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<document id="BEA1EA5280884AABD634BA6F7513444E" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="03877623626CFFC82DEFB681FC455FF3" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Crossoglossa) cruralis Andrewes, Borneo. E 1933" docType="treatment" docVersion="8" lastPageNumber="44" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="43" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="261C4C23C4E95ED2EB19C55DE9C399D7">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="FFE2DABD4DA067C7E807949A4DE5573D">Shi, Hongliang</mods:namePart>
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<subSubSection id="C33494BE626CFFC92DEFB681FCE3584E" box="[151,772,1273,1300]" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8B91C735626CFFC92DEFB681FCE3584E" blockId="42.[151,772,1273,1336]" box="[151,772,1273,1300]" pageId="42" pageNumber="43">
<emphasis id="B95A1B27626CFFC92DEFB681FCE3584E" bold="true" box="[151,772,1273,1300]" pageId="42" pageNumber="43">
[8]
<taxonomicName id="4C2EBCB6626CFFC92DB9B681FCE3584E" authority="Andrewes, 1933" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[193,772,1273,1300]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="cruralis" subGenus="Crossoglossa">
<emphasis id="B95A1B27626CFFC92DB9B681FEFE584E" bold="true" box="[193,281,1274,1300]" italics="true" pageId="42" pageNumber="43">Parena</emphasis>
(
<emphasis id="B95A1B27626CFFC92C52B682FE2F584E" bold="true" box="[298,456,1273,1300]" italics="true" pageId="42" pageNumber="43">Crossoglossa</emphasis>
)
<emphasis id="B95A1B27626CFFC92CA1B682FDDF584E" bold="true" box="[473,568,1273,1300]" italics="true" pageId="42" pageNumber="43">cruralis</emphasis>
<bibRefCitation id="EFBFBAC4626CFFC92F45B681FCE3584E" author="Andrewes, H. E." box="[573,772,1274,1300]" pageId="42" pageNumber="43" pagination="273 - 286" refId="ref109750" refString="Andrewes, H. E. (1933) On some new species of Carabidae, chiefly from Java. Treubia, 14 (2), 273 - 286." type="journal article" year="1933">Andrewes, 1933</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626CFFC92DEFB765FE785862" box="[151,415,1310,1336]" pageId="42" pageNumber="43" type="description">
<paragraph id="8B91C735626CFFC92DEFB765FE785862" blockId="42.[151,772,1273,1336]" box="[151,415,1310,1336]" pageId="42" pageNumber="43">
Habitus:
<figureCitation id="1315DBB0626CFFC92D85B765FE7C5862" box="[253,411,1310,1336]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="42" pageNumber="43">Figs 21H, 21I</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626CFFC92DEFB726FE3058CE" pageId="42" pageNumber="43" type="reference_group">
<paragraph id="8B91C735626CFFC92DEFB726FE3058CE" blockId="42.[151,1436,1373,1428]" pageId="42" pageNumber="43">
<bibRefCitation id="EFBFBAC4626CFFC92DEFB726FE96582F" author="Andrewes, H. E." box="[151,369,1373,1397]" pageId="42" pageNumber="43" pagination="273 - 286" refId="ref109750" refString="Andrewes, H. E. (1933) On some new species of Carabidae, chiefly from Java. Treubia, 14 (2), 273 - 286." type="journal article" year="1933">Andrewes, 1933: 286</bibRefCitation>
(
<taxonomicName id="4C2EBCB6626CFFC92CF8B725FD42582F" ID-CoL="7HQ4S" authorityName="Andrewes" authorityYear="1935" box="[384,677,1373,1397]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="variety" species="testacea" variety="cruralis">
<emphasis id="B95A1B27626CFFC92CF8B725FDC4582F" box="[384,547,1374,1397]" italics="true" pageId="42" pageNumber="43">Parena testacea</emphasis>
var.
<emphasis id="B95A1B27626CFFC92F2DB726FD42582F" box="[597,677,1373,1397]" italics="true" pageId="42" pageNumber="43">cruralis</emphasis>
</taxonomicName>
; type locality:
<collectingRegion id="49EA09D7626CFFC92E46B725FC88582F" box="[830,879,1374,1397]" country="Indonesia" name="Jawa Timur" pageId="42" pageNumber="43">Java</collectingRegion>
: Pengalengan;
<typeStatus id="54957997626CFFC92970B726FB86582F" box="[1032,1121,1373,1397]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
in NHML);
<bibRefCitation id="EFBFBAC4626CFFC9299AB725FA7B582F" author="Lorenz, W." box="[1250,1436,1373,1397]" pageId="42" pageNumber="43" refId="ref112064" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world (Insecta Coleoptera Geadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Tutzing: Wolfgang Lorenz, 530 pp." type="book" year="2005">Lorenz, 2005: 490</bibRefCitation>
(
<taxonomicName id="4C2EBCB6626CFFC92DA8B707FE2E58CE" ID-CoL="7HQ4S" authorityName="Andrewes" authorityYear="1935" box="[208,457,1404,1428]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="testacea" subSpecies="cruralis">
<emphasis id="B95A1B27626CFFC92DA8B707FE2E58CE" box="[208,457,1404,1428]" italics="true" pageId="42" pageNumber="43">Parena testacea cruralis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626CFFC92DEFB7C6FDBC5B88" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8B91C735626CFFC92DEFB7C6FEAF5B19" blockId="42.[151,1437,1468,2035]" pageId="42" pageNumber="43">
<emphasis id="B95A1B27626CFFC92DEFB7C6FE5C588C" bold="true" box="[151,443,1468,1495]" pageId="42" pageNumber="43">Type material examined</emphasis>
.
<emphasis id="B95A1B27626CFFC92CB3B7C6FC65588D" bold="true" box="[459,898,1468,1495]" pageId="42" pageNumber="43">
<taxonomicName id="4C2EBCB6626CFFC92CB3B7C6FCE0588C" authority="Andrewes" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[459,775,1468,1495]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B95A1B27626CFFC92CB3B7C6FD65588D" bold="true" box="[459,642,1468,1495]" italics="true" pageId="42" pageNumber="43">Parena cruralis</emphasis>
Andrewes
</taxonomicName>
:
<typeStatus id="54957997626CFFC92E6DB7C6FC65588D" box="[789,898,1469,1495]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
</emphasis>
(NHML,
<figureCitation id="1315DBB0626CFFC92E8EB7C7FBB8588D" box="[1014,1119,1468,1495]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="42" pageNumber="43">Fig. 21H</figureCitation>
): female, body length =
<quantity id="4CD66AD0626CFFC92801B7C6FF2258A1" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="42" pageNumber="43" unit="mm" value="9.4">9.4 mm</quantity>
, board mounted, &quot;
<collectingRegion id="49EA09D7626CFFC92CEDB79AFE2D58A1" box="[405,458,1505,1531]" country="Indonesia" name="Jawa Timur" pageId="42" pageNumber="43">Java</collectingRegion>
occident. / Pengalengan / 4000 1893 / H. Fruhstorfer&quot;, &quot;Ex coll. / T. Sloane&quot;, &quot;
<emphasis id="B95A1B27626CFFC92818B79BFA6D58A0" box="[1376,1418,1504,1530]" italics="true" pageId="42" pageNumber="43">169</emphasis>
&quot;, &quot;Type&quot; [red label]; &quot;
<emphasis id="B95A1B27626CFFC92C04B47EFC7D5B44" box="[380,922,1540,1567]" italics="true" pageId="42" pageNumber="43">
<taxonomicName id="4C2EBCB6626CFFC92C04B47EFE2A5B44" box="[380,461,1541,1566]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/
<taxonomicName id="4C2EBCB6626CFFC92C9DB47EFD995B44" authority="Chd." authorityName="Chd." box="[485,638,1540,1566]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="testacea">testacea Chd.</taxonomicName>
/ v. Cruralis Andr. / Type
</emphasis>
H.E. Andrewes det.&quot;, &quot;H.E. Andrewes Coll. / B.M. 1945-97&quot;.
</paragraph>
<paragraph id="8B91C735626CFFC92DBFB436FDBC5B88" blockId="42.[151,1437,1468,2035]" pageId="42" pageNumber="43">
<emphasis id="B95A1B27626CFFC92DBFB436FDCE5B3C" bold="true" box="[199,553,1612,1639]" pageId="42" pageNumber="43">Non-type material examined</emphasis>
.
<materialsCitation id="3B46CD68626CFFC92F45B436FA605BD1" ID-GBIF-Occurrence="4115871503" collectingDate="2016-05-01" county="Boreno" elevation="1182" latitude="5.443" location="Keningau district" longLatPrecision="7" longitude="116.4512" municipality="Sahah" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626CFFC92F45B436FD425B3C" box="[573,677,1612,1639]" pageId="42" pageNumber="43" type="female">1 female</specimenCount>
(IZAS), &quot;
<collectingCounty id="62F0BFB9626CFFC92E58B436FC9D5B3D" box="[800,890,1613,1639]" pageId="42" pageNumber="43">Boreno</collectingCounty>
:
<collectingMunicipality id="6BF55D4F626CFFC92EF3B437FC355B3C" box="[907,978,1612,1638]" pageId="42" pageNumber="43">Sahah</collectingMunicipality>
,
<location id="8EF191EE626CFFC92E9CB437FB575B3C" LSID="urn:lsid:plazi:treatment:03877623626CFFC82DEFB681FC455FF3:8EF191EE626CFFC92E9CB437FB575B3C" box="[996,1200,1612,1638]" county="Boreno" latitude="5.443" longLatPrecision="7" longitude="116.4512" municipality="Sahah" name="Keningau district" pageId="42" pageNumber="43">Keningau district</location>
, Jungle Girl Camp.
<geoCoordinate id="EE1AA1F2626CFFC92DEFB40BFF1B5BD1" box="[151,252,1648,1675]" degrees="5.4430" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="5" value="5.443">N5.4430</geoCoordinate>
,
<geoCoordinate id="EE1AA1F2626CFFC92C7FB40BFE635BD1" box="[263,388,1648,1675]" degrees="116.4512" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="5" value="116.4512">E116.4512</geoCoordinate>
,
<quantity id="4CD66AD0626CFFC92CF7B40AFE005BD1" box="[399,487,1649,1675]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="42" pageNumber="43" unit="m" value="1182.0">
<elevation id="00032006626CFFC92CF7B40AFE005BD1" box="[399,487,1649,1675]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="42" pageNumber="43" unit="m" value="1182.0">1182 m</elevation>
</quantity>
; Shi H.L. &amp; Liu Y. lgt.
<collectingMethod id="526FBF22626CFFC92F8BB40BFCBA5BD1" box="[755,861,1648,1675]" pageId="42" pageNumber="43">light trap</collectingMethod>
, Ins. Zoo., CAS.
<date id="FF90E1F5626CFFC9295BB40BFB655BD1" box="[1059,1154,1648,1675]" pageId="42" pageNumber="43" value="2016-05-01">
<collectingDate id="EFD4181D626CFFC9295BB40BFB655BD1" box="[1059,1154,1648,1675]" pageId="42" pageNumber="43" value="2016-05-01">2016.V.1</collectingDate>
</date>
N&quot; &lt;
<figureCitation id="1315DBB0626CFFC929C6B40BFAFC5BD1" box="[1214,1307,1648,1675]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="42" pageNumber="43">Figs 8D</figureCitation>
,
<figureCitation id="1315DBB0626CFFC92850B40AFAB35BD1" box="[1320,1364,1649,1675]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. AH Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. IL Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="42" pageNumber="43">14F</figureCitation>
,
<figureCitation id="1315DBB0626CFFC92818B40AFA605BD1" box="[1376,1415,1649,1675]" captionStart="FIGURE 21" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[219,1375,184,1823]" captionTargetId="figure-28@38.[212,1375,181,1827]" captionTargetPageId="38" captionText="FIGURE 21. Habitus of Parena (Crossoglossa) testacea species group. A Lectotype of Parena testacea (Chaudoir). B P. testacea (Chaudoir), male from Guangdong. C Holotype of Parena sulawesiensis Kirschenhofer. D P. sulawesiensis Kirschenhofer, male from Sulawesi. E Holotype of Parena mellea (Chaudoir). F P. mellea (Chaudoir), male from Maluku. G P. mellea (Chaudoir), female from Philippine. H Holotype of Parena cruralis Andrewes. I P. cruralis Andrewes, female from Borneo. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958979" httpUri="https://zenodo.org/record/7958979/files/figure.png" pageId="42" pageNumber="43">21I</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD68626CFFC92DEFB4EEFC0B5BF5" ID-GBIF-Occurrence="4115871567" box="[151,1004,1684,1711]" collectorName="N. Borneo" elevation="1150" location="Tambunan distr." pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC626CFFC92DEFB4EEFF1D5BF4" box="[151,250,1684,1711]" pageId="42" pageNumber="43" type="female">1 female</specimenCount>
(CRS), &quot;
<collectorName id="26DBA2E3626CFFC92C1CB4EEFE3A5BF5" box="[356,477,1685,1711]" pageId="42" pageNumber="43">N. Borneo</collectorName>
<quantity id="4CD66AD0626CFFC92C9EB4EFFDDB5BF5" box="[486,572,1684,1711]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="42" pageNumber="43" unit="m" value="1150.0">
<elevation id="00032006626CFFC92C9EB4EFFDDB5BF5" box="[486,572,1684,1711]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="42" pageNumber="43" unit="m" value="1150.0">1150 m</elevation>
</quantity>
<location id="8EF191EE626CFFC92F3DB4EFFD1C5BF4" LSID="urn:lsid:plazi:treatment:03877623626CFFC82DEFB681FC455FF3:8EF191EE626CFFC92F3DB4EFFD1C5BF4" box="[581,763,1684,1710]" name="Tambunan distr." pageId="42" pageNumber="43">Tambunan distr.</location>
Trus Madi m.IV.12&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68626CFFC92E80B4EEFDB05B88" ID-GBIF-Occurrence="4115871310" collectingDate="2007-03" country="Malaysia" location="Trus-Madi" municipality="Borneo" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="9D280CBC626CFFC92E80B4EEFBBC5BF4" box="[1016,1115,1684,1711]" pageId="42" pageNumber="43" type="female">1 female</specimenCount>
(CRS), &quot;
<collectingCountry id="F33987A5626CFFC929BBB4EFFACC5BF4" box="[1219,1323,1684,1710]" name="Malaysia" pageId="42" pageNumber="43">Malaysia</collectingCountry>
<collectingMunicipality id="6BF55D4F626CFFC92831B4EFFA7A5BF4" box="[1353,1437,1684,1710]" pageId="42" pageNumber="43">Borneo</collectingMunicipality>
<collectingRegion id="49EA09D7626CFFC92DEFB4C3FF395B88" box="[151,222,1720,1746]" country="Malaysia" name="Sabah" pageId="42" pageNumber="43">Sabah</collectingRegion>
,
<location id="8EF191EE626CFFC92D91B4C3FE875B88" LSID="urn:lsid:plazi:treatment:03877623626CFFC82DEFB681FC455FF3:8EF191EE626CFFC92D91B4C3FE875B88" box="[233,352,1720,1746]" country="Malaysia" municipality="Borneo" name="Trus-Madi" pageId="42" pageNumber="43" stateProvince="Sabah">Trus-Madi</location>
<date id="FF90E1F5626CFFC92C1FB4C2FE215B89" box="[359,454,1721,1747]" pageId="42" pageNumber="43" value="2007-03">
<collectingDate id="EFD4181D626CFFC92C1FB4C2FE215B89" box="[359,454,1721,1747]" pageId="42" pageNumber="43" value="2007-03">III.2007</collectingDate>
</date>
. Gromenko&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626CFFC92DBFB4A7FCEF5A39" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8B91C735626CFFC92DBFB4A7FCEF5A39" blockId="42.[151,1437,1468,2035]" pageId="42" pageNumber="43">
<emphasis id="B95A1B27626CFFC92DBFB4A7FE815BAC" bold="true" box="[199,358,1756,1782]" pageId="42" pageNumber="43">Comparisons</emphasis>
.
<taxonomicName id="4C2EBCB6626CFFC92C0AB4A6FE015BAC" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[370,486,1756,1782]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B95A1B27626CFFC92C0AB4A6FE015BAC" box="[370,486,1756,1782]" italics="true" pageId="42" pageNumber="43">P. cruralis</emphasis>
</taxonomicName>
is distinguishable from the other four species in this species group by the combination of: elytra sutural angles shortly pointed, forming short denticles; outer apical angles almost rounded; elytra without black patch; tibiae black.
<taxonomicName id="4C2EBCB6626CFFC92CC4B55EFDD55A64" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[444,562,1828,1854]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B95A1B27626CFFC92CC4B55EFDD55A64" box="[444,562,1828,1854]" italics="true" pageId="42" pageNumber="43">P. cruralis</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C2EBCB6626CFFC92E7DB55EFC8A5A64" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[773,877,1828,1854]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626CFFC92E7DB55EFC8A5A64" box="[773,877,1828,1854]" italics="true" pageId="42" pageNumber="43">P. mellea</emphasis>
</taxonomicName>
, but only different in tibiae black and elytra disc depressions absent or much shallower than in
<taxonomicName id="4C2EBCB6626CFFC92FE3B532FCE65A38" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[667,769,1864,1890]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626CFFC92FE3B532FCE65A38" box="[667,769,1864,1890]" italics="true" pageId="42" pageNumber="43">P. mellea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626CFFC82DBFB517FB175C3F" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
<paragraph id="8B91C735626CFFC82DBFB517FB175C3F" blockId="42.[151,1437,1468,2035]" lastBlockId="43.[151,1437,151,681]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="B95A1B27626CFFC92DBFB517FEB65ADC" bold="true" box="[199,337,1900,1926]" pageId="42" pageNumber="43">Description</emphasis>
. Body length
<quantity id="4CD66AD0626CFFC92C82B516FD725ADD" box="[506,661,1901,1927]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.1199999999999999" metricValueMin="0.94" pageId="42" pageNumber="43" unit="mm" value="10.3" valueMax="11.2" valueMin="9.4">9.411.2 mm</quantity>
. Dorsum pale yellow; antennomere 1 yellow, remaining ten antennomeres black; apices of mandibles black; terminal labial palpomere black, terminal and penultimate maxillary palpomeres black; tarsomeres black; tibiae black, middle portion dark brown; venter pale yellow. Pronotum strongly transverse, PW/PL = 1.631.70; much wider than head, PW/HW = 1.101.19; widest at anterior third, lateral explanations very wide. Elytra without microsculpture; striae not incised, replaced by rows of very fine punctures; intervals completely flat, with punctures sparser than in striae; disc not depressed or very shallowly depressed near middle of intervals 3 to 6; apical truncation distinct, straight or slightly rounded; outer apical angles almost rounded, not forming distinct angles; sutural angles slightly pointed, forming short denticles. Apex of abdominal sternite VII very weakly projected at middle in females (
<figureCitation id="1315DBB0626DFFC82FF5B35CFD005C1B" box="[653,743,295,321]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="43" pageNumber="44">Fig. 8D</figureCitation>
). Gonocoxite II of ovipositor dichotomous, inner branch with two ensiform setae apically, outer branch with two or three ensiform setae apically. Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626DFFC82DBFB315FABF5CAF" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8B91C735626DFFC82DBFB315FC9F5CD3" blockId="43.[151,1437,151,681]" box="[199,888,366,393]" pageId="43" pageNumber="44">
<emphasis id="B95A1B27626DFFC82DBFB315FEBE5CD2" bold="true" box="[199,345,366,392]" pageId="43" pageNumber="44">Distribution</emphasis>
(
<figureCitation id="1315DBB0626DFFC82C10B314FE535CD3" box="[360,436,367,393]" captionStart="MAP 4" captionStartId="36.[152,211,1895,1919]" captionTargetBox="[158,1430,1002,1871]" captionTargetId="figure-71@36.[158,1430,1002,1871]" captionTargetPageId="36" captionText="MAP 4. Distributions for species in the Parena (Crossoglossa) testacea group." figureDoi="http://doi.org/10.5281/zenodo.7963663" httpUri="https://zenodo.org/record/7963663/files/figure.png" pageId="43" pageNumber="44">Map 4</figureCitation>
, magenta). West Java and
<collectingCountry id="F33987A5626DFFC82FA4B314FC935CD3" box="[732,884,367,393]" name="Malaysia" pageId="43" pageNumber="44">north Borneo</collectingCountry>
.
</paragraph>
<paragraph id="8B91C735626DFFC82DBFB3E9FABF5CAF" blockId="43.[151,1437,151,681]" pageId="43" pageNumber="44">
<emphasis id="B95A1B27626DFFC82DBFB3E9FE395CF6" bold="true" box="[199,478,402,428]" pageId="43" pageNumber="44">Geographical variation</emphasis>
. Specimens from Borneo are somewhat different from the
<typeStatus id="54957997626DFFC82902B3E8FB3B5CF7" box="[1146,1244,403,429]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
from
<collectingRegion id="49EA09D7626DFFC8285AB3E8FAB05CF7" box="[1314,1367,403,429]" country="Indonesia" name="Jawa Timur" pageId="43" pageNumber="44">Java</collectingRegion>
. They are much larger (ca.
<quantity id="4CD66AD0626DFFC82C04B3CCFE2E5C8B" box="[380,457,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="43" pageNumber="44" unit="mm" value="11.0">11 mm</quantity>
vs
<quantity id="4CD66AD0626DFFC82C97B3CCFDA25C8B" box="[495,581,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="43" pageNumber="44" unit="mm" value="9.6">9.6 mm</quantity>
) and elytra disc with very shallow depressions in the middle of intervals 3 to 6 (without depression in the
<typeStatus id="54957997626DFFC82CAFB3A0FDD95CAF" box="[471,574,475,501]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
). Based on their similar leg color, we determine them all as
<emphasis id="B95A1B27626DFFC829A4B3A0FABF5CAF" box="[1244,1368,475,501]" italics="true" pageId="43" pageNumber="44">
<taxonomicName id="4C2EBCB6626DFFC829A4B3A0FAB35CAF" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[1244,1364,475,501]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="cruralis">P. cruralis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE626DFFC82DBFB384FC455FF3" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8B91C735626DFFC82DBFB384FC455FF3" blockId="43.[151,1437,151,681]" pageId="43" pageNumber="44">
<emphasis id="B95A1B27626DFFC82DBFB384FED35F43" bold="true" box="[199,308,511,537]" pageId="43" pageNumber="44">Remarks</emphasis>
.
<taxonomicName id="4C2EBCB6626DFFC82C3BB384FE5E5F43" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[323,441,511,537]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B95A1B27626DFFC82C3BB384FE5E5F43" box="[323,441,511,537]" italics="true" pageId="43" pageNumber="44">P. cruralis</emphasis>
</taxonomicName>
was originally described as a variety of
<taxonomicName id="4C2EBCB6626DFFC82EFDB384FC195F42" box="[901,1022,511,536]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B27626DFFC82EFDB384FC195F42" box="[901,1022,511,536]" italics="true" pageId="43" pageNumber="44">P. testacea</emphasis>
</taxonomicName>
, and recently (
<bibRefCitation id="EFBFBAC4626DFFC829D0B384FAA65F43" author="Lorenz, W." box="[1192,1345,510,537]" pageId="43" pageNumber="44" refId="ref112064" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world (Insecta Coleoptera Geadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Tutzing: Wolfgang Lorenz, 530 pp." type="book" year="2005">Lorenz, 2005</bibRefCitation>
) treated as a subspecific name under
<taxonomicName id="4C2EBCB6626DFFC82CA3B058FDB45F66" box="[475,595,547,572]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B27626DFFC82CA3B058FDB45F66" box="[475,595,547,572]" italics="true" pageId="43" pageNumber="44">P. testacea</emphasis>
</taxonomicName>
. However, after the examination of the
<typeStatus id="54957997626DFFC8296CB058FB9E5F67" box="[1044,1145,547,573]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
, we found that
<taxonomicName id="4C2EBCB6626DFFC82850B058FA7A5F67" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[1320,1437,547,573]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B95A1B27626DFFC82850B058FA7A5F67" box="[1320,1437,547,573]" italics="true" pageId="43" pageNumber="44">P. cruralis</emphasis>
</taxonomicName>
is closer to
<taxonomicName id="4C2EBCB6626DFFC82C62B03CFE655F3B" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[282,386,583,609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="mellea">
<emphasis id="B95A1B27626DFFC82C62B03CFE655F3B" box="[282,386,583,609]" italics="true" pageId="43" pageNumber="44">P. mellea</emphasis>
</taxonomicName>
than to
<taxonomicName id="4C2EBCB6626DFFC82CA7B03CFDBF5F3A" box="[479,600,583,608]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="B95A1B27626DFFC82CA7B03CFDBF5F3A" box="[479,600,583,608]" italics="true" pageId="43" pageNumber="44">P. testacea</emphasis>
</taxonomicName>
for the similarities in elytra apices and interval punctures. The taxonomic position of
<taxonomicName id="4C2EBCB6626DFFC82C61B010FE685FDF" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[281,399,619,645]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B95A1B27626DFFC82C61B010FE685FDF" box="[281,399,619,645]" italics="true" pageId="43" pageNumber="44">P. cruralis</emphasis>
</taxonomicName>
is still doubtful because the male is unknown. We treat
<taxonomicName id="4C2EBCB6626DFFC82975B010FB645FDF" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[1037,1155,619,645]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="cruralis">
<emphasis id="B95A1B27626DFFC82975B010FB645FDF" box="[1037,1155,619,645]" italics="true" pageId="43" pageNumber="44">P. cruralis</emphasis>
</taxonomicName>
as a separate species for now because it is markedly different from other species in coloration.
</paragraph>
</subSubSection>
</treatment>
</document>