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<document id="BECB721CE20B0F85AD3C30086B05C7D6" ID-DOI="10.11646/zootaxa.4742.1.5" ID-GBIF-Dataset="d9aaf71d-e347-4fc3-b5fd-cae0989cac05" ID-ISSN="1175-5326" ID-Zenodo-Dep="3674493" ID-ZooBank="3016F563-FDAC-44FD-B6A5-44E1E8DFCFB4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582096979764" checkinUser="plazi" docAuthor="Arimoto, Kôichi &amp; Suzuki, Wataru" docDate="2020" docId="FC3C87A4327BB31B83CEBE80BD70FF29" docLanguage="en" docName="zootaxa.4742.1.5.pdf" docOrigin="Zootaxa 4742 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limonius onyx Arimoto &amp; Suzuki 2020, sp. n." docType="treatment" docVersion="6" lastPageNumber="100" masterDocId="0005FFDC327CB3108359BB4DBF09FF9A" masterDocTitle="First record and two new species of the genus Limonius Eschscholtz (Coleoptera Elateridae, Dendrometrinae) from Taiwan" masterLastPageNumber="104" masterPageNumber="89" pageNumber="96" updateTime="1698719267030" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9B4D61FA20CC16589DDDC9CDE465404B">First record and two new species of the genus Limonius Eschscholtz (Coleoptera Elateridae, Dendrometrinae) from Taiwan</mods:title>
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<mods:name id="471B9EF1ACA46DB70FA2CC2A5A0D31CD" type="personal">
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<mods:namePart id="E15F89B7B7F92E48500D0E1AD96A890C">Arimoto, Kôichi</mods:namePart>
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<mods:namePart id="8BFFC00A91E99C0DF01F1A78DF960C12">Suzuki, Wataru</mods:namePart>
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<mods:title id="18198696BFCAC2FE02DA3237B8D69CAA">Zootaxa</mods:title>
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<mods:date id="5812CA82FA9AB7EBFA2AA276811BB5DE">2020</mods:date>
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<mods:number id="4D008F2A3AB7FCC1ED712D6AE07F2492">2020-02-19</mods:number>
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<mods:number id="8DCA8D7DCF4FCB16EDA66458D07E5856">4742</mods:number>
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<treatment id="FC3C87A4327BB31B83CEBE80BD70FF29" ID-DOI="http://doi.org/10.5281/zenodo.3681227" ID-GBIF-Taxon="162163999" ID-Zenodo-Dep="3681227" LSID="urn:lsid:plazi:treatment:FC3C87A4327BB31B83CEBE80BD70FF29" httpUri="http://treatment.plazi.org/id/FC3C87A4327BB31B83CEBE80BD70FF29" lastPageId="11" lastPageNumber="100" pageId="7" pageNumber="96">
<subSubSection id="3C8F6539327BB31783CEBE80BD84FA7D" box="[151,653,1485,1511]" pageId="7" pageNumber="96" type="nomenclature">
<paragraph id="742A36B2327BB31783CEBE80BD84FA7D" blockId="7.[151,653,1485,1547]" box="[151,653,1485,1511]" pageId="7" pageNumber="96">
<heading id="2F6281DE327BB31783CEBE80BD84FA7D" bold="true" box="[151,653,1485,1511]" fontSize="11" level="1" pageId="7" pageNumber="96" reason="1">
<emphasis id="46E1EAA0327BB31783CEBE80BD84FA7D" bold="true" box="[151,653,1485,1511]" pageId="7" pageNumber="96">
<taxonomicName id="B3954D31327BB31783CEBE80BD34FA7D" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" box="[151,573,1485,1511]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="onyx" status="sp. nov.">
<emphasis id="46E1EAA0327BB31783CEBE80BE44FA7D" bold="true" box="[151,333,1485,1511]" italics="true" pageId="7" pageNumber="96">Limonius onyx</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
,
<taxonomicNameLabel id="5DD257DB327BB3178113BE80BD84FA7D" box="[586,653,1485,1511]" pageId="7" pageNumber="96" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3C8F6539327BB31783CEBEBCBE57F991" box="[151,350,1520,1547]" pageId="7" pageNumber="96" type="description">
<paragraph id="742A36B2327BB31783CEBEBCBE57F991" blockId="7.[151,653,1485,1547]" box="[151,350,1520,1547]" pageId="7" pageNumber="96">
(
<figureCitation id="ECAE2A37327BB31783C6BEBCBE05F991" box="[159,268,1520,1547]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="7" pageNumber="96">Figures 5</figureCitation>
,
<figureCitation id="ECAE2A37327BB3178241BEBCBE21F991" box="[280,296,1521,1547]" captionStart="FIGURE 6" captionStartId="9.[151,250,1076,1101]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure@9.[151,1436,181,1050]" captionTargetPageId="9" captionText="FIGURE 6. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674505" httpUri="https://zenodo.org/record/3674505/files/figure.png" pageId="7" pageNumber="96">6</figureCitation>
,
<figureCitation id="ECAE2A37327BB317826DBEBCBE5CF991" box="[308,341,1521,1547]" captionStart="FIGURE 7" captionStartId="9.[151,250,1830,1855]" captionTargetBox="[158,1426,1166,1818]" captionTargetId="figure@9.[151,1436,1159,1826]" captionTargetPageId="9" captionText="FIGURE 7. Limonius spp., male, holotype, apical part of paramere A: Limonius sapphirus Arimoto &amp; Suzuki sp. n.; B: Limonius onyx Arimoto &amp; Suzuki sp. n." figureDoi="http://doi.org/10.5281/zenodo.3674507" httpUri="https://zenodo.org/record/3674507/files/figure.png" pageId="7" pageNumber="96">7B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3C8F6539327BB31783CEBD74BCCAF9C9" box="[151,963,1593,1619]" pageId="7" pageNumber="96" type="etymology">
<paragraph id="742A36B2327BB31783CEBD74BCCAF9C9" blockId="7.[151,1437,1593,2015]" box="[151,963,1593,1619]" pageId="7" pageNumber="96">
<emphasis id="46E1EAA0327BB31783CEBD74BE17F9C9" bold="true" box="[151,286,1593,1619]" pageId="7" pageNumber="96">Etymology.</emphasis>
In reference to the dark and shining body like onyx jewelry.
</paragraph>
</subSubSection>
<subSubSection id="3C8F6539327BB317839EBD10BC9EF845" pageId="7" pageNumber="96" type="materials_examined">
<paragraph id="742A36B2327BB317839EBD10BE90F901" blockId="7.[151,1437,1593,2015]" pageId="7" pageNumber="96">
<emphasis id="46E1EAA0327BB317839EBD10BE7EF9ED" bold="true" box="[199,375,1628,1655]" pageId="7" pageNumber="96">Type material.</emphasis>
<materialsCitation id="C4FD3CEF327BB31782D9BD10BE90F901" ID-GBIF-Occurrence="2566071263" collectingDate="1977-05-08" collectionCode="LOA" collectorName="Shusei Saito" country="Taiwan" county="Xinyi Township" location="Male" municipality="Shenmu Village" pageId="7" pageNumber="96" specimenCode="LOA001" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
<emphasis id="46E1EAA0327BB31782D9BD10BEE4F9ED" bold="true" box="[384,493,1629,1655]" pageId="7" pageNumber="96">
<typeStatus id="AB2E8810327BB31782D9BD10BEE4F9ED" box="[384,493,1629,1655]" pageId="7" pageNumber="96" type="holotype">Holotype</typeStatus>
</emphasis>
.
<location id="714A6069327BB31782A7BD10BD31F9ED" LSID="urn:lsid:plazi:treatment:FC3C87A4327BB31B83CEBE80BD70FF29:714A6069327BB31782A7BD10BD31F9ED" box="[510,568,1629,1655]" country="Taiwan" county="Xinyi Township" municipality="Shenmu Village" name="Male" pageId="7" pageNumber="96" stateProvince="Nantou">Male</location>
(
<specimenCode id="24339EC9327BB3178111BD10BDB8F9ED" box="[584,689,1629,1655]" collectionCode="LOA" pageId="7" pageNumber="96">LOA001</specimenCode>
),
<collectingMunicipality id="944EACC8327BB317819EBD10BC76F9ED" box="[711,895,1629,1655]" pageId="7" pageNumber="96">Shenmu Village</collectingMunicipality>
,
<collectingCounty id="9D4B4E3E327BB31780D5BD10BB4FF9ED" box="[908,1094,1629,1655]" pageId="7" pageNumber="96">Xinyi Township</collectingCounty>
,
<collectingRegion id="B651F850327BB317870ABD10BA0AF9ED" box="[1107,1283,1629,1655]" country="Taiwan" name="Nantou" pageId="7" pageNumber="96">Nantou County</collectingRegion>
,
<collectingCountry id="0C827622327BB3178656BD10BA6DF9ED" box="[1295,1380,1629,1655]" name="Taiwan" pageId="7" pageNumber="96">Taiwan</collectingCountry>
,
<date id="002B1072327BB317862BBD10BFDBF901" pageId="7" pageNumber="96" value="1977-05-08">
<collectingDate id="106FE99A327BB317862BBD10BFDBF901" pageId="7" pageNumber="96" value="1977-05-08">8 V 1977</collectingDate>
</date>
,
<collectorName id="D9605364327BB3178384BDCCBE60F901" box="[221,361,1665,1691]" pageId="7" pageNumber="96">Shûsei Saitô</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="742A36B2327BB317839EBDE8BC9EF845" blockId="7.[151,1437,1593,2015]" pageId="7" pageNumber="96">
<emphasis id="46E1EAA0327BB317839EBDE8BE82F925" bold="true" box="[199,395,1700,1727]" pageId="7" pageNumber="96">Male. Diagnosis.</emphasis>
Body surface without metallic luster; elytra interspaces between punctures smaller than puncture diameter. Body black. Frons in profile slightly depressed apically; supra-antennal carina in dorsal view broadly rounded, but slightly emarginate apically. Antennae extending beyond pronotum posterior lateral apices by apical two antennomeres; antennomere III slightly longer than II; IV distinctly longer than wide, longer than IIIII combined, longer than
<collectionCode id="1284AE77327BB31782C3BC78BEA7F8D5" box="[410,430,1845,1871]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="96" type="Museum">V</collectionCode>
. Prothorax widening posteriad; lateral carina of prothorax strongly prominent just behind anterior angles. Anterior lobe of prosternum in profile weakly inclined. Mesial edge of hypomeron carinate anterolaterad. Prosternal spine without distinct subapical tooth; sides at ventral view roundly expanded and anteriorly strongly prominent. Scutellum longer than wide, weakly elevated medially; posterior margin broadly rounded, but emarginate medially. Mesepisternum not reaching mesocoxal cavity.
</paragraph>
</subSubSection>
<caption id="20EA663A3274B31883CEBDA0BC58F844" ID-DOI="http://doi.org/10.5281/zenodo.3674503" ID-Zenodo-Dep="3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="8" pageNumber="97" startId="8.[151,250,1773,1798]" targetBox="[151,1436,189,1742]" targetPageId="8">
<paragraph id="742A36B23274B31883CEBDA0BC58F844" blockId="8.[151,1437,1773,2014]" pageId="8" pageNumber="97">
<emphasis id="46E1EAA03274B31883CEBDA0BE1BF89C" bold="true" box="[151,274,1773,1798]" pageId="8" pageNumber="97">FIGURE 5.</emphasis>
<taxonomicName id="B3954D313274B3188241BDA2BD65F89C" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" box="[280,620,1773,1798]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="97" phylum="Arthropoda" rank="species" species="onyx" status="sp. nov.">
<emphasis id="46E1EAA03274B3188241BDA2BEA5F89C" box="[280,428,1775,1798]" italics="true" pageId="8" pageNumber="97">Limonius onyx</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
<taxonomicNameLabel id="5DD257DB3274B318812BBDA2BDA0F89C" box="[626,681,1775,1798]" pageId="8" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
, male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V.
</paragraph>
</caption>
<caption id="20EA663A3275B31983CEBF79BC2BFBEB" ID-DOI="http://doi.org/10.5281/zenodo.3674505" ID-Zenodo-Dep="3674505" httpUri="https://zenodo.org/record/3674505/files/figure.png" pageId="9" pageNumber="98" startId="9.[151,250,1076,1101]" targetBox="[151,1436,181,1050]" targetPageId="9">
<paragraph id="742A36B23275B31983CEBF79BC2BFBEB" blockId="9.[151,1437,1076,1137]" pageId="9" pageNumber="98">
<emphasis id="46E1EAA03275B31983CEBF79BE1CFBD7" bold="true" box="[151,277,1076,1101]" pageId="9" pageNumber="98">FIGURE 6.</emphasis>
<taxonomicName id="B3954D313275B3198247BF78BD74FBD7" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" box="[286,637,1076,1101]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="98" phylum="Arthropoda" rank="species" species="onyx" status="sp. nov.">
<emphasis id="46E1EAA03275B3198247BF78BEBDFBD6" box="[286,436,1077,1100]" italics="true" pageId="9" pageNumber="98">Limonius onyx</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
<taxonomicNameLabel id="5DD257DB3275B31981DFBF78BDC9FBD6" box="[646,704,1077,1100]" pageId="9" pageNumber="98" rank="species">sp. n.</taxonomicNameLabel>
, male, holotype. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view.
</paragraph>
</caption>
<caption id="20EA663A3275B31983CEBC6BBEFEF8F9" ID-DOI="http://doi.org/10.5281/zenodo.3674507" ID-Zenodo-Dep="3674507" httpUri="https://zenodo.org/record/3674507/files/figure.png" pageId="9" pageNumber="98" startId="9.[151,250,1830,1855]" targetBox="[158,1426,1166,1818]" targetPageId="9">
<paragraph id="742A36B23275B31983CEBC6BBEFEF8F9" blockId="9.[151,1437,1830,1891]" pageId="9" pageNumber="98">
<emphasis id="46E1EAA03275B31983CEBC6BBE1AF8A5" bold="true" box="[151,275,1830,1855]" pageId="9" pageNumber="98">FIGURE 7.</emphasis>
<taxonomicName id="B3954D313275B3198243BC65BE71F8A5" authorityName="Eschscholtz" authorityYear="1829" box="[282,376,1832,1855]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="46E1EAA03275B3198243BC65BE71F8A5" box="[282,376,1832,1855]" italics="true" pageId="9" pageNumber="98">Limonius</emphasis>
</taxonomicName>
spp., male, holotype, apical part of paramere A:
<taxonomicName id="B3954D313275B3198032BC65BBFFF8A5" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" box="[875,1270,1830,1855]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="98" phylum="Arthropoda" rank="species" species="sapphirus" status="sp. nov.">
<emphasis id="46E1EAA03275B3198032BC65BB3DF8A5" box="[875,1076,1831,1855]" italics="true" pageId="9" pageNumber="98">Limonius sapphirus</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
<taxonomicNameLabel id="5DD257DB3275B31987A4BC65BA3CF8A5" box="[1277,1333,1832,1855]" pageId="9" pageNumber="98" rank="species">sp. n.</taxonomicNameLabel>
; B:
<taxonomicName id="B3954D313275B3198639BC65BEB3F8F9" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="98" phylum="Arthropoda" rank="species" species="onyx" status="sp. nov.">
<emphasis id="46E1EAA03275B3198639BC65BFF1F8F9" italics="true" pageId="9" pageNumber="98">Limonius onyx</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
<taxonomicNameLabel id="5DD257DB3275B3198298BC01BEFEF8F9" box="[449,503,1868,1891]" pageId="9" pageNumber="98" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="3C8F65393275B31A839EBCDFBC78F845" lastPageId="10" lastPageNumber="99" pageId="9" pageNumber="98" type="description">
<paragraph id="742A36B23275B319839EBCDFBD34F84A" blockId="9.[151,1437,1937,2036]" pageId="9" pageNumber="98">
<emphasis id="46E1EAA03275B319839EBCDFBE7FF836" bold="true" box="[199,374,1938,1964]" pageId="9" pageNumber="98">Measurements</emphasis>
. BL: 6.97, BW: 1.95,
<collectingRegion id="B651F8503275B3198134BCDFBDACF836" box="[621,677,1938,1964]" country="Japan" name="Mie" pageId="9" pageNumber="98">MIE</collectingRegion>
: 0.76, MAE: 1.08, OI: 141, PL: 1.85, PML: 1.78, PW: 1.72, PI: 107, EL: 4.69, EW: 1.95, EI: 240, BI: 254.
</paragraph>
<paragraph id="742A36B23275B31A839EBC94BEB7FE11" blockId="9.[151,1437,1937,2036]" lastBlockId="10.[151,1437,153,2015]" lastPageId="10" lastPageNumber="99" pageId="9" pageNumber="98">
<emphasis id="46E1EAA03275B319839EBC94BE50F86E" bold="true" box="[199,345,2009,2036]" pageId="9" pageNumber="98">Description.</emphasis>
Body surface shining and without metallic luster, generally smooth but elytra rough; dorsal inter- spaces between punctures smaller than puncture diameter; ventral interspaces between punctures generally larger than puncture diameter, but in hypomeron, anterior lobe of prosternum and prosternal spine interspaces between punctures smaller than puncture diameter (
<figureCitation id="ECAE2A373276B31A8122BBACBDC9FF61" box="[635,704,224,251]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5</figureCitation>
, F, H). Color. Body black (
<figureCitation id="ECAE2A373276B31A80ACBBACBB79FF61" box="[1013,1136,224,251]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5A, B</figureCitation>
). Mandible black, but apical part orange. Legs black, but partly with orange tinge; claws orange. Tergites and sternites VIIIX and aedeagus yellow-brown, but terigite and sternite VIII colorless in basal median area (
<figureCitation id="ECAE2A373276B31A80B8BA64BB52FED9" box="[993,1115,297,323]" captionStart="FIGURE 6" captionStartId="9.[151,250,1076,1101]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure@9.[151,1436,181,1050]" captionTargetPageId="9" captionText="FIGURE 6. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674505" httpUri="https://zenodo.org/record/3674505/files/figure.png" pageId="10" pageNumber="99">Fig. 6A, B</figureCitation>
). Pubescence. Body covered with orange setae dorsally, with white setae ventrally. Antennae with yellow to brown setae. Setae of legs white becoming yellow apically.
</paragraph>
<paragraph id="742A36B23276B31A839EBAD8BE71FD1D" blockId="10.[151,1437,153,2015]" pageId="10" pageNumber="99">
Head depressed longitudinally medially; the depression becoming narrow and shallow posteriad (
<figureCitation id="ECAE2A373276B31A864ABAD8BA84FE35" box="[1299,1421,404,431]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5C, D</figureCitation>
). Frons in profile slightly depressed apically (
<figureCitation id="ECAE2A373276B31A81C4BAF4BDF1FE49" box="[669,760,440,467]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5E</figureCitation>
: green line); supra-antennal carina in dorsal view broadly rounded, but slightly emarginate apically (
<figureCitation id="ECAE2A373276B31A812DBA90BDC5FE6D" box="[628,716,476,503]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5C</figureCitation>
), in profile strongly prominent anteriad above clypeus (
<figureCitation id="ECAE2A373276B31A8619BA90BA90FE6D" box="[1344,1433,476,503]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5E</figureCitation>
: arrow). Antennae extending beyond pronotum posterior lateral apices by apical two antennomeres (
<figureCitation id="ECAE2A373276B31A87B0B94CBA4AFD81" box="[1257,1347,512,539]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5A</figureCitation>
); antennomeres I and IVXI longer than wide; II globular, shortest; III obconical, slightly longer than II; IVX serrate; IV 1.12 x as long as wide, longer than IIIII combined, 2.06 x as long as III, longer than V; XI 2.70 x as long as wide. Mandible bidentate.
</paragraph>
<paragraph id="742A36B23276B31A839EB9DCBDE9FA95" blockId="10.[151,1437,153,2015]" pageId="10" pageNumber="99">
Prothorax trapezoidal, longer than wide, widening posteriad; side weakly rounded; lateral carina extending from anterior angles to hind angles of prothorax, strongly prominent just behind anterior angles (
<figureCitation id="ECAE2A373276B31A87BDB9F8BA36FD55" box="[1252,1343,692,719]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5C</figureCitation>
: arrow). Pronotum posteriorly with a shallow median longitudinal depression; hind angles broad, rounded, unicarinate; the carina short, extending to the base. Hypomeron: mesial edge with elevated impunctate ridge next to pronotosternal suture, carinate anterolaterad (
<figureCitation id="ECAE2A373276B31A82B4B86CBD4AFCA1" box="[493,579,800,827]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5F</figureCitation>
: black arrow); posterior margin without large emargination, weakly sinuate. Anterior lobe of prosternum slightly exceeding anterior angles of prothorax (
<figureCitation id="ECAE2A373276B31A875FB808BB69FCC5" box="[1030,1120,836,863]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5B</figureCitation>
), in profile weakly inclined (at 24 degrees against prosternal horizontal line,
<figureCitation id="ECAE2A373276B31A81E7B824BC1CFC19" box="[702,789,872,899]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5E</figureCitation>
: red line). Prosternal spine in profile strongly inclined (at 23 degrees against prosternal horizontal line,
<figureCitation id="ECAE2A373276B31A81C5B8C0BDFAFC3D" box="[668,755,908,935]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5G</figureCitation>
: red line); ventral margin in profile without distinct subapical tooth and sinuate; lateral lobe in profile with ventral margin straight apically (
<figureCitation id="ECAE2A373276B31A876DB8FCBB99FC51" box="[1076,1168,944,971]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5G</figureCitation>
: yellow line), at ventral view with sides roundly expanded and anteriorly strongly prominent (
<figureCitation id="ECAE2A373276B31A80E2B898BB11FC75" box="[955,1048,980,1007]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5H</figureCitation>
: arrows); apical margin at ventral view rounded, with weakly prominent apex (
<figureCitation id="ECAE2A373276B31A81CEB8B4BDFDFB89" box="[663,756,1016,1043]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5H</figureCitation>
). Pronotosternal sutures not grooved, anteriorly opened and appearing double (
<figureCitation id="ECAE2A373276B31A8234BF50BECAFBAD" box="[365,451,1052,1079]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5F</figureCitation>
: white arrow). Scutellum 1.30 x as long as wide (
<figureCitation id="ECAE2A373276B31A80A7BF50BB44FBAD" box="[1022,1101,1052,1079]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5I</figureCitation>
), weakly widening posteriad, widest near posterior 2/5, weakly elevated medially (
<figureCitation id="ECAE2A373276B31A81A6BF0CBC54FBC1" box="[767,861,1088,1115]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5G</figureCitation>
: arrow), moderately inclined anterior-downwards; anterior margin almost transverse; posterior margin broadly rounded, but emarginate medially (
<figureCitation id="ECAE2A373276B31A87E5BF28BA03FBE5" box="[1212,1290,1124,1151]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5I</figureCitation>
: arrow). Mesosternum with a median longitudinal carina posterior to mesosternal cavity; borders of mesosternal cavity sinuate followed by strongly declivous (
<figureCitation id="ECAE2A373276B31A815BBFE0BD55FB5D" box="[514,604,1196,1223]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5H</figureCitation>
: green line). Mesepisternum not reaching mesocoxal cavity. Mesepimeron with shallow depression for reception of femora of mid-leg. Metasternum sulcate medially and behind mesocoxae. Elytral striae distinctly grooved. Tarsomeres simple.
</paragraph>
<paragraph id="742A36B23276B31A839EBE54BA45FA59" blockId="10.[151,1437,153,2015]" pageId="10" pageNumber="99">
Abdomen with external margin micro-serrated (
<figureCitation id="ECAE2A373276B31A81BABE54BC3DFAA9" box="[739,820,1304,1331]" captionStart="FIGURE 5" captionStartId="8.[151,250,1773,1798]" captionTargetBox="[151,1436,189,1742]" captionTargetId="figure@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: habitus, dorsal view; B: habitus, ventral view; C: head, dorsal view (arrow: lateral carina weakly prominent just behind anterior angle); D: basal antennomeres; E: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); F: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); G: prosternal spine and mesosternum, lateral view (arrow: scutellum weakly elevated medially); H: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); I: scutellum (arrow: posterior margin broadly rounded, but emarginated medially); J: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674503" httpUri="https://zenodo.org/record/3674503/files/figure.png" pageId="10" pageNumber="99">Fig. 5J</figureCitation>
); ventrite V 0.57 x as long as wide. Sternite VIII with median notch rounded (
<figureCitation id="ECAE2A373276B31A82C2BE70BEF8FAC2" box="[411,497,1341,1368]" captionStart="FIGURE 6" captionStartId="9.[151,250,1076,1101]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure@9.[151,1436,181,1050]" captionTargetPageId="9" captionText="FIGURE 6. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674505" httpUri="https://zenodo.org/record/3674505/files/figure.png" pageId="10" pageNumber="99">Fig. 6B</figureCitation>
). Tergite IX 1.22 x length of tergite X (
<figureCitation id="ECAE2A373276B31A80C3BE70BCE7FAC2" box="[922,1006,1341,1368]" captionStart="FIGURE 6" captionStartId="9.[151,250,1076,1101]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure@9.[151,1436,181,1050]" captionTargetPageId="9" captionText="FIGURE 6. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674505" httpUri="https://zenodo.org/record/3674505/files/figure.png" pageId="10" pageNumber="99">Fig. 6C</figureCitation>
). Tergite X 1.12 x as long as wide (
<figureCitation id="ECAE2A373276B31A8636BE70BFB2FAE6" captionStart="FIGURE 6" captionStartId="9.[151,250,1076,1101]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure@9.[151,1436,181,1050]" captionTargetPageId="9" captionText="FIGURE 6. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674505" httpUri="https://zenodo.org/record/3674505/files/figure.png" pageId="10" pageNumber="99">Fig. 6C</figureCitation>
). Sternite IX transverse apically (
<figureCitation id="ECAE2A373276B31A816EBE2CBD9BFAE6" box="[567,658,1377,1404]" captionStart="FIGURE 6" captionStartId="9.[151,250,1076,1101]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure@9.[151,1436,181,1050]" captionTargetPageId="9" captionText="FIGURE 6. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674505" httpUri="https://zenodo.org/record/3674505/files/figure.png" pageId="10" pageNumber="99">Fig. 6D</figureCitation>
). Aedeagus (
<figureCitation id="ECAE2A373276B31A807ABE2CBC93FAE1" box="[803,922,1377,1403]" captionStart="FIGURE 6" captionStartId="9.[151,250,1076,1101]" captionTargetBox="[151,1436,181,1050]" captionTargetId="figure@9.[151,1436,181,1050]" captionTargetPageId="9" captionText="FIGURE 6. Limonius onyx Arimoto &amp; Suzuki sp. n., male, holotype. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674505" httpUri="https://zenodo.org/record/3674505/files/figure.png" pageId="10" pageNumber="99">Fig. 6E, F</figureCitation>
). Basal piece 0.38 x total length of aedeagus. Median lobe exceeding apices of parameres; basal struts 0.30 x total length of median lobe. Parameres separated ventrally; apex beyond lateral subapical barb wider sub-triangular (
<figureCitation id="ECAE2A373276B31A80D8BEE4BCD1FA59" box="[897,984,1449,1475]" captionStart="FIGURE 7" captionStartId="9.[151,250,1830,1855]" captionTargetBox="[158,1426,1166,1818]" captionTargetId="figure@9.[151,1436,1159,1826]" captionTargetPageId="9" captionText="FIGURE 7. Limonius spp., male, holotype, apical part of paramere A: Limonius sapphirus Arimoto &amp; Suzuki sp. n.; B: Limonius onyx Arimoto &amp; Suzuki sp. n." figureDoi="http://doi.org/10.5281/zenodo.3674507" httpUri="https://zenodo.org/record/3674507/files/figure.png" pageId="10" pageNumber="99">Fig. 7B</figureCitation>
), rounded apically, without setae.
</paragraph>
<paragraph id="742A36B23276B31A839EBE80BEABFA7D" blockId="10.[151,1437,153,2015]" box="[199,418,1485,1511]" pageId="10" pageNumber="99">
<emphasis id="46E1EAA03276B31A839EBE80BE2CFA7D" bold="true" box="[199,293,1485,1511]" pageId="10" pageNumber="99">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="742A36B23276B31A839EBEBDBC78F845" blockId="10.[151,1437,153,2015]" pageId="10" pageNumber="99">
<emphasis id="46E1EAA03276B31A839EBEBDBEA6F991" bold="true" box="[199,431,1520,1547]" pageId="10" pageNumber="99">Comparative notes.</emphasis>
This species is distinct from the congeners by the whole black body without metallic luster and with dense punctures. It is similar to
<taxonomicName id="B3954D313276B31A8139BD5BBDECF9B5" authorityName="Lewis" authorityYear="1894" box="[608,741,1557,1583]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="species" species="ignicollis">
<emphasis id="46E1EAA03276B31A8139BD5BBDECF9B5" box="[608,741,1557,1583]" italics="true" pageId="10" pageNumber="99">L. ignicollis</emphasis>
</taxonomicName>
in having the following characters: elytra interspaces between punctures smaller than puncture diameter; supra-antennal carina in dorsal view broadly rounded, but more or less emarginate apically, in profile strongly prominent anteriad above clypeus; antennae extending beyond pronotum posterior lateral apices by apical two antennomeres; antennomere III slightly longer than II, longer than IIIII combined, longer than V; prothorax widening posteriad; lateral carina of prothorax strongly prominent just behind anterior angles; mesial edge of hypomeron carinate anterolaterad; sides of prosternal spine at ventral view roundly expanded and anteriorly strongly prominent; scutellum longer than wide; posterior margin of scutellum broadly rounded, but emarginate medially. However it is distinguished by the following contrasting characters (
<taxonomicName id="B3954D313276B31A864EBC5FBA95F8B1" authorityName="Lewis" authorityYear="1894" box="[1303,1436,1809,1835]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="species" species="ignicollis">
<emphasis id="46E1EAA03276B31A864EBC5FBA95F8B1" box="[1303,1436,1809,1835]" italics="true" pageId="10" pageNumber="99">L. ignicollis</emphasis>
</taxonomicName>
in parentheses): body surface without metallic luster (body surface with metallic luster); frons in profile slightly depressed apically (frons in profile weakly depressed apically); antennomere IV distinctly longer than wide (antennomere IV shorter than wide); anterior lobe of prosternum in profile weakly inclined (anterior lobe of prosternum in profile strongly inclined); prosternal spine without distinct subapical tooth (prosternal spine with subapical tooth); scutellum weakly elevated medially (scutellum strongly convex).
</paragraph>
</subSubSection>
<subSubSection id="3C8F65393277B31B839EBBD5BD70FF29" box="[199,633,152,179]" pageId="11" pageNumber="100" type="distribution">
<paragraph id="742A36B23277B31B839EBBD5BD70FF29" blockId="11.[199,633,152,179]" box="[199,633,152,179]" pageId="11" pageNumber="100">
<emphasis id="46E1EAA03277B31B839EBBD5BE68FF29" bold="true" box="[199,353,152,179]" pageId="11" pageNumber="100">Distribution.</emphasis>
<collectingCountry id="0C8276223277B31B823EBBD5BEB4FF28" box="[359,445,152,178]" name="Taiwan" pageId="11" pageNumber="100">Taiwan</collectingCountry>
:
<collectingRegion id="B651F8503277B31B8291BBD4BD7CFF29" box="[456,629,153,179]" country="Taiwan" name="Nantou" pageId="11" pageNumber="100">Nantou County</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>