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<document id="C074930A86F8A7851AD8E999EF0DCCB3" ID-DOI="10.5252/geodiversitas2019v41a7" ID-GBIF-Dataset="175f115f-a52b-4b98-b183-5f3054a9ba64" ID-ISSN="1638-9395" ID-Zenodo-Dep="3699968" ID-ZooBank="urn:lsid:zoobank.org:pub:58412891-24D4-40B0-B74A-3D9743A140B8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1583514998434" checkinUser="valdenar" docAuthor="Guinot, Danièle, Carbot-Chanona, Gerardo &amp; Vega, Francisco J." docDate="2019" docId="038E3408FFF3CB1844A4FE1FD6D1F914" docLanguage="en" docName="GEODIVERSITAS.41.7.285-322.pdf.imf" docOrigin="Geodiversitas 41 (7)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Archaeochiapasidae Guinot, Carbot-Chanona &amp; Vega 2019, n. fam" docType="treatment" docUuid="07C44088-09FD-4D27-A65E-4B3D02BB2BFD" docUuidSource="ZooBank" docVersion="7" lastPageNumber="292" masterDocId="FFB74C70FFFACB114700FFC7D37EFFFA" masterDocTitle="Archaeochiapasidae n. fam., a new early Cenomanian brachyuran family from Chiapas, Mexico, new hypothesis on Lecythocaridae Schweitzer &amp; Feldmann, 2009, and phylogenetic implications (Crustacea, Decapoda, Brachyura, Eubrachyura)" masterLastPageNumber="322" masterPageNumber="285" pageNumber="292" updateTime="1698725880851" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="C01D3E18598FADE01A7EB33118DB09ED">Archaeochiapasidae n. fam., a new early Cenomanian brachyuran family from Chiapas, Mexico, new hypothesis on Lecythocaridae Schweitzer &amp; Feldmann, 2009, and phylogenetic implications (Crustacea, Decapoda, Brachyura, Eubrachyura)</mods:title>
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<mods:namePart id="39810B15DE2FD969F6063AF7F2C3A02F">Guinot, Danièle</mods:namePart>
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<mods:namePart id="9FD268E18E8C547A6159B92D06C39EE6">Carbot-Chanona, Gerardo</mods:namePart>
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<mods:namePart id="BEE39B0E569BE923DECF470121DCABB6">Vega, Francisco J.</mods:namePart>
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<mods:title id="A8B9600F5DBC1867FBD6127617D5E7B3">Geodiversitas</mods:title>
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<mods:date id="644DD194A743DAF760D52B556CCCFD46">2019</mods:date>
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<heading id="D0D03272FFF3CB1844A4FE1FD647FE08" box="[932,1337,472,499]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="292" reason="3">
Family
<taxonomicName id="4C27FE9DFFF3CB1844F5FE1FD79AFE0A" ID-CoL="29ea070a-134f-414d-910a-7e9d70bfaf8b" authority="Guinot, Carbot-Chanona &amp; Vega, 2019" authorityName="Guinot, Carbot-Chanona &amp; Vega" authorityYear="2019" box="[1013,1252,472,498]" family="Archaeochiapasidae" kingdom="Animalia" pageId="9" pageNumber="292" rank="family" status="n. fam">ARCHAEOCHIAPASIDAE</taxonomicName>
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.
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</paragraph>
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<typeStatus id="549C3BBCFFF3CB18442DFD8BD020FD98" box="[813,862,588,612]" pageId="9" pageNumber="292">TYPE</typeStatus>
GENUS. —
<emphasis id="B953590CFFF3CB1844D7FD8AD70DFD9F" box="[983,1139,589,613]" italics="true" pageId="9" pageNumber="292">Archaeochiapasa</emphasis>
n. gen. by present designation. Gender feminine.
</paragraph>
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<subSubSection id="C33DD695FFF3CB18442DFD51D627FB5F" pageId="9" pageNumber="292" type="diagnosis">
<paragraph id="8B98851EFFF3CB18442DFD51D627FB5F" blockId="9.[813,1456,662,1189]" pageId="9" pageNumber="292">DIAGNOSIS. — Cuticle very thin. Carapace with lateroposterior and posterior regions expanded, forming all around a narrow, sunken, continuous area (“flange”) bordered by a strongly rimmed, thickened, raised margin. Carapace posterior margin deeply concave. Dorsal surface with a few, small, regularly distributed tubercles; some regions strongly inflated and separated by distinct grooves. Hepatic region conspicuous, delimitated by margin forming right angle; subbhepatic and branchial regions expanded laterally; cardiac region large, extending nearly to posterior border. Cervical and postcervical grooves marked; branchiocardiac groove deep. Rostrum short, sulcate, bifurcate. Chelipeds stout, equal, homomorph, with massive merus, carpus and propodus. P2, P3 very long and slender. Only one last pereiopod (P4 or P5) preserved, straight, and only reduced approximately by about half compared to P3. Thoracic sternum extremely wide compared to the carapace width; sutures 5/6-7/8 reaching margins of pleon, complete on exposed portion of sternum, sutures underneath the folded pleon being not visible. Pleon triangular, of six free somites, plus telson; first three somites dorsal; somite 6 with two lateral prominences, probably at location of sockets for pleonal-locking mechanism; no pleotelson.</paragraph>
</subSubSection>
<subSubSection id="C33DD695FFF3CB18442DFB12D6D1F914" pageId="9" pageNumber="292" type="discussion">
<paragraph id="8B98851EFFF3CB18442DFB12D0EFFB14" blockId="9.[813,1456,1237,1774]" box="[813,913,1237,1263]" pageId="9" pageNumber="292">REMARKS</paragraph>
<paragraph id="8B98851EFFF3CB18442DFB31D6D1F914" blockId="9.[813,1456,1237,1774]" pageId="9" pageNumber="292">
In
<taxonomicName id="4C27FE9DFFF3CB18444CFB32D0C6FAF5" authorityName="Latreille" authorityYear="1802" box="[844,952,1269,1295]" class="Reptilia" family="Colubridae" genus="Brachyura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="314" phylum="Chordata" rank="genus">Brachyura</taxonomicName>
the ventral part provides a reliable complex of characters, in particular the thoracic sternum. The organisation of the thoracic sternum and pleon, remarkably well preserved in
<emphasis id="B953590CFFF3CB18442DFA92D733FA95" box="[813,1101,1365,1391]" italics="true" pageId="9" pageNumber="292">Archaeochiapasa mardoqueoi</emphasis>
n. gen., n. sp. (
<figureCitation id="131C999BFFF3CB1843F1FA92D644FA95" box="[1265,1338,1365,1391]" captionStart="FIG" captionStartId="21.[132,143,1293,1311]" captionTargetBox="[132,1455,215,1248]" captionTargetId="figure@21.[132,1455,214,1248]" captionTargetPageId="21" captionText="FIG. 11. — Archaeochiapasa mardoqueoi n. gen.,n. sp., male 10.3 × 11.1 mm, holotype (IHNFG-5616): thoracic sternum and pleon. Abbreviations:a4-a6, pleonal somites 4-6; t, telson; 3-8, thoracic sternites 3-8; 4/5, 5/6, 7/8, sternal sutures 4/5, 5/6, 7/8. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3699991" httpUri="https://zenodo.org/record/3699991/files/figure.png" pageId="9" pageNumber="292">Fig. 11</figureCitation>
), is indicative of a non-podotreme crab, in contradiction with what the carapace suggested, and provide evidence of a eubrachyuran affiliation.The new fossil from the lower Cenomanian of
<collectingCountry id="F330C58EFFF3CB184263FA72D6D1FA35" box="[1379,1455,1461,1487]" name="Mexico" pageId="9" pageNumber="292">Mexico</collectingCountry>
is therefore quite distinct from the podotreme crabs found preserved in Albian-Cenomanian deposits from southern
<collectingCountry id="F330C58EFFF3CB184257FA32D6D1F9F5" box="[1367,1455,1525,1551]" name="United Kingdom" pageId="9" pageNumber="292">England</collectingCountry>
and
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(
<bibRefCitation id="EFB6F8EFFFF3CB1844B0F9D3D7D8F9D5" author="WRIGHT C. W. &amp; COLLINS J. S. H." box="[944,1190,1556,1583]" pageId="9" pageNumber="292" pagination="1 - 113" refId="ref33698" refString="WRIGHT C. W. &amp; COLLINS J. S. H. 1972. - British Cretaceous crabs. Monograph of the Palaeontographical Society 126: 1 - 113, pls 1 - 22." type="journal article" year="1972">Wright &amp; Collins 1972</bibRefCitation>
;
<bibRefCitation id="EFB6F8EFFFF3CB1843B4F9D2D6D6F9D5" author="BRETON G. &amp; COLLINS J. S. H." box="[1204,1448,1556,1583]" pageId="9" pageNumber="292" pagination="141 - 156" refId="ref22463" refString="BRETON G. &amp; COLLINS J. S. H. 2011. - New and rare Cenomanian crabs (Crustacea, Decapoda, Brachyura) from the Paris Basin (France), and a comparison with necrocarcinids, etyids and dynomenids from Devon (England). Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen 260: 141 - 156. https: // doi. org / 10.1127 / 0077 - 7749 / 2011 / 0166" type="journal article" year="2011">Breton &amp; Collins 2011</bibRefCitation>
;
<bibRefCitation id="EFB6F8EFFFF3CB18442DF9F3D77EF9B5" author="VALENTIN X. &amp; GOMEZ B. &amp; GOMEZ V. &amp; CHARBONNIER S. &amp; FERCHAUD P. &amp; KIREJTSHUK A. G. &amp; LICHT A. &amp; NERAUDEAU D. &amp; VULLO R. &amp; GARCIA G." box="[813,1024,1588,1615]" pageId="9" pageNumber="292" pagination="443 - 454" refId="ref32299" refString="VALENTIN X., GOMEZ B., DAVIERO- GOMEZ V., CHARBONNIER S., FERCHAUD P., KIREJTSHUK A. G., LICHT A., NERAUDEAU D., VULLO R. &amp; GARCIA G. 2014. - Plant-dominated assemblage and invertebrates from the lower Cenomanian of Jaunay-Clan, western France. Comptes Rendus Palevol 13 (5): 443 - 454. https: // doi. org / 10.1016 / j. crpv. 2014.04.001" type="journal article" year="2014">
Valentin
<emphasis id="B953590CFFF3CB18448EF9F2D0BCF9B4" box="[910,962,1588,1614]" italics="true" pageId="9" pageNumber="292">et al.</emphasis>
2014
</bibRefCitation>
), from
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, northern
<collectingCountry id="F330C58EFFF3CB18421DF9F3D626F9B4" box="[1309,1368,1588,1614]" name="Spain" pageId="9" pageNumber="292">Spain</collectingCountry>
(
<bibRefCitation id="EFB6F8EFFFF3CB184269F9F3D0E4F994" author="FRAAIJE R. H. B. &amp; VAN BAKEL B. W. M. &amp; JAGT J. W. M. &amp; ARTAL P." pageId="9" pageNumber="292" pagination="193 - 208" refId="ref24323" refString="FRAAIJE R. H. B., VAN BAKEL B. W. M., JAGT J. W. M. &amp; ARTAL P. 2008. - New decapod crustaceans (Anomura, Brachyura) from mid-Cretaceous reefal deposits at Monte Orobe (Navarra, northern Spain), and comments on related type-Maastrichtian material. Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 78: 193 - 208." type="journal article" year="2008">
Fraaije
<emphasis id="B953590CFFF3CB18442DF992D020F994" box="[813,862,1620,1646]" italics="true" pageId="9" pageNumber="292">et al.</emphasis>
2008
</bibRefCitation>
;
<bibRefCitation id="EFB6F8EFFFF3CB1844A8F993D7D8F994" author="KLOMPMAKER A. A. &amp; ARTAL P. &amp; VAN BAKEL B. M. W. &amp; FRAAIJE R. H. B. &amp; JAGT J. W. M." box="[936,1190,1620,1646]" pageId="9" pageNumber="292" pagination="1199 - 1212" refId="ref27593" refString="KLOMPMAKER A. A., ARTAL P., VAN BAKEL B. M. W., FRAAIJE R. H. B. &amp; JAGT J. W. M. 2011. - Etyid crabs (Crustacea, Decapoda) from mid-cretaceous reefal strata of Navarra, Northern Spain. Palaeontology 54 (5): 1199 - 1212. https: // doi. org / 10.1111 / j. 1475 - 4983.2011.01072. x" type="journal article" year="2011">
Klompmaker
<emphasis id="B953590CFFF3CB184339F992D714F994" box="[1081,1130,1620,1646]" italics="true" pageId="9" pageNumber="292">et al.</emphasis>
2011
</bibRefCitation>
, 2013;
<bibRefCitation id="EFB6F8EFFFF3CB1843F8F993D6E1F994" author="ARTAL P. &amp; VAN BAKEL B. W. M. &amp; FRAAIJE R. H. B. &amp; JAGT J. W. M. &amp; KLOMPMAKER A. A." box="[1272,1439,1620,1646]" pageId="9" pageNumber="292" pagination="398 - 410" refId="ref21678" refString="ARTAL P., VAN BAKEL B. W. M., FRAAIJE R. H. B., JAGT J. W. M. &amp; KLOMPMAKER A. A. 2012. - New Albian-Cenomanian crabs (Crustacea, Decapoda, Podotremata) from Monte Orobe, Navarra, northern Spain. Revista Mexicana de Ciencias Geologicas 29: 398 - 410." type="journal article" year="2012">
Artal
<emphasis id="B953590CFFF3CB184232F992D61DF994" box="[1330,1379,1620,1646]" italics="true" pageId="9" pageNumber="292">et al.</emphasis>
2012
</bibRefCitation>
), from Tarragona, southern
<collectingCountry id="F330C58EFFF3CB184339F9B3D70DF974" box="[1081,1139,1652,1678]" name="Spain" pageId="9" pageNumber="292">Spain</collectingCountry>
(
<bibRefCitation id="EFB6F8EFFFF3CB184381F9B3D65AF974" author="OSSO A. &amp; VAN BAKEL B. M. W. &amp; KWEKEL F. A. &amp; BEDMAR J. A." box="[1153,1316,1652,1678]" pageId="9" pageNumber="292" pagination="94 - 107" refId="ref29490" refString="OSSO A., VAN BAKEL B. M. W., FERRATGES- KWEKEL F. A. &amp; MORENO- BEDMAR J. A. 2018. - A new decapod crustacean assemblage from the lower Aptian of La Cova del Vidre (Baix Ebre, province of Tarragona, Catalonia). Cretaceous Research 92: 94 - 107. https: // doi. org / 10.1016 / j. cretres. 2018.07.011" type="journal article" year="2018">
Ossó
<emphasis id="B953590CFFF3CB1843BAF9B2D794F974" box="[1210,1258,1652,1678]" italics="true" pageId="9" pageNumber="292">et al.</emphasis>
2018
</bibRefCitation>
), from Sicilia (Checchia-Rispoli 1917), from
<collectingCountry id="F330C58EFFF3CB184362F953D7D3F954" box="[1122,1197,1684,1710]" name="Mexico" pageId="9" pageNumber="292">Mexico</collectingCountry>
and
<collectingCountry id="F330C58EFFF3CB1843DCF953D63FF954" box="[1244,1345,1684,1710]" name="Colombia" pageId="9" pageNumber="292">Colombia</collectingCountry>
(
<bibRefCitation id="EFB6F8EFFFF3CB18424EF952D013F934" author="VEGA F. J. &amp; ALVAREZ F. &amp; CHANONA G." pageId="9" pageNumber="292" pagination="6 - 8" refId="ref32671" refString="VEGA F. J., ALVAREZ F. &amp; CARBOT- CHANONA G. 2007 a. - Albian penaeoidea [sic] (Decapoda: Dendrobranchiata) from Chiapas, Southern Mexico. Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 35 (2): 6 - 8." type="journal article" year="2007">
Vega
<emphasis id="B953590CFFF3CB184282F952D6D1F954" box="[1410,1455,1684,1710]" italics="true" pageId="9" pageNumber="292">et al.</emphasis>
2007
<bibRefCitation id="EFB6F8EFFFF3CB184462F973D013F934" author="HILLER A. &amp; VIVIANI C. A. &amp; WERDING B." box="[866,877,1716,1742]" pageId="9" pageNumber="292" pagination="95 - 102" refId="ref26474" refString="HILLER A., VIVIANI C. A. &amp; WERDING B. 2010. - Hypercarcinisation: an evolutionary novelty in thecommensal porcellanid Allopetrolisthes spinifrons (Crustacea: Decapoda: Porcellanidae). Nauplius 18 (1): 95 - 102." type="journal article" year="2010">a</bibRefCitation>
</bibRefCitation>
, 2010), from Texas (
<bibRefCitation id="EFB6F8EFFFF3CB184348F972D790F934" author="VEGA F. J. &amp; JACKSON J. &amp; OSSO A." box="[1096,1262,1716,1743]" pageId="9" pageNumber="292" pagination="1 - 7" refId="ref32730" refString="VEGA F. J., JACKSON J. &amp; OSSO A. 2014. - Exceptional preservation of a late Cenomanian (Late Cretaceous) crab from Texas, U. S. A. Boletin de la Sociedad Geologica Mexicana 66 (1): 1 - 7. https: // doi. org / 10.18268 / BSGM 2014 v 66 n 1 a 15" type="journal article" year="2014">
Vega
<emphasis id="B953590CFFF3CB184380F972D7CCF934" box="[1152,1202,1716,1742]" italics="true" pageId="9" pageNumber="292">et al.</emphasis>
2014
</bibRefCitation>
), and from a wide range of regions (
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</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>