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<document ID-DOI="http://doi.org/10.5281/zenodo.6268028" ID-GBIF-Dataset="94a0cedb-15bb-4bfd-955e-77200a536045" ID-GBIF-Taxon="120828889" ID-ISSN="1175-5334" ID-Zenodo-Dep="6268028" checkinTime="1466267330851" checkinUser="plazi" docAuthor="DAVID ADAMSKI" docDate="2005" docId="2E2A7A1DFF5EE0DFFED562CEFAD1FEF1" docLanguage="en" docName="zt00858a.pdf.imf" docOrigin="Zootaxa 858" docStyle="DocumentStyle{}" docTitle="Glyphidocera ranae ADAMSKI, 2005, new species" docType="treatment" docVersion="7" lastPageId="144" lastPageNumber="145" masterDocId="D2130265FFD0E04FFFDD6122FFC9FFC7" masterDocTitle="Review of Glyphidocera Walsingham of Costa Rica (Lepidoptera: Gelechioidea: Glyphidoceridae)" masterLastPageNumber="158" masterPageNumber="1" pageId="142" pageNumber="143" updateTime="1645749571040" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of Glyphidocera Walsingham of Costa Rica (Lepidoptera: Gelechioidea: Glyphidoceridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>DAVID ADAMSKI</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2005</mods:date>
<mods:detail type="volume">
<mods:number>858</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>158</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.170793</mods:identifier>
<mods:identifier type="GBIF-Dataset">94a0cedb-15bb-4bfd-955e-77200a536045</mods:identifier>
<mods:identifier type="ISSN">1175-5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">170793</mods:identifier>
<mods:identifier type="ZooBank">511FBF55-10F2-4A00-B15B-CBD34EAB023E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6268028" ID-GBIF-Taxon="120828889" ID-Zenodo-Dep="6268028" LSID="urn:lsid:plazi:treatment:2E2A7A1DFF5EE0DFFED562CEFAD1FEF1" httpUri="http://treatment.plazi.org/id/2E2A7A1DFF5EE0DFFED562CEFAD1FEF1" lastPageId="144" lastPageNumber="145" pageId="142" pageNumber="143">
<subSubSection pageId="142" pageNumber="143" type="nomenclature">
<paragraph blockId="142.[264,663,1004,1065]" box="[264,663,1004,1030]" pageId="142" pageNumber="143">
<heading bold="true" box="[264,663,1004,1030]" fontSize="11" level="4" pageId="142" pageNumber="143" reason="6">
<emphasis bold="true" box="[264,663,1004,1030]" pageId="142" pageNumber="143">
<taxonomicName box="[264,504,1004,1030]" class="Insecta" family="Symmocidae" genus="Glyphidocera" kingdom="Animalia" order="Lepidoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="species" species="ranae" status="new species">
<emphasis bold="true" box="[264,504,1004,1030]" italics="true" pageId="142" pageNumber="143">Glyphidocera ranae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[519,663,1004,1030]" pageId="142" pageNumber="143" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="142.[264,663,1004,1065]" box="[264,613,1039,1065]" pageId="142" pageNumber="143">
(Figs.
<number box="[339,501,1039,1065]" pageId="142" pageNumber="143" value="127311.5" valueMax="254371.0" valueMin="252.0">252–254, 371</number>
,
<figureCitation box="[513,605,1039,1065]" captionStart="MAP 46" captionStartId="143.[264,325,1824,1848]" captionTargetBox="[556,1031,1358,1806]" captionTargetId="figure@143.[553,1033,1353,1810]" captionTargetPageId="143" captionText="MAP 46. Distribution of Glyphidocera ranae () and G. sollertiae (). White triangle within a black circle indicates a shared locality." httpUri="https://zenodo.org/record/170835/files/figure.png" pageId="142" pageNumber="143">
Map
<number box="[574,605,1039,1065]" pageId="142" pageNumber="143" value="46.0">46</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="144" lastPageNumber="145" pageId="142" pageNumber="143" type="diagnosis">
<paragraph blockId="142.[264,1323,1119,1870]" pageId="142" pageNumber="143">
Diagnosis.—
<taxonomicName box="[426,666,1119,1144]" class="Insecta" family="Symmocidae" genus="Glyphidocera" kingdom="Animalia" order="Lepidoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="species" species="ranae">
<emphasis box="[426,666,1119,1144]" italics="true" pageId="142" pageNumber="143">Glyphidocera ranae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName box="[900,1031,1119,1144]" class="Insecta" family="Symmocidae" genus="Glyphidocera" kingdom="Animalia" order="Lepidoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="species" species="ancillae">
<emphasis box="[900,1031,1119,1144]" italics="true" pageId="142" pageNumber="143">G. ancillae</emphasis>
</taxonomicName>
; it can be distinguished from the latter by having a larger gnathos, a larger digitate process of the valva, a slightly recurved dorsal furca of the valval costa, and the vesica of aedeagus with an apical cornutus with a bulbous base.
</paragraph>
<paragraph blockId="142.[264,1323,1119,1870]" pageId="142" pageNumber="143">
Description.— Head: Vertex and frontoclypeus brown; outer surface of labial palpus brown intermixed with few pale­brown scales to apical margin of segment II, inner surface pale brown intermixed with few brown scales; scape of antenna brown, flagellum with alternating bands of pale brown and brown scales; male
<number box="[921,936,1402,1428]" pageId="142" pageNumber="143" value="4.0">4</number>
th flagellomere dorsally protuberant and curved, forming a deep concavity opposite slightly excavated flagellomeres
<number box="[1272,1317,1442,1468]" pageId="142" pageNumber="143" value="5.5" valueMax="6.0" valueMin="5.0">5–6</number>
; concavity nearly closed by a digitate protuberance of
<number box="[937,951,1484,1510]" pageId="142" pageNumber="143" value="4.0">4</number>
th flagellomere; proboscis pale brown.
</paragraph>
<paragraph blockId="142.[264,1323,1119,1870]" pageId="142" pageNumber="143">
Thorax: Tegula and mesonotum brown. Legs brown intermixed with pale­brown scales to apical margins of all segments and tarsomeres. Forewing (Fig.
<number box="[1111,1160,1604,1630]" pageId="142" pageNumber="143" value="371.0">371</number>
): Length
<quantity metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.2" metricValueMin="5.9" pageId="142" pageNumber="143" unit="mm" value="6.05" valueMax="6.2" valueMin="5.9">5.9– 6.2 mm</quantity>
(n =
<number box="[427,444,1644,1670]" pageId="142" pageNumber="143" value="7.0">7</number>
), pale brown intermixed with pale grayish­orange, brown, and few darkbrown scales; cell with
<number box="[549,563,1684,1710]" pageId="142" pageNumber="143" value="2.0">2</number>
brown spots,
<number box="[734,748,1684,1710]" pageId="142" pageNumber="143" value="1.0">1</number>
large spot near middle,
<number box="[1042,1057,1684,1710]" pageId="142" pageNumber="143" value="1.0">1</number>
small spot near distal end [spots are faint in some specimens]. Undersurface brown. Hindwing: Brown.
</paragraph>
<paragraph blockId="142.[264,1323,1119,1870]" pageId="142" pageNumber="143">
Abdomen (Fig.
<number box="[506,555,1764,1790]" pageId="142" pageNumber="143" value="252.0">252</number>
): Male with
<number box="[715,730,1764,1790]" pageId="142" pageNumber="143" value="3.0">3</number>
transverse rows of sex scales on intersegmental membrane between terga
<number box="[565,610,1804,1830]" pageId="142" pageNumber="143" value="2.5" valueMax="3.0" valueMin="2.0">2–3</number>
.
</paragraph>
<paragraph blockId="142.[264,1323,1119,1870]" lastBlockId="143.[264,1323,284,1310]" lastPageId="143" lastPageNumber="144" pageId="142" pageNumber="143">
Male Genitalia (Figs.
<number box="[574,680,1844,1870]" pageId="142" pageNumber="143" value="253.5" valueMax="254.0" valueMin="253.0">253–254</number>
): Uncus elongate, wide at base, broadly constricted at
<number box="[264,275,284,310]" pageId="143" pageNumber="144" value="1.0">1</number>
/
<number box="[287,303,284,310]" pageId="143" pageNumber="144" value="3.0">3</number>
, slightly widened apically, apex narrowly rounded and setose; gnathos sparsely setose, protuberant, wide throughout length, upturned apically; tegumen slightly widened basally; dorsal strut of tegumen with arms convergent, fusing from a widened base from posteroventral margin to base of uncus; costa of valva slightly sloped dorsally from shallow depression beyond a slanted, basal, digitate process to an upturned dorsal furca; apex widely bifurcate from a narrowly rounded inner margin; dorsal furca about
<number box="[1160,1175,484,510]" pageId="143" pageNumber="144" value="3.0">3</number>
times larger than ventral furca; sacculus twisted apically constricting valva near midlength, widening distoventrally forming an elongate and broadly rounded cucullus; cucullus membranous and setose; vesica of aedeagus denticulate, bearing a large, conical, apical cornutus; curnutus with a bulbous base, with one side slightly shorter, exposing a hollow, basal cavity.
</paragraph>
<paragraph blockId="143.[264,1323,284,1310]" box="[312,652,684,710]" pageId="143" pageNumber="144">Female Genitalia: Unknown.</paragraph>
<paragraph blockId="143.[264,1323,284,1310]" pageId="143" pageNumber="144">
<typeStatus box="[312,424,724,750]" pageId="143" pageNumber="144">Holotype</typeStatus>
, ɗ, “Est[ación] Pitilla,
<quantity box="[741,830,724,750]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="143" pageNumber="144" unit="m" value="700.0">700 m</quantity>
,
<quantity box="[853,924,724,750]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="143" pageNumber="144" unit="km" value="9.0">9 km</quantity>
S S[an]ta Cecilia, Prov[incia] Guan[acaste],
<collectingCountry box="[437,615,764,790]" name="Costa Rica" pageId="143" pageNumber="144">
<collectionCode box="[437,531,764,790]" pageId="143" pageNumber="144">COSTA</collectionCode>
<collectionCode box="[542,615,764,790]" pageId="143" pageNumber="144">RICA</collectionCode>
</collectingCountry>
, C. Moraga &amp; P. Rios,
<date box="[910,1032,764,790]" pageId="143" pageNumber="144" value="1990-12">
Dic.
<number box="[971,1032,764,790]" pageId="143" pageNumber="144" value="1990.0">1990</number>
</date>
, L­N­
<number box="[1108,1198,764,790]" pageId="143" pageNumber="144" value="330200.0">330200</number>
,
<number box="[1214,1307,764,790]" pageId="143" pageNumber="144" value="380200.0">380200</number>
”, “
<collectionCode box="[275,355,804,830]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="143" pageNumber="144">INBio</collectionCode>
:
<collectingCountry box="[367,545,804,830]" name="Costa Rica" pageId="143" pageNumber="144">
<collectionCode box="[367,461,804,830]" pageId="143" pageNumber="144">COSTA</collectionCode>
<collectionCode box="[470,545,804,830]" pageId="143" pageNumber="144">RICA</collectionCode>
</collectingCountry>
:
<collectionCode box="[556,606,804,830]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universit<69>rio" pageId="143" pageNumber="144">CRI</collectionCode>
000,
<number box="[665,756,804,830]" pageId="143" pageNumber="144" value="652704.0">652704</number>
” [bar code label], “
<collectionCode box="[988,1066,804,830]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="143" pageNumber="144">INBio</collectionCode>
, ɗ Genitalia Slide by D. Adamski, No.
<number box="[471,518,844,870]" pageId="143" pageNumber="144" value="776.0">776</number>
” [yellow label].
</paragraph>
<paragraph blockId="143.[264,1323,284,1310]" pageId="143" pageNumber="144">
<typeStatus box="[312,426,884,910]" pageId="143" pageNumber="144">Paratypes</typeStatus>
(
<number box="[446,461,884,910]" pageId="143" pageNumber="144" value="6.0">6</number>
ɗ):
<number box="[523,537,884,910]" pageId="143" pageNumber="144" value="3.0">3</number>
ɗ, Same label data as
<typeStatus box="[831,933,884,910]" pageId="143" pageNumber="144">holotype</typeStatus>
except, “Set.,
<number box="[1119,1182,884,910]" pageId="143" pageNumber="144" value="1991.0">1991</number>
”, “
<collectionCode box="[1220,1272,884,910]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universit<69>rio" pageId="143" pageNumber="144">CRI</collectionCode>
000,
<number box="[264,357,924,950]" pageId="143" pageNumber="144" value="460293.0">460293</number>
”, “Slide No.
<number box="[519,568,924,950]" pageId="143" pageNumber="144" value="775.0">775</number>
”; “
<collectionCode box="[604,656,924,950]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universit<69>rio" pageId="143" pageNumber="144">CRI</collectionCode>
000,
<number box="[717,810,924,950]" pageId="143" pageNumber="144" value="580151.0">580151</number>
”, “Slide No.
<number box="[972,1021,924,950]" pageId="143" pageNumber="144" value="765.0">765</number>
”; “
<date box="[1058,1183,924,950]" pageId="143" pageNumber="144" value="1990-10">
Oct.
<number box="[1120,1183,924,950]" pageId="143" pageNumber="144" value="1990.0">1990</number>
</date>
”, “
<collectionCode box="[1219,1271,924,950]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universit<69>rio" pageId="143" pageNumber="144">CRI</collectionCode>
000,
<number box="[264,357,964,990]" pageId="143" pageNumber="144" value="380795.0">380795</number>
”, “Slide No.
<number box="[529,578,964,990]" pageId="143" pageNumber="144" value="768.0">768</number>
”;
<number box="[607,621,964,990]" pageId="143" pageNumber="144" value="2.0">2</number>
ɗ, Cerro Tortuguero,
<quantity box="[906,1017,964,990]" metricMagnitude="1" metricUnit="m" metricValue="6.0" metricValueMax="12.0" metricValueMin="0.0" pageId="143" pageNumber="144" unit="m" value="60.0" valueMax="120.0" valueMin="0.0">0–120 m</quantity>
, P. N. Tortuguero, Prov. Limón,
<collectingCountry box="[368,554,1004,1030]" name="Costa Rica" pageId="143" pageNumber="144">
<collectionCode box="[368,462,1004,1030]" pageId="143" pageNumber="144">COSTA</collectionCode>
<collectionCode box="[481,554,1004,1030]" pageId="143" pageNumber="144">RICA</collectionCode>
</collectingCountry>
,
<date box="[578,714,1004,1030]" pageId="143" pageNumber="144" value="1992-11">
Nov.
<number box="[653,714,1004,1030]" pageId="143" pageNumber="144" value="1992.0">1992</number>
</date>
, R. Delgado, L­N­
<number box="[968,1058,1004,1030]" pageId="143" pageNumber="144" value="285000.0">285000</number>
,
<number box="[1082,1175,1004,1030]" pageId="143" pageNumber="144" value="588000.0">588000</number>
”, “
<collectionCode box="[1219,1271,1004,1030]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universit<69>rio" pageId="143" pageNumber="144">CRI</collectionCode>
000,
<number box="[264,357,1044,1070]" pageId="143" pageNumber="144" value="992205.0">992205</number>
”, “Slide No.
<number box="[516,565,1044,1070]" pageId="143" pageNumber="144" value="588.0">588</number>
”; “
<date box="[601,730,1044,1070]" pageId="143" pageNumber="144" value="1993-03">
Mar.
<number box="[667,730,1044,1070]" pageId="143" pageNumber="144" value="1993.0">1993</number>
</date>
”, “
<collectionCode box="[766,819,1044,1070]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universit<69>rio" pageId="143" pageNumber="144">CRI</collectionCode>
001,
<number box="[878,972,1044,1070]" pageId="143" pageNumber="144" value="353218.0">353218</number>
”; “Slide No.
<number box="[1130,1179,1044,1070]" pageId="143" pageNumber="144" value="652.0">652</number>
”;
<number box="[1205,1220,1044,1070]" pageId="143" pageNumber="144" value="1.0">1</number>
ɗ, “Est. Sirena, P. N. Corcovado,
<quantity box="[564,670,1084,1110]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="143" pageNumber="144" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100 m</quantity>
, Prov. Punt.,
<collectingCountry box="[829,1005,1084,1110]" name="Costa Rica" pageId="143" pageNumber="144">
<collectionCode box="[829,923,1084,1110]" pageId="143" pageNumber="144">COSTA</collectionCode>
<collectionCode box="[932,1005,1084,1110]" pageId="143" pageNumber="144">RICA</collectionCode>
</collectingCountry>
, G. Fonseca,
<date box="[1162,1281,1084,1110]" pageId="143" pageNumber="144" value="1990-10">
Oct.
<number box="[1220,1281,1084,1110]" pageId="143" pageNumber="144" value="1990.0">1990</number>
</date>
, L­ S­
<date box="[289,381,1124,1150]" pageId="143" pageNumber="144" value="00-05-27">
<number box="[289,381,1124,1150]" pageId="143" pageNumber="144" value="270500.0">270500</number>
</date>
,
<number box="[392,486,1124,1150]" pageId="143" pageNumber="144" value="508300.0">508300</number>
”, “
<collectionCode box="[518,570,1124,1150]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universit<69>rio" pageId="143" pageNumber="144">CRI</collectionCode>
000,
<number box="[628,721,1124,1150]" pageId="143" pageNumber="144" value="179053.0">179053</number>
”, “Slide No.
<number box="[875,923,1124,1150]" pageId="143" pageNumber="144" value="805.0">805</number>
”, (
<quantity box="[956,1001,1124,1150]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="143" pageNumber="144" unit="in" value="3.0">3 in</quantity>
<collectionCode box="[1008,1084,1124,1150]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="143" pageNumber="144">INBio</collectionCode>
,
<quantity box="[1096,1141,1124,1150]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="143" pageNumber="144" unit="in" value="3.0">3 in</quantity>
<collectionCode box="[1148,1238,1124,1150]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="143" pageNumber="144">USNM</collectionCode>
).
</paragraph>
<paragraph blockId="143.[264,1323,284,1310]" pageId="143" pageNumber="144">
Distribution (
<figureCitation box="[472,573,1164,1190]" captionStart="MAP 46" captionStartId="143.[264,325,1824,1848]" captionTargetBox="[556,1031,1358,1806]" captionTargetId="figure@143.[553,1033,1353,1810]" captionTargetPageId="143" captionText="MAP 46. Distribution of Glyphidocera ranae () and G. sollertiae (). White triangle within a black circle indicates a shared locality." httpUri="https://zenodo.org/record/170835/files/figure.png" pageId="143" pageNumber="144">
Map
<number box="[539,573,1164,1190]" pageId="143" pageNumber="144" value="46.0">46</number>
</figureCitation>
):
<taxonomicName box="[598,838,1164,1189]" class="Insecta" family="Symmocidae" genus="Glyphidocera" kingdom="Animalia" order="Lepidoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="ranae">
<emphasis box="[598,838,1164,1189]" italics="true" pageId="143" pageNumber="144">Glyphidocera ranae</emphasis>
</taxonomicName>
is known from three collecting sites in
<collectingCountry box="[264,395,1204,1230]" name="Costa Rica" pageId="143" pageNumber="144">Costa Rica</collectingCountry>
, from the north­western part along the Cordillera Guanacaste, east to the northcentral part of the country along the Caribbean Coast, south­east to a coastal site along the Pacific Ocean.
</paragraph>
<caption httpUri="https://zenodo.org/record/170835/files/figure.png" pageId="143" pageNumber="144" targetBox="[556,1031,1358,1806]" targetPageId="143">
<paragraph blockId="143.[264,1322,1824,1885]" pageId="143" pageNumber="144">
<emphasis bold="true" box="[264,368,1824,1848]" pageId="143" pageNumber="144">MAP 46.</emphasis>
Distribution of
<taxonomicName box="[548,766,1825,1848]" class="Insecta" family="Symmocidae" genus="Glyphidocera" kingdom="Animalia" order="Lepidoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="ranae">
<emphasis box="[548,766,1825,1848]" italics="true" pageId="143" pageNumber="144">Glyphidocera ranae</emphasis>
</taxonomicName>
() and
<taxonomicName box="[869,1002,1825,1848]" class="Insecta" family="Symmocidae" genus="Glyphidocera" kingdom="Animalia" order="Lepidoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="sollertiae">
<emphasis box="[869,1002,1825,1848]" italics="true" pageId="143" pageNumber="144">G. sollertiae</emphasis>
</taxonomicName>
(). White triangle within a black circle indicates a shared locality.
</paragraph>
</caption>
<paragraph blockId="144.[312,1304,284,310]" box="[312,1304,284,310]" pageId="144" pageNumber="145">
Etymology: The species epithet
<taxonomicName box="[689,758,284,309]" class="Insecta" family="Symmocidae" genus="Glyphidocera" kingdom="Animalia" order="Lepidoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="ranae">
<emphasis box="[689,758,284,309]" italics="true" pageId="144" pageNumber="145">ranae</emphasis>
</taxonomicName>
is derived from the Latin word for frog ­ rana.
</paragraph>
</subSubSection>
</treatment>
</document>
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