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<document id="44756AC1E202257D93CCDBEF7BDBEFE6" ID-CLB-Dataset="7373" ID-DOI="10.11646/zootaxa.4952.1.7" ID-GBIF-Dataset="024ed081-5dff-4cd4-a12a-201e4cc78f2c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4671878" ID-ZooBank="F276FC72-7FFD-4813-9817-B677C20284FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617874253906" checkinUser="plazi" docAuthor="Michelsen, Verner" docDate="2021" docId="039C4C261D6FBE20FF46A84BFCE63879" docLanguage="en" docName="zootaxa.4952.1.7.pdf" docOrigin="Zootaxa 4952 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spilogona maderensis Michelsen 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="133" masterDocId="FFA5345E1D6EBE25FFD1AF0AFFAA3C07" masterDocTitle="Macaronesian Muscidae (Diptera). III. First record of the genus Spilogona Schnabl based on two new endemic species from the Azores and Madeira" masterLastPageNumber="134" masterPageNumber="128" pageNumber="129" updateTime="1698936198281" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E95AE7CC0D500021735DDA3FB0E00FB1">Macaronesian Muscidae (Diptera). III. First record of the genus Spilogona Schnabl based on two new endemic species from the Azores and Madeira</mods:title>
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<mods:namePart id="F878486ADBC1ED6B70633CFEA099804B">Michelsen, Verner</mods:namePart>
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<mods:title id="95CFBD5EA95141462DD1DD1711045F76">Zootaxa</mods:title>
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<mods:date id="5838E04AAB8A7EF52BAC9822D4177DF0">2021</mods:date>
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<mods:number id="0E538424DB4BF17006938FA8B9CA3E62">2021-04-08</mods:number>
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<mods:number id="3E7AA0CBB6751E98F52D552EF90696BD">4952</mods:number>
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<mods:identifier id="BDB7BDE099C0856E4545802B48493A8F" type="DOI">10.11646/zootaxa.4952.1.7</mods:identifier>
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<treatment id="039C4C261D6FBE20FF46A84BFCE63879" ID-DOI="http://doi.org/10.5281/zenodo.4685797" ID-GBIF-Taxon="181003823" ID-Zenodo-Dep="4685797" LSID="urn:lsid:plazi:treatment:039C4C261D6FBE20FF46A84BFCE63879" httpUri="http://treatment.plazi.org/id/039C4C261D6FBE20FF46A84BFCE63879" lastPageId="5" lastPageNumber="133" pageId="1" pageNumber="129">
<subSubSection id="C32FAEBB1D6FBE24FF46A84BFDAD3B5B" box="[151,519,1857,1884]" pageId="1" pageNumber="129" type="nomenclature">
<paragraph id="8B8AFD301D6FBE24FF46A84BFDAD3B5B" blockId="1.[151,519,1857,1920]" box="[151,519,1857,1884]" pageId="1" pageNumber="129">
<heading id="D0C24A5C1D6FBE24FF46A84BFDAD3B5B" bold="true" box="[151,519,1857,1884]" fontSize="11" level="1" pageId="1" pageNumber="129" reason="1">
<emphasis id="B94121221D6FBE24FF46A84BFDAD3B5B" bold="true" box="[151,519,1857,1884]" pageId="1" pageNumber="129">
<taxonomicName id="4C3586B31D6FBE24FF46A84BFE083B5B" ID-CoL="9HW3H" authority="Michelsen, 2021" authorityName="Michelsen" authorityYear="2021" box="[151,418,1857,1884]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="129" phylum="Arthropoda" rank="species" species="maderensis" status="sp. nov.">
<emphasis id="B94121221D6FBE24FF46A84BFE083B5B" bold="true" box="[151,418,1857,1884]" italics="true" pageId="1" pageNumber="129">Spilogona maderensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2729C591D6FBE24FE78A848FDAD3B5B" box="[425,519,1858,1884]" pageId="1" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32FAEBB1D6FBE24FF46A86CFE8E3B87" box="[151,292,1894,1920]" pageId="1" pageNumber="129" type="description">
<paragraph id="8B8AFD301D6FBE24FF46A86CFE8E3B87" blockId="1.[151,519,1857,1920]" box="[151,292,1894,1920]" pageId="1" pageNumber="129">
(
<figureCitation id="130EE1B51D6FBE24FF4EA86CFEB13B87" box="[159,283,1894,1920]" captionStart="FIGURE 3" captionStartId="4.[151,250,1914,1938]" captionTargetBox="[190,1410,195,1879]" captionTargetId="figure-15@4.[151,1436,181,1889]" captionTargetPageId="4" captionText="FIGURE 3. Spilogona maderensis sp. nov. Female holotype: lateral habitus (A); head and anterior mesonotum in dorsal view (B)." figureDoi="http://doi.org/10.5281/zenodo.4671884" httpUri="https://zenodo.org/record/4671884/files/figure.png" pageId="1" pageNumber="129">Figs 3A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32FAEBB1D6FBE24FF46A8A7FA373BEB" pageId="1" pageNumber="129" type="materials_examined">
<paragraph id="8B8AFD301D6FBE24FF46A8A7FA373BEB" blockId="1.[151,1437,1965,2029]" pageId="1" pageNumber="129">
<emphasis id="B94121221D6FBE24FF46A8A7FE2A3BC0" bold="true" box="[151,384,1965,1991]" pageId="1" pageNumber="129">Material examined.</emphasis>
<emphasis id="B94121221D6FBE24FE57A8A5FDB53BCF" box="[390,543,1966,1992]" italics="true" pageId="1" pageNumber="129">Type material</emphasis>
[
<materialsCitation id="3B5DF76D1D6FBE24FDFDA8A7FA373BEB" ID-GBIF-Occurrence="3071307303" collectingDate="1957-06-27" collectingDateMax="1957-06-29" collectingDateMin="1957-06-27" collectionCode="MZH" collectorName="H. Lindberg &amp; Condition" latitude="32.76" location="Pico Ruivo" longLatPrecision="725" longitude="-16.94" pageId="1" pageNumber="129" specimenCount="1" specimenCount-female="1" stateProvince="Madeira" typeStatus="holotype">
<collectionCode id="ED2465F51D6FBE24FDFDA8A7FDD93BCF" box="[556,627,1965,1992]" country="Finland" httpUri="http://grbio.org/cool/2vmj-fp0v" name="Finnish Museum of Natural History" pageId="1" pageNumber="129" type="Museum">MZH</collectionCode>
]:
<typeStatus id="548E43921D6FBE24FD55A8A4FD463BCF" box="[644,748,1966,1992]" pageId="1" pageNumber="129" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3336B91D6FBE24FD23A8A4FCA33BCF" box="[754,777,1966,1992]" pageId="1" pageNumber="129" type="female"></specimenCount>
:
<collectingRegion id="49F133D21D6FBE24FCC4A8A4FC363BCF" box="[789,924,1966,1992]" country="Portugal" name="Madeira" pageId="1" pageNumber="129">MADEIRA</collectingRegion>
:
<location id="8EEAABEB1D6FBE24FC77A8A4FB8B3BCF" LSID="urn:lsid:plazi:treatment:039C4C261D6FBE20FF46A84BFCE63879:8EEAABEB1D6FBE24FC77A8A4FB8B3BCF" box="[934,1057,1966,1992]" latitude="32.76" longLatPrecision="725" longitude="-16.94" name="Pico Ruivo" pageId="1" pageNumber="129" stateProvince="Madeira">Pico Ruivo</location>
[
<geoCoordinate id="EE019BF71D6FBE24FBFFA8A4FB2F3BCF" box="[1070,1157,1966,1992]" degrees="32.76" direction="north" orientation="latitude" pageId="1" pageNumber="129" precision="555" value="32.76">32.76N</geoCoordinate>
<geoCoordinate id="EE019BF71D6FBE24FB5AA8A7FB433BCF" box="[1163,1257,1965,1992]" degrees="16.94" direction="west" orientation="longitude" pageId="1" pageNumber="129" precision="555" value="-16.94">16.94W</geoCoordinate>
],
<date id="FF8BDBF01D6FBE24FB2BA8A4FA363BCE" box="[1274,1436,1966,1993]" pageId="1" pageNumber="129" value="1957-06-27" valueMax="1957-06-29" valueMin="1957-06-27">
<collectingDate id="EFCF22181D6FBE24FB2BA8A4FA363BCE" box="[1274,1436,1966,1993]" pageId="1" pageNumber="129" value="1957-06-27" valueMax="1957-06-29" valueMin="1957-06-27">2729.vi.1957</collectingDate>
</date>
(
<collectorName id="26C098E61D6FBE24FF71A8D8FE853BEA" box="[160,303,2002,2029]" pageId="1" pageNumber="129">H. Lindberg</collectorName>
).
<collectorName id="26C098E61D6FBE24FE91A8D8FE1A3BEB" box="[320,432,2002,2028]" pageId="1" pageNumber="129">Condition</collectorName>
excellent except for the missing left postpedicel, left fore and mid legs and right hind leg.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF4AADB81D6CBE27FF46A892FF6A3BD3" ID-DOI="http://doi.org/10.5281/zenodo.4671880" ID-Zenodo-Dep="4671880" httpUri="https://zenodo.org/record/4671880/files/figure.png" pageId="2" pageNumber="130" startId="2.[151,250,1944,1968]" targetBox="[347,1247,194,1861]" targetPageId="2">
<paragraph id="8B8AFD301D6CBE27FF46A892FF6A3BD3" blockId="2.[151,1436,1944,2004]" pageId="2" pageNumber="130">
<emphasis id="B94121221D6CBE27FF46A892FDE23BB7" bold="true" box="[151,584,1944,1968]" pageId="2" pageNumber="130">
FIGURE 1.
<taxonomicName id="4C3586B31D6CBE27FECDA892FE443BB7" authority="Michelsen, 2021" authorityName="Michelsen" authorityYear="2021" box="[284,494,1944,1968]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="130" phylum="Arthropoda" rank="species" species="azorensis" status="sp. nov.">
<emphasis id="B94121221D6CBE27FECDA892FE443BB7" bold="true" box="[284,494,1944,1968]" italics="true" pageId="2" pageNumber="130">Spilogona azorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2729C591D6CBE27FE27A892FDE23BB7" box="[502,584,1944,1968]" pageId="2" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female holotype: lateral habitus (A); head and anterior mesonotum in dorsal view (B).
</paragraph>
</caption>
<caption id="DF4AADB81D6DBE26FF46A892FB9E3BB7" ID-DOI="http://doi.org/10.5281/zenodo.4671882" ID-Zenodo-Dep="4671882" box="[151,1076,1944,1968]" httpUri="https://zenodo.org/record/4671882/files/figure.png" pageId="3" pageNumber="131" startId="3.[151,250,1944,1968]" targetBox="[187,1349,228,1890]" targetPageId="3">
<paragraph id="8B8AFD301D6DBE26FF46A892FB9E3BB7" blockId="3.[151,1076,1944,1968]" box="[151,1076,1944,1968]" pageId="3" pageNumber="131">
<emphasis id="B94121221D6DBE26FF46A892FDEB3BB7" bold="true" box="[151,577,1944,1968]" pageId="3" pageNumber="131">
FIGURE 2.
<taxonomicName id="4C3586B31D6DBE26FEC8A892FE403BB7" authority="Michelsen, 2021" authorityName="Michelsen" authorityYear="2021" box="[281,490,1944,1968]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="species" species="azorensis" status="sp. nov.">
<emphasis id="B94121221D6DBE26FEC8A892FE403BB7" bold="true" box="[281,490,1944,1968]" italics="true" pageId="3" pageNumber="131">Spilogona azorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2729C591D6DBE26FE20A892FDEB3BB7" box="[497,577,1944,1968]" pageId="3" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Ovipositor in lateral (A) and ventral (B) aspects.
</paragraph>
</caption>
<caption id="DF4AADB81D6ABE21FF46A870FF6A3BB1" ID-DOI="http://doi.org/10.5281/zenodo.4671884" ID-Zenodo-Dep="4671884" httpUri="https://zenodo.org/record/4671884/files/figure.png" pageId="4" pageNumber="132" startId="4.[151,250,1914,1938]" targetBox="[190,1410,195,1879]" targetPageId="4">
<paragraph id="8B8AFD301D6ABE21FF46A870FF6A3BB1" blockId="4.[151,1436,1913,1974]" pageId="4" pageNumber="132">
<emphasis id="B94121221D6ABE21FF46A870FDFC3B95" bold="true" box="[151,598,1913,1938]" pageId="4" pageNumber="132">
FIGURE 3.
<taxonomicName id="4C3586B31D6ABE21FEC8A873FE553B96" authority="Michelsen, 2021" authorityName="Michelsen" authorityYear="2021" box="[281,511,1913,1937]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="132" phylum="Arthropoda" rank="species" species="maderensis" status="sp. nov.">
<emphasis id="B94121221D6ABE21FEC8A873FE553B96" bold="true" box="[281,511,1913,1937]" italics="true" pageId="4" pageNumber="132">Spilogona maderensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2729C591D6ABE21FDD7A870FDFC3B95" box="[518,598,1914,1938]" pageId="4" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female holotype: lateral habitus (A); head and anterior mesonotum in dorsal view (B).
</paragraph>
</caption>
<subSubSection id="C32FAEBB1D6BBE20FF16AF9CFBBE385E" pageId="5" pageNumber="133" type="description">
<paragraph id="8B8AFD301D6BBE20FF16AF9CFED83FA2" blockId="5.[151,1437,150,1150]" pageId="5" pageNumber="133">
<emphasis id="B94121221D6BBE20FF16AF9CFEF33CB7" bold="true" box="[199,345,150,176]" pageId="5" pageNumber="133">Description.</emphasis>
<emphasis id="B94121221D6BBE20FEB5AF9DFE123CB6" box="[356,440,151,177]" italics="true" pageId="5" pageNumber="133">Female</emphasis>
(
<figureCitation id="130EE1B51D6BBE20FE1DAF9DFDF93CB6" box="[460,595,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1914,1938]" captionTargetBox="[190,1410,195,1879]" captionTargetId="figure-15@4.[151,1436,181,1889]" captionTargetPageId="4" captionText="FIGURE 3. Spilogona maderensis sp. nov. Female holotype: lateral habitus (A); head and anterior mesonotum in dorsal view (B)." figureDoi="http://doi.org/10.5281/zenodo.4671884" httpUri="https://zenodo.org/record/4671884/files/figure.png" pageId="5" pageNumber="133">Figs 3A, B</figureCitation>
). Medium sized, wing length
<quantity id="4CCD50D51D6BBE20FC68AF9CFBA23CB6" box="[953,1032,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="133" unit="mm" value="5.5">5.5mm</quantity>
(n=1). Head and prementum grey pruinose; entire frons suffused with brown, frontal vitta barely darker than frontal triangle; parafacial plates silvery white with a golden brown tinge. Thorax covered in dense grey pruinosity except extensively dark brown on dorsum of mesonotum including scutellar disc and on upper part of anepisternum. Wing membrane rather intensely smoky brown, yet both crossveins distinctly darker shadowed; calypteres dirty whitish with brown margins, broadly so on lower calypter; halter deep yellow. Legs black, femora grey pruinose. Abdomen grey pruinose except for very large paired trapezoid marks on tergites III and IV and a median field on tergite V being shiny dark brown. Postpedicel about 2 times as long as wide, anterior apical corner forming a very obtuse angle; arista short-pubescent, longest branches barely as long as its greatest basal diameter. Haustellum conical, short and thick. Broad frons widening towards lunule; frontal vitta parallel-sided, frontal triangle relatively small, extended to middle of frontal vitta. Frons and lower facial margin slightly and equally projecting. Fronto-orbital plate with 1 reclinate orbital seta about same size as outer postvertical seta, 4 robust inclinate frontal setae with a few interspatial setulae and bordered laterally by an irregular row of setulae. Mesonotal ground setulae abundant. Dorsocentral setae 2 + 3; presutural acrostichal field with numerous setulae arranged in six irregular rows, outermost ones standing close to adjacent dorsocentral lines and separated from median rows by narrow bare stripes. Notopleuron without setulae. Anepisternum without interspatial setula(e). Katepisternal setae 1 + 2, but lower posterior seta much weaker than both upper ones. Scutellum with 1 setula on upper, preapical border of declivity. Vein C distal to subcostal break with extensive row of dorsal setulae; spines at subcostal break inconspicuous. Fore tibia with submedian
<emphasis id="B94121221D6BBE20FC0EADF1FC473F13" box="[991,1005,763,788]" italics="true" pageId="5" pageNumber="133">p</emphasis>
seta below middle. Mid femur without subapical
<emphasis id="B94121221D6BBE20FED9AC15FE8E3F3E" box="[264,292,799,825]" italics="true" pageId="5" pageNumber="133">ad</emphasis>
seta and without distinct
<emphasis id="B94121221D6BBE20FD94AC15FDF53F3F" box="[581,607,799,824]" italics="true" pageId="5" pageNumber="133">pv</emphasis>
setae; mid tibia before apex with 2 strong
<emphasis id="B94121221D6BBE20FB91AC15FBF63F3E" box="[1088,1116,799,825]" italics="true" pageId="5" pageNumber="133">pd</emphasis>
setae only. Hind femur with 3
<emphasis id="B94121221D6BBE20FF7CAC49FF6D3F5B" box="[173,199,835,860]" italics="true" pageId="5" pageNumber="133">av</emphasis>
setae on distal third, without
<emphasis id="B94121221D6BBE20FDC7AC49FD9A3F5B" box="[534,560,835,860]" italics="true" pageId="5" pageNumber="133">pv</emphasis>
setae; hind tibia with 1
<emphasis id="B94121221D6BBE20FC90AC49FCF13F5B" box="[833,859,835,860]" italics="true" pageId="5" pageNumber="133">av</emphasis>
and 3
<emphasis id="B94121221D6BBE20FC79AC49FC6E3F5A" box="[936,964,835,861]" italics="true" pageId="5" pageNumber="133">ad</emphasis>
setae. Abdomen with modest-sized discal and hind marginal setae. Sternite I bare. Ovipositor: Not dissected, but exposed distal part appears very similar to that of
<taxonomicName id="4C3586B31D6BBE20FF37AC81FEC13FA3" authorityName="Michelsen" authorityYear="2021" box="[230,363,907,933]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="133" phylum="Arthropoda" rank="species" species="azorensis">
<emphasis id="B94121221D6BBE20FF37AC81FEC13FA3" box="[230,363,907,933]" italics="true" pageId="5" pageNumber="133">S. azorensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8AFD301D6BBE20FF16ACA5FE2F3FCE" blockId="5.[151,1437,150,1150]" box="[199,389,943,969]" pageId="5" pageNumber="133">
<emphasis id="B94121221D6BBE20FF16ACA5FEAB3FCE" box="[199,257,943,969]" italics="true" pageId="5" pageNumber="133">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B8AFD301D6BBE20FF16ACD8FBBE385E" blockId="5.[151,1437,150,1150]" pageId="5" pageNumber="133">
<emphasis id="B94121221D6BBE20FF16ACD8FEDA3FEB" bold="true" box="[199,368,978,1004]" pageId="5" pageNumber="133">Relationships.</emphasis>
The shadowed crossveins and presence of setulae dorsally on vein C may readily distinguish female
<taxonomicName id="4C3586B31D6BBE20FF3CACFDFE213816" authorityName="Michelsen" authorityYear="2021" box="[237,395,1015,1041]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="133" phylum="Arthropoda" rank="species" species="maderensis">
<emphasis id="B94121221D6BBE20FF3CACFDFE213816" box="[237,395,1015,1041]" italics="true" pageId="5" pageNumber="133">S. maderensis</emphasis>
</taxonomicName>
from females of other species of
<taxonomicName id="4C3586B31D6BBE20FCC6ACFDFC233816" authorityName="Schnabl" authorityYear="1911" box="[791,905,1015,1041]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="B94121221D6BBE20FCC6ACFDFC233816" box="[791,905,1015,1041]" italics="true" pageId="5" pageNumber="133">Spilogona</emphasis>
</taxonomicName>
. Apart from these obvious autapomorphies, it shares many characters with
<taxonomicName id="4C3586B31D6BBE20FE3BAB11FDD93833" authorityName="Michelsen" authorityYear="2021" box="[490,627,1051,1077]" class="Insecta" family="Muscidae" genus="Spilogona" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="133" phylum="Arthropoda" rank="species" species="azorensis">
<emphasis id="B94121221D6BBE20FE3BAB11FDD93833" box="[490,627,1051,1077]" italics="true" pageId="5" pageNumber="133">S. azorensis</emphasis>
</taxonomicName>
. However, a qualified assessment of relationships is difficult without access to the male sex of both species allowing comparisons of their terminalia.
</paragraph>
</subSubSection>
<subSubSection id="C32FAEBB1D6BBE20FF16AB68FCE63879" box="[199,844,1122,1150]" pageId="5" pageNumber="133" type="distribution">
<paragraph id="8B8AFD301D6BBE20FF16AB68FCE63879" blockId="5.[151,1437,150,1150]" box="[199,844,1122,1150]" pageId="5" pageNumber="133">
<emphasis id="B94121221D6BBE20FF16AB68FECB387B" bold="true" box="[199,353,1122,1148]" pageId="5" pageNumber="133">Distribution.</emphasis>
Apparently endemic to
<collectingRegion id="49F133D21D6BBE20FDBEAB69FD66387A" box="[623,716,1123,1149]" country="Portugal" name="Madeira" pageId="5" pageNumber="133">Madeira</collectingRegion>
(
<collectingCountry id="F322BDA01D6BBE20FD0AAB69FCEA3879" box="[731,832,1123,1150]" name="Portugal" pageId="5" pageNumber="133">Portugal</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>