326 lines
31 KiB
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326 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1159.99218" ID-Pensoft-Pub="1313-2970-1159-87" ID-Pensoft-UUID="ECCECB868071523DB7628E304850BD62" ID-ZooBank="CD97DD120549412F976508D8DE21605B" ModsDocID="1313-2970-1159-87" checkinTime="1682458941538" checkinUser="pensoft" docAuthor="Okada, Ryohei, Jaitrong, Weeyawat & Wewalka, Guenther" docDate="2023" docId="E3744614B5C25BF8985606CE116951EF" docLanguage="en" docName="ZooKeys 1159: 87-119" docOrigin="ZooKeys 1159" docPubDate="2023-04-25" docSource="http://dx.doi.org/10.3897/zookeys.1159.99218" docTitle="Microdytes eliasi Wewalka & Okada 2023, sp. nov." docType="treatment" docUuid="8C12C23B-3B90-4F46-9D48-F5ABD6625734" docUuidSource="ZooBank" docVersion="1" id="ECCECB868071523DB7628E304850BD62" lastPageNumber="87" masterDocId="ECCECB868071523DB7628E304850BD62" masterDocTitle="A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae)" masterLastPageNumber="119" masterPageNumber="87" pageNumber="87" updateTime="1682458941538" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Okada, Ryohei</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8488-0660</mods:nameIdentifier>
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<mods:affiliation>Thailand Natural History Museum, National Science Museum, Technopolis, Pathum Thani, Thailand & Coleopterological Society of Japan, National Museum of Nature and Science, Tsukuba, Japan</mods:affiliation>
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<mods:nameIdentifier type="email">wasserinsekt@kub.biglobe.ne.jp</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jaitrong, Weeyawat</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1362-0754</mods:nameIdentifier>
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<mods:affiliation>Thailand Natural History Museum, National Science Museum, Technopolis, Pathum Thani, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wewalka, Guenther</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2994-8001</mods:nameIdentifier>
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<mods:affiliation>Starkfriedgasse 16 / 1 / 3, A - 1190 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1159</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>87</mods:start>
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<mods:end>119</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1159.99218</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1159.99218</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1159-87</mods:identifier>
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<mods:identifier type="ZooBank">CD97DD120549412F976508D8DE21605B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">ECCECB868071523DB7628E304850BD62</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:8C12C23B-3B90-4F46-9D48-F5ABD6625734" httpUri="http://treatment.plazi.org/id/E3744614B5C25BF8985606CE116951EF" lastPageNumber="87" pageId="0" pageNumber="87">
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<subSubSection pageId="0" pageNumber="87" type="nomenclature">
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<paragraph pageId="0" pageNumber="87">
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<taxonomicName LSID="https://zoobank.org/8C12C23B-3B90-4F46-9D48-F5ABD6625734" authority="Wewalka & Okada" authorityName="Wewalka & Okada" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes eliasi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="eliasi" status="sp. nov.">Microdytes eliasi Wewalka & Okada</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="87">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="87" type="description">
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<paragraph pageId="0" pageNumber="87">
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<figureCitation captionStart="Figures 2–6" captionStartId="F2" captionText="Figures 2 - 6. Habitus of 2 Microdytes eliasi sp. nov., female, paratype 3 M. jeenthongi sp. nov., male, holotype 4 M. maximiliani sp. nov., male, holotype 5 M. sekaensis sp. nov., male, holotype 6 M. ubonensis sp. nov., male, holotype. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures2-6" httpUri="https://binary.pensoft.net/fig/842221" pageId="0" pageNumber="87">Figs 2</figureCitation>
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<figureCitation captionStart="Figures 7–11" captionStartId="F3" captionText="Figures 7 - 11. Variation of markings of 7 Microdytes eliasi sp. nov. 8 M. jeenthongi sp. nov 9 M. maximiliani sp. nov. 10 M. sekaensis sp. nov. 11 M. ubonensis sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures7-11" httpUri="https://binary.pensoft.net/fig/842222" pageId="0" pageNumber="87">, 7</figureCitation>
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<figureCitation captionStart="Figures 12–14" captionStartId="F4" captionText="Figures 12 - 14. 12 Microdytes eliasi sp. nov. 13 M. jeenthongi sp. nov. 14 M. maximiliani sp. nov. A median lobe in ventral aspect B median lobe in lateral aspect C left paramere in lateral aspect D right paramere in medial aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1159.99218.figures12-14" httpUri="https://binary.pensoft.net/fig/842223" pageId="0" pageNumber="87">, 12</figureCitation>
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<figureCitation captionStart="Figures 41–43" captionStartId="F9" captionText="Figures 41 - 43. 41 Lectotype of Microdytes maculatus (photograph by Christophe Rivier, Museum national d'histoire naturelle, Paris) 42 median lobe of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov. 43 paramere of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov." figureDoi="10.3897/zookeys.1159.99218.figures41-43" httpUri="https://binary.pensoft.net/fig/842229" pageId="0" pageNumber="87">, 42C</figureCitation>
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<figureCitation captionStart="Figures 41–43" captionStartId="F9" captionText="Figures 41 - 43. 41 Lectotype of Microdytes maculatus (photograph by Christophe Rivier, Museum national d'histoire naturelle, Paris) 42 median lobe of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov. 43 paramere of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov." figureDoi="10.3897/zookeys.1159.99218.figures41-43" httpUri="https://binary.pensoft.net/fig/842229" pageId="0" pageNumber="87">, 43C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="87" type="type locality">
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<paragraph pageId="0" pageNumber="87">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="87">
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Cambodia, Koh Hong Province, 20 km SW Koh Hong, Tatai River,
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<geoCoordinate degrees="11" direction="north" minutes="34" orientation="latitude" precision="925" value="11.566667">11°34'N</geoCoordinate>
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,
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<geoCoordinate degrees="103" direction="east" minutes="07" orientation="longitude" precision="925" value="103.11667">103°07'E</geoCoordinate>
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, alt. 50-300 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="87" type="type material">
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<paragraph pageId="0" pageNumber="87">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Holotype</emphasis>
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(NMB): ♂ "Cambodia SW, Tatai riv. 20 km SE Koh Hong
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<geoCoordinate degrees="11" direction="north" minutes="34" orientation="latitude" precision="925" value="11.566667">11°34'N</geoCoordinate>
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,
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<geoCoordinate degrees="103" direction="east" minutes="07" orientation="longitude" precision="925" value="103.11667">103°07'E</geoCoordinate>
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, 3.-19.v.2005, 50-300 m, E. Jendek, O.
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<normalizedToken originalValue="Šauša">Sausa</normalizedToken>
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leg." [printed white label] \ "HOLOTYPUS
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<taxonomicName authorityName="Wewalka & Okada" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes eliasi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="eliasi">Microdytes eliasi</taxonomicName>
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sp. nov. Wewalka & Okada 2022" [printed red label].
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<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Paratypes</emphasis>
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: (47 exs.): 7♂♂, 14♀♀, with same data as the holotype (CGW, CRO, NMB, NMP); 2♂♂, Thailand, Saraburi Province, Kaeng Khoi District, Ched Khot St. 122 (alt. 140 m),
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<geoCoordinate degrees="14" direction="north" minutes="28" orientation="latitude" precision="15" seconds="26" value="14.473888">14°28'26"N</geoCoordinate>
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,
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<geoCoordinate degrees="101" direction="east" minutes="09" orientation="longitude" precision="15" seconds="59" value="101.16639">101°09'59"E</geoCoordinate>
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, 30.V.2020, R. Okada leg. (CRO, THNHM); 4♂♂, 4♀♀, Thailand, Sa Kaeo Province, Mueang Sa Kaeo District, Ban Kaeng St. 277 (alt. 80 m),
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<geoCoordinate degrees="13" direction="north" minutes="58" orientation="latitude" precision="15" seconds="47" value="13.979722">13°58'47"N</geoCoordinate>
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,
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<geoCoordinate degrees="102" direction="east" minutes="11" orientation="longitude" precision="15" seconds="29" value="102.19139">102°11'29"E</geoCoordinate>
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, 30.I.2022, R. Okada leg. (CGW, CRO, THNHM); 2♂♂, 3♀♀, Thailand, Chonburi Province, Bo Thong District, That Thong St. 107 (alt. 100 m),
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<geoCoordinate degrees="13" direction="north" minutes="15" orientation="latitude" precision="15" seconds="01" value="13.2502775">13°15'01"N</geoCoordinate>
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,
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<geoCoordinate degrees="101" direction="east" minutes="22" orientation="longitude" precision="15" seconds="34" value="101.376114">101°22'34"E</geoCoordinate>
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, 14.III.2020, R. Okada leg. (CGW, CRO, THNHM); 2♂♂, 2♀♀, same locality St. 298 (alt. 100 m), 13.VII.2022, R. Okada leg. (CGW, CRO); 1♀, same province, Si Racha District, Bang Phra St. 269 (alt. 90 m),
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<geoCoordinate degrees="13" direction="north" minutes="14" orientation="latitude" precision="15" seconds="39" value="13.244167">13°14'39"N</geoCoordinate>
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,
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<geoCoordinate degrees="101" direction="east" minutes="02" orientation="longitude" precision="15" seconds="20" value="101.03889">101°02'20"E</geoCoordinate>
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, 26.XII.2021, R. Okada leg. (CRO); 3♂♂, 2♀♀, Thailand, Chanthaburi Province, Khaeng Hong Maeo District, Kaeng Hong waterfall St. 257 (alt. 160 m),
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<geoCoordinate degrees="13" direction="north" minutes="02" orientation="latitude" precision="15" seconds="57" value="13.049167">13°02'57"N</geoCoordinate>
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,
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<geoCoordinate degrees="101" direction="east" minutes="45" orientation="longitude" precision="15" seconds="35" value="101.75972">101°45'35"E</geoCoordinate>
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, 26.VI.2021, R. Okada leg. (CGW, CRO, THNHM); 1♀, same district, Kha riv. St. 258 (alt. 90 m),
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<geoCoordinate degrees="12" direction="north" minutes="57" orientation="latitude" precision="15" seconds="38" value="12.960555">12°57'38"N</geoCoordinate>
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,
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<geoCoordinate degrees="101" direction="east" minutes="46" orientation="longitude" precision="15" seconds="30" value="101.775">101°46'30"E</geoCoordinate>
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, 26.VI.2021, R. Okada leg. (CRO). All paratypes are provided with printed red paratype labels.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="87" type="diagnosis">
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<paragraph pageId="0" pageNumber="87">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="87">
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<taxonomicName authorityName="Wewalka & Okada" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes eliasi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="eliasi">
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<emphasis italics="true" pageId="0" pageNumber="87">Microdytes eliasi</emphasis>
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</taxonomicName>
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sp. nov. very closely resembles
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<taxonomicName lsidName="M. maculatus" pageId="0" pageNumber="87" rank="species" species="maculatus">
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<emphasis italics="true" pageId="0" pageNumber="87">M. maculatus</emphasis>
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</taxonomicName>
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(Motschulsky, 1860) (Figs
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<figureCitation captionStart="Figures 18–37" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">25</figureCitation>
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,
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<figureCitation captionStart="Figures 41–43" captionStartId="F9" captionText="Figures 41 - 43. 41 Lectotype of Microdytes maculatus (photograph by Christophe Rivier, Museum national d'histoire naturelle, Paris) 42 median lobe of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov. 43 paramere of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov." figureDoi="10.3897/zookeys.1159.99218.figures41-43" httpUri="https://binary.pensoft.net/fig/842229" pageId="0" pageNumber="87">41</figureCitation>
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) in size and coloration but differs from this species by the more regularly oval habitus, the laterally expanded median lobe at apex and constricted tips of paramere (Figs
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<figureCitation captionStart="Figures 12–14" captionStartId="F4" captionText="Figures 12 - 14. 12 Microdytes eliasi sp. nov. 13 M. jeenthongi sp. nov. 14 M. maximiliani sp. nov. A median lobe in ventral aspect B median lobe in lateral aspect C left paramere in lateral aspect D right paramere in medial aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1159.99218.figures12-14" httpUri="https://binary.pensoft.net/fig/842223" pageId="0" pageNumber="87">12</figureCitation>
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,
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<figureCitation captionStart="Figures 41–43" captionStartId="F9" captionText="Figures 41 - 43. 41 Lectotype of Microdytes maculatus (photograph by Christophe Rivier, Museum national d'histoire naturelle, Paris) 42 median lobe of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov. 43 paramere of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov." figureDoi="10.3897/zookeys.1159.99218.figures41-43" httpUri="https://binary.pensoft.net/fig/842229" pageId="0" pageNumber="87">42C</figureCitation>
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,
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<figureCitation captionStart="Figures 41–43" captionStartId="F9" captionText="Figures 41 - 43. 41 Lectotype of Microdytes maculatus (photograph by Christophe Rivier, Museum national d'histoire naturelle, Paris) 42 median lobe of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov. 43 paramere of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov." figureDoi="10.3897/zookeys.1159.99218.figures41-43" httpUri="https://binary.pensoft.net/fig/842229" pageId="0" pageNumber="87">43C</figureCitation>
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) (see also comments in
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<taxonomicName lsidName="M. maculatus" pageId="0" pageNumber="87" rank="species" species="maculatus">
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<emphasis italics="true" pageId="0" pageNumber="87">M. maculatus</emphasis>
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</taxonomicName>
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). It is also similar to
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<taxonomicName lsidName="M. feryi" pageId="0" pageNumber="87" rank="species" species="feryi">
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<emphasis italics="true" pageId="0" pageNumber="87">M. feryi</emphasis>
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</taxonomicName>
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Wewalka, 2011 (Fig.
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<figureCitation captionStart="Figures 18–37" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">21</figureCitation>
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) in habitus and coloration, but it is smaller and can be distinguished by male genitalia [compared with a male paratype from Myanmar, "Tenasserim, Birma Coll. V. Helfer National Museum Prague" (CGW)].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="87" type="description">
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<paragraph pageId="0" pageNumber="87">Description.</paragraph>
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" pageId="0" pageNumber="87">
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<emphasis italics="true" pageId="0" pageNumber="87">Measurements</emphasis>
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.
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</emphasis>
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TL = 1.64-1.85 mm (1.76 mm), TL-H = 1.49-1.65 mm (1.55 mm), MW = 1.12-1.25 mm (1.24 mm), TL/MW = 1.33-1.44 (1.42). Body regularly oval, moderately convex (Fig.
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<figureCitation captionStart="Figures 2–6" captionStartId="F2" captionText="Figures 2 - 6. Habitus of 2 Microdytes eliasi sp. nov., female, paratype 3 M. jeenthongi sp. nov., male, holotype 4 M. maximiliani sp. nov., male, holotype 5 M. sekaensis sp. nov., male, holotype 6 M. ubonensis sp. nov., male, holotype. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures2-6" httpUri="https://binary.pensoft.net/fig/842221" pageId="0" pageNumber="87">2</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" pageId="0" pageNumber="87">
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<emphasis italics="true" pageId="0" pageNumber="87">Coloration</emphasis>
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.
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</emphasis>
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Head reddish brown. Pronotum reddish brown, narrowly dark brown along anterior and posterior margins. Elytron dark brown with yellowish brown markings forming a distinct transverse band near base not reaching suture connected along lateral margin with a post-median transverse lateral band, with a small post-median spot near suture, and a triangular spot near apex (Figs
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<figureCitation captionStart="Figures 2–6" captionStartId="F2" captionText="Figures 2 - 6. Habitus of 2 Microdytes eliasi sp. nov., female, paratype 3 M. jeenthongi sp. nov., male, holotype 4 M. maximiliani sp. nov., male, holotype 5 M. sekaensis sp. nov., male, holotype 6 M. ubonensis sp. nov., male, holotype. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures2-6" httpUri="https://binary.pensoft.net/fig/842221" pageId="0" pageNumber="87">2</figureCitation>
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,
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<figureCitation captionStart="Figures 7–11" captionStartId="F3" captionText="Figures 7 - 11. Variation of markings of 7 Microdytes eliasi sp. nov. 8 M. jeenthongi sp. nov 9 M. maximiliani sp. nov. 10 M. sekaensis sp. nov. 11 M. ubonensis sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures7-11" httpUri="https://binary.pensoft.net/fig/842222" pageId="0" pageNumber="87">7</figureCitation>
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). Ventral surface of head, prothorax and elytral epipleuron yellowish brown; thorax and abdominal ventrites reddish brown to dark brown. Legs, antennae and palpi yellowish brown.
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</paragraph>
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<caption doi="10.3897/zookeys.1159.99218.figures2-6" httpUri="https://binary.pensoft.net/fig/842221" pageId="0" pageNumber="87" start="Figures 2–6" startId="F2">
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<paragraph pageId="0" pageNumber="87">
|
||
<emphasis bold="true" pageId="0" pageNumber="87">Figures 2-6.</emphasis>
|
||
Habitus of
|
||
<emphasis bold="true" pageId="0" pageNumber="87">2</emphasis>
|
||
<taxonomicName authorityName="Wewalka & Okada" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes eliasi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="eliasi">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">Microdytes eliasi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., female, paratype
|
||
<emphasis bold="true" pageId="0" pageNumber="87">3</emphasis>
|
||
<taxonomicName lsidName="M. jeenthongi" pageId="0" pageNumber="87" rank="species" species="jeenthongi">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. jeenthongi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male, holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="87">4</emphasis>
|
||
<taxonomicName lsidName="M. maximiliani" pageId="0" pageNumber="87" rank="species" species="maximiliani">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. maximiliani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male, holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="87">5</emphasis>
|
||
<taxonomicName lsidName="M. sekaensis" pageId="0" pageNumber="87" rank="species" species="sekaensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. sekaensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male, holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="87">6</emphasis>
|
||
<taxonomicName lsidName="M. ubonensis" pageId="0" pageNumber="87" rank="species" species="ubonensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. ubonensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male, holotype. Scale bar: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1159.99218.figures7-11" httpUri="https://binary.pensoft.net/fig/842222" pageId="0" pageNumber="87" start="Figures 7–11" startId="F3">
|
||
<paragraph pageId="0" pageNumber="87">
|
||
<emphasis bold="true" pageId="0" pageNumber="87">Figures 7-11.</emphasis>
|
||
Variation of markings of
|
||
<emphasis bold="true" pageId="0" pageNumber="87">7</emphasis>
|
||
<taxonomicName authorityName="Wewalka & Okada" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes eliasi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="eliasi">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">Microdytes eliasi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="87">8</emphasis>
|
||
<taxonomicName lsidName="M. jeenthongi" pageId="0" pageNumber="87" rank="species" species="jeenthongi">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. jeenthongi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov
|
||
<emphasis bold="true" pageId="0" pageNumber="87">9</emphasis>
|
||
<taxonomicName lsidName="M. maximiliani" pageId="0" pageNumber="87" rank="species" species="maximiliani">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. maximiliani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="87">10</emphasis>
|
||
<taxonomicName lsidName="M. sekaensis" pageId="0" pageNumber="87" rank="species" species="sekaensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. sekaensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="87">11</emphasis>
|
||
<taxonomicName lsidName="M. ubonensis" pageId="0" pageNumber="87" rank="species" species="ubonensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. ubonensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Scale bar: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="87">
|
||
<emphasis bold="true" pageId="0" pageNumber="87">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">Sculpture and structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Head finely, sparsely, and relatively regularly punctured; anterior half to two-thirds finely microreticulate; clypeus not bordered. Pronotum quite regularly, sparsely, and fairly strongly punctured, with coarser punctures along posterior margin; without microreticulation; lateral margins finely bordered, regularly rounded. Elytron quite regularly, moderately densely and fairly strongly punctured, progressively finer and sparser towards lateral margin; without longitudinal rows of stronger punctures; highly polished and shining; without microreticulation. Ventral surface: metacoxae and metasternum strongly but sparsely punctured, abdomen finely and sparsely punctured; without microreticulation.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="87">
|
||
<emphasis bold="true" pageId="0" pageNumber="87">Male.</emphasis>
|
||
The two parts of the median lobe expanded laterally at apex in ventral aspect (Figs
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F4" captionText="Figures 12 - 14. 12 Microdytes eliasi sp. nov. 13 M. jeenthongi sp. nov. 14 M. maximiliani sp. nov. A median lobe in ventral aspect B median lobe in lateral aspect C left paramere in lateral aspect D right paramere in medial aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1159.99218.figures12-14" httpUri="https://binary.pensoft.net/fig/842223" pageId="0" pageNumber="87">12A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 41–43" captionStartId="F9" captionText="Figures 41 - 43. 41 Lectotype of Microdytes maculatus (photograph by Christophe Rivier, Museum national d'histoire naturelle, Paris) 42 median lobe of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov. 43 paramere of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov." figureDoi="10.3897/zookeys.1159.99218.figures41-43" httpUri="https://binary.pensoft.net/fig/842229" pageId="0" pageNumber="87">42A</figureCitation>
|
||
); slightly curved in lateral aspect (Fig.
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F4" captionText="Figures 12 - 14. 12 Microdytes eliasi sp. nov. 13 M. jeenthongi sp. nov. 14 M. maximiliani sp. nov. A median lobe in ventral aspect B median lobe in lateral aspect C left paramere in lateral aspect D right paramere in medial aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1159.99218.figures12-14" httpUri="https://binary.pensoft.net/fig/842223" pageId="0" pageNumber="87">12B</figureCitation>
|
||
). The tips of parameres twisted in lateral aspect (Fig.
|
||
<figureCitation captionStart="Figures 12–14" captionStartId="F4" captionText="Figures 12 - 14. 12 Microdytes eliasi sp. nov. 13 M. jeenthongi sp. nov. 14 M. maximiliani sp. nov. A median lobe in ventral aspect B median lobe in lateral aspect C left paramere in lateral aspect D right paramere in medial aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1159.99218.figures12-14" httpUri="https://binary.pensoft.net/fig/842223" pageId="0" pageNumber="87">12C, D</figureCitation>
|
||
); constricted in ventral aspect (Fig.
|
||
<figureCitation captionStart="Figures 41–43" captionStartId="F9" captionText="Figures 41 - 43. 41 Lectotype of Microdytes maculatus (photograph by Christophe Rivier, Museum national d'histoire naturelle, Paris) 42 median lobe of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov. 43 paramere of A M. maculatus from northern Thailand B from northeastern Thailand C M. eliasi sp. nov." figureDoi="10.3897/zookeys.1159.99218.figures41-43" httpUri="https://binary.pensoft.net/fig/842229" pageId="0" pageNumber="87">43C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1159.99218.figures12-14" httpUri="https://binary.pensoft.net/fig/842223" pageId="0" pageNumber="87" start="Figures 12–14" startId="F4">
|
||
<paragraph pageId="0" pageNumber="87">
|
||
<emphasis bold="true" pageId="0" pageNumber="87">Figures 12-14.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="87">12</emphasis>
|
||
<taxonomicName authorityName="Wewalka & Okada" authorityYear="2023" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes eliasi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="eliasi">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">Microdytes eliasi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="87">13</emphasis>
|
||
<taxonomicName lsidName="M. jeenthongi" pageId="0" pageNumber="87" rank="species" species="jeenthongi">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. jeenthongi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="87">14</emphasis>
|
||
<taxonomicName lsidName="M. maximiliani" pageId="0" pageNumber="87" rank="species" species="maximiliani">
|
||
<emphasis italics="true" pageId="0" pageNumber="87">M. maximiliani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="87">A</emphasis>
|
||
median lobe in ventral aspect
|
||
<emphasis bold="true" pageId="0" pageNumber="87">B</emphasis>
|
||
median lobe in lateral aspect
|
||
<emphasis bold="true" pageId="0" pageNumber="87">C</emphasis>
|
||
left paramere in lateral aspect
|
||
<emphasis bold="true" pageId="0" pageNumber="87">D</emphasis>
|
||
right paramere in medial aspect. Scale bar: 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="87">
|
||
<emphasis bold="true" pageId="0" pageNumber="87">Female.</emphasis>
|
||
Without secondary sexual characters. Sclerotized spermatheca not found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="87" type="variation">
|
||
<paragraph pageId="0" pageNumber="87">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="87">
|
||
Variation of markings is shown in Fig.
|
||
<figureCitation captionStart="Figures 7–11" captionStartId="F3" captionText="Figures 7 - 11. Variation of markings of 7 Microdytes eliasi sp. nov. 8 M. jeenthongi sp. nov 9 M. maximiliani sp. nov. 10 M. sekaensis sp. nov. 11 M. ubonensis sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures7-11" httpUri="https://binary.pensoft.net/fig/842222" pageId="0" pageNumber="87">7</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="87" type="etymology">
|
||
<paragraph pageId="0" pageNumber="87">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="87">
|
||
This species is dedicated to Elias
|
||
<normalizedToken originalValue="Bramböck">Bramboeck</normalizedToken>
|
||
, Vienna, Austria. The species epithet is a name in the genitive singular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="87" type="habitat">
|
||
<paragraph pageId="0" pageNumber="87">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="87">
|
||
In Thailand, this species was collected in small streams at low altitude lower than 200 m (Fig.
|
||
<figureCitation captionStart="Figures 45–48" captionStartId="F11" captionText="Figures 45 - 48. Collecting localities of Microdytes species in Thailand 45 Ban Kaeng, Sa Kaeo Province, one of the localities of M. eliasi sp. nov. 46 Tha Pha, Chiang Mai Province, type locality of M. jeenthongi sp. nov. 47 Ban Tong, Bueng Kan Province, type locality of M. sekaensis sp. nov. 48 Nam Thaeng, Ubon Ratchathani Province, type locality of M. ubonensis sp. nov." figureDoi="10.3897/zookeys.1159.99218.figures45-48" httpUri="https://binary.pensoft.net/fig/842231" pageId="0" pageNumber="87">45</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="87" type="distribution">
|
||
<paragraph pageId="0" pageNumber="87">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="87">Thailand: Saraburi, Sa Kaeo, Chonburi, and Chanthaburi provinces; Cambodia: Koh Hong Province.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |