181 lines
17 KiB
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181 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.962.52759" ID-GBIF-Dataset="0aa621ef-de45-41be-8c10-73ba877577b9" ID-PMC="PMC7471473" ID-Pensoft-Pub="1313-2970-962-1" ID-Pensoft-UUID="AD14C60754A7579492E50291C659E3C6" ID-PubMed="32943974" ID-ZooBank="CA7DEA7CBFEB4BC8A598A802941FD59F" ModsDocID="1313-2970-962-1" checkinTime="1598026755870" checkinUser="pensoft" docAuthor="Laudee, Pongsak, Seetapan, Kriengkrai, Vongsombath, Chanda & Malicky, Hans" docDate="2020" docId="BCFCC471AED159DAAE84C09B8E4BF563" docLanguage="en" docName="ZooKeys 962: 1-11" docOrigin="ZooKeys 962" docSource="http://dx.doi.org/10.3897/zookeys.962.52759" docTitle="Ecnomus boonsawaengae Malicky & Vongsombath 2020, sp. nov." docType="treatment" docUuid="A970D2D6-D505-4A72-B693-D2C6DC131325" docUuidSource="ZooBank" docVersion="4" id="AD14C60754A7579492E50291C659E3C6" lastPageNumber="1" masterDocId="AD14C60754A7579492E50291C659E3C6" masterDocTitle="New species of caddisflies (Trichoptera, Ecnomidae, Polycentropodidae, Psychomyiidae) from Mekong tributaries, Laos" masterLastPageNumber="11" masterPageNumber="1" pageNumber="1" updateTime="1668169335156" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of caddisflies (Trichoptera, Ecnomidae, Polycentropodidae, Psychomyiidae) from Mekong tributaries, Laos</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Laudee, Pongsak</mods:namePart>
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<mods:affiliation>Department of Fishery and Costal Resources, Faculty of Science and Industrial Technology, Prince of Songkla University, Surat Thani Campus, Muang District, Surat Thani Province, Thailand</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-3819-7980</mods:nameIdentifier>
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<mods:nameIdentifier type="email">pongsak.l@psu.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Seetapan, Kriengkrai</mods:namePart>
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<mods:affiliation>School of Agriculture and Natural Resources, University of Phayao, Tumbol Maeka, Muang District, Phayao Province, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vongsombath, Chanda</mods:namePart>
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<mods:affiliation>Faculty of Environmental Sciences, National University of Laos, Dong Dok Campus, Vientiane, Laos</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Malicky, Hans</mods:namePart>
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<mods:affiliation>Sonnengasse 13, A- 3293 Lunz am See, Austria</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>962</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>11</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.962.52759</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.962.52759</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-962-1</mods:identifier>
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<mods:identifier type="ZooBank">CA7DEA7CBFEB4BC8A598A802941FD59F</mods:identifier>
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<mods:identifier type="Pensoft-UUID">AD14C60754A7579492E50291C659E3C6</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="167287971" LSID="urn:lsid:zoobank.org:act:A970D2D6-D505-4A72-B693-D2C6DC131325" httpUri="http://treatment.plazi.org/id/BCFCC471AED159DAAE84C09B8E4BF563" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="BCFCC471-AED1-59DA-AE84-C09B8E4BF563" authority="Malicky & Vongsombath" authorityName="Malicky & Vongsombath" authorityYear="2020" class="Insecta" family="Ecnomidae" genus="Ecnomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecnomus boonsawaengae" order="Trichoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boonsawaengae" status="sp. nov.">Ecnomus boonsawaengae Malicky & Vongsombath</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">Figure 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The male genitalia of
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<taxonomicName lsidName="E. boonsawaengae" pageId="0" pageNumber="1" rank="species" species="boonsawaengae">
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<emphasis italics="true" pageId="0" pageNumber="1">E. boonsawaengae</emphasis>
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</taxonomicName>
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sp. nov. are similar to
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<taxonomicName lsidName="E. aktaion" pageId="0" pageNumber="1" rank="species" species="aktaion">
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<emphasis italics="true" pageId="0" pageNumber="1">E. aktaion</emphasis>
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</taxonomicName>
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Malicky & Chantaramongkol, 1997 and
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<taxonomicName lsidName="E. uttu" pageId="0" pageNumber="1" rank="species" species="uttu">
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<emphasis italics="true" pageId="0" pageNumber="1">E. uttu</emphasis>
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</taxonomicName>
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Malicky & Chantaramongkol, 1993. In these species, the superior appendages are particularly long and slender with a basoventral process on the superior appendage. However,
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<taxonomicName lsidName="E. boonsawaengae" pageId="0" pageNumber="1" rank="species" species="boonsawaengae">
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<emphasis italics="true" pageId="0" pageNumber="1">E. boonsawaengae</emphasis>
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</taxonomicName>
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sp. nov. can be easily distinguished by the shape of the inferior appendages. In lateral view, the inferior appendages are tubular with a subapical concave incision and truncated apex in
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<taxonomicName lsidName="E. boonsawaengae" pageId="0" pageNumber="1" rank="species" species="boonsawaengae">
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<emphasis italics="true" pageId="0" pageNumber="1">E. boonsawaengae</emphasis>
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</taxonomicName>
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sp. nov., but in
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<taxonomicName lsidName="E. aktaion" pageId="0" pageNumber="1" rank="species" species="aktaion">
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<emphasis italics="true" pageId="0" pageNumber="1">E. aktaion</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="E. uttu" pageId="0" pageNumber="1" rank="species" species="uttu">
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<emphasis italics="true" pageId="0" pageNumber="1">E. uttu</emphasis>
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</taxonomicName>
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the inferior appendages are somewhat triangular and trapezoidal, respectively, and with a pointed apex. In addition, each inferior appendage in
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<taxonomicName lsidName="E. boonsawaengae" pageId="0" pageNumber="1" rank="species" species="boonsawaengae">
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<emphasis italics="true" pageId="0" pageNumber="1">E. boonsawaengae</emphasis>
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</taxonomicName>
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sp. nov. has a process, in ventral view, which is lacking in
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<taxonomicName lsidName="E. aktaion" pageId="0" pageNumber="1" rank="species" species="aktaion">
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<emphasis italics="true" pageId="0" pageNumber="1">E. aktaion</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="E. uttu" pageId="0" pageNumber="1" rank="species" species="uttu">
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<emphasis italics="true" pageId="0" pageNumber="1">E. uttu</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Adult, male, length of each male forewing 4.0 mm; color in alcohol of head, thorax, forewings, abdomen, and legs grayish brown. Male genitalia as in Figure
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2A-D</figureCitation>
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. Tergum IX in dorsal view bilobed posteriorly, U-shaped
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<normalizedToken originalValue="½">1/2</normalizedToken>
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incision anteriorly (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2A</figureCitation>
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); in lateral view, tergum IX narrow, expanded dorsally (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2B</figureCitation>
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). Superior appendages tubular, base with lateral lobe, slightly bent apically to form beak-like apex in dorsal view (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2A</figureCitation>
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); in lateral view, tubular, truncated apically (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2B</figureCitation>
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). Basoventral projection of superior appendage tubular with setae apically in lateral view. Sternum IX in lateral view chicken-drumstick-like and rounded apically (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2B</figureCitation>
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); in ventral view, trapezoid, slightly expanded apically, with shallow U-shaped incision anteriorly and shallow V-shaped incision posteriorly (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2C</figureCitation>
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). In lateral view, the inferior appendages tubular, with concave incision subapically, truncated apex (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2B</figureCitation>
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); in ventral view, claw-like, with process basodorsally (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2C</figureCitation>
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). Phallus sickle-like, with lobe mesodosally in lateral view (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ecnomus boonsawaengae, sp. nov. Male genitalia. A Segment IX and superior appendages, dorsal B segment IX and superior appendages, left lateral C segment IX and inferior appendages, ventral D phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf = inferior appendage." figureDoi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1">2D</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.962.52759.figure2" httpUri="https://binary.pensoft.net/fig/443671" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
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<taxonomicName authorityName="Malicky & Vongsombath" authorityYear="2020" class="Insecta" family="Ecnomidae" genus="Ecnomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ecnomus boonsawaengae" order="Trichoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="boonsawaengae">
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<emphasis italics="true" pageId="0" pageNumber="1">Ecnomus boonsawaengae</emphasis>
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</taxonomicName>
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, sp. nov. Male genitalia.
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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Segment IX and superior appendages, dorsal
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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segment IX and superior appendages, left lateral
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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segment IX and inferior appendages, ventral
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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phallus, lateral. Ter IX = tergum IX, Ste IX = sternum IX, Sup = superior appendage, Inf=inferior appendage.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Male. Laos: Pakse Province
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</emphasis>
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: Paksong, E-Tu Waterfall,
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<geoCoordinate degrees="15" direction="north" minutes="11" orientation="latitude" precision="15" seconds="25" value="15.190278">15°11'25"N</geoCoordinate>
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,
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<geoCoordinate degrees="106" direction="east" minutes="06" orientation="longitude" precision="15" seconds="14" value="106.10389">106°06'14"E</geoCoordinate>
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, elev. 900 m, 7.iv.2019, Pongsak Laudee. (PSUNHM).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: same data as holotype, 2 males: 1 male (PSUNHM), 1 male (CHM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The species epithet honors Mrs Wasana Boonsawaeng, Vice Dean of Faculty of Science and Industrial Technology, Prince of Songkla University, Surat Thani Campus.</paragraph>
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</subSubSection>
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</treatment>
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</document> |