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<mods:title id="21BEDF6D83802C831B0DC25113E318D3">A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region</mods:title>
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<treatment id="038DBF6E5403CC76FF06FB3CFAE4FC97" LSID="urn:lsid:plazi:treatment:038DBF6E5403CC76FF06FB3CFAE4FC97" httpUri="http://treatment.plazi.org/id/038DBF6E5403CC76FF06FB3CFAE4FC97" lastPageId="41" lastPageNumber="192" pageId="38" pageNumber="189">
<subSubSection id="C33E5DF35403CC79FF06FB3CFD64FB7C" box="[151,700,1252,1279]" pageId="38" pageNumber="189" type="nomenclature">
<paragraph id="8B9B0E785403CC79FF06FB3CFD64FB7C" blockId="38.[151,700,1252,1315]" box="[151,700,1252,1279]" pageId="38" pageNumber="189">
<heading id="D0D3B9145403CC79FF06FB3CFD64FB7C" bold="true" box="[151,700,1252,1279]" fontSize="11" level="1" pageId="38" pageNumber="189" reason="1">
<emphasis id="B950D26A5403CC79FF06FB3CFD64FB7C" bold="true" box="[151,700,1252,1279]" pageId="38" pageNumber="189">
3.2.5.2.2.
<taxonomicName id="4C2475FB5403CC79FE9CFB3CFD64FB7C" ID-CoL="499SD" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[269,700,1252,1279]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A5403CC79FE9CFB3CFE2EFB7C" bold="true" box="[269,502,1252,1279]" italics="true" pageId="38" pageNumber="189">Omalium cariosum</emphasis>
<bibRefCitation id="EFB573895403CC79FE6FFB3CFD64FB7C" author="Cameron, M." box="[510,700,1252,1278]" pageId="38" pageNumber="189" pagination="160 - 198" refId="ref32716" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF35403CC79FF06FAD0FEA3FAA0" box="[151,379,1288,1315]" pageId="38" pageNumber="189" type="description">
<paragraph id="8B9B0E785403CC79FF06FAD0FEA3FAA0" blockId="38.[151,700,1252,1315]" box="[151,379,1288,1315]" pageId="38" pageNumber="189">
(
<figureCitation id="131F12FD5403CC79FF0EFAD0FF2EFAA1" box="[159,246,1288,1315]" captionStart="FIGURE 19" captionStartId="7.[151,250,1891,1915]" captionTargetBox="[151,1435,957,1868]" captionTargetId="figure-299@7.[151,1435,957,1868]" captionTargetPageId="7" captionText="FIGURE 19. Distribution of Omalium in the Himalayan region: O. acutangulum sp. n. (circles), O. amplissimum sp. n. (star), O. beesoni (squares), O. cariosum (diamonds), O. flavotestaceum sp. n. (triangles)." figureDoi="http://doi.org/10.5281/zenodo.10146178" httpUri="https://zenodo.org/record/10146178/files/figure.png" pageId="38" pageNumber="189">Figs 19</figureCitation>
,
<figureCitation id="131F12FD5403CC79FE93FAD1FEF9FAA1" box="[258,289,1289,1315]" captionStart="FIGURES 70–72" captionStartId="39.[151,264,1085,1109]" captionTargetBox="[159,1419,187,1052]" captionTargetId="figure-16@39.[151,1436,181,1061]" captionTargetPageId="39" captionText="FIGURES 70–72. Habitus of Omalium: 70—O. cariosum (lectotype), 71—O. flavotestaceum (paratype), 72—O. kumari (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="38" pageNumber="189">70</figureCitation>
,
<figureCitation id="131F12FD5403CC79FEBDFAD0FEACFAA0" box="[300,372,1288,1314]" captionStart-0="FIGURES 73–75" captionStart-1="FIGURES 76–83" captionStartId-0="40.[151,264,1430,1454]" captionStartId-1="41.[151,264,1651,1675]" captionTargetBox-0="[169,1433,187,1403]" captionTargetBox-1="[154,1432,816,1625]" captionTargetId-0="figure-22@40.[151,1436,181,1406]" captionTargetId-1="figure-275@41.[151,1435,813,1628]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 73–75. Aedeagus of Omalium cariosum (paralectotype): 73—aedeagus, parameral view, 74—aedeagus, lateral view, 75—median lobe, lateral view. Scale bar: 0.1 mm." captionText-1="FIGURES 76–83. Female accessory sclerite (Figs 76–77, 79, 81–82) and spermatheca (Figs 78, 80, 83) of Omalium: 76— O.cariosum, 77–78—O. flavotestaceum, 79–80—O. kumari, 81—O. latocavum, 82–83—O. secretum. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10146222" figureDoi-1="http://doi.org/10.5281/zenodo.10146224" httpUri-0="https://zenodo.org/record/10146222/files/figure.png" httpUri-1="https://zenodo.org/record/10146224/files/figure.png" pageId="38" pageNumber="189">73–76</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF35403CC79FF06FA95FCA1FAE7" box="[151,889,1357,1381]" pageId="38" pageNumber="189" type="reference_group">
<paragraph id="8B9B0E785403CC79FF06FA95FCA1FAE7" blockId="38.[151,889,1357,1381]" box="[151,889,1357,1381]" pageId="38" pageNumber="189">
<taxonomicName id="4C2475FB5403CC79FF06FA95FDA7FAE7" authority="Cameron, 1924: 167" authorityName="Cameron" authorityPageNumber="167" authorityYear="1924" box="[151,639,1357,1381]" class="Insecta" family="Staphylinidae" genus="Omalium" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A5403CC79FF06FA95FF2DFAE7" box="[151,245,1357,1381]" italics="true" pageId="38" pageNumber="189">Omalium</emphasis>
(s. str.)
<emphasis id="B950D26A5403CC79FED9FA96FE7EFAE7" box="[328,422,1358,1381]" italics="true" pageId="38" pageNumber="189">cariosum</emphasis>
<bibRefCitation id="EFB573895403CC79FE3DFA95FDA7FAE7" author="Cameron, M." box="[428,639,1357,1381]" pageId="38" pageNumber="189" pagination="160 - 198" refId="ref32716" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924: 167</bibRefCitation>
</taxonomicName>
; Scheerpeltz 1933: 1044
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF35403CC79FF06FA4BFBCBF8BB" pageId="38" pageNumber="189" type="materials_examined">
<paragraph id="8B9B0E785403CC79FF06FA4BFEDDF9E3" blockId="38.[151,1437,1427,1958]" pageId="38" pageNumber="189">
<emphasis id="B950D26A5403CC79FF06FA4BFE9CFA2F" bold="true" box="[151,324,1427,1453]" pageId="38" pageNumber="189">Type material.</emphasis>
<materialsCitation id="3B4C04255403CC79FED8FA4CFD72F99B" collectorName="Dr. Cameron &amp; M. Cameron." location="Dun" pageId="38" pageNumber="189" specimenCount="1" specimenCount-female="1" stateProvince="Down" typeStatus="lectotype">
<typeStatus id="549FB0DA5403CC79FED8FA4CFE61FA2C" box="[329,441,1428,1454]" pageId="38" pageNumber="189" type="lectotype">Lectotype</typeStatus>
(
<emphasis id="B950D26A5403CC79FE59FA4BFD5AFA2F" bold="true" box="[456,642,1427,1453]" pageId="38" pageNumber="189">here designated</emphasis>
) of
<taxonomicName id="4C2475FB5403CC79FD3CFA4BFB5CFA2F" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[685,1156,1427,1453]" class="Insecta" family="Staphylinidae" genus="Omalium" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A5403CC79FD3CFA4BFCCDFA2F" box="[685,789,1427,1453]" italics="true" pageId="38" pageNumber="189">Omalium</emphasis>
(s. str.)
<emphasis id="B950D26A5403CC79FCFDFA4CFC0CFA2F" box="[876,980,1428,1453]" italics="true" pageId="38" pageNumber="189">cariosum</emphasis>
<bibRefCitation id="EFB573895403CC79FC48FA4BFB5CFA2F" author="Cameron, M." box="[985,1156,1427,1453]" pageId="38" pageNumber="189" pagination="160 - 198" refId="ref32716" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
<specimenCount id="9D22C5F15403CC79FB1BFA4CFB47FA2C" box="[1162,1183,1428,1454]" count="1" pageId="38" pageNumber="189" type="female">♀</specimenCount>
(
<figureCitation id="131F12FD5403CC79FB3CFA4BFAD9FA2F" box="[1197,1281,1427,1453]" captionStart="FIGURES 70–72" captionStartId="39.[151,264,1085,1109]" captionTargetBox="[159,1419,187,1052]" captionTargetId="figure-16@39.[151,1436,181,1061]" captionTargetPageId="39" captionText="FIGURES 70–72. Habitus of Omalium: 70—O. cariosum (lectotype), 71—O. flavotestaceum (paratype), 72—O. kumari (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="38" pageNumber="189">Fig. 70</figureCitation>
): ‘Type | H.T.’ &lt;round label with red margins, printed&gt;,‘Dehra Dun |
<collectorName id="26D16BAE5403CC79FD77FA60FCADFA53" box="[742,885,1463,1490]" pageId="38" pageNumber="189">Dr. Cameron</collectorName>
| 21. 3. [handwritten] 1922.’ &lt;printed&gt;, ‘
<taxonomicName id="4C2475FB5403CC79FABCFA6FFA44FA53" authorityName="Gravenhorst" authorityYear="1802" box="[1325,1436,1463,1489]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="genus">Omalium</taxonomicName>
| cariosum | Cam..’ &lt;handwritten&gt;, ‘
<collectorName id="26D16BAE5403CC79FDABFA03FD15FA77" box="[570,717,1499,1525]" pageId="38" pageNumber="189">M.Cameron.</collectorName>
| Bequest. | B.M.1955-147.’ &lt;printed&gt;, ‘NHMUK014663259’ &lt;printed label, with barcode on the right side
</materialsCitation>
&gt;, ‘
<typeStatus id="549FB0DA5403CC79FD5CFA27FCAAF99B" box="[717,882,1535,1561]" pageId="38" pageNumber="189" type="lectotype">LECTOTYPE</typeStatus>
|
<taxonomicName id="4C2475FB5403CC79FC1DFA27FC2CF99B" authorityName="Gravenhorst" authorityYear="1802" box="[908,1012,1535,1561]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="genus">
<emphasis id="B950D26A5403CC79FC1DFA27FC2CF99B" box="[908,1012,1535,1561]" italics="true" pageId="38" pageNumber="189">Omalium</emphasis>
</taxonomicName>
(s. str.) |
<taxonomicName id="4C2475FB5403CC79FBF8F9D8FA54F99B" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[1129,1420,1535,1561]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A5403CC79FBF8F9D8FB09F99B" box="[1129,1233,1536,1561]" italics="true" pageId="38" pageNumber="189">cariosum</emphasis>
<bibRefCitation id="EFB573895403CC79FB4DFA27FA54F99B" author="Cameron, M." box="[1244,1420,1535,1561]" pageId="38" pageNumber="189" pagination="160 - 198" refId="ref32716" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
| Shavrin A.V. des. 2023 &lt;red, printed&gt;‘, ‘
<taxonomicName id="4C2475FB5403CC79FDF8F9FBFD09F9BF" authorityName="Gravenhorst" authorityYear="1802" box="[617,721,1571,1597]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="genus">
<emphasis id="B950D26A5403CC79FDF8F9FBFD09F9BF" box="[617,721,1571,1597]" italics="true" pageId="38" pageNumber="189">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C2475FB5403CC79FD76F9FCFBDDF9BF" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[743,1029,1571,1597]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A5403CC79FD76F9FCFC97F9BF" box="[743,847,1572,1597]" italics="true" pageId="38" pageNumber="189">cariosum</emphasis>
<bibRefCitation id="EFB573895403CC79FCC6F9FBFBDDF9BF" author="Cameron, M." box="[855,1029,1571,1597]" pageId="38" pageNumber="189" pagination="160 - 198" refId="ref32716" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
| Shavrin A.V. det. 2023’ &lt;printed&gt; (BMNH).
</paragraph>
<paragraph id="8B9B0E785403CC79FF56F9B3FEDDF94F" blockId="38.[151,1437,1427,1958]" pageId="38" pageNumber="189">
<materialsCitation id="3B4C04255403CC79FF56F9B3FED9F94F" collectorName="Dr. Cameron &amp; M. Cameron." determinerName="Shavrin A. V." location="Dun" pageId="38" pageNumber="189" specimenCount="1" stateProvince="Down" typeStatus="paralectotype">
<typeStatus id="549FB0DA5403CC79FF56F9B3FEBCF907" box="[199,356,1643,1669]" pageId="38" pageNumber="189" type="paralectotype">Paralectotype</typeStatus>
: 1 ♁ (dissected): ‘Dehra
<collectingRegion id="49E0C09A5403CC79FDECF9B4FD69F904" box="[637,689,1644,1670]" country="United Kingdom" name="Down" pageId="38" pageNumber="189">Dun</collectingRegion>
. |
<collectorName id="26D16BAE5403CC79FD59F9B4FC81F907" box="[712,857,1643,1670]" pageId="38" pageNumber="189">Dr. Cameron</collectorName>
| 4. 3. [handwritten] 1922.’ &lt;printed&gt;,
<collectorName id="26D16BAE5403CC79FA9EF9B3FA45F907" box="[1295,1437,1643,1669]" pageId="38" pageNumber="189">M.Cameron.</collectorName>
| Bequest. | B.M.1955-147.’ &lt;printed&gt;, ‘
<taxonomicName id="4C2475FB5403CC79FDF3F957FD12F92B" authorityName="Gravenhorst" authorityYear="1802" box="[610,714,1679,1705]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="genus">
<emphasis id="B950D26A5403CC79FDF3F957FD12F92B" box="[610,714,1679,1705]" italics="true" pageId="38" pageNumber="189">Omalium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C2475FB5403CC79FD73F948FBDAF92B" authority="Cameron, 1924" authorityName="Cameron" authorityYear="1924" box="[738,1026,1679,1705]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A5403CC79FD73F948FC92F92B" box="[738,842,1680,1705]" italics="true" pageId="38" pageNumber="189">cariosum</emphasis>
<bibRefCitation id="EFB573895403CC79FCC2F957FBDAF92B" author="Cameron, M." box="[851,1026,1679,1705]" pageId="38" pageNumber="189" pagination="160 - 198" refId="ref32716" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
</taxonomicName>
|
<determinerName id="7E47E2BC5403CC79FB88F957FB73F92B" box="[1049,1195,1679,1705]" pageId="38" pageNumber="189">Shavrin A.V.</determinerName>
det. 2023’ &lt;printed&gt; (BMNH)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9B0E785403CC79FF56F90FFBCBF8BB" blockId="38.[151,1437,1427,1958]" pageId="38" pageNumber="189">
<emphasis id="B950D26A5403CC79FF56F90FFDE9F973" bold="true" box="[199,561,1751,1777]" pageId="38" pageNumber="189">Additional material examined.</emphasis>
<materialsCitation id="3B4C04255403CC79FDA7F90FFBD7F8BB" collectingDate="1921-03-27" collectorName="Dr. Cameron" country="India" location="Woodstock Falls" municipality="United Prov." pageId="38" pageNumber="189" specimenCount="1" specimenCount-female="1" stateProvince="Uttarakhand">
<emphasis id="B950D26A5403CC79FDA7F90FFC93F973" bold="true" box="[566,843,1751,1777]" pageId="38" pageNumber="189">
<collectingCountry id="F3334EE85403CC79FDA7F90FFD55F973" box="[566,653,1751,1777]" name="India" pageId="38" pageNumber="189">INDIA</collectingCountry>
:
<collectingRegion id="49E0C09A5403CC79FD06F90FFC9FF973" box="[663,839,1751,1777]" country="India" name="Uttarakhand" pageId="38" pageNumber="189">UTTARAKHAND</collectingRegion>
:
</emphasis>
<specimenCount id="9D22C5F15403CC79FCC1F90FFCA2F970" box="[848,890,1751,1778]" count="1" pageId="38" pageNumber="189" type="female">1 ♀</specimenCount>
: ‘
<collectorName id="26D16BAE5403CC79FC1BF90FFC77F973" box="[906,943,1751,1777]" pageId="38" pageNumber="189">Co</collectorName>
- | type’, ‘
<collectorName id="26D16BAE5403CC79FB89F90FFA8EF970" box="[1048,1366,1751,1778]" pageId="38" pageNumber="189">
<collectingMunicipality id="6BFF94025403CC79FB89F90FFB74F970" box="[1048,1196,1751,1778]" pageId="38" pageNumber="189">United Prov.</collectingMunicipality>
, N. India. B.M.
</collectorName>
1924- 335.’, ‘
<location id="8EFB58A35403CC79FF72F923FE7DF897" LSID="urn:lsid:plazi:treatment:038DBF6E5403CC76FF06FB3CFAE4FC97:8EFB58A35403CC79FF72F923FE7DF897" box="[227,421,1787,1813]" country="India" municipality="United Prov." name="Woodstock Falls" pageId="38" pageNumber="189" stateProvince="Uttarakhand">Woodstock Falls</location>
,
<collectorName id="26D16BAE5403CC79FE20F923FD15F897" box="[433,717,1787,1814]" pageId="38" pageNumber="189">Mussoorie. Dr. Cameron.</collectorName>
<date id="FF9A28B85403CC79FD43F923FC82F897" box="[722,858,1787,1813]" pageId="38" pageNumber="189" value="1921-03-27">
<collectingDate id="EFDED1505403CC79FD43F923FC82F897" box="[722,858,1787,1813]" pageId="38" pageNumber="189" value="1921-03-27">27.III.1921</collectingDate>
</date>
.’, ‘
<taxonomicName id="4C2475FB5403CC79FCE2F923FB1FF897" authority=", Cam. Co." authorityName="Cam. Co." box="[883,1223,1787,1813]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="cariosum">Omalium cariosum, Cam. Co.</taxonomicName>
type’, ‘locality not mentioned in description!
<collectorName id="26D16BAE5403CC79FE2FF8F8FD97F8BB" box="[446,591,1823,1850]" pageId="38" pageNumber="189">M.K. Thayer</collectorName>
1989’, ‘NHMUK014663258’ (BMNH)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF35403CC76FF56F89BFDD7FE0B" lastPageId="41" lastPageNumber="192" pageId="38" pageNumber="189" type="description">
<paragraph id="8B9B0E785403CC79FF56F89BFB60F827" blockId="38.[151,1437,1427,1958]" pageId="38" pageNumber="189">
<emphasis id="B950D26A5403CC79FF56F89BFEADF8DF" bold="true" box="[199,373,1859,1885]" pageId="38" pageNumber="189">Redescription.</emphasis>
Measurements (n=3): HW: 0.42–0.46; HL: 0.31–0.33; OL: 0.14; TL: 0.06; AL (
<typeStatus id="549FB0DA5403CC79FAB2F89BFA49F8DF" box="[1315,1425,1859,1885]" pageId="38" pageNumber="189" type="lectotype">lectotype</typeStatus>
): 0.83; PL: 0.40; PWmax: 0.57; PWmin: 0.49; ESL: 0.72; EW: 0.72–0.75; MTbL (
<typeStatus id="549FB0DA5403CC79FB84F8BFFB5BF803" box="[1045,1155,1895,1921]" pageId="38" pageNumber="189" type="lectotype">lectotype</typeStatus>
): 0.35; MTrL (
<typeStatus id="549FB0DA5403CC79FAB2F8BFFA49F803" box="[1315,1425,1895,1921]" pageId="38" pageNumber="189" type="lectotype">lectotype</typeStatus>
): 0.17 (MTrL 1–4: 0.07; MTrL 5: 0.10); AW: 0.72–0.75; AedL: 0.52; BL: 2.85–3.15 (
<typeStatus id="549FB0DA5403CC79FBAEF853FB74F827" box="[1087,1196,1931,1957]" pageId="38" pageNumber="189" type="lectotype">lectotype</typeStatus>
).
</paragraph>
<caption id="DF5B5EF05402CC78FF06FBE5FE0DFBFA" ID-DOI="http://doi.org/10.5281/zenodo.10146220" ID-Zenodo-Dep="10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="39" pageNumber="190" startId="39.[151,264,1085,1109]" targetBox="[159,1419,187,1052]" targetPageId="39" targetType="figure">
<paragraph id="8B9B0E785402CC78FF06FBE5FE0DFBFA" blockId="39.[151,1436,1085,1145]" pageId="39" pageNumber="190">
<emphasis id="B950D26A5402CC78FF06FBE5FE81FBD7" bold="true" box="[151,345,1085,1109]" pageId="39" pageNumber="190">FIGURES 70–72.</emphasis>
Habitus of
<taxonomicName id="4C2475FB5402CC78FE4CFBE5FDE3FBD7" authorityName="Gravenhorst" authorityYear="1802" box="[477,571,1085,1109]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="B950D26A5402CC78FE4CFBE5FDE3FBD7" box="[477,571,1085,1109]" italics="true" pageId="39" pageNumber="190">Omalium</emphasis>
</taxonomicName>
: 70—
<taxonomicName id="4C2475FB5402CC78FD11FBE5FCDAFBD7" box="[640,770,1085,1109]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="190" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A5402CC78FD11FBE5FCDAFBD7" box="[640,770,1085,1109]" italics="true" pageId="39" pageNumber="190">O. cariosum</emphasis>
</taxonomicName>
(lectotype), 71—
<taxonomicName id="4C2475FB5402CC78FC2FFBE5FBA2FBD7" authorityName="Shavrin" authorityYear="2023" box="[958,1146,1085,1109]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="190" phylum="Arthropoda" rank="species" species="flavotestaceum">
<emphasis id="B950D26A5402CC78FC2FFBE5FBA2FBD7" box="[958,1146,1085,1109]" italics="true" pageId="39" pageNumber="190">O. flavotestaceum</emphasis>
</taxonomicName>
(paratype), 72—
<taxonomicName id="4C2475FB5402CC78FAA0FBE5FA44FBD7" authorityName="Shavrin" authorityYear="2023" box="[1329,1436,1085,1109]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="190" phylum="Arthropoda" rank="species" species="kumari">
<emphasis id="B950D26A5402CC78FAA0FBE5FA9BFBD6" box="[1329,1347,1085,1108]" italics="true" pageId="39" pageNumber="190">O</emphasis>
.
<emphasis id="B950D26A5402CC78FAC5FBE5FA44FBD7" box="[1364,1436,1085,1109]" italics="true" pageId="39" pageNumber="190">kumari</emphasis>
</taxonomicName>
(paratype). Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B9B0E785402CC78FF56FB13FB29F9AB" blockId="39.[151,1437,1227,2009]" pageId="39" pageNumber="190">
Habitus as in
<figureCitation id="131F12FD5402CC78FEE1FB13FE1FFB67" box="[368,455,1227,1253]" captionStart="FIGURES 70–72" captionStartId="39.[151,264,1085,1109]" captionTargetBox="[159,1419,187,1052]" captionTargetId="figure-16@39.[151,1436,181,1061]" captionTargetPageId="39" captionText="FIGURES 70–72. Habitus of Omalium: 70—O. cariosum (lectotype), 71—O. flavotestaceum (paratype), 72—O. kumari (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="39" pageNumber="190">Fig. 70</figureCitation>
. Head, pronotum and abdomen reddish-brown (head slightly darker; apical tergites of abdomen and paratergites distinctly paler); lateral and basal portions of pronotum and elytra yellow-brown; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellowish. Punctation of head dense and moderately large, denser in middle and on infraorbital portions (
<typeStatus id="549FB0DA5402CC78FCE7FAEFFBD7FAD3" box="[886,1039,1335,1361]" pageId="39" pageNumber="190" type="paralectotype">paralectotype</typeStatus>
with slightly sparser punctation in middle); punctation of pronotum about as that in middle portion of head or slightly larger and deeper, distinctly finer and sparser in mediobasal portion; scutellum without punctures; punctation of elytra dense, about as that on head or slightly coarser, slightly finer and sparser along suture; abdominal tergites without or with very indistinct, sparse and fine punctation. Apical portion of clypeus with fine trasverse meshes, latero-apical portion of clypeus between middle and supra-antennal portions with subdiagonal meshes, middle portion of head between eyes and infraorbital portions without distinct microsculpture; pronotum, scutellum and elytra without microreticulation.
</paragraph>
<paragraph id="8B9B0E785402CC78FF56F9EBFE98F85B" blockId="39.[151,1437,1227,2009]" pageId="39" pageNumber="190">
Head with supra-antennal elevations indistinctly elevated and moderately narrow clypeus stretching posteriad toward level of middle length of eyes. Lateral portions of clypeus with moderately shallow and long anteriomedian depressions. Middle portion between eyes slightly and posterior parts of infraorbital portions distinctly elevated. Middle surface between eyes with narrow diagonal and infraorbital portions with irregular longitudinal elevations between punctures (
<typeStatus id="549FB0DA5402CC78FE10F91BFDC2F95F" box="[385,538,1731,1757]" pageId="39" pageNumber="190" type="paralectotype">paralectotype</typeStatus>
without sculpture); each lateral portion near posterior margins of eyes with indistinct one to two elongate wrinkles. Anteocellar foveae deep and relatively long, slightly convergent latero-apicad toward level of middle length of eyes. Temples very short, distinctly more than twice shorter than longitudinal length of eye, strongly narrowed posteriad toward nuchal constriction. Ocelli located about at level of postocular ridges, distance between ocelli slightly less than twice as distance between ocellus and posterior margin of eye. Antennomere 3 about as long as 2, 4 small, about twice shorter than 3, 5 slightly broader than 4, 6 slightly broader and longer than 5, 7 slightly broader than 6, 8 indistinctly broader and longer than 7, 9–10 distinctly longer and broader than 8.
</paragraph>
<caption id="DF5B5EF0540DCC77FF06FA4EFD4DFA50" ID-DOI="http://doi.org/10.5281/zenodo.10146222" ID-Zenodo-Dep="10146222" httpUri="https://zenodo.org/record/10146222/files/figure.png" pageId="40" pageNumber="191" startId="40.[151,264,1430,1454]" targetBox="[169,1433,187,1403]" targetPageId="40" targetType="figure">
<paragraph id="8B9B0E78540DCC77FF06FA4EFD4DFA50" blockId="40.[151,1436,1430,1490]" pageId="40" pageNumber="191">
<emphasis id="B950D26A540DCC77FF06FA4EFE8BFA2C" bold="true" box="[151,339,1430,1454]" pageId="40" pageNumber="191">FIGURES 73–75.</emphasis>
Aedeagus of
<taxonomicName id="4C2475FB540DCC77FE4AFA4EFD44FA2C" box="[475,668,1430,1454]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="191" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A540DCC77FE4AFA4EFD44FA2C" box="[475,668,1430,1454]" italics="true" pageId="40" pageNumber="191">Omalium cariosum</emphasis>
</taxonomicName>
(paralectotype): 73—aedeagus, parameral view, 74—aedeagus, lateral view, 75—median lobe, lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B9B0E78540DCC77FF56F9FDFEAFF928" blockId="40.[151,1436,1572,2031]" pageId="40" pageNumber="191">Pronotum 1.4 times as broad as long, 1.2–1.3 times as broad as head, widest in middle, gradually narrowed posteriad, with widely rounded apical angles not protruded anteriad. Surface of disc with two indistinct wide longitudinal depressions. Surface with distinct diagonal elevations in middle and between longitudinal depression and lateral portions.</paragraph>
<paragraph id="8B9B0E78540DCC77FF56F96CFCE7F970" blockId="40.[151,1436,1572,2031]" pageId="40" pageNumber="191">
Elytra disitinctly broader than long, 1.8 times as long as pronotum, subparallel. Medioapical portion of each elytron with slightly elevated between punctures (
<typeStatus id="549FB0DA540DCC77FD57F900FCEBF970" box="[710,819,1752,1778]" pageId="40" pageNumber="191" type="lectotype">lectotype</typeStatus>
).
</paragraph>
<paragraph id="8B9B0E78540DCC77FF56F924FDA0F894" blockId="40.[151,1436,1572,2031]" box="[199,632,1788,1814]" pageId="40" pageNumber="191">Metatarsi twice shorter than metatibia.</paragraph>
<paragraph id="8B9B0E78540DCC77FF56F8F8FD95F8B8" blockId="40.[151,1436,1572,2031]" box="[199,589,1824,1851]" pageId="40" pageNumber="191">Abdomen about as broad as elytra.</paragraph>
<paragraph id="8B9B0E78540DCC76FF56F89CFDC9FE9F" blockId="40.[151,1436,1572,2031]" lastBlockId="41.[151,1437,151,789]" lastPageId="41" lastPageNumber="192" pageId="40" pageNumber="191">
Male. Antennomeres 9–10 strongly transverse. Posterior margins of abdominal tergite VIII slightly and sternite VIII distinctly and widely sinuate. Aedeagus with wide basal portion, gradually narrowed toward middle, with elongate and moderately narrow median lobe, from basal portion significantly narrowed apicad and from about middle forming elongate spear-shaped apical part; mediolateral portions with elongate sclerotized accessory plates slightly curved and rounded apically; parameres very wide, significantly shorter than apex of median lobe, with narrow and elongate apical portions and sclerotized inner lateral margins, each paramere with two long apical and one short preapical setae; internal sac moderately narrow and long, with narrow elongate sclerotized structure in middle of basal part (
<figureCitation id="131F12FD540CCC76FE1BFF07FE07FF7B" box="[394,479,223,249]" captionStart="FIGURES 73–75" captionStartId="40.[151,264,1430,1454]" captionTargetBox="[169,1433,187,1403]" captionTargetId="figure-22@40.[151,1436,181,1406]" captionTargetPageId="40" captionText="FIGURES 73–75. Aedeagus of Omalium cariosum (paralectotype): 73—aedeagus, parameral view, 74—aedeagus, lateral view, 75—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146222" httpUri="https://zenodo.org/record/10146222/files/figure.png" pageId="41" pageNumber="192">Fig. 73</figureCitation>
). Lateral aspect of aedeagus as in
<figureCitation id="131F12FD540CCC76FCF5FF07FC60FF7B" box="[868,952,223,249]" captionStart="FIGURES 73–75" captionStartId="40.[151,264,1430,1454]" captionTargetBox="[169,1433,187,1403]" captionTargetId="figure-22@40.[151,1436,181,1406]" captionTargetPageId="40" captionText="FIGURES 73–75. Aedeagus of Omalium cariosum (paralectotype): 73—aedeagus, parameral view, 74—aedeagus, lateral view, 75—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146222" httpUri="https://zenodo.org/record/10146222/files/figure.png" pageId="41" pageNumber="192">Fig. 74</figureCitation>
; median lobe with subacute hook in apical part, with rounded apex (
<figureCitation id="131F12FD540CCC76FE21FEDBFDDDFE9F" box="[432,517,258,285]" captionStart="FIGURES 73–75" captionStartId="40.[151,264,1430,1454]" captionTargetBox="[169,1433,187,1403]" captionTargetId="figure-22@40.[151,1436,181,1406]" captionTargetPageId="40" captionText="FIGURES 73–75. Aedeagus of Omalium cariosum (paralectotype): 73—aedeagus, parameral view, 74—aedeagus, lateral view, 75—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146222" httpUri="https://zenodo.org/record/10146222/files/figure.png" pageId="41" pageNumber="192">Fig. 75</figureCitation>
).
</paragraph>
<paragraph id="8B9B0E78540CCC76FF56FEFFFDD7FE0B" blockId="41.[151,1437,151,789]" pageId="41" pageNumber="192">
Female. Antennomeres 9–10 less transverse. Posterior margins of abdominal tergite VIII and sternite VIII truncate. Accessory sclerite with very wide basal portion, gradually narrowed apicad toward subacute angle (
<figureCitation id="131F12FD540CCC76FAFEFE93FF6FFE0B" captionStart="FIGURES 76–83" captionStartId="41.[151,264,1651,1675]" captionTargetBox="[154,1432,816,1625]" captionTargetId="figure-275@41.[151,1435,813,1628]" captionTargetPageId="41" captionText="FIGURES 76–83. Female accessory sclerite (Figs 76–77, 79, 81–82) and spermatheca (Figs 78, 80, 83) of Omalium: 76— O.cariosum, 77–78—O. flavotestaceum, 79–80—O. kumari, 81—O. latocavum, 82–83—O. secretum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146224" httpUri="https://zenodo.org/record/10146224/files/figure.png" pageId="41" pageNumber="192">Fig. 76</figureCitation>
). Spermatheca not recognized.
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540CCC76FF56FE4AFDD6FD9B" pageId="41" pageNumber="192" type="diagnosis">
<paragraph id="8B9B0E78540CCC76FF56FE4AFDD6FD9B" blockId="41.[151,1437,151,789]" pageId="41" pageNumber="192">
<emphasis id="B950D26A540CCC76FF56FE4AFE68FE2F" bold="true" box="[199,432,402,429]" pageId="41" pageNumber="192">Comparative notes.</emphasis>
Based on the general shape of temples, gradually narrowed toward nuchal constriction,
<taxonomicName id="4C2475FB540CCC76FA10FE4BFF27FE52" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="192" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A540CCC76FA10FE4BFA4DFE2F" box="[1409,1429,403,429]" italics="true" pageId="41" pageNumber="192">O</emphasis>
.
<emphasis id="B950D26A540CCC76FF06FE6FFF27FE52" box="[151,255,439,464]" italics="true" pageId="41" pageNumber="192">cariosum</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C2475FB540CCC76FE1EFE6FFD80FE53" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[399,600,439,465]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="192" phylum="Arthropoda" rank="species" species="flavotestaceum" status="sp. n.">
<emphasis id="B950D26A540CCC76FE1EFE6FFE7BFE53" box="[399,419,439,465]" italics="true" pageId="41" pageNumber="192">O</emphasis>
.
<emphasis id="B950D26A540CCC76FE21FE6FFD80FE53" box="[432,600,439,465]" italics="true" pageId="41" pageNumber="192">flavotestaceum</emphasis>
</taxonomicName>
<emphasis id="B950D26A540CCC76FDCFFE6FFD45FE53" bold="true" box="[606,669,439,465]" pageId="41" pageNumber="192">
<taxonomicNameLabel id="A2636F11540CCC76FDCFFE6FFD45FE53" box="[606,669,439,465]" pageId="41" pageNumber="192" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from which it can be distinguished by the distinctly shorter temples, the shape of the pronotum widest in the middle portion, slightly narrower elytra, and details of the morphology of the aedeagus (see the key above).
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540CCC76FF56FDFAFB34FDBF" box="[199,1260,546,573]" pageId="41" pageNumber="192" type="distribution">
<paragraph id="8B9B0E78540CCC76FF56FDFAFB34FDBF" blockId="41.[151,1437,151,789]" box="[199,1260,546,573]" pageId="41" pageNumber="192">
<emphasis id="B950D26A540CCC76FF56FDFAFEB9FDBE" bold="true" box="[199,353,546,572]" pageId="41" pageNumber="192">Distribution.</emphasis>
<taxonomicName id="4C2475FB540CCC76FEF6FDFBFDE7FDBE" box="[359,575,547,573]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="192" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A540CCC76FEF6FDFBFDE7FDBE" box="[359,575,547,573]" italics="true" pageId="41" pageNumber="192">Omalium cariosum</emphasis>
</taxonomicName>
is known from two localities in
<collectingRegion id="49E0C09A540CCC76FC3BFDFBFBEFFDBF" box="[938,1079,547,573]" country="India" name="Uttarakhand" pageId="41" pageNumber="192">Uttarakhand</collectingRegion>
,
<collectingCountry id="F3334EE8540CCC76FBD3FDFBFBA4FDBF" box="[1090,1148,547,573]" name="India" pageId="41" pageNumber="192">India</collectingCountry>
(
<figureCitation id="131F12FD540CCC76FB1AFDFBFB07FDBF" box="[1163,1247,547,573]" captionStart="FIGURE 19" captionStartId="7.[151,250,1891,1915]" captionTargetBox="[151,1435,957,1868]" captionTargetId="figure-299@7.[151,1435,957,1868]" captionTargetPageId="7" captionText="FIGURE 19. Distribution of Omalium in the Himalayan region: O. acutangulum sp. n. (circles), O. amplissimum sp. n. (star), O. beesoni (squares), O. cariosum (diamonds), O. flavotestaceum sp. n. (triangles)." figureDoi="http://doi.org/10.5281/zenodo.10146178" httpUri="https://zenodo.org/record/10146178/files/figure.png" pageId="41" pageNumber="192">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540CCC76FF56FD9EFEDFFD07" pageId="41" pageNumber="192" type="biology_ecology">
<paragraph id="8B9B0E78540CCC76FF56FD9EFEDFFD07" blockId="41.[151,1437,151,789]" pageId="41" pageNumber="192">
<emphasis id="B950D26A540CCC76FF56FD9EFE93FDE2" bold="true" box="[199,331,582,608]" pageId="41" pageNumber="192">Bionomics.</emphasis>
<typeStatus id="549FB0DA540CCC76FEC2FD9FFE53FDE3" box="[339,395,583,609]" pageId="41" pageNumber="192">Type</typeStatus>
specimens were collected at elevation about
<quantity id="4CDCA39D540CCC76FC04FD9FFC34FDE3" box="[917,1004,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="41" pageNumber="192" unit="m" value="1980.0">1980 m</quantity>
a.s.l. The detailed binomical data are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33E5DF3540CCC76FF56FD57FAE4FC97" pageId="41" pageNumber="192" type="discussion">
<paragraph id="8B9B0E78540CCC76FF56FD57FAE4FC97" blockId="41.[151,1437,151,789]" pageId="41" pageNumber="192">
<emphasis id="B950D26A540CCC76FF56FD57FEE3FD2B" bold="true" box="[199,315,655,681]" pageId="41" pageNumber="192">Remarks.</emphasis>
<taxonomicName id="4C2475FB540CCC76FED2FD57FDAAFD2A" box="[323,626,655,681]" class="Insecta" family="Staphylinidae" genus="Omalium" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="192" phylum="Arthropoda" rank="species" species="cariosum">
<emphasis id="B950D26A540CCC76FED2FD57FE73FD2B" box="[323,427,655,681]" italics="true" pageId="41" pageNumber="192">Omalium</emphasis>
(s. str.)
<emphasis id="B950D26A540CCC76FD9BFD57FDAAFD2A" box="[522,626,655,680]" italics="true" pageId="41" pageNumber="192">cariosum</emphasis>
</taxonomicName>
was originally described from “Dehra
<collectingRegion id="49E0C09A540CCC76FBBEFD57FBBCFD2B" box="[1071,1124,655,681]" country="United Kingdom" name="Down" pageId="41" pageNumber="192">Dun</collectingRegion>
: Mussorie,
<quantity id="4CDCA39D540CCC76FB76FD56FA89FD2B" box="[1255,1361,654,681]" metricMagnitude="3" metricUnit="m" metricValue="1.9812" pageId="41" pageNumber="192" unit="ft" value="6500.0">6500 feet</quantity>
above the sea” based on the unspecified number of the
<typeStatus id="549FB0DA540CCC76FD22FD6BFCC1FD4F" box="[691,793,691,717]" pageId="41" pageNumber="192" type="syntype">syntypes</typeStatus>
. I designate here the female (
<figureCitation id="131F12FD540CCC76FBCCFD6BFB76FD4F" box="[1117,1198,691,717]" captionStart="FIGURES 70–72" captionStartId="39.[151,264,1085,1109]" captionTargetBox="[159,1419,187,1052]" captionTargetId="figure-16@39.[151,1436,181,1061]" captionTargetPageId="39" captionText="FIGURES 70–72. Habitus of Omalium: 70—O. cariosum (lectotype), 71—O. flavotestaceum (paratype), 72—O. kumari (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10146220" httpUri="https://zenodo.org/record/10146220/files/figure.png" pageId="41" pageNumber="192">Fig. 70</figureCitation>
) with the round label “Type H.T.” as the
<typeStatus id="549FB0DA540CCC76FEE3FD0FFE01FD73" box="[370,473,727,753]" pageId="41" pageNumber="192" type="lectotype">lectotype</typeStatus>
in order to fix the identity of the name. The female with the round label with yellow margins (from “Woodstock Falls, Mussoorie”) does not belong to the type series due the different locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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