEA951228393DFD81FB74357DB648" box="[572,605,1258,1281]" italics="true" pageId="6" pageNumber="67">em</emphasis>
1,
<emphasis id="B94FEA951228393DFDCEFB7435B4B648" box="[627,660,1258,1281]" italics="true" pageId="6" pageNumber="67">em</emphasis>
2,
<emphasis id="B94FEA951228393DFD17FB7735EFB648" box="[682,719,1257,1281]" italics="true" pageId="6" pageNumber="67">esb</emphasis>
1,
<emphasis id="B94FEA951228393DFD5BFB77342BB648" box="[742,779,1257,1281]" italics="true" pageId="6" pageNumber="67">esb</emphasis>
2,
<emphasis id="B94FEA951228393DFC9FFB77341DB648" box="[802,829,1257,1281]" italics="true" pageId="6" pageNumber="67">eb</emphasis>
1
<emphasis id="B94FEA951228393DFCEEFB77344DB648" box="[851,877,1257,1281]" italics="true" pageId="6" pageNumber="67">eb</emphasis>
5); two dorsal (
<emphasis id="B94FEA951228393DFBA8FB773303B648" box="[1045,1059,1257,1281]" italics="true" pageId="6" pageNumber="67">d</emphasis>
1,
<emphasis id="B94FEA951228393DFB87FB773368B648" box="[1082,1096,1257,1281]" italics="true" pageId="6" pageNumber="67">d</emphasis>
2); one internal (
<emphasis id="B94FEA951228393DFABCFB743229B648" box="[1281,1289,1258,1281]" italics="true" pageId="6" pageNumber="67">i</emphasis>
). Chela with 26 trichobothria: 16 on manus, four ventral (
<date id="FF8510471228393DFD2CFA8F35F1B664" box="[657,721,1297,1325]" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFD2CFA8F3582B661" box="[657,674,1297,1320]" italics="true" pageId="6" pageNumber="67">V</emphasis>
1
<emphasis id="B94FEA951228393DFD17FA8F35E9B661" box="[682,713,1297,1320]" italics="true" pageId="6" pageNumber="67">V</emphasis>
4
</date>
), 10 external (
<emphasis id="B94FEA951228393DFCC5FA8F34B1B661" box="[888,913,1297,1320]" italics="true" pageId="6" pageNumber="67">Et</emphasis>
1
<emphasis id="B94FEA951228393DFC1AFA8F34E0B661" box="[935,960,1297,1320]" italics="true" pageId="6" pageNumber="67">Et</emphasis>
5,
<emphasis id="B94FEA951228393DFC6BFA8F34DAB661" box="[982,1018,1297,1320]" italics="true" pageId="6" pageNumber="67">Est</emphasis>
,
<emphasis id="B94FEA951228393DFBB4FA8E3313B661" box="[1033,1075,1296,1320]" italics="true" pageId="6" pageNumber="67">Esb</emphasis>
,
<emphasis id="B94FEA951228393DFBFFFA8E3341B661" box="[1090,1121,1296,1320]" italics="true" pageId="6" pageNumber="67">Eb</emphasis>
1
<emphasis id="B94FEA951228393DFBCAFA8E33B6B661" box="[1143,1174,1296,1320]" italics="true" pageId="6" pageNumber="67">Eb</emphasis>
3), two dorsal (
<emphasis id="B94FEA951228393DFA80FA8E327FB661" box="[1341,1375,1296,1320]" italics="true" pageId="6" pageNumber="67">Db</emphasis>
,
<emphasis id="B94FEA951228393DFAD2FA8F32ABB661" box="[1391,1419,1297,1320]" italics="true" pageId="6" pageNumber="67">Dt</emphasis>
); 10 on fixed finger, four external (
<emphasis id="B94FEA951228393DFDB0FAA63501B606" box="[525,545,1336,1359]" italics="true" pageId="6" pageNumber="67">et</emphasis>
,
<emphasis id="B94FEA951228393DFD92FAA6356EB606" box="[559,590,1336,1359]" italics="true" pageId="6" pageNumber="67">est</emphasis>
,
<emphasis id="B94FEA951228393DFDE1FAA935A1B606" box="[604,641,1335,1359]" italics="true" pageId="6" pageNumber="67">esb</emphasis>
,
<emphasis id="B94FEA951228393DFD2DFAA9358BB606" box="[656,683,1335,1359]" italics="true" pageId="6" pageNumber="67">eb</emphasis>
), four dorsal (
<emphasis id="B94FEA951228393DFCF1FAA93442B606" box="[844,866,1335,1359]" italics="true" pageId="6" pageNumber="67">dt</emphasis>
,
<emphasis id="B94FEA951228393DFCCDFAA934B1B606" box="[880,913,1335,1359]" italics="true" pageId="6" pageNumber="67">dst</emphasis>
,
<emphasis id="B94FEA951228393DFC23FAA934CFB606" box="[926,1007,1335,1359]" italics="true" pageId="6" pageNumber="67">dsb, db</emphasis>
), two internal (
<emphasis id="B94FEA951228393DFB20FAA6338CB606" box="[1181,1196,1336,1359]" italics="true" pageId="6" pageNumber="67">it</emphasis>
,
<emphasis id="B94FEA951228393DFB06FAA933F1B606" box="[1211,1233,1335,1359]" italics="true" pageId="6" pageNumber="67">ib</emphasis>
).
</paragraph>
<paragraph id="8B8436871228393DFF74FAC236BFB696" blockId="6.[151,1436,1221,2008]" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFF74FAC237DEB63A" box="[201,254,1372,1395]" italics="true" pageId="6" pageNumber="67">Legs</emphasis>
: femur, patella and tibia with pro- and retrolateral surfaces smooth; tibial spurs absent; basitarsi IIV scarcely setose; prolateral and retrolateral pedal spurs present; telotarsi IIV scarcely setose, ventral surface with a median row of spinules, comprising six spinules on I, six on II, eight on III, and nine on IV; ungues well-developed, curved, equal in length.
</paragraph>
<paragraph id="8B8436871228393DFF74FA7234C7B56E" blockId="6.[151,1436,1221,2008]" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFF74FA72363FB54A" box="[201,287,1516,1539]" italics="true" pageId="6" pageNumber="67">Tergites</emphasis>
: Pre-tergites smooth. Post-tergites IVI, surfaces smooth. Post-tergite VII with four vestigial carinae, comprising few posterior small granules; posterior margin slightly granular.
</paragraph>
<paragraph id="8B8436871228393DFF74F9AD333AB526" blockId="6.[151,1436,1221,2008]" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFF74F9AD3605B503" box="[201,293,1587,1610]" italics="true" pageId="6" pageNumber="67">Sternum</emphasis>
(
<figureCitation id="13002A021228393DFE8BF9AC36AFB503" box="[310,399,1586,1610]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="6" pageNumber="67">Fig. 1B</figureCitation>
): Shape subpentagonal with two well developed lateral lobes; lateral margins sub-parallel; posterior width slightly greater than length; posterior depression deeply marked.
</paragraph>
<paragraph id="8B8436871228393DFF74F9E2366AB593" blockId="6.[151,1436,1221,2008]" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFF74F9E23608B5DA" box="[201,296,1660,1683]" italics="true" pageId="6" pageNumber="67">Pectines</emphasis>
(
<figureCitation id="13002A021228393DFE84F9E436B1B5DB" box="[313,401,1658,1682]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="6" pageNumber="67">Fig. 1B</figureCitation>
): Pectinal plate with a median sulcus, lateral margins sub-parallel. Lamella comprising six plates; marginal and basal plates occupying more than half of the lamella. Tooth count 910; proximal teeth slightly smaller.
</paragraph>
<paragraph id="8B8436871228393DFF74F97935E0B46B" blockId="6.[151,1436,1221,2008]" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFF74F9793609B5B7" box="[201,297,1767,1790]" italics="true" pageId="6" pageNumber="67">Sternites</emphasis>
: Sternites IIIVI, surfaces smooth, each with pair of small, rounded spiracles, situated lateromedially (
<figureCitation id="13002A021228393DFF22F89437D6B46B" box="[159,246,1802,1826]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="6" pageNumber="67">Fig. 1B</figureCitation>
); sternite VII, surface smooth, acarinate.
</paragraph>
<paragraph id="8B8436871228393CFF74F8AE3392B2CE" blockId="6.[151,1436,1221,2008]" lastBlockId="7.[151,1436,151,789]" lastPageId="7" lastPageNumber="68" pageId="6" pageNumber="67">
<emphasis id="B94FEA951228393DFF74F8AE361EB40E" box="[201,318,1840,1863]" italics="true" pageId="6" pageNumber="67">Metasoma</emphasis>
(
<figureCitation id="13002A021228393DFEF2F8B136FBB40E" box="[335,475,1839,1863]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="6" pageNumber="67">Fig. 1C, 1D</figureCitation>
): Segments IIV: DSM carinae absent, comprising only fine granulation on median third of segment IV; DL carinae complete with the posterior granule bigger, obsolete and sparsely granular on segment I, slightly more developed in segments IIIV; LSM carinae absent; LM carinae complete, coarsely granular on segments IIV; LIM carinae complete and obsolete on I, absent on segments II-IV; VL carinae absent on I and II, obsolete on III and IV with scarce granules, slightly more developed in IV; VSM carinae absent on segments IIII, few granules medially on IV; dorsal intercarinal surfaces smooth; lateral intercarinal surfaces mostly smooth, with small scattered granules, ventral intercarinal surfaces smooth on IIII, with small scattered granules on segment IV. Segment V: dorsal intercarinal surfaces sparsely and finely granular; lateral and ventral surfaces coarsely granular; DL carinae complete, granular; LM carinae granular on anterior two-thirds; VL and VM carinae complete, well developed, costate granular; VSM carinae with sparse granules more evident on anterior half. Segment I with one pair of LM setae, one pair of VL and VSM setae; segments IIIV with one pair of LM setae, two pairs of VL and VSM setae; segment V with three pairs of LM and VL setae, two pairs of VSM setae.
</paragraph>
<paragraph id="8B8436871229393CFF74FE0A3548B286" blockId="7.[151,1436,151,789]" pageId="7" pageNumber="68">
<emphasis id="B94FEA951229393CFF74FE0A362FB2E5" box="[201,271,404,428]" italics="true" pageId="7" pageNumber="68">Telson</emphasis>
(
<figureCitation id="13002A021229393CFE9CFE0A368AB2E5" box="[289,426,404,428]" captionStart="FIGURE 1 A D" captionStartId="4.[151,250,1699,1722]" captionTargetBox="[163,1422,185,1675]" captionTargetId="figure@4.[163,1423,185,1676]" captionTargetPageId="4" captionText="FIGURE 1 A D. Adult female holotype of Teuthraustes camposi (Mello-Leitão, 1939) comb. n.. A, carapace; B, coxosternal region, genital operculum and pectines; C, metasomal segment V and telson, ventral view; D, metasomal segments II V and telson, lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/215451/files/figure.png" pageId="7" pageNumber="68">Fig. 1C, 1D</figureCitation>
): Vesicle globose; ventral and lateral surfaces weakly granular, dorsal surface smooth and flat; aculeus elongated, shallowly curved.
</paragraph>
</subSubSection>
<subSubSection id="C321650C1229393CFF74FE453493B05D" pageId="7" pageNumber="68" type="distribution">
<paragraph id="8B8436871229393CFF74FE453493B05D" blockId="7.[151,1436,151,789]" pageId="7" pageNumber="68">
<emphasis id="B94FEA951229393CFF74FE453642B2BD" bold="true" box="[201,354,475,500]" pageId="7" pageNumber="68">Distribution.</emphasis>
<taxonomicName id="4C3B4D041229393CFED1FE423541B2BD" box="[364,609,476,500]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA951229393CFED1FE423541B2BD" box="[364,609,476,500]" italics="true" pageId="7" pageNumber="68">Teuthraustes camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE1229393CFDD6FE4235EFB2BD" box="[619,719,476,500]" pageId="7" pageNumber="68">comb. n.</taxonomicNameLabel>
is currently known only from the Cañar Province of
<collectingCountry id="F32C76171229393CFA82FE4232BCB2BD" box="[1343,1436,476,500]" name="Ecuador" pageId="7" pageNumber="68">Ecuador</collectingCountry>
(
<figureCitation id="13002A021229393CFF22FE6137C3B15E" box="[159,227,511,535]" captionStart="FIGURE 3" captionStartId="6.[151,249,1159,1182]" captionTargetBox="[245,1341,193,1138]" captionTargetId="figure@6.[245,1341,193,1138]" captionTargetPageId="6" captionText="FIGURE 3. Map of Ecuador plotting the type locality of Teuthraustes camposi (Mello-Leitão, 1939) comb. n." httpUri="https://zenodo.org/record/215453/files/figure.png" pageId="7" pageNumber="68">Fig. 3</figureCitation>
). The label of the specimen refers “montañas de Cañar”, but in the original description
<bibRefCitation id="EFAA4B761229393CFB0AFE6132BCB15E" author="Mello-Leitao" box="[1207,1436,511,535]" pageId="7" pageNumber="68" refString="Mello-Leitao, C. de. (1939) Una nueva especie de alacran del genero Chactas. Physis, 17, 147 - 148." type="journal article" year="1939">Mello-Leitão (1939)</bibRefCitation>
indicates only “
<collectingCountry id="F32C76171229393CFEFBFDBA3688B175" box="[326,424,548,572]" name="Ecuador" pageId="7" pageNumber="68">Ecuador</collectingCountry>
” as the
<typeStatus id="548088251229393CFDBEFDBB3513B174" box="[515,563,549,573]" pageId="7" pageNumber="68">type</typeStatus>
locality. The province Cañar is situated in the Central Andes of
<collectingCountry id="F32C76171229393CFAB2FDBA324EB175" box="[1295,1390,548,572]" name="Ecuador" pageId="7" pageNumber="68">Ecuador</collectingCountry>
, the region comprises inter-Andean valleys, currently the largest percentage of its land is dedicated to agriculture; the specimen probably was collected around the village Cañar at
<quantity id="4CC39B621229393CFCEAFDF2348EB1CD" box="[855,942,620,644]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="68" unit="m" value="3000.0">3000 m</quantity>
approximately.
<bibRefCitation id="EFAA4B761229393CFBDBFDF2326FB1CD" author="Mello-Leitao" box="[1126,1359,620,644]" pageId="7" pageNumber="68" refString="Mello-Leitao, C. de. (1945) Escorpioes sul-americanos. Arquivos do Museu Nacional, Rio de Janeiro, 40, 7 - 468." type="journal article" year="1945">Mello-Leitão (1945)</bibRefCitation>
, in his monograph of South American scorpions, erroneously mentioned Guayaquil as the
<typeStatus id="548088251229393CFBE0FD0E33ADB1E1" box="[1117,1165,656,680]" pageId="7" pageNumber="68">type</typeStatus>
locality for
<taxonomicName id="4C3B4D041229393CFA9DFD0F32BCB1E1" box="[1312,1436,657,680]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="camposi" status="comb. nov.">
<emphasis id="B94FEA951229393CFA9DFD0F32BCB1E1" box="[1312,1436,657,680]" italics="true" pageId="7" pageNumber="68">T. camposi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C57EE1229393CFF2AFD2A37DCB185" box="[151,252,692,716]" pageId="7" pageNumber="68">comb. n.</taxonomicNameLabel>
The Ecuadorian species of the genus
<taxonomicName id="4C3B4D041229393CFD00FD2A346BB185" box="[701,843,692,716]" class="Arachnida" family="Chactidae" genus="Teuthraustes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEA951229393CFD00FD2A346BB185" box="[701,843,692,716]" italics="true" pageId="7" pageNumber="68">Teuthraustes</emphasis>
</taxonomicName>
are distributed along the inter-Andean valleys and mountain rainforests in eastern and western slopes of the Andes (
<bibRefCitation id="EFAA4B761229393CFC2EFD49336CB1A6" author="Lourenco" box="[915,1100,727,751]" pageId="7" pageNumber="68" refString="Lourenco, W. R. (1995) Les scorpions (Chelicerata, Scorpiones) de l'Equateur avec quelques considerations sur la biogeographie et la diversite des especes. Revue Suisse de Zoologie, 102 (1), 61 - 88." type="journal article" year="1995">Lourenço, 1995</bibRefCitation>
), and therefore it is doubtful whether this genus occurs in Guayaquil, situated near the Pacific coast.
</paragraph>
</subSubSection>
</treatment>
</document>