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<mods:title id="F3C12BDB27A6962E0DCD276B55C7AF2C">First Potwarmus from the Miocene of Saudi Arabia and the early phylogeny of murines (Rodentia: Muroidea)</mods:title>
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<heading id="D0C26D17FF8AFFA3D6EBC6A94B61FA55" allCaps="true" box="[906,1361,1506,1527]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="672" reason="9">
<taxonomicName id="4C35A1F8FF8AFFA3D6EBC6A94B61FA55" authority="DEBRUIJNI JACOBS, 1978" authorityName="DEBRUIJNI JACOBS" authorityYear="1978" box="[906,1361,1506,1527]" class="Mammalia" family="Muridae" genus="Progonomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="672" phylum="Chordata" rank="genus">
<emphasis id="B9410669FF8AFFA3D6EBC6A94A98FA54" box="[906,1192,1506,1527]" italics="true" pageId="8" pageNumber="672">PROGONOMYS DEBRUIJNI</emphasis>
<bibRefCitation id="EFA4A78AFF8AFFA3D1D3C6A94B61FA55" author="Jacobs LL" box="[1202,1361,1506,1527]" pageId="8" pageNumber="672" pagination="1 - 103" refId="ref11113" refString="Jacobs LL. 1978. Fossil Rodents (Rhizomyidae and Muridae) from Neogene Siwalik deposits, Pakistan. Museum of Northern Arizona Bulletin 52: 1 - 103." type="journal article" year="1978">JACOBS, 1978</bibRefCitation>
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<paragraph id="8B8ADA7BFF8AFFA2D65BC5434C16FB33" blockId="8.[826,1441,1544,1903]" lastBlockId="9.[144,759,963,1169]" lastPageId="9" lastPageNumber="673" pageId="8" pageNumber="672">
This species was erected by
<bibRefCitation id="EFA4A78AFF8AFFA3D118C5434B23F9BC" author="Jacobs LL" box="[1145,1299,1544,1566]" pageId="8" pageNumber="672" pagination="1 - 103" refId="ref11113" refString="Jacobs LL. 1978. Fossil Rodents (Rhizomyidae and Muridae) from Neogene Siwalik deposits, Pakistan. Museum of Northern Arizona Bulletin 52: 1 - 103." type="journal article" year="1978">Jacobs (1978)</bibRefCitation>
on the basis of 50 isolated teeth from the Late Miocene YGSP locality 182A, Dhok Pathan Formation,
<collectingRegion id="49F11499FF8AFFA3D028C50E4BADF9F9" box="[1353,1437,1605,1627]" country="Pakistan" name="Punjab" pageId="8" pageNumber="672">Punjab</collectingRegion>
,
<collectingCountry id="F3229AEBFF8AFFA3D65BC52F4D93F9D8" box="[826,931,1636,1658]" name="Pakistan" pageId="8" pageNumber="672">Pakistan</collectingCountry>
. The
<typeStatus id="548E64D9FF8AFFA3D68EC52F4A60F9D8" box="[1007,1104,1636,1658]" pageId="8" pageNumber="672" type="holotype">holotype</typeStatus>
(YGSP 7739,
<bibRefCitation id="EFA4A78AFF8AFFA3D062C52F4BADF9DB" author="Jacobs LL" box="[1283,1437,1636,1658]" pageId="8" pageNumber="672" pagination="1 - 103" refId="ref11113" refString="Jacobs LL. 1978. Fossil Rodents (Rhizomyidae and Muridae) from Neogene Siwalik deposits, Pakistan. Museum of Northern Arizona Bulletin 52: 1 - 103." type="journal article" year="1978">Jacobs, 1978</bibRefCitation>
: figs. 1517) is a left M1. The morphology of the teeth of
<taxonomicName id="4C35A1F8FF8AFFA3D636C5E94A61F914" box="[855,1105,1697,1719]" class="Mammalia" family="Muridae" genus="Progonomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="672" phylum="Chordata" rank="species" species="debruijni">
<emphasis id="B9410669FF8AFFA3D636C5E94A61F914" box="[855,1105,1697,1719]" italics="true" pageId="8" pageNumber="672">Progonomys debruijni</emphasis>
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is very different from that of
<taxonomicName id="4C35A1F8FF8AFFA3D65BC58B4A3CF977" authority="Antoñanzas, 2009" authorityName="Antoñanzas" authorityYear="2009" box="[826,1036,1728,1749]" class="Mammalia" family="Muridae" genus="Potwarmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="672" phylum="Chordata" rank="species" species="flynni" status="sp. nov.">
<emphasis id="B9410669FF8AFFA3D65BC58B4A3CF977" box="[826,1036,1728,1749]" italics="true" pageId="8" pageNumber="672">Potwarmus flynni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272BB12FF8AFFA3D17BC58B4A47F977" box="[1050,1143,1728,1749]" pageId="8" pageNumber="672" rank="species">sp. nov.</taxonomicNameLabel>
The M1 of
<taxonomicName id="4C35A1F8FF8AFFA3D074C58B4B91F977" box="[1301,1441,1728,1749]" class="Mammalia" family="Muridae" genus="Progonomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="672" phylum="Chordata" rank="species" species="debruijni">
<emphasis id="B9410669FF8AFFA3D074C58B4B91F977" box="[1301,1441,1728,1749]" italics="true" pageId="8" pageNumber="672">P. debruijni</emphasis>
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shows the basic murine pattern of three chevrons, lacking in AJ-7, which has a much smaller anterocone and enterostyle, the cusps more alternate, and lacks a well-developed anterostyle. The M2s of
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<emphasis id="B9410669FF8AFFA3D07AC4704B91F8ED" box="[1307,1441,1850,1872]" italics="true" pageId="8" pageNumber="672">P. debruijni</emphasis>
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are also very distinct, having a prominent anterostyle, which is lacking in AJ11 and AJ12, and a large enterostyle connected to the protocone, which is smaller and isolated in AJ11 and AJ12. The m1 of
<taxonomicName id="4C35A1F8FF8BFFA2D783C74A4EC9FB96" class="Mammalia" family="Muridae" genus="Progonomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="673" phylum="Chordata" rank="species" species="debruijni">
<emphasis id="B9410669FF8BFFA2D783C74A4EC9FB96" italics="true" pageId="9" pageNumber="673">P. debruijni</emphasis>
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differs from AJ13, for example, in having a well-developed bilobed anteroconid with a central X-shaped connection pattern, and a variable number of cingular cusps (absent in AJ13).
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