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<document id="16EE4539662DA631A5559DE1E356C439" ID-DOI="10.11646/zootaxa.1817.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5125134" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627013020755" checkinUser="felipe" docAuthor="Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang &amp; Wang, Xinhua" docDate="2008" docId="03926A1E9E5E7030FF58FF51FE2F92D6" docLanguage="en" docName="zt01817p017.pdf" docOrigin="Zootaxa 1817 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Boreoheptagyia sasai Makarchenko et Endo 2008, sp. n." docType="treatment" docVersion="4" lastPageNumber="11" masterDocId="FFAB12669E54703AFFCFFFC6FF8B9563" masterDocTitle="A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species" masterLastPageNumber="17" masterPageNumber="1" pageNumber="11" updateTime="1699014933596" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="AAD946D9CE51F30E2A0F1A1C7EAFB428" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BEB15C3D3F8D0EF2FA4333D39F4145FF">
<mods:title id="B14DDA1CB95303FFE0A7A876CF85EC0B">A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species</mods:title>
</mods:titleInfo>
<mods:name id="EAD7E7D88B7CB7D9C6BACDFC59215C95" type="personal">
<mods:role id="29DA84FA3B533CF8CD17C07166893091">
<mods:roleTerm id="2146ED38807C15F655BAC7C92639FB20">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9C3F50FDE48265B277BB0A100DFF6FF3">Makarchenko, Eugenyi A.</mods:namePart>
<mods:affiliation id="6BB9D6B3512CCF511702405BF20F3AA9">Institute of Biology and Soil Sciences, Russian Academy of Sciences, Far East Branch, 100 let Vladivostoku Avenue, Vladivostok 690022 Russia. E-mail: makarchenko @ biosoil. ru Koporasu-Tamura-C, Minami-machi Higashi- 5 - 1 - 14, Obihiro, Hokkaido 080 - 0855, Japan. E-mail: endito @ snow. plala. or. jp Life Science College of Nankai University, Tianjin, 300071 China. E-mail: xhwang @ nankai. edu. cn Corresponding author</mods:affiliation>
</mods:name>
<mods:name id="192E5F6682EE5976C603CF1CA0DD7D9F" type="personal">
<mods:role id="B9029767E6160046AA9154C019F48AF9">
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<mods:namePart id="610C4D91DB49DA2E06159F17359B0CF5">Endo, Kazuo</mods:namePart>
</mods:name>
<mods:name id="7F58A33ADFB29010D80C9C26D66E8C64" type="personal">
<mods:role id="E08954785ECB74421950FB8F79CE7F81">
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<mods:namePart id="2153F709CCDDE7CE9675D44F622667E4">Wu, Jingyang</mods:namePart>
</mods:name>
<mods:name id="D6FCE7AA0C108FD06547B4EF072AC8DA" type="personal">
<mods:role id="E48B340B2716109CC37175428EE85EB1">
<mods:roleTerm id="43AF0A6A57BCD91ACAD2843D3348EDB8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="23604504C34D8AAC792E28C603EDAA38">Wang, Xinhua</mods:namePart>
</mods:name>
<mods:typeOfResource id="BF6CB4C70EECE8DB5DD16F0A46645A97">text</mods:typeOfResource>
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<mods:titleInfo id="2B119BDB33122CDD621AAECF53040BA3">
<mods:title id="FCA3324286B14AB066B85B879108027B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BA2F33435D41B6FF0BFB92231C10A683">
<mods:date id="EAC69E0036BE82BE4FEE3A5B6C019F8C">2008</mods:date>
<mods:detail id="ECAA8B1D78E122340FB8846A95E562FA" type="pubDate">
<mods:number id="4C73ECDB6F2EA88434E632878A621576">2008-07-07</mods:number>
</mods:detail>
<mods:detail id="7590E52AD0B3FD493D719AE20AF89184" type="volume">
<mods:number id="57BDE771FAFCACECA8ABF213A5536AA4">1817</mods:number>
</mods:detail>
<mods:detail id="C426120BAB2EB4712E85C819F10C48F9" type="issue">
<mods:number id="1455E9BF7F5231E129D00447A1B1C5A1">1</mods:number>
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<mods:start id="C7421884DD48F0E76F5F8898627FA221">1</mods:start>
<mods:end id="AA2461DEA9F2B35AA0D68D103B2638C4">17</mods:end>
</mods:extent>
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<mods:location id="FD4CE13AB36B26E70BB9BA7564749B90">
<mods:url id="61513DE61F7B7F1573B799184E4C2F93">https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1</mods:url>
</mods:location>
<mods:classification id="491EB5B69C23DE4ECA6C4AFC8F953623">journal article</mods:classification>
<mods:identifier id="6F3A22752E31894352262815FF2DA89F" type="DOI">10.11646/zootaxa.1817.1.1</mods:identifier>
<mods:identifier id="0A1C347405E330696DFECD8EEBEA4F6D" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="22E3A48BF6E22AFD2D27C8B14C9812B3" type="Zenodo-Dep">5125134</mods:identifier>
</mods:mods>
<treatment id="03926A1E9E5E7030FF58FF51FE2F92D6" ID-DOI="http://doi.org/10.5281/zenodo.5126518" ID-Zenodo-Dep="5126518" LSID="urn:lsid:plazi:treatment:03926A1E9E5E7030FF58FF51FE2F92D6" httpUri="http://treatment.plazi.org/id/03926A1E9E5E7030FF58FF51FE2F92D6" lastPageNumber="11" pageId="10" pageNumber="11">
<subSubSection id="C32188839E5E7030FF58FF51FC8D95D2" box="[151,774,151,177]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8B84DB089E5E7030FF58FF51FC8D95D2" blockId="10.[151,774,151,212]" box="[151,774,151,177]" pageId="10" pageNumber="11">
<heading id="D0CC6C649E5E7030FF58FF51FC8D95D2" bold="true" box="[151,774,151,177]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B94F071A9E5E7030FF58FF51FC8D95D2" bold="true" box="[151,774,151,177]" pageId="10" pageNumber="11">
<taxonomicName id="4C3BA08B9E5E7030FF58FF51FD3C95D2" ID-CoL="9HZNP" authority="Makarchenko et Endo" authorityName="Makarchenko et Endo" authorityYear="2008" box="[151,695,151,177]" class="Insecta" family="Chironomidae" genus="Boreoheptagyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sasai" status="sp. nov.">
<emphasis id="B94F071A9E5E7030FF58FF51FE1395D2" bold="true" box="[151,408,151,177]" italics="true" pageId="10" pageNumber="11">Boreoheptagyia sasai</emphasis>
Makarchenko
<emphasis id="B94F071A9E5E7030FD99FF51FDE095D2" bold="true" box="[598,619,151,177]" italics="true" pageId="10" pageNumber="11">et</emphasis>
Endo
</taxonomicName>
,
<taxonomicNameLabel id="A27CBA619E5E7030FD0BFF51FC8D95D2" box="[708,774,151,177]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32188839E5E7030FF58FF7CFEA695B7" box="[151,301,186,212]" pageId="10" pageNumber="11" type="description">
<paragraph id="8B84DB089E5E7030FF58FF7CFEA695B7" blockId="10.[151,774,151,212]" box="[151,301,186,212]" pageId="10" pageNumber="11">
(
<figureCitation id="1300C78D9E5E7030FF50FF7CFEAE95B7" box="[159,293,186,212]" captionStart="FIGURES 26–33" captionStartId="11.[151,269,1611,1635]" captionTargetBox="[156,1431,187,1587]" captionTargetId="figure-120@11.[156,1431,181,1587]" captionTargetPageId="11" captionText="FIGURES 26–33. Boreoheptagyia sasai Makarchenko et Endo, sp. n., male from Magadan region (26–30) and Hokkaido (31–33). 26. Scutum, dorsal view. 27. Scutum, lateral view. 28, 30, 32. Hypopygium with tergite IX removed, dorsal view. 29, 31. Tergite IX. 33. Transverse sternapodeme. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5125150" httpUri="https://zenodo.org/record/5125150/files/figure.png" pageId="10" pageNumber="11">Figs 26–33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32188839E5E7030FF58FECCFBFE9442" box="[151,1141,266,289]" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8B84DB089E5E7030FF58FECCFBFE9442" blockId="10.[151,1141,266,289]" box="[151,1141,266,289]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB2BFC269E5E7030FF58FECCFBFE9442" box="[151,1141,266,289]" pageId="10" pageNumber="11">
<taxonomicName id="4C3BA08B9E5E7030FF58FECCFC649442" ID-CoL="MJSF" authority="(Tokunaga) sensu Makarchenko (1981: 89" authorityName="sensu Makarchenko" authorityYear="1981" baseAuthorityName="Tokunaga" baseAuthorityYear="1936" box="[151,1007,266,289]" class="Insecta" family="Chironomidae" genus="Boreoheptagyia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brevitarsis">
<treatmentCitation id="0A9AFD199E5E7030FF58FECCFC0F9442" author="Makarchenko, E. A." box="[151,900,266,289]" page="89" pageId="10" pageNumber="11" year="1981">
<emphasis id="B94F071A9E5E7030FF58FECCFE339442" box="[151,440,266,289]" italics="true" pageId="10" pageNumber="11">Boreoheptagyia brevitarsis</emphasis>
(Tokunaga)
<emphasis id="B94F071A9E5E7030FDF0FECCFDF19442" box="[575,634,266,289]" italics="true" pageId="10" pageNumber="11">sensu</emphasis>
<bibRefCitation id="EFAAA6F99E5E7030FD4EFECCFC0F9442" author="Makarchenko, E. A." box="[641,900,266,289]" pageId="10" pageNumber="11" pagination="45 - 51" refId="ref8837" refString="Makarchenko, E. A. (1981) Taxonomy and distribution of some chironomids of subfamily Diamesinae (Diptera, Chironomidae) from the Soviet Far East. In: Invertebrate animals in ecosystems of salmon rivers of the Far East. DVNC AN SSSR Press, Vladivostok, pp. 45 - 51. [In Russian.]" type="book chapter" year="1981">Makarchenko (1981: 89</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9AFD199E5E7030FC40FECCFC649442" author="Makarchenko, E. A." box="[911,1007,266,289]" page="38" pageId="10" pageNumber="11" year="1985">
<bibRefCitation id="EFAAA6F99E5E7030FC40FECCFC649442" author="Makarchenko, E. A." box="[911,1007,266,289]" pageId="10" pageNumber="11" refId="ref8898" refString="Makarchenko, E. A. (1985) Chironomids of the Soviet Far East. Subfamilies Podonominae, Diamesinae and Prodiamesinae (Diptera, Chironomidae). DVNC AN SSSR Press, Vladivostok, 208 pp. [In Russian.]" type="book" year="1985">1985: 38</bibRefCitation>
</treatmentCitation>
</taxonomicName>
,
<treatmentCitation id="0A9AFD199E5E7030FC35FECCFBE29442" author="Makarchenko, E. A." box="[1018,1129,266,289]" page="259" pageId="10" pageNumber="11" year="2006">
<bibRefCitation id="EFAAA6F99E5E7030FC35FECCFBE29442" author="Makarchenko, E. A." box="[1018,1129,266,289]" pageId="10" pageNumber="11" pagination="607 - 621" refId="ref9028" refString="Makarchenko, E. A. (2006) Subfamily Diamesinae. In: Key to the Insects of Russian Far East. Vol. 6. Diptera and Sipphonaptera. Pt 4. Dal'nauka, Vladivostok, pp. 253 - 276, 468 - 480, 607 - 621. [In Russian.]" type="journal article" year="2006">2006: 259</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32188839E5E7030FF58FE94FAF5973F" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8B84DB089E5E7030FF58FE94FAF5973F" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF58FE94FEDD940F" bold="true" box="[151,342,338,364]" pageId="10" pageNumber="11">Type material.</emphasis>
<materialsCitation id="3B53D1559E5E7030FEACFE94FC3D94DF" collectingDate="1977-07-11" collectingDateMax="1977-07-27" collectingDateMin="1977-07-11" collectionCode="IBSS" collectorName="E. Makarchenko &amp; S. Kocharina" country="Russia" county="Ten'kinsky County" elevation="1" location="Kolyma River Basin" municipality="North-Eastern" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Magadan" typeStatus="holotype">
<typeStatus id="548065AA9E5E7030FEACFE94FE5B940F" box="[355,464,338,364]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F32C9B989E5E7030FDE1FE94FD10940F" box="[558,667,338,364]" name="Russia" pageId="10" pageNumber="11">RUSSIA</collectingCountry>
:
<collectingMunicipality id="6BE041729E5E7030FD61FE94FCD8940F" box="[686,851,338,364]" pageId="10" pageNumber="11">North-Eastern</collectingMunicipality>
Russia,
<collectingCounty id="62E5A3849E5E7030FC08FE94FB3B940F" box="[967,1200,338,364]" pageId="10" pageNumber="11">Ten’kinsky County</collectingCounty>
,
<collectingRegion id="49FF15EA9E5E7030FB0BFE94FA1C940F" box="[1220,1431,338,364]" country="Russia" name="Magadan" pageId="10" pageNumber="11">Magadan Region</collectingRegion>
,
<location id="8EE48DD39E5E7030FF58FEBCFE1B94F7" LSID="urn:lsid:plazi:treatment:03926A1E9E5E7030FF58FF51FE2F92D6:8EE48DD39E5E7030FF58FEBCFE1B94F7" box="[151,400,378,404]" country="Russia" county="Ten'kinsky County" municipality="North-Eastern" name="Kolyma River Basin" pageId="10" pageNumber="11" stateProvince="Magadan">
<location id="8EE48DD39E5E7030FF58FEBCFECB94F7" LSID="urn:lsid:plazi:treatment:03926A1E9E5E7030FF58FF51FE2F92D6:8EE48DD39E5E7030FF58FEBCFECB94F7" box="[151,320,378,404]" country="Russia" county="Ten'kinsky County" municipality="North-Eastern" name="Kolyma River" pageId="10" pageNumber="11" stateProvince="Magadan">
Kolyma
<location id="8EE48DD39E5E7030FF30FEBCFECB94F7" LSID="urn:lsid:plazi:treatment:03926A1E9E5E7030FF58FF51FE2F92D6:8EE48DD39E5E7030FF30FEBCFECB94F7" box="[255,320,378,404]" country="Russia" county="Ten'kinsky County" municipality="North-Eastern" name="River Basin" pageId="10" pageNumber="11" stateProvince="Magadan">River</location>
</location>
Basin
</location>
,
<location id="8EE48DD39E5E7030FE51FEBCFDC894F7" LSID="urn:lsid:plazi:treatment:03926A1E9E5E7030FF58FF51FE2F92D6:8EE48DD39E5E7030FE51FEBCFDC894F7" box="[414,579,378,404]" country="Russia" county="Ten'kinsky County" municipality="North-Eastern" name="Vlastnyi Peak" pageId="10" pageNumber="11" stateProvince="Magadan">Vlastnyi Peak</location>
not far from
<location id="8EE48DD39E5E7030FD26FEBCFC6894F7" LSID="urn:lsid:plazi:treatment:03926A1E9E5E7030FF58FF51FE2F92D6:8EE48DD39E5E7030FD26FEBCFC6894F7" box="[745,995,378,404]" country="Russia" county="Ten'kinsky County" municipality="North-Eastern" name="Sibit-Tyellah Village" pageId="10" pageNumber="11" stateProvince="Magadan">Sibit-Tyellah Village</location>
,
<location id="8EE48DD39E5E7030FC3EFEBCFB1F94F7" LSID="urn:lsid:plazi:treatment:03926A1E9E5E7030FF58FF51FE2F92D6:8EE48DD39E5E7030FC3EFEBCFB1F94F7" box="[1009,1172,378,404]" country="Russia" county="Ten'kinsky County" municipality="North-Eastern" name="Olen' Stream" pageId="10" pageNumber="11" stateProvince="Magadan">Olen’ Stream</location>
,
<elevation id="00163C3B9E5E7030FB6EFEBCFA1D94F7" box="[1185,1430,378,404]" metricMagnitude="0" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="10" pageNumber="11" unit="m" value="1.2" valueMax="1.3" valueMin="1.1">
<quantity id="4CC376ED9E5E7030FB6EFEBCFADF94F7" box="[1185,1364,378,404]" metricMagnitude="0" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="10" pageNumber="11" unit="m" value="1.2" valueMax="1.3" valueMin="1.1">1.100–1.300 m</quantity>
a.s.l.
</elevation>
,
<date id="FF85FDC89E5E7030FF58FE64FEC194DF" box="[151,330,418,444]" pageId="10" pageNumber="11" value="1977-07-11" valueMax="1977-07-27" valueMin="1977-07-11">
<collectingDate id="EFC104209E5E7030FF58FE64FEC194DF" box="[151,330,418,444]" pageId="10" pageNumber="11" value="1977-07-11" valueMax="1977-07-27" valueMin="1977-07-11">11–27.vii.1977</collectingDate>
</date>
,
<collectorName id="26CEBEDE9E5E7030FE97FE64FD9094DF" box="[344,539,418,444]" pageId="10" pageNumber="11">E. Makarchenko</collectorName>
&amp;
<collectorName id="26CEBEDE9E5E7030FD8AFE64FD6B94DF" box="[581,736,418,444]" pageId="10" pageNumber="11">S. Kocharina</collectorName>
(
<collectionCode id="ED2A43CD9E5E7030FD3EFE64FCBB94DF" box="[753,816,418,444]" pageId="10" pageNumber="11">IBSS</collectionCode>
FEBRAS)
</materialsCitation>
.
<typeStatus id="548065AA9E5E7030FC0CFE64FBB294DF" box="[963,1081,418,444]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3D10819E5E7030FB89FE64FB3894DF" box="[1094,1203,418,444]" pageId="10" pageNumber="11" type="male">11 males</specimenCount>
,
<specimenCount id="9D3D10819E5E7030FB0FFE64FAB894DF" box="[1216,1331,418,444]" pageId="10" pageNumber="11" type="female">4 females</specimenCount>
as
<typeStatus id="548065AA9E5E7030FA91FE64FF469487" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
.
<materialsCitation id="3B53D1559E5E7030FF15FE0CFF75976F" collectingDate="1999-06-11" collectingDateMax="1999-06-21" collectingDateMin="1999-06-11" collectorName="K. Endo" country="Japan" elevation="700" location="Yutomuraushi River" municipality="Shintoku" pageId="10" pageNumber="11" specimenCount="2" specimenCount-male="2" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCount id="9D3D10819E5E7030FF15FE0CFEB29487" box="[218,313,458,484]" pageId="10" pageNumber="11" type="male">2 males</specimenCount>
,
<collectingCountry id="F32C9B989E5E7030FE89FE0CFE289487" box="[326,419,458,484]" name="Japan" pageId="10" pageNumber="11">JAPAN</collectingCountry>
:
<collectingRegion id="49FF15EA9E5E7030FE7FFE0CFDAD9487" box="[432,550,458,484]" country="Japan" name="Hokkaido" pageId="10" pageNumber="11">Hokkaido</collectingRegion>
,
<collectingMunicipality id="6BE041729E5E7030FDFBFE0CFD2A9487" box="[564,673,458,484]" pageId="10" pageNumber="11">Shintoku</collectingMunicipality>
,
<location id="8EE48DD39E5E7030FD61FE0CFC149487" LSID="urn:lsid:plazi:treatment:03926A1E9E5E7030FF58FF51FE2F92D6:8EE48DD39E5E7030FD61FE0CFC149487" box="[686,927,458,484]" country="Japan" municipality="Shintoku" name="Yutomuraushi River" pageId="10" pageNumber="11" stateProvince="Hokkaido">Yutomuraushi River</location>
,
<elevation id="00163C3B9E5E7030FC63FE0CFBB29487" box="[940,1081,458,484]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="11" unit="m" value="700.0">
<quantity id="4CC376ED9E5E7030FC63FE0CFC729487" box="[940,1017,458,484]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="11" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
,
<collectingMethod id="527AA31F9E5E7030FB87FE0CFB559487" box="[1096,1246,458,484]" pageId="10" pageNumber="11">Malaise trap</collectingMethod>
,
<date id="FF85FDC89E5E7030FB22FE0CFA1C9487" box="[1261,1431,458,484]" pageId="10" pageNumber="11" value="1999-06-11" valueMax="1999-06-21" valueMin="1999-06-11">
<collectingDate id="EFC104209E5E7030FB22FE0CFA1C9487" box="[1261,1431,458,484]" pageId="10" pageNumber="11" value="1999-06-11" valueMax="1999-06-21" valueMin="1999-06-11">11–21.vi.1999</collectingDate>
</date>
,
<collectorName id="26CEBEDE9E5E7030FF58FE34FF75976F" box="[151,254,498,524]" pageId="10" pageNumber="11">K. Endo</collectorName>
</materialsCitation>
;
<specimenCount id="9D3D10819E5E7030FEC3FE34FEEB976F" box="[268,352,498,524]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
, same as previous except
<date id="FF85FDC89E5E7030FD55FE34FCD6976F" box="[666,861,498,524]" pageId="10" pageNumber="11" value="1999-06-21" valueMax="1999-07-02" valueMin="1999-06-21">21.vi–2.vii.1999</date>
, Malaise trap, K. Endo;
<materialsCitation id="3B53D1559E5E7030FB4AFE34FBF19757" collectingDate="2000-06-05" collectingDateMax="2000-06-15" collectingDateMin="2000-06-05" collectorName="K. Endo" country="Japan" elevation="620" location="Opirineppu River" municipality="Obihiro" pageId="10" pageNumber="11" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCount id="9D3D10819E5E7030FB4AFE34FB51976F" box="[1157,1242,498,524]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
,
<specimenCount id="9D3D10819E5E7030FB27FE34FAD8976F" box="[1256,1363,498,524]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
,
<collectingRegion id="49FF15EA9E5E7030FAAFFE34FF509757" country="Japan" name="Hokkaido" pageId="10" pageNumber="11">Hokkaido</collectingRegion>
,
<collectingMunicipality id="6BE041729E5E7030FF28FDDCFECE9757" box="[231,325,538,564]" pageId="10" pageNumber="11">Obihiro</collectingMunicipality>
,
<location id="8EE48DD39E5E7030FE9DFDDCFDA89757" LSID="urn:lsid:plazi:treatment:03926A1E9E5E7030FF58FF51FE2F92D6:8EE48DD39E5E7030FE9DFDDCFDA89757" box="[338,547,538,564]" country="Japan" municipality="Obihiro" name="Opirineppu River" pageId="10" pageNumber="11" stateProvince="Hokkaido">Opirineppu River</location>
,
<elevation id="00163C3B9E5E7030FDE0FDDCFD319757" box="[559,698,538,564]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="11" unit="m" value="620.0">
<quantity id="4CC376ED9E5E7030FDE0FDDCFDF19757" box="[559,634,538,564]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="11" unit="m" value="620.0">620 m</quantity>
a.s.l.
</elevation>
,
<date id="FF85FDC89E5E7030FD07FDDCFCEE9757" box="[712,869,538,564]" pageId="10" pageNumber="11" value="2000-06-05" valueMax="2000-06-15" valueMin="2000-06-05">
<collectingDate id="EFC104209E5E7030FD07FDDCFCEE9757" box="[712,869,538,564]" pageId="10" pageNumber="11" value="2000-06-05" valueMax="2000-06-15" valueMin="2000-06-05">5–15.vi.2000</collectingDate>
</date>
,
<collectingMethod id="527AA31F9E5E7030FCBDFDDCFB8C9757" box="[882,1031,538,564]" pageId="10" pageNumber="11">Malaise trap</collectingMethod>
,
<collectorName id="26CEBEDE9E5E7030FBDBFDDCFBF19757" box="[1044,1146,538,564]" pageId="10" pageNumber="11">K. Endo</collectorName>
</materialsCitation>
;
<specimenCount id="9D3D10819E5E7030FB48FDDCFB6F9757" box="[1159,1252,538,564]" pageId="10" pageNumber="11" type="male">2 males</specimenCount>
, same as previous except
<date id="FF85FDC89E5E7030FED5FD84FE46973F" box="[282,461,578,604]" pageId="10" pageNumber="11" value="1999-09-13" valueMax="1999-10-01" valueMin="1999-09-13">13.ix–1.x.1999</date>
, K. Endo;
<specimenCount id="9D3D10819E5E7030FD84FD84FD16973F" box="[587,669,578,604]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
, same as previous except
<date id="FF85FDC89E5E7030FC04FD84FBE3973F" box="[971,1128,578,604]" pageId="10" pageNumber="11" value="2000-06-05" valueMax="2000-06-15" valueMin="2000-06-05">5–15.vi.2000</date>
, Malaise trap, K. Endo.
</paragraph>
</subSubSection>
<subSubSection id="C32188839E5E7030FF0AFDACFE349276" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8B84DB089E5E7030FF0AFDACFEB097CF" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AFDACFED997E7" bold="true" box="[197,338,618,644]" pageId="10" pageNumber="11">Etymology.</emphasis>
Named in honor and memory of the famous Japanese chironomidologist and colleague Prof. Manabu Sasa.
</paragraph>
<paragraph id="8B84DB089E5E7030FF0AFD7CFE3F9647" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AFD7CFE5597B7" bold="true" box="[197,478,698,724]" pageId="10" pageNumber="11">Diagnostic characters.</emphasis>
The male is well separated from other species of the genus by the presence of setae on mesanepisternum II and preepisternum, and by the field of small spinules distal of transverse sternapodeme (
<figureCitation id="1300C78D9E5E7030FED0FCCCFE2C9647" box="[287,423,778,804]" captionStart="FIGURES 26–33" captionStartId="11.[151,269,1611,1635]" captionTargetBox="[156,1431,187,1587]" captionTargetId="figure-120@11.[156,1431,181,1587]" captionTargetPageId="11" captionText="FIGURES 26–33. Boreoheptagyia sasai Makarchenko et Endo, sp. n., male from Magadan region (26–30) and Hokkaido (31–33). 26. Scutum, dorsal view. 27. Scutum, lateral view. 28, 30, 32. Hypopygium with tergite IX removed, dorsal view. 29, 31. Tergite IX. 33. Transverse sternapodeme. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5125150" httpUri="https://zenodo.org/record/5125150/files/figure.png" pageId="10" pageNumber="11">Figs 30, 32</figureCitation>
).
</paragraph>
<paragraph id="8B84DB089E5E7030FF0AFCF4FED7962F" blockId="10.[151,1437,338,1973]" box="[197,348,818,844]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AFCF4FE8F962F" bold="true" box="[197,260,818,844]" pageId="10" pageNumber="11">Male</emphasis>
(n = 9)
</paragraph>
<paragraph id="8B84DB089E5E7030FF0AFC9CFB7C9617" blockId="10.[151,1437,338,1973]" box="[197,1271,858,884]" pageId="10" pageNumber="11">
Total length
<quantity id="4CC376ED9E5E7030FE96FC9CFE629617" box="[345,489,858,884]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="10" pageNumber="11" unit="mm" value="2.55" valueMax="2.7" valueMin="2.4">2.4–2.7 mm</quantity>
. Wing length
<quantity id="4CC376ED9E5E7030FD43FC9CFCB19617" box="[652,826,858,884]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" metricValueMax="2.13" metricValueMin="1.61" pageId="10" pageNumber="11" unit="mm" value="1.87" valueMax="2.13" valueMin="1.61">1.61–2.13 mm</quantity>
. Total length / wing length 1.17–1.29.
</paragraph>
<paragraph id="8B84DB089E5E7030FF0AFC44FB4296FF" blockId="10.[151,1437,338,1973]" box="[197,1225,898,924]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AFC44FEC696F8" box="[197,333,898,923]" italics="true" pageId="10" pageNumber="11">Coloration.</emphasis>
Head, thorax and legs brown to dark brown, abdomen grayish to yellowish.
</paragraph>
<paragraph id="8B84DB089E5E7030FF0AFC6CFA1291EF" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AFC6CFE8696A0" box="[197,269,938,963]" italics="true" pageId="10" pageNumber="11">Head.</emphasis>
Temporal setae includes 6–7 postorbitals, 30–31 outer verticals, and 26–31 inner verticals. Clypeus with 11–12 setae. Antenna with 13 flagellomeres, antennal plume reduced, pedicel with 4 setae. Lengths (in µm) of flagellomeres 2–13: 17–29, 25–34, 31–42, 35–46, 39–42, 34–46, 36–46, 38–55, 38–42, 38–46, 34–42, 88–101. Number of setae on flagellomeres 2–13 as: 4–5, 6, 8, 9–10, 8–9, 9–10, 8, 8–9, 8, 8, 7–8, 4. Subapical seta of terminal flagellomere
<quantity id="4CC376ED9E5E7030FE3EFB8CFDE39107" box="[497,616,1098,1124]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="10" pageNumber="11" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">11–15 µm</quantity>
long. AR 0.11–0.20. Lengths (in µm) of palpomeres 1–5: 27–30, 63– 80, 101–109, 92–118, 160–193. Antennal length / palp length 1.35–1.40. Head width / palp length 1.04–1.16.
</paragraph>
<paragraph id="8B84DB089E5E7030FF0AFB5DFACB9067" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AFB5DFEA991D7" box="[197,290,1179,1204]" italics="true" pageId="10" pageNumber="11">Thorax.</emphasis>
Antepronotum with 6–11 lateral setae. Acrostichals 33–36; dorsocentrals
<quantity id="4CC376ED9E5E7030FB55FB5CFA8F91D7" box="[1178,1284,1178,1204]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" metricValueMax="6.604" metricValueMin="6.096" pageId="10" pageNumber="11" unit="in" value="25.0" valueMax="26.0" valueMin="24.0">24–26 in</quantity>
anterior and posterior groups; prealars
<quantity id="4CC376ED9E5E7030FE06FB04FDBB91BF" box="[457,560,1218,1244]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="6.604" metricValueMin="4.826" pageId="10" pageNumber="11" unit="in" value="22.5" valueMax="26.0" valueMin="19.0">19–26 in</quantity>
2 rows; supraalars 2; preepisternals 6–13; median anepisternum II with 4– 6 setae. Scutellum with 58–62 setae. Longest and shortest acrostichals
<quantity id="4CC376ED9E5E7030FC1BFB2CFBC69067" box="[980,1101,1258,1284]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="4.2" metricValueMin="3.6" pageId="10" pageNumber="11" unit="mm" value="39.0" valueMax="42.0" valueMin="36.0">36–42 µm</quantity>
and
<quantity id="4CC376ED9E5E7030FB49FB2CFB769067" box="[1158,1277,1258,1284]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.1" metricValueMin="1.1" pageId="10" pageNumber="11" unit="mm" value="16.0" valueMax="21.0" valueMin="11.0">11–21 µm</quantity>
long.
</paragraph>
<paragraph id="8B84DB089E5E7030FF0AFAD5FC23901C" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AFAD5FE8A904F" box="[197,257,1299,1324]" italics="true" pageId="10" pageNumber="11">Wing</emphasis>
<quantity id="4CC376ED9E5E7030FEC2FAD4FE30904F" box="[269,443,1298,1324]" metricMagnitude="-4" metricUnit="m" metricValue="5.7" metricValueMax="6.2" metricValueMin="5.2" pageId="10" pageNumber="11" unit="mm" value="0.57" valueMax="0.62" valueMin="0.52">0.52–0.62 mm</quantity>
wide. Costal extension
<quantity id="4CC376ED9E5E7030FD2BFAD4FCEA904F" box="[740,865,1298,1324]" metricMagnitude="-2" metricUnit="m" metricValue="7.2" metricValueMax="8.0" metricValueMin="6.4" pageId="10" pageNumber="11" unit="mm" value="72.0" valueMax="80.0" valueMin="64.0">64–80 µm</quantity>
long. Anal lobe well developed, fully fringed. Alula without setae. Squama with 32–48 setae. R with 14 setae, R
<subScript id="17BFD94D9E5E7030FC50FA8CFC2C903B" attach="left" box="[927,935,1354,1368]" fontSize="6" pageId="10" pageNumber="11">1</subScript>
with 13–17 setae, R
<subScript id="17BFD94D9E5E7030FB56FA8CFB3A903B" attach="left" box="[1177,1201,1354,1368]" fontSize="6" pageId="10" pageNumber="11">4+5</subScript>
with 3–7 setae, R
<subScript id="17BFD94D9E5E7030FA4CFA8CFA10903B" attach="left" box="[1411,1435,1354,1368]" fontSize="6" pageId="10" pageNumber="11">2+3</subScript>
usually reduced, often only basal part visible. RM/MCu 1.80–2.25.
</paragraph>
<paragraph id="8B84DB089E5E7030FF0AFA48FF54932E" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AFA48FE8F90C4" box="[197,260,1422,1447]" italics="true" pageId="10" pageNumber="11">Legs.</emphasis>
Spur of foretibia
<quantity id="4CC376ED9E5E7030FE1BFA4BFDC690C4" box="[468,589,1421,1447]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="4.2" metricValueMin="3.4" pageId="10" pageNumber="11" unit="mm" value="38.0" valueMax="42.0" valueMin="34.0">34–42 µm</quantity>
long; spurs of midtibia
<quantity id="4CC376ED9E5E7030FCAAFA4BFC5590C4" box="[869,990,1421,1447]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="4.2" metricValueMin="3.4" pageId="10" pageNumber="11" unit="mm" value="38.0" valueMax="42.0" valueMin="34.0">34–42 µm</quantity>
and
<quantity id="4CC376ED9E5E7030FBD8FA4BFB0490C4" box="[1047,1167,1421,1447]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.8" metricValueMin="3.4" pageId="10" pageNumber="11" unit="mm" value="36.0" valueMax="38.0" valueMin="34.0">34–38 µm</quantity>
long, of hind tibia
<quantity id="4CC376ED9E5E7030FAA0FA4BFF6890AC" metricMagnitude="-2" metricUnit="m" metricValue="4.45" metricValueMax="5.5" metricValueMin="3.4" pageId="10" pageNumber="11" unit="mm" value="44.5" valueMax="55.0" valueMin="34.0">34– 55 µm</quantity>
and
<quantity id="4CC376ED9E5E7030FED4FA73FE1890AC" box="[283,403,1461,1487]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.8" metricValueMin="3.4" pageId="10" pageNumber="11" unit="mm" value="36.0" valueMax="38.0" valueMin="34.0">34–38 µm</quantity>
long. Hind tibial comb with 10–12 setae. Mid ta
<subScript id="17BFD94D9E5E7030FC04FA03FC5890B0" attach="left" box="[971,979,1477,1491]" fontSize="6" pageId="10" pageNumber="11">1</subScript>
with 4–7 pseudospurs, ta
<subScript id="17BFD94D9E5E7030FB33FA03FA8F90B0" attach="left" box="[1276,1284,1477,1491]" fontSize="6" pageId="10" pageNumber="11">2</subScript>
with 1 pseudospur,
<quantity id="4CC376ED9E5E7030FF3BFA26FEE69099" box="[244,365,1504,1530]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="10" pageNumber="11" unit="mm" value="25.0" valueMax="27.0" valueMin="23.0">23–27 µm</quantity>
long; hind ta
<subScript id="17BFD94D9E5E7030FDC8FA36FD84909D" attach="left" box="[519,527,1520,1534]" fontSize="6" pageId="10" pageNumber="11">1</subScript>
with 10–12 pseudospurs, ta
<subScript id="17BFD94D9E5E7030FC98FA36FCD4909D" attach="left" box="[855,863,1520,1534]" fontSize="6" pageId="10" pageNumber="11">2</subScript>
with 1 pseudospur,
<quantity id="4CC376ED9E5E7030FB85FA26FB489099" box="[1098,1219,1504,1530]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="10" pageNumber="11" unit="mm" value="25.0" valueMax="27.0" valueMin="23.0">23–27 µm</quantity>
long. Lengths and proportions of legs of specimens from Kolyma River Basin and from
<collectingRegion id="49FF15EA9E5E7030FC1EF9CDFBCF9346" box="[977,1092,1547,1573]" country="Japan" name="Hokkaido" pageId="10" pageNumber="11">Hokkaido</collectingRegion>
as in
<tableCitation id="C6B9EEB39E5E7030FB43F9CDFB649346" box="[1164,1263,1547,1573]" captionStart="TABLE 5" captionStartId="12.[151,239,514,538]" captionText="TABLE 5. Lengths (in µm) and proportions of legs of Boreoheptagyia sasai Makarchenko et Endo, sp. n., males from the Kolyma River Basin, Northeastern Russia (n = 6)." pageId="10" pageNumber="11">Tables 5</tableCitation>
and
<tableCitation id="C6B9EEB39E5E7030FAE6F9CDFAB29346" box="[1321,1337,1547,1573]" captionStart="TABLE 6" captionStartId="12.[151,239,1106,1130]" captionText="TABLE 6. Lengths (in µm) and proportions of legs of Boreoheptagyia sasai Makarchenko et Endo, sp. n., males from Hokkaido, Japan (n = 3)." pageId="10" pageNumber="11">6</tableCitation>
, respectively.
</paragraph>
<paragraph id="8B84DB089E5E7030FF0AF99DFE349276" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AF99DFED19317" box="[197,346,1627,1652]" italics="true" pageId="10" pageNumber="11">Hypopygium</emphasis>
(
<figureCitation id="1300C78D9E5E7030FEA2F99DFE739316" box="[365,504,1627,1653]" captionStart="FIGURES 26–33" captionStartId="11.[151,269,1611,1635]" captionTargetBox="[156,1431,187,1587]" captionTargetId="figure-120@11.[156,1431,181,1587]" captionTargetPageId="11" captionText="FIGURES 26–33. Boreoheptagyia sasai Makarchenko et Endo, sp. n., male from Magadan region (26–30) and Hokkaido (31–33). 26. Scutum, dorsal view. 27. Scutum, lateral view. 28, 30, 32. Hypopygium with tergite IX removed, dorsal view. 29, 31. Tergite IX. 33. Transverse sternapodeme. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5125150" httpUri="https://zenodo.org/record/5125150/files/figure.png" pageId="10" pageNumber="11">Figs 28–33</figureCitation>
). Tergite IX with 20–27 setae. Laterosternite IX with 12–17 setae. Transverse sternapodeme
<quantity id="4CC376ED9E5E7030FE8EF945FE3193FE" box="[321,442,1667,1693]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" metricValueMax="6.0" metricValueMin="4.6" pageId="10" pageNumber="11" unit="mm" value="53.0" valueMax="60.0" valueMin="46.0">46–60 µm</quantity>
wide. Aedeagal lobe long, narrow; phallapodeme narrow, well visible. With field of small spinules distal of transverse sternapodeme. Gonocoxite
<quantity id="4CC376ED9E5E7030FCA6F96DFC7493A6" box="[873,1023,1707,1733]" metricMagnitude="-1" metricUnit="m" metricValue="2.03" metricValueMax="2.05" metricValueMin="2.01" pageId="10" pageNumber="11" unit="mm" value="203.0" valueMax="205.0" valueMin="201.0">201–205 µm</quantity>
long; basal lobe rounded, with 1–3 short setae. Gonostylus
<quantity id="4CC376ED9E5E7030FE7FF915FDB3938E" box="[432,568,1747,1773]" metricMagnitude="-2" metricUnit="m" metricValue="9.9" metricValueMax="10.3" metricValueMin="9.5" pageId="10" pageNumber="11" unit="mm" value="99.0" valueMax="103.0" valueMin="95.0">95–103 µm</quantity>
long, with wide median part and short narrow apical part, ending in short megaseta. HR 2.20–2.73.
</paragraph>
</subSubSection>
<subSubSection id="C32188839E5E7030FF0AF8E5FA179206" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8B84DB089E5E7030FF0AF8E5FA179206" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AF8E5FEB5925E" bold="true" box="[197,318,1827,1853]" pageId="10" pageNumber="11">Remarks.</emphasis>
The males from Kolyma River Basin and from
<collectingRegion id="49FF15EA9E5E7030FCB8F8E5FC60925E" box="[887,1003,1827,1853]" country="Japan" name="Hokkaido" pageId="10" pageNumber="11">Hokkaido</collectingRegion>
differ in the lengths and proportions of the leg segments (see
<tableCitation id="C6B9EEB39E5E7030FE7AF88DFD9C9206" box="[437,535,1867,1893]" captionStart="TABLE 5" captionStartId="12.[151,239,514,538]" captionText="TABLE 5. Lengths (in µm) and proportions of legs of Boreoheptagyia sasai Makarchenko et Endo, sp. n., males from the Kolyma River Basin, Northeastern Russia (n = 6)." pageId="10" pageNumber="11">Tables 5</tableCitation>
&amp;
<tableCitation id="C6B9EEB39E5E7030FDF4F88DFDC09206" box="[571,587,1867,1893]" captionStart="TABLE 6" captionStartId="12.[151,239,1106,1130]" captionText="TABLE 6. Lengths (in µm) and proportions of legs of Boreoheptagyia sasai Makarchenko et Endo, sp. n., males from Hokkaido, Japan (n = 3)." pageId="10" pageNumber="11">6</tableCitation>
), while in other characters both populations are very similar or identical.
</paragraph>
</subSubSection>
<subSubSection id="C32188839E5E7030FF0AF8B5FE2F92D6" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8B84DB089E5E7030FF0AF8B5FE2F92D6" blockId="10.[151,1437,338,1973]" pageId="10" pageNumber="11">
<emphasis id="B94F071A9E5E7030FF0AF8B5FEED92EE" bold="true" box="[197,358,1907,1933]" pageId="10" pageNumber="11">Distribution.</emphasis>
The species is known from localities in the Kolyma River Basin in Northeastern
<collectingCountry id="F32C9B989E5E7030FAD3F8B5FAE192EE" box="[1308,1386,1907,1933]" name="Russia" pageId="10" pageNumber="11">Russia</collectingCountry>
and from
<collectingRegion id="49FF15EA9E5E7030FF18F85DFEC592D6" box="[215,334,1947,1973]" country="Japan" name="Hokkaido" pageId="10" pageNumber="11">Hokkaido</collectingRegion>
,
<collectingCountry id="F32C9B989E5E7030FE96F85DFE1492D6" box="[345,415,1947,1973]" name="Japan" pageId="10" pageNumber="11">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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