239 lines
35 KiB
XML
239 lines
35 KiB
XML
<document id="C9EA0E1CC2B2FCB875E3040CEE5AEC67" ID-DOI="10.5281/zenodo.157264" ID-GBIF-Dataset="45961c86-477a-4d6c-a3de-e18d51aa2251" ID-ISSN="11755326" ID-Zenodo-Dep="157264" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461021445387" checkinUser="plazi" docAuthor="Mesibov, Robert" docDate="2004" docId="9A6F4F12FFFCFFEC21409552FBEAFCE2" docLanguage="en" docName="zt00558.pdf" docOrigin="Zootaxa 558" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Bromodesmus Mesibov, 2004, n. gen." docType="treatment" docVersion="10" lastPageNumber="8" masterDocId="6656376AFFFFFFEB20489304FFEDFFDC" masterDocTitle="A new genus and four new species of millipedes from Tasmania, Australia (Diplopoda: Polydesmida: Dalodesmidae), with notes on male leg setae in some Tasmanian dalodesmids" masterLastPageNumber="19" masterPageNumber="1" pageNumber="4" updateTime="1698570320343" updateUser="plazi">
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<mods:title id="FA92E35D492B0D0B26D50AAC0F3F1CA8">A new genus and four new species of millipedes from Tasmania, Australia (Diplopoda: Polydesmida: Dalodesmidae), with notes on male leg setae in some Tasmanian dalodesmids</mods:title>
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<mods:namePart id="98E148F0CECC6072EE7954B5544783EC">Mesibov, Robert</mods:namePart>
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<mods:title id="8D8E4409A45684036E41BC0B86578A4E">Zootaxa</mods:title>
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<mods:part id="5B9C657D5CC6B409AE2244C1F45055C5">
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<mods:date id="A393D58632D14F504C69EA4BB95385CA">2004</mods:date>
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<mods:identifier id="01677793DA2A70AAAD6D3C986AFFE47F" type="DOI">10.5281/zenodo.157264</mods:identifier>
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<treatment id="9A6F4F12FFFCFFEC21409552FBEAFCE2" ID-DOI="http://doi.org/10.5281/zenodo.6269963" ID-GBIF-Taxon="119602998" ID-Zenodo-Dep="6269963" LSID="urn:lsid:plazi:treatment:9A6F4F12FFFCFFEC21409552FBEAFCE2" httpUri="http://treatment.plazi.org/id/9A6F4F12FFFCFFEC21409552FBEAFCE2" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4">
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<subSubSection id="5ADCAD8FFFFCFFE821409552FDE3F9AC" box="[264,526,1622,1648]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="1279FE04FFFCFFE821409552FDE3F9AC" blockId="3.[264,526,1622,1648]" box="[264,526,1622,1648]" pageId="3" pageNumber="4">
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<heading id="49314968FFFCFFE821409552FDE3F9AC" bold="true" box="[264,526,1622,1648]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis id="20B22216FFFCFFE821409552FDE3F9AC" bold="true" box="[264,526,1622,1648]" pageId="3" pageNumber="4">
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<taxonomicName id="D5C68587FFFCFFE821409552FE59F9AC" ID-CoL="7NV5L" box="[264,436,1622,1648]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="20B22216FFFCFFE821409552FE59F9AC" bold="true" box="[264,436,1622,1648]" italics="true" pageId="3" pageNumber="4">Bromodesmus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3B819F6DFFFCFFE821F49552FDE3F9AC" box="[444,526,1622,1648]" pageId="3" pageNumber="4" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="5ADCAD8FFFFCFFE8214095A5FDF6F8D7" pageId="3" pageNumber="4" type="materials_examined">
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<paragraph id="1279FE04FFFCFFE8214095A5FBC1F967" blockId="3.[264,1323,1697,1883]" box="[264,1068,1697,1723]" pageId="3" pageNumber="4">
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<emphasis id="20B22216FFFCFFE8214095A5FE75F966" box="[264,408,1697,1722]" italics="true" pageId="3" pageNumber="4">
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<typeStatus id="CD7D40A6FFFCFFE8214095A5FED1F966" box="[264,316,1697,1722]" pageId="3" pageNumber="4">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="D5C68587FFFCFFE821E295A5FD2AF966" box="[426,711,1697,1722]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="catrionae" status="sp. nov.">
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<emphasis id="20B22216FFFCFFE821E295A5FD2AF966" box="[426,711,1697,1722]" italics="true" pageId="3" pageNumber="4">Bromodesmus catrionae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3B819F6DFFFCFFE8228695A5FCE2F967" box="[718,783,1697,1723]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
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, by present designation.
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</paragraph>
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<paragraph id="1279FE04FFFCFFE8217095CDFDF6F8D7" blockId="3.[264,1323,1697,1883]" pageId="3" pageNumber="4">
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<emphasis id="20B22216FFFCFFE8217095CDFDB8F93E" box="[312,597,1737,1762]" italics="true" pageId="3" pageNumber="4">Other assigned species</emphasis>
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:
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<taxonomicName id="D5C68587FFFCFFE8222595CDFC69F93E" box="[621,900,1737,1762]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="militaris">
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<emphasis id="20B22216FFFCFFE8222595CDFC69F93E" box="[621,900,1737,1762]" italics="true" pageId="3" pageNumber="4">Bromodesmus militaris</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="D5C68587FFFCFFE823D395CDFB43F93E" box="[923,1198,1737,1762]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="riparius">
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<emphasis id="20B22216FFFCFFE823D395CDFB43F93E" box="[923,1198,1737,1762]" italics="true" pageId="3" pageNumber="4">Bromodesmus riparius</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="D5C68587FFFCFFE824BE95CDFE28F8D6" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="rufus">
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<emphasis id="20B22216FFFCFFE824BE95CDFE28F8D6" italics="true" pageId="3" pageNumber="4">Bromodesmus rufus</emphasis>
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</taxonomicName>
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n. spp.
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</paragraph>
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</subSubSection>
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<subSubSection id="5ADCAD8FFFFCFFEF2170941DFD65FE5A" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="1279FE04FFFCFFEF2170941DFD65FE5A" blockId="3.[264,1323,1697,1883]" lastBlockId="4.[264,1325,284,1870]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<emphasis id="20B22216FFFCFFE82170941DFE42F8EE" box="[312,431,1817,1842]" italics="true" pageId="3" pageNumber="4">Diagnosis</emphasis>
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: Small to mediumsized burrowing dalodesmids (
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<figureCitation id="8AFDE281FFFCFFE82458941DFBB6F8EF" box="[1040,1115,1817,1843]" captionStart="FIGURE 2" captionStartId="5.[264,368,806,830]" captionTargetBox="[368,1219,310,781]" captionTargetId="figure@5.[368,1219,307,783]" captionTargetPageId="5" captionText="FIGURE 2. Bromodesmus spp. females. (A) B. catrionae n. sp. (QVM 23: 22082), (B) B. militaris n. sp. (QVM 23: 15150), (C) B. riparius n. sp. (QVM 23: 45813) and (D) B. rufus n. sp. (QVM 23: 40519). Scale bar = 10 mm." httpUri="https://zenodo.org/record/169085/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
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) with head + 20 segments and greatly reduced paranota. Gonopod telopodite straight, slender, greatly expanded at distal end into a posteriorly concave ‘hood’ fringed with teeth; solenomerite arising mesally onethird to halfway along the telopodite, curving anteriorly, then posteriorly, and tapering to a fine point.
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</paragraph>
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</subSubSection>
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<subSubSection id="5ADCAD8FFFFBFFEE21709290FD61F93B" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
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<paragraph id="1279FE04FFFBFFEF21709290FBB7FA62" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
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<emphasis id="20B22216FFFBFFEF21709290FE2FFE71" box="[312,450,404,429]" italics="true" pageId="4" pageNumber="5">Description</emphasis>
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: Head with moderately deep impressions ventrolateral of antennal bases; bases separated by about twice a socket diameter; vertigial sulcus not deeply impressed but marked by dark pigment; head moderately setose with moderately long setae on clypeus and frons, and a few long setae on the lower portion of the vertex. Antennae (
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<figureCitation id="8AFDE281FFFBFFEF24B39108FEFAFD92" captionStart="FIGURE 4" captionStartId="6.[264,368,867,891]" captionTargetBox="[288,1282,327,824]" captionTargetId="figure@6.[234,1332,308,854]" captionTargetPageId="6" captionText="FIGURE 4. Bromodesmus catrionae n. sp., male and female from the same site on Ballroom Hill (male from QVM 23: 22072, female from QVM 23: 22082). (A) male antenna; (B) dorsal and (C) posterior outlines of segment 12 of male (top) and female (bottom); (D) posterior leg on segment 6 of male (top) and female (bottom); (E) posterior leg on segment 12 of male (top) and female (bottom). Setation and antennal cones not shown. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/157267/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
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A) slender, with antennomeres 2 and 3 subequal in length, longer than 4 and 5 and slightly longer than 6; antennomere 6 the widest. Collum slightly narrower than head and second segment; anterior and posterior margins straight, corners rounded; sparsely setose, principally near the margins. Segments with welldefined waist (= constriction incorporating posterior portion of prozonite, suture and anterior portion of metazonite) (
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<figureCitation id="8AFDE281FFFBFFEF24F491D0FAEAFD32" box="[1212,1287,724,750]" captionStart="FIGURE 4" captionStartId="6.[264,368,867,891]" captionTargetBox="[288,1282,327,824]" captionTargetId="figure@6.[234,1332,308,854]" captionTargetPageId="6" captionText="FIGURE 4. Bromodesmus catrionae n. sp., male and female from the same site on Ballroom Hill (male from QVM 23: 22072, female from QVM 23: 22082). (A) male antenna; (B) dorsal and (C) posterior outlines of segment 12 of male (top) and female (bottom); (D) posterior leg on segment 6 of male (top) and female (bottom); (E) posterior leg on segment 12 of male (top) and female (bottom). Setation and antennal cones not shown. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/157267/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
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B); paranotal development variable (see species descriptions) but lateral margin of paranotum on segment 2 always well below collum corner, lateral margin of paranota on segment 3 just above level of collum corner. Segment surfaces smooth; a few small setae near the waist and the posterior margin of the metazonite; limbus a comb of fine, straight teeth. Ozopore small, opening midlaterally near posterior edge of metazonite, at the level of the paranotal thickening on segments 5, 7, 9, 10, 12, 13, 15–19. Sternites a little longer than wide, cross impressions welldefined. Anterior legs (
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<figureCitation id="8AFDE281FFFBFFEF23C290E8FC3EFBDA" box="[906,979,1004,1030]" captionStart="FIGURE 4" captionStartId="6.[264,368,867,891]" captionTargetBox="[288,1282,327,824]" captionTargetId="figure@6.[234,1332,308,854]" captionTargetPageId="6" captionText="FIGURE 4. Bromodesmus catrionae n. sp., male and female from the same site on Ballroom Hill (male from QVM 23: 22072, female from QVM 23: 22082). (A) male antenna; (B) dorsal and (C) posterior outlines of segment 12 of male (top) and female (bottom); (D) posterior leg on segment 6 of male (top) and female (bottom); (E) posterior leg on segment 12 of male (top) and female (bottom). Setation and antennal cones not shown. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/157267/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
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D) with swollen podomeres, prefemur strongly arched dorsally; posterior legs (
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<figureCitation id="8AFDE281FFFBFFEF232D9710FC43FBF2" box="[869,942,1044,1070]" captionStart="FIGURE 4" captionStartId="6.[264,368,867,891]" captionTargetBox="[288,1282,327,824]" captionTargetId="figure@6.[234,1332,308,854]" captionTargetPageId="6" captionText="FIGURE 4. Bromodesmus catrionae n. sp., male and female from the same site on Ballroom Hill (male from QVM 23: 22072, female from QVM 23: 22082). (A) male antenna; (B) dorsal and (C) posterior outlines of segment 12 of male (top) and female (bottom); (D) posterior leg on segment 6 of male (top) and female (bottom); (E) posterior leg on segment 12 of male (top) and female (bottom). Setation and antennal cones not shown. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/157267/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
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E) less swollen; tarsus about as long as next longest podomere, the femur. Sphaerotrichomes on tibia and tarsus of leg 3 to penultimate leg; sphaerotrichome shafts with expanded tips (
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<figureCitation id="8AFDE281FFFBFFEF23909760FBF1FBA2" box="[984,1052,1124,1150]" captionStart="FIGURE 1" captionStartId="1.[264,367,1723,1747]" captionText="FIGURE 1. SEM views of setae on ventral side of podomeres of male leg 10. (A – D) sphaerotrichomes, (E – F) ‘ brush’ setae. (A) Atrophotergum silvaticum tibia (QVM 23: 25279); (B) tarsus and (E) femur of Bromodesmus catrionae n. sp. (QVM 23: 22072); (C) tibia and (F) femur of Gasterogramma psi (QVM 23: 25577); (D) tarsus and (G) femur of Lissodesmus modestus (QVM 23: 41975); (H) femur of Tasmanodesmus hardyi (QVM 23: 45800). All scale bars = 0.020 mm." pageId="4" pageNumber="5">Fig. 1</figureCitation>
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B). Dense ‘brushes’ of setae ventrally on prefemur, femur, postfemur and tibia of most legs; ‘brush’ setae with expanded tips (
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<figureCitation id="8AFDE281FFFBFFEF21F297B0FE11FB12" box="[442,508,1204,1230]" captionStart="FIGURE 1" captionStartId="1.[264,367,1723,1747]" captionText="FIGURE 1. SEM views of setae on ventral side of podomeres of male leg 10. (A – D) sphaerotrichomes, (E – F) ‘ brush’ setae. (A) Atrophotergum silvaticum tibia (QVM 23: 25279); (B) tarsus and (E) femur of Bromodesmus catrionae n. sp. (QVM 23: 22072); (C) tibia and (F) femur of Gasterogramma psi (QVM 23: 25577); (D) tarsus and (G) femur of Lissodesmus modestus (QVM 23: 41975); (H) femur of Tasmanodesmus hardyi (QVM 23: 45800). All scale bars = 0.020 mm." pageId="4" pageNumber="5">Fig. 1</figureCitation>
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E). Leg 2 coxa somewhat extended mesally, genital opening a simple pit on this extension. Legpair 4 not separated at bases, legpair 5 slightly separated, legpairs 6 and 7 wellseparated; flexed gonopod telopodites reaching to legpair 5. Spiracle simple in structure with slightly raised rim; on diplosegments, anterior spiracle just above and anterior to anterior leg, posterior spiracle just above and posterior to anterior leg. Preanal ring moderately setose; epiproct tapering to blunt, somewhat rounded tip extending well past anal valves; hypoproct paraboloid or (in
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<taxonomicName id="D5C68587FFFBFFEF235496A0FC95FA61" box="[796,888,1444,1469]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rufus" status="sp. nov.">
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<emphasis id="20B22216FFFBFFEF235496A0FC95FA61" box="[796,888,1444,1469]" italics="true" pageId="4" pageNumber="5">B. rufus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3B819F6DFFFBFFEF23C896A0FC52FA62" box="[896,959,1444,1470]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
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) trapezoidal.
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</paragraph>
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<paragraph id="1279FE04FFFBFFEF217096C8FC35F892" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
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Gonopod aperture ovoid, wider than long, about onethird the width of segment 7 prozonite, with posterolateral portions of rim projecting ventrally. Gonocoxa more or less conical, tapering distally, but with broad, shallow excavation on posteromesal surface (
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<figureCitation id="8AFDE281FFFBFFEF21599540FEB0F982" box="[273,349,1604,1630]" captionStart="FIGURE 6" captionStartId="7.[264,367,1684,1708]" captionTargetBox="[603,984,1201,1660]" captionTargetId="figure@7.[603,984,1201,1660]" captionTargetPageId="7" captionText="FIGURE 6. Bromodesmus catrionae n. sp. SEM view of gonopods removed from segment 7, showing dorsal (posterior) aspect of gonocoxae (QVM 23: 17836); c = cannula. Scale bar = 0.50 mm." httpUri="https://zenodo.org/record/169088/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
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), excavation lined with short setae; gonocoxae very lightly joined mesally near base. Cannula prominent (
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<figureCitation id="8AFDE281FFFBFFEF220A9568FD61F95A" box="[578,652,1644,1670]" captionStart="FIGURE 6" captionStartId="7.[264,367,1684,1708]" captionTargetBox="[603,984,1201,1660]" captionTargetId="figure@7.[603,984,1201,1660]" captionTargetPageId="7" captionText="FIGURE 6. Bromodesmus catrionae n. sp. SEM view of gonopods removed from segment 7, showing dorsal (posterior) aspect of gonocoxae (QVM 23: 17836); c = cannula. Scale bar = 0.50 mm." httpUri="https://zenodo.org/record/169088/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
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). Telopodite (
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<figureCitation id="8AFDE281FFFBFFEF237B9568FC96F95A" box="[819,891,1644,1670]" captionStart="FIGURE 5" captionStartId="6.[264,367,1699,1723]" captionTargetBox="[481,1093,1084,1643]" captionTargetPageId="6" captionText="FIGURE 5. Bromodesmus catrionae n. sp. (Left) SEM view of gonopods in situ (QVM 23: 22076). Note that the shapes of the ‘ hoods’ and the solenomerite tips have been somewhat distorted in preparing the specimen for electron microscopy. Scale bar = 0.50 mm. (Right) Mesal view of distal portion of right telopodite (QVM 23: 16185). Dashed line shows course of prostatic groove; for clarity the continuation of the groove to the tip of the solenomerite is not indicated." httpUri="https://zenodo.org/record/169087/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
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) straight, slender; tip expanded into posteriorly concave ‘hood’ with toothed rim; moderately setose with long setae on posterior surface to onethird to onehalf the telopodite length; solenomerite arising mesally at onethird to half the telopodite length, curved anteriorly, then posteriorly, tapering to fine point, terminating within or just mesal to ‘hood’; prostatic groove running more or less directly on anteromesal side of telopodite to solenomerite tip.
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</paragraph>
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<caption id="46B9AE8CFFFAFFEE21409022FDB9FC5F" httpUri="https://zenodo.org/record/169085/files/figure.png" pageId="5" pageNumber="6" targetBox="[368,1219,310,781]" targetPageId="5">
|
||
<paragraph id="1279FE04FFFAFFEE21409022FDB9FC5F" blockId="5.[264,1323,806,899]" pageId="5" pageNumber="6">
|
||
<emphasis id="20B22216FFFAFFEE21409022FE66FCE2" bold="true" box="[264,395,806,830]" pageId="5" pageNumber="6">FIGURE 2.</emphasis>
|
||
<taxonomicName id="D5C68587FFFAFFEE21DB9023FDC4FCE2" box="[403,553,807,830]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="20B22216FFFAFFEE21DB9023FDC4FCE2" box="[403,553,807,830]" italics="true" pageId="5" pageNumber="6">Bromodesmus</emphasis>
|
||
</taxonomicName>
|
||
spp. females. (A)
|
||
<taxonomicName id="D5C68587FFFAFFEE22B89023FC99FCE2" box="[752,884,807,830]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="catrionae" status="sp. nov.">
|
||
<emphasis id="20B22216FFFAFFEE22B89023FC99FCE2" box="[752,884,807,830]" italics="true" pageId="5" pageNumber="6">B. catrionae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFFAFFEE23349023FC5BFCE2" box="[892,950,807,830]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
(QVM 23:22082), (B)
|
||
<taxonomicName id="D5C68587FFFAFFEE24FA9023FAC6FCE2" box="[1202,1323,807,830]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="militaris" status="sp. nov.">
|
||
<emphasis id="20B22216FFFAFFEE24FA9023FAC6FCE2" box="[1202,1323,807,830]" italics="true" pageId="5" pageNumber="6">B. militaris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFFAFFEE2140904EFEA9FCBD" box="[264,324,842,865]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
(QVM 23:15150), (C)
|
||
<taxonomicName id="D5C68587FFFAFFEE2204904EFD2FFCBD" box="[588,706,842,865]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="20B22216FFFAFFEE2204904EFD2FFCBD" box="[588,706,842,865]" italics="true" pageId="5" pageNumber="6">B. riparius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFFAFFEE2284904EFCE4FCBD" box="[716,777,842,865]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
(QVM 23:45813) and (D)
|
||
<taxonomicName id="D5C68587FFFAFFEE2473904EFB7EFCBD" box="[1083,1171,842,865]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rufus" status="sp. nov.">
|
||
<emphasis id="20B22216FFFAFFEE2473904EFB7EFCBD" box="[1083,1171,842,865]" italics="true" pageId="5" pageNumber="6">B. rufus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFFAFFEE24D5904EFB37FCBD" box="[1181,1242,842,865]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
(QVM 23:40519). Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="46B9AE8CFFFAFFEE214096DBFB06F9C2" httpUri="https://zenodo.org/record/169086/files/figure.png" pageId="5" pageNumber="6" targetBox="[266,1322,972,1481]" targetPageId="5">
|
||
<paragraph id="1279FE04FFFAFFEE214096DBFB06F9C2" blockId="5.[264,1322,1503,1566]" pageId="5" pageNumber="6">
|
||
<emphasis id="20B22216FFFAFFEE214096DBFE60FA2B" bold="true" box="[264,397,1503,1527]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
|
||
<taxonomicName id="D5C68587FFFAFFEE21DE96E4FD66FA2B" box="[406,651,1504,1527]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
|
||
<emphasis id="20B22216FFFAFFEE21DE96E4FD66FA2B" box="[406,651,1504,1527]" italics="true" pageId="5" pageNumber="6">Bromodesmus riparius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFFAFFEE22DD96E4FD3CFA2B" box="[661,721,1504,1527]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
Right lateral views of males with gonocoxae retracted into aperture (left) and extruded (right). Both specimens from type locality (QVM 23:45812).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1279FE04FFFAFFEE21709579FD61F93B" blockId="5.[264,1323,1661,1847]" pageId="5" pageNumber="6">
|
||
Females (
|
||
<figureCitation id="8AFDE281FFFAFFEE21E49579FDEFF94B" box="[428,514,1661,1687]" captionStart="FIGURE 2" captionStartId="5.[264,368,806,830]" captionTargetBox="[368,1219,310,781]" captionTargetId="figure@5.[368,1219,307,783]" captionTargetPageId="5" captionText="FIGURE 2. Bromodesmus spp. females. (A) B. catrionae n. sp. (QVM 23: 22082), (B) B. militaris n. sp. (QVM 23: 15150), (C) B. riparius n. sp. (QVM 23: 45813) and (D) B. rufus n. sp. (QVM 23: 40519). Scale bar = 10 mm." httpUri="https://zenodo.org/record/169085/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="8AFDE281FFFAFFEE225A9579FDCFF94B" box="[530,546,1661,1687]" captionStart="FIGURE 4" captionStartId="6.[264,368,867,891]" captionTargetBox="[288,1282,327,824]" captionTargetId="figure@6.[234,1332,308,854]" captionTargetPageId="6" captionText="FIGURE 4. Bromodesmus catrionae n. sp., male and female from the same site on Ballroom Hill (male from QVM 23: 22072, female from QVM 23: 22082). (A) male antenna; (B) dorsal and (C) posterior outlines of segment 12 of male (top) and female (bottom); (D) posterior leg on segment 6 of male (top) and female (bottom); (E) posterior leg on segment 12 of male (top) and female (bottom). Setation and antennal cones not shown. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/157267/files/figure.png" pageId="5" pageNumber="6">4</figureCitation>
|
||
) larger than males, with same coloring; legs smaller than corresponding male legs and not swollen (
|
||
<figureCitation id="8AFDE281FFFAFFEE228B95A1FCE0F963" box="[707,781,1701,1727]" captionStart="FIGURE 4" captionStartId="6.[264,368,867,891]" captionTargetBox="[288,1282,327,824]" captionTargetId="figure@6.[234,1332,308,854]" captionTargetPageId="6" captionText="FIGURE 4. Bromodesmus catrionae n. sp., male and female from the same site on Ballroom Hill (male from QVM 23: 22072, female from QVM 23: 22082). (A) male antenna; (B) dorsal and (C) posterior outlines of segment 12 of male (top) and female (bottom); (D) posterior leg on segment 6 of male (top) and female (bottom); (E) posterior leg on segment 12 of male (top) and female (bottom). Setation and antennal cones not shown. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/157267/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
|
||
); posterior rim of epigynum raised slightly in center; cyphopods not examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5ADCAD8FFFFAFFEE217095F1FCBEF8EB" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="1279FE04FFFAFFEE217095F1FCBEF8EB" blockId="5.[264,1323,1661,1847]" pageId="5" pageNumber="6">
|
||
<emphasis id="20B22216FFFAFFEE217095F1FE58F8D2" box="[312,437,1781,1806]" italics="true" pageId="5" pageNumber="6">Etymology</emphasis>
|
||
: Greek
|
||
<emphasis id="20B22216FFFAFFEE225695F1FD98F8D2" box="[542,629,1781,1806]" italics="true" pageId="5" pageNumber="6">bromos</emphasis>
|
||
, stench, +
|
||
<emphasis id="20B22216FFFAFFEE234695F1FC89F8D2" box="[782,868,1781,1806]" italics="true" pageId="5" pageNumber="6">desmus</emphasis>
|
||
, commonly used combining form in generic names of
|
||
<taxonomicName id="D5C68587FFFAFFEE219F9419FD9FF8EB" box="[471,626,1821,1847]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
|
||
; gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="46B9AE8CFFF9FFED21409067FC26FBDA" httpUri="https://zenodo.org/record/157267/files/figure.png" pageId="6" pageNumber="7" targetBox="[288,1282,327,824]" targetPageId="6">
|
||
<paragraph id="1279FE04FFF9FFED21409067FC26FBDA" blockId="6.[264,1323,867,1030]" pageId="6" pageNumber="7">
|
||
<emphasis id="20B22216FFF9FFED21409067FE66FCA7" bold="true" box="[264,395,867,891]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
|
||
<taxonomicName id="D5C68587FFF9FFED21DC9060FD7AFCA7" box="[404,663,868,891]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="catrionae" status="sp. nov.">
|
||
<emphasis id="20B22216FFF9FFED21DC9060FD7AFCA7" box="[404,663,868,891]" italics="true" pageId="6" pageNumber="7">Bromodesmus catrionae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFF9FFED22D79060FD36FCA7" box="[671,731,868,891]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
|
||
, male and female from the same site on Ballroom Hill (male from QVM 23:22072, female from QVM 23:22082). (A) male antenna; (B) dorsal and (C) posterior outlines of segment 12 of male (top) and female (bottom); (D) posterior leg on segment 6 of male (top) and female (bottom); (E) posterior leg on segment 12 of male (top) and female (bottom). Setation and antennal cones not shown. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="46B9AE8CFFF9FFED214095A7FB6DF88E" httpUri="https://zenodo.org/record/169087/files/figure.png" pageId="6" pageNumber="7" targetBox="[481,1093,1084,1643]" targetPageId="6">
|
||
<paragraph id="1279FE04FFF9FFED214095A7FB6DF88E" blockId="6.[264,1323,1699,1874]" pageId="6" pageNumber="7">
|
||
<emphasis id="20B22216FFF9FFED214095A7FE64F967" bold="true" box="[264,393,1699,1723]" pageId="6" pageNumber="7">FIGURE 5.</emphasis>
|
||
<taxonomicName id="D5C68587FFF9FFED21D995A0FD7EF967" box="[401,659,1700,1723]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="catrionae" status="sp. nov.">
|
||
<emphasis id="20B22216FFF9FFED21D995A0FD7EF967" box="[401,659,1700,1723]" italics="true" pageId="6" pageNumber="7">Bromodesmus catrionae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFF9FFED22D295A0FD3EF967" box="[666,723,1700,1723]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
|
||
(Left) SEM view of gonopods
|
||
<emphasis id="20B22216FFF9FFED246895A0FB8FF967" box="[1056,1122,1700,1723]" italics="true" pageId="6" pageNumber="7">in situ</emphasis>
|
||
(QVM 23:22076). Note that the shapes of the ‘hoods’ and the solenomerite tips have been somewhat distorted in preparing the specimen for electron microscopy. Scale bar = 0.50 mm. (Right) Mesal view of distal portion of right telopodite (QVM 23:16185). Dashed line shows course of prostatic groove; for clarity the continuation of the groove to the tip of the solenomerite is not indicated.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="5ADCAD8FFFF8FFEC21709218FBEAFCE2" pageId="7" pageNumber="8" type="discussion">
|
||
<paragraph id="1279FE04FFF8FFEC21709218FC75FD42" blockId="7.[264,1323,284,830]" pageId="7" pageNumber="8">
|
||
<emphasis id="20B22216FFF8FFEC21709218FE73FEE9" box="[312,414,284,309]" italics="true" pageId="7" pageNumber="8">Remarks</emphasis>
|
||
: The muscles and some subcuticular tissues of specimens of
|
||
<taxonomicName id="D5C68587FFF8FFEC24329218FAE1FEE9" box="[1146,1292,284,309]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="catrionae" status="sp. nov.">
|
||
<emphasis id="20B22216FFF8FFEC24329218FAE1FEE9" box="[1146,1292,284,309]" italics="true" pageId="7" pageNumber="8">B. catrionae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFF8FFEC255C9218FEC6FE82" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="D5C68587FFF8FFEC21709240FE51FE81" box="[312,444,324,349]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="militaris" status="sp. nov.">
|
||
<emphasis id="20B22216FFF8FFEC21709240FE51FE81" box="[312,444,324,349]" italics="true" pageId="7" pageNumber="8">B. militaris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFF8FFEC218C9240FDEEFE82" box="[452,515,324,350]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="D5C68587FFF8FFEC22749240FD74FE81" box="[572,665,324,349]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="rufus" status="sp. nov.">
|
||
<emphasis id="20B22216FFF8FFEC22749240FD74FE81" box="[572,665,324,349]" italics="true" pageId="7" pageNumber="8">B. rufus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B819F6DFFF8FFEC22E99240FD0DFE82" box="[673,736,324,350]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
||
are stained a dark redbrown after long storage in alcohol. It seems likely that the staining is due to a chemically reactive component or components in the defensive secretion. Similar staining can occur in alcoholpreserved specimens of Julida, Spirobolida and Spirostreptida as a result of the tanning action of 1,4 benzoquinones in the defensive secretion (
|
||
<bibRefCitation id="765783F5FFF8FFEC234E92E0FC60FE22" author="Huth" box="[774,909,484,510]" pageId="7" pageNumber="15" refString="Huth, A. (2000) Defensive secretions of millipedes: more than just a product of melting point decrease? In: Wytwer, J. & Golovatch, S. (Eds.), Progress in Studies on Myriapoda and Onychophora. Fragmenta Faunistica (Warsaw) 43 (Supplement 2000). Museum and Institute of Zoology, Polish Academy of Sciences, Warsaw, pp. 191 - 200." type="journal article" year="2000">Huth 2000</bibRefCitation>
|
||
). However, the
|
||
<taxonomicName id="D5C68587FFF8FFEC240492E0FB1CFE21" box="[1100,1265,484,509]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="20B22216FFF8FFEC240492E0FB1CFE21" box="[1100,1265,484,509]" italics="true" pageId="7" pageNumber="8">Bromodesmus</emphasis>
|
||
</taxonomicName>
|
||
spp. secretion is unlikely to be a benzoquinone, as it is far more volatile than the secretion of Tasmanian spirostreptidans and has a very different odor. Further, the supernatant alcohol in vials containing longpreserved
|
||
<taxonomicName id="D5C68587FFF8FFEC22E39158FCBDFDA9" box="[683,848,604,629]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="20B22216FFF8FFEC22E39158FCBDFDA9" box="[683,848,604,629]" italics="true" pageId="7" pageNumber="8">Bromodesmus</emphasis>
|
||
</taxonomicName>
|
||
spp. is colorless, whereas the liquid in Spirostreptidacontaining vials is a deep reddishpurple.
|
||
</paragraph>
|
||
<paragraph id="1279FE04FFF8FFEC217091A8FBEAFCE2" blockId="7.[264,1323,284,830]" pageId="7" pageNumber="8">
|
||
Alcoholpreserved
|
||
<taxonomicName id="D5C68587FFF8FFEC225091A8FD50FD19" box="[536,701,684,709]" class="Diplopoda" family="Dalodesmidae" genus="Bromodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="20B22216FFF8FFEC225091A8FD50FD19" box="[536,701,684,709]" italics="true" pageId="7" pageNumber="8">Bromodesmus</emphasis>
|
||
</taxonomicName>
|
||
specimens are also noticeably more brittle than other Tasmanian dalodesmids preserved in the same way for the same length of time. This brittleness is likely to be due to greater calcification of the cuticle, as the cuticle of preserved specimens can be made flexible by soaking in household vinegar.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |