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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.878.37845" ID-GBIF-Dataset="faa91fff-ad2f-44bc-8309-388d6755f640" ID-PMC="PMC6791898" ID-Pensoft-Pub="1313-2970-878-33" ID-Pensoft-UUID="C0F938DD0EFE5A18BB267539EC83EFDB" ID-PubMed="31632176" ID-ZooBank="BCE3960BE7C6418FB8802978DF9F099E" ModsDocID="1313-2970-878-33" checkinTime="1570497201903" checkinUser="pensoft" docAuthor="Hopkins, Tapani, Roininen, Heikki, Noort, Simon van, Broad, Gavin R., Kaunisto, Kari &amp; Saeaeksjaervi, Ilari E." docDate="2019" docId="7E4997D9C6695140B12221AF66A1328B" docLanguage="en" docName="ZooKeys 878: 33-71" docOrigin="ZooKeys 878" docSource="http://dx.doi.org/10.3897/zookeys.878.37845" docTitle="Epirhyssa uelensis Benoit 1951" docType="treatment" docVersion="5" id="C0F938DD0EFE5A18BB267539EC83EFDB" lastPageNumber="33" masterDocId="C0F938DD0EFE5A18BB267539EC83EFDB" masterDocTitle="Extensive sampling and thorough taxonomic assessment of Afrotropical Rhyssinae (Hymenoptera, Ichneumonidae) reveals two new species and demonstrates the limitations of previous sampling efforts" masterLastPageNumber="71" masterPageNumber="33" pageNumber="33" updateTime="1668167851423" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Extensive sampling and thorough taxonomic assessment of Afrotropical Rhyssinae (Hymenoptera, Ichneumonidae) reveals two new species and demonstrates the limitations of previous sampling efforts</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hopkins, Tapani</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Roininen, Heikki</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Noort, Simon van</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Broad, Gavin R.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kaunisto, Kari</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saeaeksjaervi, Ilari E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>878</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>33</mods:start>
<mods:end>71</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.878.37845</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.878.37845</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-878-33</mods:identifier>
<mods:identifier type="ZooBank">BCE3960BE7C6418FB8802978DF9F099E</mods:identifier>
<mods:identifier type="Pensoft-UUID">C0F938DD0EFE5A18BB267539EC83EFDB</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159654855" LSID="urn:lsid:plazi:treatment:7E4997D9C6695140B12221AF66A1328B" httpUri="http://treatment.plazi.org/id/7E4997D9C6695140B12221AF66A1328B" lastPageNumber="33" pageId="0" pageNumber="33">
<subSubSection pageId="0" pageNumber="33" type="nomenclature">
<paragraph pageId="0" pageNumber="33">
<taxonomicName LSID="7E4997D9-C669-5140-B122-21AF66A1328B" authority="Benoit, 1951" authorityName="Benoit" authorityYear="1951" class="Insecta" family="Ichneumonidae" genus="Epirhyssa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epirhyssa uelensis" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="uelensis">Epirhyssa uelensis Benoit, 1951</taxonomicName>
<figureCitation captionStart="Figures 5763" captionStartId="F13" captionText="Figures 57 - 63. Epirhyssa uelensis female (http: // mus. utu. fi / ZMUT. 2520), a species found in Uganda. 57 Habitus 58 face and clypeus 59 frons 60 hypostomal flange 61 mesopleuron dorsal margin 62 apical horn of metasoma 63 tergites 1 - 7. Scale bars: 0.5 mm (58 - 63), 1 mm (57)." figureDoi="10.3897/zookeys.878.37845.figures57-63" httpUri="https://binary.pensoft.net/fig/345029" pageId="0" pageNumber="33" tableDoi="10.3897/zookeys.878.37845.figures57-63">Figs 57-63</figureCitation>
,
<figureCitation captionStart="Figures 6467" captionStartId="F14" captionText="Figures 64 - 67. Epirhyssa gavinbroadi female (holotype). We treat this species as a synonym of E. uelensis in this work. 64 Habitus 65 hypostomal flange 66 mesopleuron dorsal margin 67 apical horn of metasoma. Figure 64 is from van Noort (2019)." figureDoi="10.3897/zookeys.878.37845.figures64-67" httpUri="https://binary.pensoft.net/fig/345030" pageId="0" pageNumber="33" tableDoi="10.3897/zookeys.878.37845.figures64-67">64-67</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="reference_group">
<paragraph pageId="0" pageNumber="33">
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Epirhyssa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epirhyssa gavinbroadi" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="gavinbroadi">Epirhyssa gavinbroadi</taxonomicName>
Rousse &amp; van Noort, 2014, syn. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="materials_examined">
<paragraph pageId="0" pageNumber="33">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Type material</emphasis>
: CAMEROON:
</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♀, holotype of
<taxonomicName lsidName="E. gavinbroadi" pageId="0" pageNumber="33" rank="species" species="gavinbroadi">
<emphasis italics="true" pageId="0" pageNumber="33">E. gavinbroadi</emphasis>
</taxonomicName>
; Nkoemvon [
<normalizedToken originalValue="Nkoemvon">Nko'emvon</normalizedToken>
];
<geoCoordinate degrees="02" direction="north" minutes="48" orientation="latitude" precision="925" value="2.8">02°48'N</geoCoordinate>
,
<geoCoordinate degrees="11" direction="east" minutes="08" orientation="longitude" precision="925" value="11.133333">11°08'E</geoCoordinate>
; 16 Mar. 1980-4 May 1980; NHMUK type number HYM 3b.2832.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Non-type material</emphasis>
: UGANDA
</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♀; Kibale National Park, Kanyawara, Site K31, Malaise trap K31T4;
<geoCoordinate degrees="0.5362" direction="north" orientation="latitude" precision="5" value="0.5362">0.5362N</geoCoordinate>
,
<geoCoordinate degrees="30.3486" direction="east" orientation="longitude" precision="5" value="30.3486">30.3486E</geoCoordinate>
(WGS84); alt. 1460 m (GPS, WGS84); 16 Jul. 2015-30 Jul. 2015; Tapani Hopkins leg.; ZMUThttp://mus.utu.fi/ZMUT.2292
</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♀; same data as preceding; Site K30, Malaise trap K30T3;
<geoCoordinate degrees="0.5590" direction="north" orientation="latitude" precision="5" value="0.559">0.5590N</geoCoordinate>
,
<geoCoordinate degrees="30.3617" direction="east" orientation="longitude" precision="5" value="30.3617">30.3617E</geoCoordinate>
(WGS84); alt. 1540 m (GPS, WGS84); 21 Apr. 2015-5 May 2015; ZMUThttp://mus.utu.fi/ZMUT.2520
</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♀; same data as preceding; Site K31, Malaise trap K31T4;
<geoCoordinate degrees="0.5362" direction="north" orientation="latitude" precision="5" value="0.5362">0.5362N</geoCoordinate>
,
<geoCoordinate degrees="30.3486" direction="east" orientation="longitude" precision="5" value="30.3486">30.3486E</geoCoordinate>
(WGS84); alt. 1460 m (GPS, WGS84); 30 Jan. 2015-13 Feb. 2015; ZMUThttp://mus.utu.fi/ZMUT.2878
</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♀; same data as preceding; Site K13, Malaise trap K13T1;
<geoCoordinate degrees="0.5932" direction="north" orientation="latitude" precision="5" value="0.5932">0.5932N</geoCoordinate>
,
<geoCoordinate degrees="30.3598" direction="east" orientation="longitude" precision="5" value="30.3598">30.3598E</geoCoordinate>
(WGS84); alt. 1460 m (GPS, WGS84); 13 Jul. 2015-27 Jul. 2015; ZMUThttp://mus.utu.fi/ZMUT.5665
</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♀; same data as preceding; Site K30, Malaise trap K30T2;
<geoCoordinate degrees="0.5566" direction="north" orientation="latitude" precision="5" value="0.5566">0.5566N</geoCoordinate>
,
<geoCoordinate degrees="30.3633" direction="east" orientation="longitude" precision="5" value="30.3633">30.3633E</geoCoordinate>
(WGS84); alt. 1490 m (GPS, WGS84); 2 Dec. 2014-15 Dec. 2014; ZMUThttp://mus.utu.fi/ZMUT.5689
</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♀; same data as preceding; Site CC, Malaise trap CCT1;
<geoCoordinate degrees="0.5497" direction="north" orientation="latitude" precision="5" value="0.5497">0.5497N</geoCoordinate>
,
<geoCoordinate degrees="30.3673" direction="east" orientation="longitude" precision="5" value="30.3673">30.3673E</geoCoordinate>
(WGS84); alt. 1450 m (GPS, WGS84); 27 Jan. 2015-10 Feb. 2015; ZMUThttp://mus.utu.fi/ZMUT.5732
</paragraph>
<paragraph pageId="0" pageNumber="33">• 1 ♀; same data as preceding; 24 Feb. 2015-10 Mar. 2015; ZMUThttp://mus.utu.fi/ZMUT.5739</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♀; same data as preceding; Site K30S, Malaise trap K30ST4;
<geoCoordinate degrees="0.5414" direction="north" orientation="latitude" precision="5" value="0.5414">0.5414N</geoCoordinate>
,
<geoCoordinate degrees="30.3755" direction="east" orientation="longitude" precision="5" value="30.3755">30.3755E</geoCoordinate>
(WGS84); alt. 1420 m (GPS, WGS84); 30 Jun. 2015-14 Jul. 2015; ZMUThttp://mus.utu.fi/ZMUT.5844
</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♂; Kibale National Park, Kanyawara, Site R93, Malaise trap R93T2;
<geoCoordinate degrees="0.5654" direction="north" orientation="latitude" precision="5" value="0.5654">0.5654N</geoCoordinate>
,
<geoCoordinate degrees="30.3593" direction="east" orientation="longitude" precision="5" value="30.3593">30.3593E</geoCoordinate>
(WGS84); alt. 1510 m (GPS, WGS84); 20 May 2015-1 Jun. 2015; Tapani Hopkins leg.; ZMUThttp://mus.utu.fi/ZMUT.5765
</paragraph>
<paragraph pageId="0" pageNumber="33">
• 1 ♂; same data as preceding; Site K13, Malaise trap K13T1;
<geoCoordinate degrees="0.5932" direction="north" orientation="latitude" precision="5" value="0.5932">0.5932N</geoCoordinate>
,
<geoCoordinate degrees="30.3598" direction="east" orientation="longitude" precision="5" value="30.3598">30.3598E</geoCoordinate>
(WGS84); alt. 1460 m (GPS, WGS84); 4 May 2015-20 May 2015; ZMUThttp://mus.utu.fi/ZMUT.5807.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Non-type material</emphasis>
(only diagnostic characters checked): UGANDA:
</paragraph>
<paragraph pageId="0" pageNumber="33">• 150 ♀; Kibale National Park, Kanyawara; Tapani Hopkins leg.; ZMUThttp://mus.utu.fi/ZMUT.1265, http://mus.utu.fi/ZMUT.1266, http://mus.utu.fi/ZMUT.1267, http://mus.utu.fi/ZMUT.1268, http://mus.utu.fi/ZMUT.1272, http://mus.utu.fi/ZMUT.1334, http://mus.utu.fi/ZMUT.1366, http://mus.utu.fi/ZMUT.1367, http://mus.utu.fi/ZMUT.1375, http://mus.utu.fi/ZMUT.1499, http://mus.utu.fi/ZMUT.1673, http://mus.utu.fi/ZMUT.1677, http://mus.utu.fi/ZMUT.1763, http://mus.utu.fi/ZMUT.1764, http://mus.utu.fi/ZMUT.1807, http://mus.utu.fi/ZMUT.2015, http://mus.utu.fi/ZMUT.2152, http://mus.utu.fi/ZMUT.2162, http://mus.utu.fi/ZMUT.2165, http://mus.utu.fi/ZMUT.2172, http://mus.utu.fi/ZMUT.2299, http://mus.utu.fi/ZMUT.2317, http://mus.utu.fi/ZMUT.2340, http://mus.utu.fi/ZMUT.2361, http://mus.utu.fi/ZMUT.2362, http://mus.utu.fi/ZMUT.2364, http://mus.utu.fi/ZMUT.2473, http://mus.utu.fi/ZMUT.2573, http://mus.utu.fi/ZMUT.2640, http://mus.utu.fi/ZMUT.2642, http://mus.utu.fi/ZMUT.2643, http://mus.utu.fi/ZMUT.2644, http://mus.utu.fi/ZMUT.2646, http://mus.utu.fi/ZMUT.2648, http://mus.utu.fi/ZMUT.2887, http://mus.utu.fi/ZMUT.2910, http://mus.utu.fi/ZMUT.3022, http://mus.utu.fi/ZMUT.3090, http://mus.utu.fi/ZMUT.3091, http://mus.utu.fi/ZMUT.3093, http://mus.utu.fi/ZMUT.3094, http://mus.utu.fi/ZMUT.3096, http://mus.utu.fi/ZMUT.3102, http://mus.utu.fi/ZMUT.3106, http://mus.utu.fi/ZMUT.3135, http://mus.utu.fi/ZMUT.3225, http://mus.utu.fi/ZMUT.3259, http://mus.utu.fi/ZMUT.3262, http://mus.utu.fi/ZMUT.3334, http://mus.utu.fi/ZMUT.3438, http://mus.utu.fi/ZMUT.3452, http://mus.utu.fi/ZMUT.3619, http://mus.utu.fi/ZMUT.3698, http://mus.utu.fi/ZMUT.3743, http://mus.utu.fi/ZMUT.3745, http://mus.utu.fi/ZMUT.3747, http://mus.utu.fi/ZMUT.3947, http://mus.utu.fi/ZMUT.4424, http://mus.utu.fi/ZMUT.4512, http://mus.utu.fi/ZMUT.4517, http://mus.utu.fi/ZMUT.5086, http://mus.utu.fi/ZMUT.5315, http://mus.utu.fi/ZMUT.5381, http://mus.utu.fi/ZMUT.5542, http://mus.utu.fi/ZMUT.5589, http://mus.utu.fi/ZMUT.5593, http://mus.utu.fi/ZMUT.5597, http://mus.utu.fi/ZMUT.5601, http://mus.utu.fi/ZMUT.5605, http://mus.utu.fi/ZMUT.5606, http://mus.utu.fi/ZMUT.5607, http://mus.utu.fi/ZMUT.5608, http://mus.utu.fi/ZMUT.5613, http://mus.utu.fi/ZMUT.5614, http://mus.utu.fi/ZMUT.5616, http://mus.utu.fi/ZMUT.5618, http://mus.utu.fi/ZMUT.5619, http://mus.utu.fi/ZMUT.5620, http://mus.utu.fi/ZMUT.5623, http://mus.utu.fi/ZMUT.5625, http://mus.utu.fi/ZMUT.5629, http://mus.utu.fi/ZMUT.5631, http://mus.utu.fi/ZMUT.5632, http://mus.utu.fi/ZMUT.5635, http://mus.utu.fi/ZMUT.5637, http://mus.utu.fi/ZMUT.5641, http://mus.utu.fi/ZMUT.5642, http://mus.utu.fi/ZMUT.5643, http://mus.utu.fi/ZMUT.5644, http://mus.utu.fi/ZMUT.5646, http://mus.utu.fi/ZMUT.5648, http://mus.utu.fi/ZMUT.5650, http://mus.utu.fi/ZMUT.5653, http://mus.utu.fi/ZMUT.5655, http://mus.utu.fi/ZMUT.5657, http://mus.utu.fi/ZMUT.5662, http://mus.utu.fi/ZMUT.5668, http://mus.utu.fi/ZMUT.5669, http://mus.utu.fi/ZMUT.5670, http://mus.utu.fi/ZMUT.5673, http://mus.utu.fi/ZMUT.5678, http://mus.utu.fi/ZMUT.5681, http://mus.utu.fi/ZMUT.5683, http://mus.utu.fi/ZMUT.5684, http://mus.utu.fi/ZMUT.5687, http://mus.utu.fi/ZMUT.5688, http://mus.utu.fi/ZMUT.5690, http://mus.utu.fi/ZMUT.5692, http://mus.utu.fi/ZMUT.5693, http://mus.utu.fi/ZMUT.5694, http://mus.utu.fi/ZMUT.5698, http://mus.utu.fi/ZMUT.5699, http://mus.utu.fi/ZMUT.5702, http://mus.utu.fi/ZMUT.5708, http://mus.utu.fi/ZMUT.5709, http://mus.utu.fi/ZMUT.5719, http://mus.utu.fi/ZMUT.5722, http://mus.utu.fi/ZMUT.5725, http://mus.utu.fi/ZMUT.5726, http://mus.utu.fi/ZMUT.5733, http://mus.utu.fi/ZMUT.5734, http://mus.utu.fi/ZMUT.5740, http://mus.utu.fi/ZMUT.5748, http://mus.utu.fi/ZMUT.5750, http://mus.utu.fi/ZMUT.5754, http://mus.utu.fi/ZMUT.5755, http://mus.utu.fi/ZMUT.5757, http://mus.utu.fi/ZMUT.5762, http://mus.utu.fi/ZMUT.5763, http://mus.utu.fi/ZMUT.5764, http://mus.utu.fi/ZMUT.5772, http://mus.utu.fi/ZMUT.5774, http://mus.utu.fi/ZMUT.5775, http://mus.utu.fi/ZMUT.5778, http://mus.utu.fi/ZMUT.5780, http://mus.utu.fi/ZMUT.5794, http://mus.utu.fi/ZMUT.5797, http://mus.utu.fi/ZMUT.5806, http://mus.utu.fi/ZMUT.5808, http://mus.utu.fi/ZMUT.5811, http://mus.utu.fi/ZMUT.5816, http://mus.utu.fi/ZMUT.5817, http://mus.utu.fi/ZMUT.5821, http://mus.utu.fi/ZMUT.5823, http://mus.utu.fi/ZMUT.5824, http://mus.utu.fi/ZMUT.5825, http://mus.utu.fi/ZMUT.5829, http://mus.utu.fi/ZMUT.5834, http://mus.utu.fi/ZMUT.5845, http://mus.utu.fi/ZMUT.5846.</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Known material</emphasis>
: 167 specimens (160 Ugandan, 7 other):
</paragraph>
<paragraph pageId="0" pageNumber="33">
158 ♀, 2 ♂; Ugandan specimens, data above and also in supplementary material (
<bibRefCitation DOI="https://doi.org/10.5281/zenodo.2554877" author="Hopkins, T" journalOrPublisher="African Entomology" pageId="0" pageNumber="33" refId="B8" refString="Hopkins, T, Roininen, H, van Noort, S, Broad, GR, Kaunisto, K, Saeaeksjaervi, IE, 2019c. . https://doi.org/10.5281/zenodo.2554877" url="https://doi.org/10.5281/zenodo.2554877" year="2019 c">Hopkins et al. 2019c</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="33">
1 ♀, holotype of
<taxonomicName lsidName="E. gavinbroadi" pageId="0" pageNumber="33" rank="species" species="gavinbroadi">
<emphasis italics="true" pageId="0" pageNumber="33">E. gavinbroadi</emphasis>
</taxonomicName>
; see
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2014.91" author="Rousse, P" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="33" pagination="1 - 42" refId="B15" refString="Rousse, P, van Noort, S, 2014. A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91: 1 - 42" title="A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species." url="https://doi.org/10.5852/ejt.2014.91" volume="91" year="2014">Rousse and van Noort (2014)</bibRefCitation>
, data above in material examined.
</paragraph>
<paragraph pageId="0" pageNumber="33">
1 ♀, holotype of
<taxonomicName lsidName="E. uelensis" pageId="0" pageNumber="33" rank="species" species="uelensis">
<emphasis italics="true" pageId="0" pageNumber="33">E. uelensis</emphasis>
</taxonomicName>
; see
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2014.91" author="Rousse, P" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="33" pagination="1 - 42" refId="B15" refString="Rousse, P, van Noort, S, 2014. A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91: 1 - 42" title="A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species." url="https://doi.org/10.5852/ejt.2014.91" volume="91" year="2014">Rousse and van Noort (2014)</bibRefCitation>
; Democratic Republic of Congo, Haut-Uele, Paulis [Isiro,
<geoCoordinate degrees="03" direction="north" minutes="28" orientation="latitude" precision="925" value="3.4666667">03°28'N</geoCoordinate>
<geoCoordinate degrees="25" direction="east" minutes="43" orientation="longitude" precision="925" value="25.716667">25°43'E</geoCoordinate>
]; Dec. 1947; &quot;P.L.G. Benoit&quot;; MRAC.
</paragraph>
<paragraph pageId="0" pageNumber="33">
1 ♂, paratype of
<taxonomicName lsidName="E. uelensis" pageId="0" pageNumber="33" rank="species" species="uelensis">
<emphasis italics="true" pageId="0" pageNumber="33">E. uelensis</emphasis>
</taxonomicName>
; see
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2014.91" author="Rousse, P" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="33" pagination="1 - 42" refId="B15" refString="Rousse, P, van Noort, S, 2014. A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91: 1 - 42" title="A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species." url="https://doi.org/10.5852/ejt.2014.91" volume="91" year="2014">Rousse and van Noort (2014)</bibRefCitation>
; Democratic Republic of Congo, Bambesa; Jul. 1933; &quot;H.J. Bredo&quot;; MRAC.
</paragraph>
<paragraph pageId="0" pageNumber="33">
1 ♀; see
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2014.91" author="Rousse, P" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="33" pagination="1 - 42" refId="B15" refString="Rousse, P, van Noort, S, 2014. A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91: 1 - 42" title="A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species." url="https://doi.org/10.5852/ejt.2014.91" volume="91" year="2014">Rousse and van Noort (2014)</bibRefCitation>
; Cameroon, Nkoemvon; 30 Mar. 1980-19 Apr. 1980; &quot;Ms. D. Jackson&quot;; NHMUK.
</paragraph>
<paragraph pageId="0" pageNumber="33">
2 ♀; see
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2014.91" author="Rousse, P" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="33" pagination="1 - 42" refId="B15" refString="Rousse, P, van Noort, S, 2014. A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91: 1 - 42" title="A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species." url="https://doi.org/10.5852/ejt.2014.91" volume="91" year="2014">Rousse and van Noort (2014)</bibRefCitation>
, same data as previous except 13 Jul. 1980-4 Aug. 1980.
</paragraph>
<paragraph pageId="0" pageNumber="33">
1 ♀; see
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2014.91" author="Rousse, P" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="33" pagination="1 - 42" refId="B15" refString="Rousse, P, van Noort, S, 2014. A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91: 1 - 42" title="A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species." url="https://doi.org/10.5852/ejt.2014.91" volume="91" year="2014">Rousse and van Noort (2014)</bibRefCitation>
, same data as previous except Oct. 1980-Nov. 1980.
</paragraph>
<caption doi="10.3897/zookeys.878.37845.figures57-63" httpUri="https://binary.pensoft.net/fig/345029" pageId="0" pageNumber="33" start="Figures 5763" startId="F13">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figures 57-63.</emphasis>
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Epirhyssa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epirhyssa uelensis" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="uelensis">
<emphasis italics="true" pageId="0" pageNumber="33">Epirhyssa uelensis</emphasis>
</taxonomicName>
female (http://mus.utu.fi/ZMUT.2520), a species found in Uganda.
<emphasis bold="true" pageId="0" pageNumber="33">57</emphasis>
Habitus
<emphasis bold="true" pageId="0" pageNumber="33">58</emphasis>
face and clypeus
<emphasis bold="true" pageId="0" pageNumber="33">59</emphasis>
frons
<emphasis bold="true" pageId="0" pageNumber="33">60</emphasis>
hypostomal flange
<emphasis bold="true" pageId="0" pageNumber="33">61</emphasis>
mesopleuron dorsal margin
<emphasis bold="true" pageId="0" pageNumber="33">62</emphasis>
apical horn of metasoma
<emphasis bold="true" pageId="0" pageNumber="33">63</emphasis>
tergites 1-7. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="33">58-63</emphasis>
), 1 mm (
<emphasis bold="true" pageId="0" pageNumber="33">57</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="diagnosis">
<paragraph pageId="0" pageNumber="33">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="33">
This species can be distinguished from other Afrotropical
<taxonomicName lsidName="" pageId="0" pageNumber="33" rank="subfamily" subfamily="Rhyssinae">Rhyssinae</taxonomicName>
by its subalar prominence having a lateral flange.
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Epirhyssa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epirhyssa tombeaodiba" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="tombeaodiba">
<emphasis italics="true" pageId="0" pageNumber="33">Epirhyssa tombeaodiba</emphasis>
</taxonomicName>
is also predominantly yellow with black spots but lacks the flange.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Head</emphasis>
: frons with median carinae converging before continuing towards median ocellus, with lateral carinae curving towards lateral ocelli; hypostomal carina raised into an elevated flange, its height greater than the maximum width of the second maxillary palp segment.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Mesosoma</emphasis>
: subalar prominence with a lateral flange; mesopleuron with a raised flange along the dorsal margin; epicnemial carina reaches the approximate height of the mesopleural pit.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Metasoma</emphasis>
: tip of apical horn elliptical in posterior view; tergite 3 mostly smooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="description">
<paragraph pageId="0" pageNumber="33">Additional or updated characters.</paragraph>
<paragraph pageId="0" pageNumber="33">
Apart from the diagnosis, we provide the following additional or updated character traits to the description in
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2014.91" author="Rousse, P" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="33" pagination="1 - 42" refId="B15" refString="Rousse, P, van Noort, S, 2014. A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species. European Journal of Taxonomy 91: 1 - 42" title="A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species." url="https://doi.org/10.5852/ejt.2014.91" volume="91" year="2014">Rousse and van Noort (2014)</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Female.</emphasis>
Body length 9.8
<normalizedToken originalValue="mm">mm-</normalizedToken>
15.7 mm (15.7 mm
<taxonomicName lsidName="E. gavinbroadi" pageId="0" pageNumber="33" rank="species" species="gavinbroadi">
<emphasis italics="true" pageId="0" pageNumber="33">E. gavinbroadi</emphasis>
</taxonomicName>
holotype, 9.8
<normalizedToken originalValue="mm">mm-</normalizedToken>
15.1 mm other specimens). Frons with more or less distinct rugae parallel to the lateral carinae, often as distinct as the lateral carinae. Clypeus sparsely punctate or smooth. Antenna with 30-35+ flagellar segments (30-33 in Ugandan specimens, at least 35 in
<taxonomicName lsidName="E. gavinbroadi" pageId="0" pageNumber="33" rank="species" species="gavinbroadi">
<emphasis italics="true" pageId="0" pageNumber="33">E. gavinbroadi</emphasis>
</taxonomicName>
type, 30-34 in other specimens). Fore wing with 2m-cu varying from clearly distal to only just distal to rs-m. Tip of apical horn with an extra dorsal projection in one aberrant female (http://mus.utu.fi/ZMUT.5844). Tergites mostly smooth, except with varied amount of punctation on tergites 3-7 (anteriorly punctate 3-6 in many Ugandan specimens, punctate 3-6 in most non-Ugandan specimens, punctate apex of 5 and 6-7 in
<taxonomicName lsidName="E. gavinbroadi" pageId="0" pageNumber="33" rank="species" species="gavinbroadi">
<emphasis italics="true" pageId="0" pageNumber="33">E. gavinbroadi</emphasis>
</taxonomicName>
holotype), anterior margins of 3-5 or 3-6 often striate, tergite 1 1.2-1.7 times as long as apically wide (1.7 in
<taxonomicName lsidName="E. gavinbroadi" pageId="0" pageNumber="33" rank="species" species="gavinbroadi">
<emphasis italics="true" pageId="0" pageNumber="33">E. gavinbroadi</emphasis>
</taxonomicName>
holotype, 1.2-1.5 in other specimens). The Ugandan specimens have black posterior margins of tergites 2-7 (sometimes faint or patchy, especially on anterior tergites) and median stripes on 4-7 (strongest on 5-6). Non-Ugandan specimens sometimes have black median stripes on 5-7, or 3-6 in the
<taxonomicName lsidName="E. gavinbroadi" pageId="0" pageNumber="33" rank="species" species="gavinbroadi">
<emphasis italics="true" pageId="0" pageNumber="33">E. gavinbroadi</emphasis>
</taxonomicName>
holotype.
</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Male.</emphasis>
Body length 8.5
<normalizedToken originalValue="mm">mm-</normalizedToken>
8.7 mm. Antenna with 30 flagellar segments. Tergites 4-7 faintly pubescent, tergite anterior margins smooth, or lightly striate and punctate, tergite 1 1.5-2.0 times as long as apically wide. The two Ugandan males are smaller than females on average.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="distribution">
<paragraph pageId="0" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="33">Democratic Republic of Congo, Cameroon. New record: Uganda.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="biology_ecology">
<paragraph pageId="0" pageNumber="33">Biology.</paragraph>
<paragraph pageId="0" pageNumber="33">
In Uganda, this species was most abundantly caught in primary forest near decaying wood, during the dry season (
<bibRefCitation DOI="https://doi.org/10.1098/rsos.190913" author="Hopkins, T" journalOrPublisher="African Entomology" pageId="0" pageNumber="33" refId="B7" refString="Hopkins, T, Roininen, H, Saeaeksjaervi, IE, 2019b. . https://doi.org/10.1098/rsos.190913" url="https://doi.org/10.1098/rsos.190913" year="2019 b">Hopkins et al. 2019b</bibRefCitation>
). It has not been caught outside the forest.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="remarks">
<paragraph pageId="0" pageNumber="33">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="33">
This species is quite variable, but we are unable to find morphological characters that would reliably split it into more than one species. We propose that
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Epirhyssa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epirhyssa gavinbroadi" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="gavinbroadi">
<emphasis italics="true" pageId="0" pageNumber="33">Epirhyssa gavinbroadi</emphasis>
</taxonomicName>
(of which there is only one specimen) is a synonym of
<taxonomicName lsidName="E. uelensis" pageId="0" pageNumber="33" rank="species" species="uelensis">
<emphasis italics="true" pageId="0" pageNumber="33">E. uelensis</emphasis>
</taxonomicName>
. It was mainly distinguished from
<taxonomicName lsidName="E. uelensis" pageId="0" pageNumber="33" rank="species" species="uelensis">
<emphasis italics="true" pageId="0" pageNumber="33">E. uelensis</emphasis>
</taxonomicName>
by the punctate clypeus and slender tergite 1, but the Ugandan specimens generally have a punctate clypeus and a stout tergite 1. These two characters also vary considerably in the Ugandan specimens and seem to represent intraspecific variation. The Ugandan material has a very skewed sex ratio with 158 female specimens collected and only two males.
</paragraph>
<caption doi="10.3897/zookeys.878.37845.figures64-67" httpUri="https://binary.pensoft.net/fig/345030" pageId="0" pageNumber="33" start="Figures 6467" startId="F14">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figures 64-67.</emphasis>
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Epirhyssa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epirhyssa gavinbroadi" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="gavinbroadi">
<emphasis italics="true" pageId="0" pageNumber="33">Epirhyssa gavinbroadi</emphasis>
</taxonomicName>
female (holotype). We treat this species as a synonym of
<taxonomicName lsidName="E. uelensis" pageId="0" pageNumber="33" rank="species" species="uelensis">
<emphasis italics="true" pageId="0" pageNumber="33">E. uelensis</emphasis>
</taxonomicName>
in this work.
<emphasis bold="true" pageId="0" pageNumber="33">64</emphasis>
Habitus
<emphasis bold="true" pageId="0" pageNumber="33">65</emphasis>
hypostomal flange
<emphasis bold="true" pageId="0" pageNumber="33">66</emphasis>
mesopleuron dorsal margin
<emphasis bold="true" pageId="0" pageNumber="33">67</emphasis>
apical horn of metasoma. Figure
<emphasis bold="true" pageId="0" pageNumber="33">64</emphasis>
is from
<bibRefCitation author="van Noort, S" journalOrPublisher="California Academy of Sciences Memoir" pageId="0" pageNumber="33" publicationUrl="www.waspweb.org" refId="B22" refString="van Noort, S, 2019. . www.waspweb.org" url="www.waspweb.org" year="2019">van Noort (2019)</bibRefCitation>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>