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<mods:title id="E2E079FEEA6BB41D9F5696810DEAFE14">The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea)</mods:title>
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<paragraph id="9B0B36F9FF8BFFBAFFDC5DB1FDB0FD54" blockId="34.[151,591,687,714]" box="[151,591,687,714]" pageId="34" pageNumber="35">
<heading id="C0438195FF8BFFBAFFDC5DB1FDB0FD54" bold="true" box="[151,591,687,714]" fontSize="11" level="2" pageId="34" pageNumber="35" reason="4">
<taxonomicName id="5CB44D7AFF8BFFBAFFDC5DB1FDB0FD54" ID-CoL="7NR8C" authority="Loomis &amp; Schmitt, 1971" authorityName="Loomis &amp; Schmitt" authorityYear="1971" box="[151,591,687,714]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="A9C0EAEBFF8BFFBAFFDC5DB1FDB0FD54" bold="true" box="[151,591,687,714]" pageId="34" pageNumber="35">
<emphasis id="A9C0EAEBFF8BFFBAFFDC5DB1FEE7FD54" bold="true" box="[151,280,688,714]" italics="true" pageId="34" pageNumber="35">Brunsonia</emphasis>
<bibRefCitation id="FF254B08FF8BFFBAFE545DAEFDB0FD54" author="Loomis, H. F. &amp; Schmitt, R." box="[287,591,687,714]" pageId="34" pageNumber="35" pagination="107 - 131" refId="ref43738" refString="Loomis, H. F. &amp; Schmitt, R. (1971) The ecology, distribution and taxonomy of the millipeds of Montana west of the continental divide. Northwest Science, 45, 107 - 131." type="journal article" year="1971">Loomis &amp; Schmitt, 1971</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="9B0B36F9FF8BFFBAFFDC5DEEFCA2FCBB" blockId="34.[151,1436,750,836]" pageId="34" pageNumber="35">
<taxonomicName id="5CB44D7AFF8BFFBAFFDC5DEEFDCAFC99" ID-CoL="7NR8C" authority="Loomis &amp; Schmitt, 1971: 122" authorityName="Loomis &amp; Schmitt" authorityPageNumber="122" authorityYear="1971" box="[151,565,750,775]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="A9C0EAEBFF8BFFBAFFDC5DEEFEFEFC98" box="[151,257,751,774]" italics="true" pageId="34" pageNumber="35">Brunsonia</emphasis>
<bibRefCitation id="FF254B08FF8BFFBAFE435DEFFDCAFC99" author="Loomis, H. F. &amp; Schmitt, R." box="[264,565,750,775]" pageId="34" pageNumber="35" pagination="107 - 131" refId="ref43738" refString="Loomis, H. F. &amp; Schmitt, R. (1971) The ecology, distribution and taxonomy of the millipeds of Montana west of the continental divide. Northwest Science, 45, 107 - 131." type="journal article" year="1971">Loomis &amp; Schmitt, 1971:122</bibRefCitation>
</taxonomicName>
.
<typeStatus id="440F885BFF8BFFBAFD745DEEFD8EFC99" box="[575,625,751,775]" pageId="34" pageNumber="35">Type</typeStatus>
species
<taxonomicName id="5CB44D7AFF8BFFBAFD835DEEFB93FC99" ID-CoL="NFZJ" authority="(Chamberlin, 1920)" baseAuthorityName="Chamberlin" baseAuthorityYear="1920" box="[712,1132,750,775]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="albertana">
<emphasis id="A9C0EAEBFF8BFFBAFD835DEEFC64FC99" box="[712,923,751,775]" italics="true" pageId="34" pageNumber="35">Brunsonia albertana</emphasis>
(
<bibRefCitation id="FF254B08FF8BFFBAFCE35DEFFB9AFC99" author="Chamberlin, R. V." box="[936,1125,750,775]" pageId="34" pageNumber="35" pagination="94 - 95" refId="ref43219" refString="Chamberlin, R. V. (1920) Some records of Canadian myriopods. The Canadian Entomologist, 52 (4), 94 - 95." type="journal article" year="1920">Chamberlin, 1920</bibRefCitation>
)
</taxonomicName>
, by original designation of its junior synonym
<taxonomicName id="5CB44D7AFF8BFFBAFE265C0FFCA5FCBB" ID-CoL="NFZL" authority="Loomis &amp; Schmitt 1971" authorityName="Loomis &amp; Schmitt" authorityYear="1971" box="[365,858,781,806]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="complexipes">
<emphasis id="A9C0EAEBFF8BFFBAFE265C0FFDA5FCBB" box="[365,602,781,805]" italics="true" pageId="34" pageNumber="35">Brunsonia complexipes</emphasis>
<bibRefCitation id="FF254B08FF8BFFBAFD2A5C0CFCA5FCBB" author="Loomis, H. F. &amp; Schmitt, R." box="[609,858,781,806]" pageId="34" pageNumber="35" pagination="107 - 131" refId="ref43738" refString="Loomis, H. F. &amp; Schmitt, R. (1971) The ecology, distribution and taxonomy of the millipeds of Montana west of the continental divide. Northwest Science, 45, 107 - 131." type="journal article" year="1971">Loomis &amp; Schmitt 1971</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="9B0B36F9FF8BFFBAFFDC5C2DFBB4FCDA" blockId="34.[151,1436,750,836]" box="[151,1099,811,836]" pageId="34" pageNumber="35">
<taxonomicName id="5CB44D7AFF8BFFBAFFDC5C2DFE86FCDA" ID-CoL="7NTGJ" authority=", Shear, 1971" authorityName=", Shear" authorityYear="1971" box="[151,377,811,836]" class="Diplopoda" family="Conotylidae" genus="Conotyla" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="A9C0EAEBFF8BFFBAFFDC5C2DFF04FCDA" box="[151,251,812,836]" italics="true" pageId="34" pageNumber="35">Conotyla,</emphasis>
<bibRefCitation id="FF254B08FF8BFFBAFE4A5C2AFE86FCDA" author="Shear, W. A." box="[257,377,811,836]" pageId="34" pageNumber="35" pagination="55 - 96" refId="ref44018" refString="Shear, W. A. (1971) The millipede family Conotylidae in North America, with a description of the new family Adritylidae (Diplopoda: Chordeumida). Bulletin of the Museum of Comparative Zoology, 141, 55 - 96." type="journal article" year="1971">Shear, 1971</bibRefCitation>
</taxonomicName>
(“Atrolineata group,” not
<taxonomicName id="5CB44D7AFF8BFFBAFDCE5C2DFC39FCDA" ID-CoL="7NTGJ" authority="Cook &amp; Collins 1895" authorityName="Cook &amp; Collins" authorityYear="1895" box="[645,966,811,836]" class="Diplopoda" family="Conotylidae" genus="Conotyla" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="A9C0EAEBFF8BFFBAFDCE5C2DFD1DFCDA" box="[645,738,812,836]" italics="true" pageId="34" pageNumber="35">Conotyla</emphasis>
<bibRefCitation id="FF254B08FF8BFFBAFDA35C2AFC39FCDA" author="Cook, O. F. &amp; Collins, G." box="[744,966,811,836]" pageId="34" pageNumber="35" pagination="1 - 100" refId="ref43361" refString="Cook, O. F. &amp; Collins, G. (1895) The Craspedosomatidae of North America. Annals of the New York Academy of Sciences, 9, 1 - 100. https: // doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55430. x" type="journal article" year="1895">Cook &amp; Collins 1895</bibRefCitation>
</taxonomicName>
): 80; 1976:5.
</paragraph>
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<subSubSection id="D3AE6572FF8BFFBAFFDC5C70FD42F8E9" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="9B0B36F9FF8BFFBAFFDC5C70FB14FC2E" blockId="34.[151,1437,881,1809]" pageId="34" pageNumber="35">
<emphasis id="A9C0EAEBFF8BFFBAFFDC5C70FEF5FC12" bold="true" box="[151,266,881,908]" pageId="34" pageNumber="35">Diagnosis</emphasis>
: Moderate-sized conotylids (
<quantity id="5C4C9B1CFF8BFFBAFD1A5C70FD34FC13" box="[593,715,881,909]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="34" pageNumber="35" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
long) distinct from other genera of the subfamily
<taxonomicName id="5CB44D7AFF8BFFBAFBBC5C73FA7FFC12" authorityName="Cook" authorityYear="1896" box="[1271,1408,882,908]" class="Diplopoda" family="Conotylidae" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subFamily" subFamily="Conotylinae">Conotylinae</taxonomicName>
in the mostly reduced anterior gonopods coupled with the complex coxites of the posterior gonopods.
</paragraph>
<paragraph id="9B0B36F9FF8BFFBAFF8C5CBBFADEFB4F" blockId="34.[151,1437,881,1809]" pageId="34" pageNumber="35">
<emphasis id="A9C0EAEBFF8BFFBAFF8C5CBBFEECFC4A" bold="true" box="[199,275,954,980]" pageId="34" pageNumber="35">Notes:</emphasis>
The genus includes the previously described species
<taxonomicName id="5CB44D7AFF8BFFBAFC335CBAFAB5FC4A" authority="(Chamberlin, 1920)" baseAuthorityName="Chamberlin" baseAuthorityYear="1920" box="[888,1354,954,980]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="albertana">
<emphasis id="A9C0EAEBFF8BFFBAFC335CBAFB9CFC4A" box="[888,1123,954,980]" italics="true" pageId="34" pageNumber="35">Brunsonia albertana</emphasis>
(
<bibRefCitation id="FF254B08FF8BFFBAFB385CBBFABDFC4A" author="Chamberlin, R. V." box="[1139,1346,954,980]" pageId="34" pageNumber="35" pagination="94 - 95" refId="ref43219" refString="Chamberlin, R. V. (1920) Some records of Canadian myriopods. The Canadian Entomologist, 52 (4), 94 - 95." type="journal article" year="1920">Chamberlin, 1920</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="5CB44D7AFF8BFFBAFACF5CBAFE27FC66" authority="(Bollman, 1893)" baseAuthorityName="Bollman" baseAuthorityYear="1893" class="Diplopoda" family="Conotylidae" genus="Conotyla" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="atrolineata">
<emphasis id="A9C0EAEBFF8BFFBAFACF5CBAFEEDFC66" italics="true" pageId="34" pageNumber="35">B. atrolineata</emphasis>
(
<bibRefCitation id="FF254B08FF8BFFBAFE6F5CDFFE2FFC66" author="Bollman, C. H." box="[292,464,990,1016]" pageId="34" pageNumber="35" pagination="1 - 210" refId="ref43018" refString="Bollman, C. H. (1893) The Myriapoda of the United States. United States National Museum Bulletin, 43, 1 - 210." type="journal article" year="1893">Bollman, 1893</bibRefCitation>
)
</taxonomicName>
. The latter species is known from western
<collectingRegion id="5970F81BFF8BFFBAFC855CDFFB6AFC66" box="[974,1173,990,1016]" country="Canada" name="British Columbia" pageId="34" pageNumber="35">British Columbia</collectingRegion>
,
<collectingCountry id="E3A37669FF8BFFBAFBE85CDFFB04FC66" box="[1187,1275,990,1016]" name="Canada" pageId="34" pageNumber="35">Canada</collectingCountry>
;
<bibRefCitation id="FF254B08FF8BFFBAFA435CDFFA63FC66" author="Shear, W. A." box="[1288,1436,990,1016]" pageId="34" pageNumber="35" pagination="55 - 96" refId="ref44018" refString="Shear, W. A. (1971) The millipede family Conotylidae in North America, with a description of the new family Adritylidae (Diplopoda: Chordeumida). Bulletin of the Museum of Comparative Zoology, 141, 55 - 96." type="journal article" year="1971">Shear (1971)</bibRefCitation>
failed to give specific collection data but mentioned Yoho, Glacier, and Mt. Robson National Parks. We have no new records of this species.
<taxonomicName id="5CB44D7AFF8BFFBAFE855B26FD4AFBDE" baseAuthorityName="Chamberlin" baseAuthorityYear="1920" box="[462,693,1062,1088]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="albertana">
<emphasis id="A9C0EAEBFF8BFFBAFE855B26FD4AFBDE" box="[462,693,1062,1088]" italics="true" pageId="34" pageNumber="35">Brunsonia albertana</emphasis>
</taxonomicName>
is a senior synonym of the designated
<typeStatus id="440F885BFF8BFFBAFB2B5B26FB6FFBDF" box="[1120,1168,1063,1089]" pageId="34" pageNumber="35">type</typeStatus>
species,
<taxonomicName id="5CB44D7AFF8BFFBAFBBF5B26FED3FBFA" authority="(Shear 1971)" baseAuthorityName="Shear" baseAuthorityYear="1971" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="complexipes">
<emphasis id="A9C0EAEBFF8BFFBAFBBF5B26FA63FBDE" box="[1268,1436,1062,1088]" italics="true" pageId="34" pageNumber="35">B. complexipes</emphasis>
(
<bibRefCitation id="FF254B08FF8BFFBAFFD45B4BFEDBFBFA" author="Shear, W. A." box="[159,292,1098,1124]" pageId="34" pageNumber="35" pagination="55 - 96" refId="ref44018" refString="Shear, W. A. (1971) The millipede family Conotylidae in North America, with a description of the new family Adritylidae (Diplopoda: Chordeumida). Bulletin of the Museum of Comparative Zoology, 141, 55 - 96." type="journal article" year="1971">Shear 1971</bibRefCitation>
)
</taxonomicName>
, and occurs in southern
<collectingRegion id="5970F81BFF8BFFBAFD0A5B4BFD6AFBFA" box="[577,661,1098,1124]" country="Canada" name="Alberta" pageId="34" pageNumber="35">Alberta</collectingRegion>
and northwestern
<collectingRegion id="5970F81BFF8BFFBAFC225B4AFC32FBFB" box="[873,973,1099,1125]" country="United States of America" name="Montana" pageId="34" pageNumber="35">Montana</collectingRegion>
(
<bibRefCitation id="FF254B08FF8BFFBAFC975B4BFB09FBFA" author="Loomis, H. F. &amp; Schmitt, R." box="[988,1270,1098,1124]" pageId="34" pageNumber="35" pagination="107 - 131" refId="ref43738" refString="Loomis, H. F. &amp; Schmitt, R. (1971) The ecology, distribution and taxonomy of the millipeds of Montana west of the continental divide. Northwest Science, 45, 107 - 131." type="journal article" year="1971">Loomis &amp; Schmitt 1971</bibRefCitation>
). Again,
<bibRefCitation id="FF254B08FF8BFFBAFA175B4BFF1DFB16" author="Shear, W. A." pageId="34" pageNumber="35" pagination="55 - 96" refId="ref44018" refString="Shear, W. A. (1971) The millipede family Conotylidae in North America, with a description of the new family Adritylidae (Diplopoda: Chordeumida). Bulletin of the Museum of Comparative Zoology, 141, 55 - 96." type="journal article" year="1971">Shear (1971)</bibRefCitation>
does not provide detailed collection records but mentions Banff and Jasper National Parks. Records given below extend the distribution to western
<collectingRegion id="5970F81BFF8BFFBAFD275B93FD0EFB32" box="[620,753,1170,1196]" country="United States of America" name="Washington" pageId="34" pageNumber="35">Washington</collectingRegion>
and
<collectingRegion id="5970F81BFF8BFFBAFC675B93FC7FFB32" box="[812,896,1170,1196]" country="United States of America" name="Oregon" pageId="34" pageNumber="35">Oregon</collectingRegion>
in the
<collectingCountry id="E3A37669FF8BFFBAFC985B93FB91FB32" box="[979,1134,1170,1196]" name="United States of America" pageId="34" pageNumber="35">United States</collectingCountry>
. The four new species described herein are all from the
<collectingCountry id="E3A37669FF8BFFBAFEA45BB7FD7AFB4E" box="[495,645,1206,1232]" name="United States of America" pageId="34" pageNumber="35">United States</collectingCountry>
and represent considerable extensions of the generic range.
</paragraph>
<paragraph id="9B0B36F9FF8BFFBAFF8C5BDBFC8DF98A" blockId="34.[151,1437,881,1809]" pageId="34" pageNumber="35">
The gonopods of
<taxonomicName id="5CB44D7AFF8BFFBAFEC55BDAFDFCFB6A" authorityName="Loomis &amp; Schmitt" authorityYear="1971" box="[398,515,1243,1268]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="A9C0EAEBFF8BFFBAFEC55BDAFDFCFB6A" box="[398,515,1243,1268]" italics="true" pageId="34" pageNumber="35">Brunsonia</emphasis>
</taxonomicName>
species show variations on a common theme. The anterior gonopods are small in some species, but nearly the size of the posterior gonopod coxites in at least one (
<emphasis id="A9C0EAEBFF8BFFBAFB6D5BFEFB12FA86" bold="true" box="[1062,1261,1277,1305]" pageId="34" pageNumber="35">
<taxonomicName id="5CB44D7AFF8BFFBAFB6D5BFEFB5FFA86" authority="Shear &amp; Richart &amp; Wong, 2020" authorityName="Shear &amp; Richart &amp; Wong" authorityYear="2020" box="[1062,1184,1277,1305]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pulchra" status="sp. nov.">
<emphasis id="A9C0EAEBFF8BFFBAFB6D5BFEFB5FFA86" bold="true" box="[1062,1184,1277,1305]" italics="true" pageId="34" pageNumber="35">B. pulchra</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="B2F35790FF8BFFBAFBE55BFFFB12FA86" box="[1198,1261,1278,1304]" pageId="34" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
), in which they also approach the form of the anterior gonopods of
<taxonomicName id="5CB44D7AFF8BFFBAFDAB5A23FCB8FAA2" box="[736,839,1314,1340]" class="Diplopoda" family="Conotylidae" genus="Taiyutyla" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="A9C0EAEBFF8BFFBAFDAB5A23FCB8FAA2" box="[736,839,1314,1340]" italics="true" pageId="34" pageNumber="35">Taiyutyla</emphasis>
</taxonomicName>
species. However, the posterior gonopod coxites are far more complicated in this species than in almost any other conotylid, fitting the
<taxonomicName id="5CB44D7AFF8BFFBAFB0C5A46FB43FAFE" authorityName="Loomis &amp; Schmitt" authorityYear="1971" box="[1095,1212,1351,1376]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="A9C0EAEBFF8BFFBAFB0C5A46FB43FAFE" box="[1095,1212,1351,1376]" italics="true" pageId="34" pageNumber="35">Brunsonia</emphasis>
</taxonomicName>
plan. Except for
<emphasis id="A9C0EAEBFF8BFFBAFAC85A46FEC5FA1A" bold="true" pageId="34" pageNumber="35">
<taxonomicName id="5CB44D7AFF8BFFBAFAC85A46FF11FA1A" authority="Shear &amp; Richart &amp; Wong, 2020" authorityName="Shear &amp; Richart &amp; Wong" authorityYear="2020" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="digitata" status="sp. nov.">
<emphasis id="A9C0EAEBFF8BFFBAFAC85A46FF11FA1A" bold="true" italics="true" pageId="34" pageNumber="35">B. digitata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="B2F35790FF8BFFBAFFB75A6BFEC5FA1A" box="[252,314,1386,1412]" pageId="34" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the anterior gonopods have an anteriolateral lobe set with small nodules, and all species have a deep pit or pore posteriolaterally in the anterior gonopods, with a narrow channel running from it to the tip of the gonopod; in
<emphasis id="A9C0EAEBFF8BFFBAFFA55AB2FE05FA52" bold="true" box="[238,506,1457,1485]" pageId="34" pageNumber="35">
<taxonomicName id="5CB44D7AFF8BFFBAFFA55AB2FE53FA52" authority="Shear &amp; Richart &amp; Wong, 2020" authorityName="Shear &amp; Richart &amp; Wong" authorityYear="2020" box="[238,428,1457,1485]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="chelanoparva" status="sp. nov.">
<emphasis id="A9C0EAEBFF8BFFBAFFA55AB2FE53FA52" bold="true" box="[238,428,1457,1485]" italics="true" pageId="34" pageNumber="35">B. chelanoparva</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="B2F35790FF8BFFBAFEF05AB3FE05FA52" box="[443,506,1458,1484]" pageId="34" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, this channel seems also to be occupied by a hair-like, grooved filament. We think this pit or pore is homologous with pores and gland openings that appear on the posterior gonopod coxites and tenth legpair, and which are also present in other species of conotylids.
</paragraph>
<paragraph id="9B0B36F9FF8BFFBAFF8C591FFAC1F88E" blockId="34.[151,1437,881,1809]" pageId="34" pageNumber="35">
The posterior gonopod coxites bear multiple features. The body of the coxa is globular, even swollen on the posterior side, and with one or two fimbriate branches on the lateral side. The coxite itself is massive, with various terminalia differing in each species; in
<taxonomicName id="5CB44D7AFF8BFFBAFD035966FD2BF91E" baseAuthorityName="Chamberlin" baseAuthorityYear="1920" box="[584,724,1638,1664]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="albertana">
<emphasis id="A9C0EAEBFF8BFFBAFD035966FD2BF91E" box="[584,724,1638,1664]" italics="true" pageId="34" pageNumber="35">B. albertana</emphasis>
</taxonomicName>
and
<emphasis id="A9C0EAEBFF8BFFBAFC425966FBEDF91E" bold="true" box="[777,1042,1637,1665]" pageId="34" pageNumber="35">
<taxonomicName id="5CB44D7AFF8BFFBAFC425966FC39F91E" authority="Shear &amp; Richart &amp; Wong, 2020" authorityName="Shear &amp; Richart &amp; Wong" authorityYear="2020" box="[777,966,1637,1665]" class="Diplopoda" family="Conotylidae" genus="Brunsonia" kingdom="Animalia" order="Chordeumatida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="chelanoparva" status="sp. nov.">
<emphasis id="A9C0EAEBFF8BFFBAFC425966FC39F91E" bold="true" box="[777,966,1637,1665]" italics="true" pageId="34" pageNumber="35">B. chelanoparva</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="B2F35790FF8BFFBAFC9F5967FBEDF91E" box="[980,1042,1638,1664]" pageId="34" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, there is a region set with irregular, distinct nodules. A long process (process
<emphasis id="A9C0EAEBFF8BFFBAFD14598AFD92F93B" bold="true" box="[607,621,1675,1701]" italics="true" pageId="34" pageNumber="35">a</emphasis>
) extends laterally, and just dorsal to it is another process (
<emphasis id="A9C0EAEBFF8BFFBAFBA0598BFB08F93A" bold="true" box="[1259,1271,1674,1700]" pageId="34" pageNumber="35">c</emphasis>
) with a pore at its base and a long channel leading from the pore along the process. Process
<emphasis id="A9C0EAEBFF8BFFBAFCA359ACFC09F956" bold="true" box="[1000,1014,1709,1736]" italics="true" pageId="34" pageNumber="35">b</emphasis>
may be multiple, and may or may not parallel
<emphasis id="A9C0EAEBFF8BFFBAFFB859D2FEFEF973" bold="true" box="[243,257,1747,1773]" italics="true" pageId="34" pageNumber="35">a</emphasis>
, but when present it is shorter than
<emphasis id="A9C0EAEBFF8BFFBAFDDD59D2FD5BF973" bold="true" box="[662,676,1747,1773]" italics="true" pageId="34" pageNumber="35">a</emphasis>
and is often “fuzzy” and set with many small cuticular projections. Modifications of the femora of the male pregonopodal legs range from none to knobs on pairs three to six.
</paragraph>
<paragraph id="9B0B36F9FF8BFFBAFFDC585CFD42F8E9" blockId="34.[151,701,1885,1912]" box="[151,701,1885,1912]" pageId="34" pageNumber="35">
<emphasis id="A9C0EAEBFF8BFFBAFFDC585CFD42F8E9" bold="true" box="[151,701,1885,1912]" italics="true" pageId="34" pageNumber="35">New records of a previously described species</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>