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<document id="C559FA019CC37E801480D20AEA7E6520" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang &amp; Liang, Hongbin" docDate="2023" docId="03877623621FFFBE2DEFB48CFE925AD0" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Parena (Parena) fasciata" docType="treatment" docVersion="8" lastPageNumber="94" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="90" updateTime="1699506634024" updateUser="plazi">
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<mods:title id="9AC4FD670BA01B78633E9BF5BC96F9EB">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<treatment id="03877623621FFFBE2DEFB48CFE925AD0" ID-DOI="http://doi.org/10.5281/zenodo.7963999" ID-GBIF-Taxon="209628999" ID-Zenodo-Dep="7963999" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0" httpUri="http://treatment.plazi.org/id/03877623621FFFBE2DEFB48CFE925AD0" lastPageId="93" lastPageNumber="94" pageId="89" pageNumber="90">
<subSubSection id="C33494BE621FFFBA2DEFB48CFD3D5A4B" box="[151,730,1783,1810]" pageId="89" pageNumber="90" type="nomenclature">
<paragraph id="8B91C735621FFFBA2DEFB48CFD3D5A4B" blockId="89.[151,1079,1783,1846]" box="[151,730,1783,1810]" pageId="89" pageNumber="90">
<emphasis id="B95A1B27621FFFBA2DEFB48CFD3D5A4B" bold="true" box="[151,730,1783,1810]" pageId="89" pageNumber="90">
[26]
<taxonomicName id="4C2EBCB6621FFFBA2DB7B483FD3D5A4B" ID-CoL="4DKPB" authority="(Chaudoir, 1872)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[207,730,1783,1810]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="fasciata" subGenus="Parena">
<emphasis id="B95A1B27621FFFBA2DB7B483FEC05A48" bold="true" box="[207,295,1784,1810]" italics="true" pageId="89" pageNumber="90">Parena</emphasis>
(
<emphasis id="B95A1B27621FFFBA2C40B483FE775A48" bold="true" box="[312,400,1784,1810]" italics="true" pageId="89" pageNumber="90">Parena</emphasis>
)
<emphasis id="B95A1B27621FFFBA2CDAB48CFDE65A4B" bold="true" box="[418,513,1783,1809]" italics="true" pageId="89" pageNumber="90">fasciata</emphasis>
(
<bibRefCitation id="EFBFBAC4621FFFBA2F77B48CFD355A4B" author="Chaudoir, B. M." box="[527,722,1783,1809]" pageId="89" pageNumber="90" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872</bibRefCitation>
)
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621FFFBA2DEFB560FBD05A6C" box="[151,1079,1819,1846]" pageId="89" pageNumber="90" type="description">
<paragraph id="8B91C735621FFFBA2DEFB560FBD05A6C" blockId="89.[151,1079,1783,1846]" box="[151,1079,1819,1846]" pageId="89" pageNumber="90">
Habitus:
<figureCitation id="1315DBB0621FFFBA2D85B560FEA85A6C" box="[253,335,1819,1846]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="89" pageNumber="90">Fig. 53</figureCitation>
. Male genitalia:
<figureCitation id="1315DBB0621FFFBA2F72B560FDB85A6C" box="[522,607,1819,1846]" captionStart="FIGURE 54" captionStartId="93.[151,250,1150,1174]" captionTargetBox="[162,1422,193,1116]" captionTargetId="figure-19@93.[151,1436,181,1126]" captionTargetPageId="93" captionText="FIGURE 54. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Philippine, the &quot;hastata&quot; form [h]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959047" httpUri="https://zenodo.org/record/7959047/files/figure.png" pageId="89" pageNumber="90">Figs 54</figureCitation>
,
<figureCitation id="1315DBB0621FFFBA2F13B560FD6D5A6C" box="[619,650,1819,1846]" captionStart="FIGURE 55" captionStartId="94.[151,250,1504,1528]" captionTargetBox="[163,1420,191,1453]" captionTargetId="figure-28@94.[151,1436,181,1481]" captionTargetPageId="94" captionText="FIGURE 55. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir) from different localities and in different elytra pattern forms, left-lateral (l) and dorsal (d) views: A Taiwan, the &quot;sellata&quot; form [s]. B Tonkin, the rhombic form [r]. C Java, the &quot;unicolor&quot; form [u]. D Papua New Guinea, the melanic form [m]. E Queensland, the &quot;fasciata&quot; form [f]. F Philippine, the &quot;sellata&quot; form [s]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959049" httpUri="https://zenodo.org/record/7959049/files/figure.png" pageId="89" pageNumber="90">55</figureCitation>
. Gonocoxites of ovipositor:
<figureCitation id="1315DBB0621FFFBA2EB1B560FBD45A6C" box="[969,1075,1819,1846]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="89" pageNumber="90">
Fig.
<specimenCount id="9D280CBC621FFFBA2E84B567FBD45A6C" box="[1020,1075,1820,1846]" pageId="89" pageNumber="90" type="generic">11M</specimenCount>
</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621FFFB82DEFB51BFE345FA6" lastPageId="91" lastPageNumber="92" pageId="89" pageNumber="90" type="reference_group">
<paragraph id="8B91C735621FFFBA2DEFB51BFBBA5ACC" blockId="89.[151,1436,1888,2035]" pageId="89" pageNumber="90">
<materialsCitation id="3B46CD68621FFFBA2DEFB51BFBBF5ACC" ID-GBIF-Occurrence="4115871442" country="Papua New Guinea" county="Darlington" location="Palawan" municipality="Sumbawa" pageId="89" pageNumber="90" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<bibRefCitation id="EFBFBAC4621FFFBA2DEFB51BFE825A22" author="Chaudoir, B. M." box="[151,357,1888,1912]" pageId="89" pageNumber="90" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872: 179</bibRefCitation>
(original:
<taxonomicName id="4C2EBCB6621FFFBA2CB6B51BFDB15A22" ID-CoL="8KWFM" authority="Chaudoir, 1873" box="[462,598,1888,1912]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27621FFFBA2CB6B51BFDB15A22" box="[462,598,1888,1912]" italics="true" pageId="89" pageNumber="90">Crossoglossa</emphasis>
</taxonomicName>
; type locality: Moluques;
<typeStatus id="54957997621FFFBA2E25B51BFC515A22" box="[861,950,1888,1912]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
in MNHN);
<bibRefCitation id="EFBFBAC4621FFFBA294EB51BFB1D5A22" author="Jedlicka, A." box="[1078,1274,1888,1912]" pageId="89" pageNumber="90" pagination="269 - 579" refId="ref111456" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article" year="1963">Jedlička, 1963: 443</bibRefCitation>
(
<location id="8EF191EE621FFFBA287EB51BFA875A22" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621FFFBA287EB51BFA875A22" box="[1286,1376,1888,1912]" country="Papua New Guinea" county="Darlington" municipality="Sumbawa" name="Palawan" pageId="89" pageNumber="90" stateProvince="Jawa Timur">Palawan</location>
,
<collectingRegion id="49EA09D7621FFFBA2811B51BFA7F5A2D" box="[1385,1432,1888,1911]" country="Indonesia" name="Jawa Timur" pageId="89" pageNumber="90">Java</collectingRegion>
,
<location id="8EF191EE621FFFBA2DBFB504FEF35ACC" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621FFFBA2DBFB504FEF35ACC" box="[199,276,1919,1942]" country="Papua New Guinea" county="Darlington" municipality="Sumbawa" name="Borneo" pageId="89" pageNumber="90" stateProvince="Jawa Timur">Borneo</location>
,
<collectingMunicipality id="6BF55D4F621FFFBA2C66B505FE605ACC" box="[286,391,1918,1942]" pageId="89" pageNumber="90">Sumbawa</collectingMunicipality>
);
<collectingCounty id="62F0BFB9621FFFBA2CE0B505FDEF5ACC" box="[408,520,1918,1942]" pageId="89" pageNumber="90">Darlington</collectingCounty>
, 1969: 139 (
<collectingCountry id="F33987A5621FFFBA2FFEB504FCB75ACC" box="[646,848,1918,1943]" name="Papua New Guinea" pageId="89" pageNumber="90">Papua New Guinea</collectingCountry>
);
<bibRefCitation id="EFBFBAC4621FFFBA2E19B505FBBF5ACC" author="Kirschenhofer, E." box="[865,1112,1918,1943]" pageId="89" pageNumber="90" pagination="61 - 75" refId="ref111662" refString="Kirschenhofer, E. (2011) Beschreibungen neuer truncatipenner Carabiden aus Ost- und Sudostasien; eine neue Art der Gattung Craspedophorus Hope, 1838 (Panageaeini) aus Sudostasien; Bemerkungen zu zwei japanischen Arten der Gattung Parena Motschulsky, 1859. Acta Coleopterologica, 27 (1), 61 - 75." type="journal article" year="2011">Kirschenhofer, 2011: 68</bibRefCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8B91C735621FFFBA2DEFB5E6FC4D5AA8" blockId="89.[151,1436,1888,2035]" pageId="89" pageNumber="90">
<taxonomicName id="4C2EBCB6621FFFBA2DEFB5E6FD8B5AEF" ID-CoL="4GGCV" authority="Macleay, 1877: 167" authorityName="Macleay" authorityPageNumber="167" authorityYear="1877" box="[151,620,1949,1973]" class="Insecta" family="Carabidae" genus="Phloeodromius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="B95A1B27621FFFBA2DEFB5E6FE7A5AEF" bold="true" box="[151,413,1949,1973]" italics="true" pageId="89" pageNumber="90">Phloeodromius plagiatus</emphasis>
Macleay, 1877: 167
</taxonomicName>
(type locality: Yule Island;
<typeStatus id="54957997621FFFBA2EFFB5E6FC075AEF" box="[903,992,1949,1973]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
in MAMU: University of Sydney, Macleay Museum, Sydney,
<collectingCountry id="F33987A5621FFFBA2CFFB5C7FE0E5A8E" box="[391,489,1980,2004]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
);
<bibRefCitation id="EFBFBAC4621FFFBA2C85B5C7FD5E5A8E" author="Sloane, T. G." box="[509,697,1980,2004]" pageId="89" pageNumber="90" pagination="346 - 381" refId="ref113054" refString="Sloane, T. G. (1907) Studies in Australian Entomology. No. XV. New genera and species of Carabidae, with some notes on synonymy (Clivinini, Scaritini, Cunipectini, Trigonotomini, Lebiini). The Proceedings of the Linnean Society of New South Wales, 32, 346 - 381. https: // doi. org / 10.5962 / bhl. part. 19572" type="journal article" year="1907">Sloane, 1907: 373</bibRefCitation>
(
<taxonomicName id="4C2EBCB6621FFFBA2FB3B5C7FC825A8E" ID-CoL="8LSMH" authorityName="Macleay" authorityYear="1871" box="[715,869,1980,2004]" class="Insecta" family="Carabidae" genus="Phloeodromius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27621FFFBA2FB3B5C7FC825A8E" box="[715,869,1980,2004]" italics="true" pageId="89" pageNumber="90">Phloeodromius</emphasis>
</taxonomicName>
;
<collectingCountry id="F33987A5621FFFBA2E0BB5C7FC325A8E" box="[883,981,1980,2004]" name="Australia" pageId="89" pageNumber="90">Australia</collectingCountry>
); Darlington, 1969: 139 (synonymized with
<taxonomicName id="4C2EBCB6621FFFBA2DBFB5A0FE805AA9" ID-CoL="4DKPB" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[199,359,2011,2035]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621FFFBA2DBFB5A0FE805AA9" box="[199,359,2011,2035]" italics="true" pageId="89" pageNumber="90">Parena fasciata</emphasis>
</taxonomicName>
). Secondary homonym of
<taxonomicName id="4C2EBCB6621FFFBA2F0EB5A0FC4D5AA8" ID-CoL="4DKQ9" authority="Motschulsky." authorityName="Motschulsky." box="[630,938,2010,2034]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="plagiata">
<emphasis id="B95A1B27621FFFBA2F0EB5A0FCFD5AA8" box="[630,794,2010,2034]" italics="true" pageId="89" pageNumber="90">Parena plagiata</emphasis>
Motschulsky.
</taxonomicName>
</paragraph>
<caption id="DF5197BD621DFFB82DEFB2EDFEBC5C38" ID-DOI="http://doi.org/10.5281/zenodo.7963961" ID-Zenodo-Dep="7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="91" pageNumber="92" startId="91.[151,250,150,174]" targetBox="[151,1436,181,2031]" targetPageId="90" targetType="figure">
<paragraph id="8B91C735621DFFB82DEFB2EDFEBC5C38" blockId="91.[151,1436,150,355]" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82DEFB2EDFEFB5DF4" bold="true" box="[151,284,150,174]" pageId="91" pageNumber="92">FIGURE 53</emphasis>
. Habitus of
<taxonomicName id="4C2EBCB6621DFFB82CDCB2ECFCCF5DF5" ID-CoL="4DKPB" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[420,808,151,175]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata" subGenus="Parena">
<emphasis id="B95A1B27621DFFB82CDCB2ECFE095DF4" box="[420,494,151,174]" italics="true" pageId="91" pageNumber="92">Parena</emphasis>
(
<emphasis id="B95A1B27621DFFB82F78B2ECFDAD5DF4" box="[512,586,151,174]" italics="true" pageId="91" pageNumber="92">Parena</emphasis>
)
<emphasis id="B95A1B27621DFFB82F24B2ECFD4B5DF5" box="[604,684,151,175]" italics="true" pageId="91" pageNumber="92">fasciata</emphasis>
(Chaudoir)
</taxonomicName>
.
<emphasis id="B95A1B27621DFFB82E4DB2EDFCA05DF4" bold="true" box="[821,839,150,174]" pageId="91" pageNumber="92">A</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6621DFFB82EA0B2ECFB1F5DF5" ID-CoL="4DKPB" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[984,1272,151,175]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621DFFB82EA0B2ECFB9C5DF5" box="[984,1147,151,175]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
(Chaudoir)
</taxonomicName>
.
<emphasis id="B95A1B27621DFFB8287CB2ECFAF25DF5" bold="true" box="[1284,1301,151,175]" pageId="91" pageNumber="92">B</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6621DFFB82DEFB2C0FE255D89" authority="Heller" authorityName="Heller" authorityYear="1921" box="[151,450,187,211]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatus">
<emphasis id="B95A1B27621DFFB82DEFB2C0FE9A5D89" box="[151,381,187,211]" italics="true" pageId="91" pageNumber="92">Phloeodromus sellatus</emphasis>
Heller
</taxonomicName>
[=
<taxonomicName id="4C2EBCB6621DFFB82C9DB2C0FCE75D89" ID-CoL="4DKPB" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[485,768,187,211]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621DFFB82C9DB2C0FD625D89" box="[485,645,187,211]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
(Chaudoir)
</taxonomicName>
].
<emphasis id="B95A1B27621DFFB82E77B2C1FCC65D88" bold="true" box="[783,801,186,210]" pageId="91" pageNumber="92">C</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6621DFFB82EDFB2C0FB3E5D89" authority="Heller" authorityName="Heller" authorityYear="1921" box="[935,1241,187,211]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B95A1B27621DFFB82EDFB2C0FB735D89" box="[935,1172,187,211]" italics="true" pageId="91" pageNumber="92">Phloeodromus hastatus</emphasis>
Heller
</taxonomicName>
[=
<taxonomicName id="4C2EBCB6621DFFB82984B2C0FEEB5DAD" ID-CoL="4DKPB" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621DFFB82984B2C0FA7B5D89" box="[1276,1436,187,211]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
(Chaudoir)
</taxonomicName>
].
<emphasis id="B95A1B27621DFFB82C64B2A4FEC95DAD" bold="true" box="[284,302,223,247]" pageId="91" pageNumber="92">D</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6621DFFB82CC0B2A4FCE95DAD" ID-CoL="ZMF3" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1940" box="[440,782,223,247]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatoides">
<emphasis id="B95A1B27621DFFB82CC0B2A4FD545DAD" box="[440,691,223,247]" italics="true" pageId="91" pageNumber="92">Crossoglossa sellatoides</emphasis>
Jedlička
</taxonomicName>
[=
<taxonomicName id="4C2EBCB6621DFFB82E4BB2A4FBB75DAD" ID-CoL="4DKPB" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[819,1104,223,247]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621DFFB82E4BB2A4FC335DAD" box="[819,980,223,247]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
(Chaudoir)
</taxonomicName>
].
<emphasis id="B95A1B27621DFFB82919B2A4FB955DAD" bold="true" box="[1121,1138,223,247]" pageId="91" pageNumber="92">E</emphasis>
Holotype of
<taxonomicName id="4C2EBCB6621DFFB82983B2A4FE765C40" ID-CoL="4DKQT" authority="Louwerens" authorityName="Louwerens" authorityYear="1949" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="variety" species="fasciata" variety="unicolor">
<emphasis id="B95A1B27621DFFB82983B2A4FA7B5DAD" box="[1275,1436,223,247]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
var.
<emphasis id="B95A1B27621DFFB82DBBB378FEFE5C41" box="[195,281,259,283]" italics="true" pageId="91" pageNumber="92">unicolor</emphasis>
Louwerens
</taxonomicName>
[=
<taxonomicName id="4C2EBCB6621DFFB82CCCB378FD375C41" ID-CoL="4DKPB" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[436,720,259,283]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621DFFB82CCCB378FDB25C41" box="[436,597,259,283]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
(Chaudoir)
</taxonomicName>
].
<emphasis id="B95A1B27621DFFB82F98B378FD085C41" bold="true" box="[736,751,259,283]" pageId="91" pageNumber="92">F</emphasis>
Male from Java, the &quot;unicolor&quot; form [u].
<emphasis id="B95A1B27621DFFB829E3B379FB485C40" bold="true" box="[1179,1199,258,282]" pageId="91" pageNumber="92">G</emphasis>
Male from Papua New Guinea, the melanic form [m].
<emphasis id="B95A1B27621DFFB82CA0B35CFE0B5C65" bold="true" box="[472,492,295,319]" pageId="91" pageNumber="92">H</emphasis>
Male from Philippine, the rhombic form [r].
<emphasis id="B95A1B27621DFFB82EBAB35CFC2B5C65" bold="true" box="[962,972,295,319]" pageId="91" pageNumber="92">I</emphasis>
Female from Philippine, the trident form [t]. Scale bars = 2 mm.
</paragraph>
</caption>
<paragraph id="8B91C735621DFFB82DEFB3CAFBE55CB2" blockId="91.[151,1437,433,764]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DEFB3CAFDD35C93" authority="Heller, 1921: 526" authorityName="Heller" authorityPageNumber="526" authorityYear="1921" box="[151,564,433,457]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatus">
<emphasis id="B95A1B27621DFFB82DEFB3CAFE645C93" bold="true" box="[151,387,433,457]" italics="true" pageId="91" pageNumber="92">Phloeodromus sellatus</emphasis>
<materialsCitation id="3B46CD68621DFFB82CFFB3CAFDD35C93" ID-GBIF-Occurrence="4115871366" box="[391,564,433,457]" country="Philippines" location="Davao" pageId="91" pageNumber="92" specimenCount="1" stateProvince="Davao" typeStatus="holotype">
<bibRefCitation id="EFBFBAC4621DFFB82CFFB3CAFDD35C93" author="Heller, K. M." box="[391,564,433,457]" pageId="91" pageNumber="92" pagination="523 - 627" refId="ref111042" refString="Heller, K. M. (1921) New Philippine Coleoptera. The Philippine Journal of Science, 19, 523 - 627. https: // doi. org / 10.5962 / bhl. part. 1235" type="journal article" year="1921">Heller, 1921: 526</bibRefCitation>
</materialsCitation>
</taxonomicName>
(type locality: the
<collectingCountry id="F33987A5621DFFB82F94B3CAFC855C93" box="[748,866,433,457]" name="Philippines" pageId="91" pageNumber="92">Philippines</collectingCountry>
:
<collectingRegion id="49EA09D7621DFFB82E12B3C9FC485C93" box="[874,943,434,457]" country="Philippines" name="Davao" pageId="91" pageNumber="92">Davao</collectingRegion>
;
<typeStatus id="54957997621DFFB82EC0B3CAFBF65C93" box="[952,1041,433,457]" pageId="91" pageNumber="92" type="holotype">holotype</typeStatus>
in SNSD);
<bibRefCitation id="EFBFBAC4621DFFB829FAB3CAFADD5C93" author="Jedlicka, A." box="[1154,1338,433,457]" pageId="91" pageNumber="92" refId="ref111358" refString="Jedlicka, A. (1946) Description des Carabides nouveaux de 1 ' Asie orient. Praha: A. Jedlicka, 16 pp." type="book" year="1946">Jedlička, 1946: 10</bibRefCitation>
;
<bibRefCitation id="EFBFBAC4621DFFB8283BB3CAFED75CB2" author="Jedlicka, A." pageId="91" pageNumber="92" pagination="269 - 579" refId="ref111456" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article" year="1963">Jedlička, 1963: 444</bibRefCitation>
;
<bibRefCitation id="EFBFBAC4621DFFB82C43B3ABFDD45CB2" author="Kirschenhofer, E." box="[315,563,464,488]" pageId="91" pageNumber="92" pagination="87 - 103" refId="ref111612" refString="Kirschenhofer, E. (2006) Neue Arten der Gattung Parena Motschulsky aus Japan, China Und Indonesien (Sulawesi) (Coleoptera: Carabidae). Mitteilungen des Internationalen Entomologischen Vereins E. V. Frankfurt, 31 (3 - 4), 87 - 103." type="journal article" year="2006">Kirschenhofer, 2006: 89</bibRefCitation>
;
<bibRefCitation id="EFBFBAC4621DFFB82F46B3ABFCC55CB2" author="Liang, H. B. &amp; Liu, Y. Z." box="[574,802,464,488]" pageId="91" pageNumber="92" pagination="4 - 39" refId="ref111903" refString="Liang, H. B. &amp; Liu, Y. Z. (2018) Carabidae. In: Wu, H., et al. (eds.), Fauna of Tianmu Mountain, Volume VI: Insecta, Coleoptera- Adephaga, Coleoptera-Polyphaga. Hangzhou: Zhejiang University Press, 266 pp. [pp. 4 - 39.]" type="book chapter" year="2018">Liang &amp; Liu, 2018: 32</bibRefCitation>
(Zhejiang).
<emphasis id="B95A1B27621DFFB82ED8B3ABFBE55CB2" bold="true" box="[928,1026,464,488]" pageId="91" pageNumber="92">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8B91C735621DFFB82DEFB395FE7C5F7F" blockId="91.[151,1437,433,764]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DEFB395FDD95F5C" authority="Heller, 1921: 528" authorityName="Heller" authorityPageNumber="528" authorityYear="1921" box="[151,574,494,519]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B95A1B27621DFFB82DEFB395FE6C5F5C" bold="true" box="[151,395,494,518]" italics="true" pageId="91" pageNumber="92">Phloeodromus hastatus</emphasis>
<materialsCitation id="3B46CD68621DFFB82CE8B394FDD95F5C" ID-GBIF-Occurrence="4115871553" box="[400,574,494,519]" country="Philippines" location="Davao" pageId="91" pageNumber="92" specimenCount="1" stateProvince="Davao" typeStatus="holotype">
<bibRefCitation id="EFBFBAC4621DFFB82CE8B394FDD95F5C" author="Heller, K. M." box="[400,574,494,519]" pageId="91" pageNumber="92" pagination="523 - 627" refId="ref111042" refString="Heller, K. M. (1921) New Philippine Coleoptera. The Philippine Journal of Science, 19, 523 - 627. https: // doi. org / 10.5962 / bhl. part. 1235" type="journal article" year="1921">Heller, 1921: 528</bibRefCitation>
</materialsCitation>
</taxonomicName>
(type locality: the
<collectingCountry id="F33987A5621DFFB82F8FB394FC8A5F5D" box="[759,877,495,519]" name="Philippines" pageId="91" pageNumber="92">Philippines</collectingCountry>
:
<collectingRegion id="49EA09D7621DFFB82E0DB394FC5D5F5C" box="[885,954,495,518]" country="Philippines" name="Davao" pageId="91" pageNumber="92">Davao</collectingRegion>
;
<typeStatus id="54957997621DFFB82EBBB394FBFB5F5D" box="[963,1052,495,519]" pageId="91" pageNumber="92" type="holotype">holotype</typeStatus>
in SNSD);
<bibRefCitation id="EFBFBAC4621DFFB829F6B394FADE5F5D" author="Jedlicka, A." box="[1166,1337,495,519]" pageId="91" pageNumber="92" refId="ref111358" refString="Jedlicka, A. (1946) Description des Carabides nouveaux de 1 ' Asie orient. Praha: A. Jedlicka, 16 pp." type="book" year="1946">Jedlička, 1946: 9</bibRefCitation>
;
<bibRefCitation id="EFBFBAC4621DFFB8283BB394FED75F7F" author="Jedlicka, A." pageId="91" pageNumber="92" pagination="269 - 579" refId="ref111456" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article" year="1963">Jedlička, 1963: 442</bibRefCitation>
.
<emphasis id="B95A1B27621DFFB82C42B076FE7C5F7F" bold="true" box="[314,411,525,549]" pageId="91" pageNumber="92">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8B91C735621DFFB82DEFB057FD4B5F39" blockId="91.[151,1437,433,764]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DEFB057FE1F5F1E" ID-CoL="4DKQK" authority="Csiki, 1932: 1455" authorityName="Csiki" authorityPageNumber="1455" authorityYear="1932" box="[151,504,555,580]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sloanei">
<emphasis id="B95A1B27621DFFB82DEFB057FED05F19" bold="true" box="[151,311,555,579]" italics="true" pageId="91" pageNumber="92">Parena sloanei</emphasis>
<bibRefCitation id="EFBFBAC4621DFFB82C46B057FE1F5F1E" author="Csiki, E." box="[318,504,556,580]" pageId="91" pageNumber="92" pagination="1279 - 1598" refId="ref110330" refString="Csiki, E. (1932) Carabidae: Harpalinae VII (Pars 124). In: Junk, W. &amp; Schenkling, S. (eds.), Coleopterorum catalogus. Volumen III. Carabidae III. Berlin: W. Junk, 1933 pp. [pp. 1279 - 1598.]" type="book chapter" year="1932">Csiki, 1932: 1455</bibRefCitation>
</taxonomicName>
[replacement name for
<taxonomicName id="4C2EBCB6621DFFB82F89B056FBEA5F1E" ID-CoL="4DKQ9" authority="(Macleay)" baseAuthorityName="Macleay" baseAuthorityYear="1876" box="[753,1037,556,580]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="plagiata">
<emphasis id="B95A1B27621DFFB82F89B056FC705F1E" box="[753,919,556,580]" italics="true" pageId="91" pageNumber="92">Parena plagiata</emphasis>
(Macleay)
</taxonomicName>
];
<collectingRegion id="49EA09D7621DFFB82958B057FB775F1E" box="[1056,1168,556,580]" country="United Kingdom" name="Darlington" pageId="91" pageNumber="92">Darlington</collectingRegion>
, 1969: 139 (synonymized with
<taxonomicName id="4C2EBCB6621DFFB82D83B030FE7C5F39" ID-CoL="4DKPB" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[251,411,587,611]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621DFFB82D83B030FE7C5F39" box="[251,411,587,611]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFBFBAC4621DFFB82CC9B030FD4F5F39" author="Kirschenhofer, E." box="[433,680,587,611]" pageId="91" pageNumber="92" pagination="61 - 75" refId="ref111662" refString="Kirschenhofer, E. (2011) Beschreibungen neuer truncatipenner Carabiden aus Ost- und Sudostasien; eine neue Art der Gattung Craspedophorus Hope, 1838 (Panageaeini) aus Sudostasien; Bemerkungen zu zwei japanischen Arten der Gattung Parena Motschulsky, 1859. Acta Coleopterologica, 27 (1), 61 - 75." type="journal article" year="2011">Kirschenhofer, 2011: 68</bibRefCitation>
.
</paragraph>
<paragraph id="8B91C735621DFFB82DEFB012FB765FFA" blockId="91.[151,1437,433,764]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DEFB012FDB25FDB" ID-CoL="ZMF3" authority="Jedlicka, 1940: 14" authorityName="Jedlicka" authorityPageNumber="14" authorityYear="1940" box="[151,597,617,641]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatoides">
<emphasis id="B95A1B27621DFFB82DEFB012FE745FDB" bold="true" box="[151,403,617,641]" italics="true" pageId="91" pageNumber="92">Crossoglossa sellatoides</emphasis>
<bibRefCitation id="EFBFBAC4621DFFB82CE3B012FDB25FDB" author="Jedlicka, A." box="[411,597,617,641]" pageId="91" pageNumber="92" refId="ref111324" refString="Jedlicka, A. (1940) Neue Carabiden aus Ostasien. (Hauptsachlich Von der Insel formosa) (XIII. Teil). Praha: A. Jedlicka, 18 pp." type="book" year="1940">Jedlička, 1940: 14</bibRefCitation>
</taxonomicName>
(type locality:
<collectingCountry id="F33987A5621DFFB82F8AB012FCA75FDB" box="[754,832,617,641]" name="Taiwan" pageId="91" pageNumber="92">Taiwan</collectingCountry>
: Kuraru;
<typeStatus id="54957997621DFFB82ED8B012FC1E5FDB" box="[928,1017,617,641]" pageId="91" pageNumber="92" type="holotype">holotype</typeStatus>
in NMPC);
<bibRefCitation id="EFBFBAC4621DFFB82902B012FAD05FDB" author="Jedlicka, A." box="[1146,1335,617,641]" pageId="91" pageNumber="92" refId="ref111358" refString="Jedlicka, A. (1946) Description des Carabides nouveaux de 1 ' Asie orient. Praha: A. Jedlicka, 16 pp." type="book" year="1946">Jedlička, 1946: 10</bibRefCitation>
;
<bibRefCitation id="EFBFBAC4621DFFB8283BB012FED75FFA" author="Jedlicka, A." pageId="91" pageNumber="92" pagination="269 - 579" refId="ref111456" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article" year="1963">Jedlička, 1963: 444</bibRefCitation>
;
<bibRefCitation id="EFBFBAC4621DFFB82C43B0F3FE305FFA" author="Habu, A." box="[315,471,648,672]" pageId="91" pageNumber="92" pagination="83 - 88" refId="ref110742" refString="Habu, A. (1965) Some Carabid-beetles from Formosa. Special Bulletin of the Lepidopteralogical Society of Japan, 1, 83 - 88." type="journal article" year="1965">Habu, 1965: 86</bibRefCitation>
(synonymized with
<taxonomicName id="4C2EBCB6621DFFB82FDFB0F2FCDC5FFA" ID-CoL="4DKQG" baseAuthorityName="Heller" baseAuthorityYear="1921" box="[679,827,648,672]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellata">
<emphasis id="B95A1B27621DFFB82FDFB0F2FCDC5FFA" box="[679,827,648,672]" italics="true" pageId="91" pageNumber="92">Parena sellata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFBFBAC4621DFFB82E28B0F3FBC15FFA" author="Xie, W. P. &amp; Yu, P. Y." box="[848,1062,648,672]" pageId="91" pageNumber="92" pagination="185 - 195" refId="ref113326" refString="Xie, W. P. &amp; Yu, P. Y. (1993) On the Chinese species of Parena Motschulsky (Coleoptera: Carabidae). Sinozoologia, 10, 185 - 195." type="journal article" year="1993">Xie &amp; Yu, 1993: 188</bibRefCitation>
.
<emphasis id="B95A1B27621DFFB82948B0F3FB765FFA" bold="true" box="[1072,1169,648,672]" pageId="91" pageNumber="92">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8B91C735621DFFB82DEFB0DCFAE75FE5" blockId="91.[151,1437,433,764]" box="[151,1280,678,703]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DEFB0DCFD415FE4" ID-CoL="4DKQT" authority="Louwerens, 1949: 51" authorityName="Louwerens" authorityPageNumber="51" authorityYear="1949" box="[151,678,678,703]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="variety" species="fasciata" variety="unicolor">
<emphasis id="B95A1B27621DFFB82DEFB0DCFE2E5FE4" bold="true" box="[151,457,678,703]" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82DEFB0DCFEDB5FE4" bold="true" box="[151,316,678,702]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
var.
<emphasis id="B95A1B27621DFFB82C08B0DDFE2E5FE4" bold="true" box="[368,457,678,702]" italics="true" pageId="91" pageNumber="92">unicolor</emphasis>
</emphasis>
<materialsCitation id="3B46CD68621DFFB82CB7B0DCFD415FE4" ID-GBIF-Occurrence="4115871395" box="[463,678,678,703]" location="Bodjo" pageId="91" pageNumber="92" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<bibRefCitation id="EFBFBAC4621DFFB82CB7B0DCFD415FE4" author="Louwerens, C. J." box="[463,678,678,703]" pageId="91" pageNumber="92" pagination="45 - 53" refId="ref112108" refString="Louwerens, C. J. (1949) Some notes on the Carabidae, collected by Mr. P. H. van Doesburg in the Malay Archipelago with descriptions of new species. Tijdschrift voor Entomologie, 90, 45 - 53." type="journal article" year="1949">Louwerens, 1949: 51</bibRefCitation>
</materialsCitation>
</taxonomicName>
(type locality:
<collectingRegion id="49EA09D7621DFFB82E38B0DCFC965FE4" box="[832,881,679,702]" country="Indonesia" name="Jawa Timur" pageId="91" pageNumber="92">Java</collectingRegion>
:
<location id="8EF191EE621DFFB82E03B0DCFC5B5FE5" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621DFFB82E03B0DCFC5B5FE5" box="[891,956,679,703]" name="Bodjo" pageId="91" pageNumber="92" stateProvince="Jawa Timur">Bodjo</location>
;
<typeStatus id="54957997621DFFB82EBEB0DCFBF85FE5" box="[966,1055,679,703]" pageId="91" pageNumber="92" type="holotype">holotype</typeStatus>
in NNML).
<emphasis id="B95A1B27621DFFB829E6B0DDFAE75FE5" bold="true" box="[1182,1280,678,703]" pageId="91" pageNumber="92">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8B91C735621DFFB82DEFB0BDFE345FA6" blockId="91.[151,1437,433,764]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DEFB0BDFDB05F87" ID-CoL="4DKQP" authority="Kirschenhofer, 2011: 67" authorityName="Kirschenhofer" authorityPageNumber="67" authorityYear="2011" box="[151,599,709,733]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sumatrana">
<emphasis id="B95A1B27621DFFB82DEFB0BDFEBC5F87" bold="true" box="[151,347,710,733]" italics="true" pageId="91" pageNumber="92">Parena sumatrana</emphasis>
<bibRefCitation id="EFBFBAC4621DFFB82C19B0BEFDB05F87" author="Kirschenhofer, E." box="[353,599,709,733]" pageId="91" pageNumber="92" pagination="61 - 75" refId="ref111662" refString="Kirschenhofer, E. (2011) Beschreibungen neuer truncatipenner Carabiden aus Ost- und Sudostasien; eine neue Art der Gattung Craspedophorus Hope, 1838 (Panageaeini) aus Sudostasien; Bemerkungen zu zwei japanischen Arten der Gattung Parena Motschulsky, 1859. Acta Coleopterologica, 27 (1), 61 - 75." type="journal article" year="2011">Kirschenhofer, 2011: 67</bibRefCitation>
</taxonomicName>
(type locality:
<collectingRegion id="49EA09D7621DFFB82F8BB0BEFCAC5F87" box="[755,843,709,733]" country="Indonesia" name="Sumatera Utara" pageId="91" pageNumber="92">Sumatra</collectingRegion>
: Payakumbuh;
<typeStatus id="54957997621DFFB82E9FB0BEFBA75F87" box="[999,1088,709,733]" pageId="91" pageNumber="92" type="holotype">holotype</typeStatus>
in CRK: Collection of R. Kmeco, Litovel, Czech).
<emphasis id="B95A1B27621DFFB82C09B09FFE345FA6" bold="true" box="[369,467,740,764]" pageId="91" pageNumber="92">Syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621DFFB82DEFB15EFD655801" pageId="91" pageNumber="92" type="materials_examined">
<paragraph id="8B91C735621DFFB82DEFB15EFB9F5EF1" blockId="91.[151,1437,804,2020]" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82DEFB15EFE5B5E65" bold="true" box="[151,444,805,831]" pageId="91" pageNumber="92">Type material examined</emphasis>
.
<taxonomicName id="4C2EBCB6621DFFB82CB4B15FFCA45E65" authority="Chaudoir" authorityName="Chaudoir" box="[460,835,804,831]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621DFFB82CB4B15FFCA45E65" bold="true" box="[460,835,804,831]" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82CB4B15FFD215E64" bold="true" box="[460,710,804,831]" italics="true" pageId="91" pageNumber="92">Crossoglossa fasciata</emphasis>
Chaudoir
</emphasis>
</taxonomicName>
:
<emphasis id="B95A1B27621DFFB82E2CB15EFC265E65" bold="true" box="[852,961,805,831]" pageId="91" pageNumber="92">
<typeStatus id="54957997621DFFB82E2CB15EFC265E65" box="[852,961,805,831]" pageId="91" pageNumber="92" type="holotype">Holotype</typeStatus>
</emphasis>
(MNHN,
<figureCitation id="1315DBB0621DFFB82941B15EFB445E1A" box="[1081,1187,805,832]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="91" pageNumber="92">Fig. 53A</figureCitation>
): female, body length =
<quantity id="4CD66AD0621DFFB82DC8B132FEED5E39" box="[176,266,841,867]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="91" pageNumber="92" unit="mm" value="8.8">8.8 mm</quantity>
, pin mounted, &quot;TYPE&quot; [red label], &quot;
<taxonomicName id="4C2EBCB6621DFFB82FD4B132FCE25E39" box="[684,773,841,867]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621DFFB82FD4B132FCE25E39" box="[684,773,841,867]" italics="true" pageId="91" pageNumber="92">fasciata</emphasis>
</taxonomicName>
/
<emphasis id="B95A1B27621DFFB82E66B132FC8E5E39" box="[798,873,841,867]" italics="true" pageId="91" pageNumber="92">Chaud</emphasis>
. /
<emphasis id="B95A1B27621DFFB82EF1B132FC105E39" box="[905,1015,841,867]" italics="true" pageId="91" pageNumber="92">
<collectingCountry id="F33987A5621DFFB82EF1B132FC105E39" box="[905,1015,841,867]" name="Indonesia" pageId="91" pageNumber="92">Moluques</collectingCountry>
</emphasis>
/
<emphasis id="B95A1B27621DFFB82969B131FB8D5E39" box="[1041,1130,842,867]" italics="true" pageId="91" pageNumber="92">Lorquin</emphasis>
&quot; [ex Chaudoir's box label, pinned under specimen now], &quot;COLL. DE CHAUDOIR 1874&quot;, &quot;MUSEUM PARIS / 1952 / Coll. R. OBERTHUR&quot;, &quot;
<typeStatus id="54957997621DFFB82DE7B1EAFED05EF1" box="[159,311,913,939]" pageId="91" pageNumber="92" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C2EBCB6621DFFB82C34B1EAFDDA5EF1" box="[332,573,913,939]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">Crossoglossa fasciata</taxonomicName>
/
<bibRefCitation id="EFBFBAC4621DFFB82F2AB1EAFD185EF1" author="Chaudoir, B. M." box="[594,767,913,939]" pageId="91" pageNumber="92" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872</bibRefCitation>
/ det. SHI H.L. 2011&quot; [red label].
</paragraph>
<paragraph id="8B91C735621DFFB82DBFB1CFFA7A5EA9" blockId="91.[151,1437,804,2020]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DBFB1CFFDFE5E95" authority="Heller" authorityName="Heller" authorityYear="1921" box="[199,537,948,975]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatus">
<emphasis id="B95A1B27621DFFB82DBFB1CFFDFE5E95" bold="true" box="[199,537,948,975]" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82DBFB1CFFE2D5E95" bold="true" box="[199,458,948,975]" italics="true" pageId="91" pageNumber="92">Phloeodromus sellatus</emphasis>
Heller
</emphasis>
</taxonomicName>
:
<emphasis id="B95A1B27621DFFB82F5CB1CEFD765E95" bold="true" box="[548,657,949,975]" pageId="91" pageNumber="92">
<typeStatus id="54957997621DFFB82F5CB1CEFD765E95" box="[548,657,949,975]" pageId="91" pageNumber="92" type="holotype">Holotype</typeStatus>
</emphasis>
(SNSD,
<figureCitation id="1315DBB0621DFFB82F89B1CEFCB25E8A" box="[753,853,949,976]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="91" pageNumber="92">Fig.53B</figureCitation>
): female, body length =
<quantity id="4CD66AD0621DFFB8292EB1CEFB4C5E95" box="[1110,1195,949,975]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="91" pageNumber="92" unit="mm" value="8.3">8.3 mm</quantity>
, pin mounted, &quot;
<collectingRegion id="49EA09D7621DFFB82828B1CEFA7A5E95" box="[1360,1437,949,975]" country="Philippines" name="Davao" pageId="91" pageNumber="92">Davao</collectingRegion>
/Mindanao /Baker&quot;, &quot;
<taxonomicName id="4C2EBCB6621DFFB82CFFB1A2FE3A5EA9" authorityName="Heller" authorityYear="1921" box="[391,477,985,1011]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatus">
<emphasis id="B95A1B27621DFFB82CFFB1A2FE3A5EA9" box="[391,477,985,1011]" italics="true" pageId="91" pageNumber="92">sellatus</emphasis>
</taxonomicName>
/ typus&quot;[red label], &quot;1920 /
<emphasis id="B95A1B27621DFFB82E7CB1A2FCF55EA9" box="[772,786,985,1011]" italics="true" pageId="91" pageNumber="92">3</emphasis>
&quot; [yellow label]; &quot;Staatl.Museum für/Tierkunde, Dresden&quot;.
</paragraph>
<paragraph id="8B91C735621DFFB82DBFB187FDC05905" blockId="91.[151,1437,804,2020]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DBFB187FDC9594D" authority="Heller" authorityName="Heller" authorityYear="1921" box="[199,558,1020,1047]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B95A1B27621DFFB82DBFB187FDC9594D" bold="true" box="[199,558,1020,1047]" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82DBFB187FE3E594D" bold="true" box="[199,473,1020,1047]" italics="true" pageId="91" pageNumber="92">Phloeodromus hastatus</emphasis>
Heller
</emphasis>
</taxonomicName>
:
<emphasis id="B95A1B27621DFFB82F47B186FD4B594D" bold="true" box="[575,684,1021,1047]" pageId="91" pageNumber="92">
<typeStatus id="54957997621DFFB82F47B186FD4B594D" box="[575,684,1021,1047]" pageId="91" pageNumber="92" type="holotype">Holotype</typeStatus>
</emphasis>
(SNSD,
<figureCitation id="1315DBB0621DFFB82E6FB186FC675942" box="[791,896,1021,1048]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="91" pageNumber="92">Fig. 53C</figureCitation>
): female, body length =
<quantity id="4CD66AD0621DFFB829E5B186FB1F594D" box="[1181,1272,1021,1047]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="91" pageNumber="92" unit="mm" value="7.7">7.7 mm</quantity>
, pin mounted, &quot;
<collectingRegion id="49EA09D7621DFFB82DE7B65AFF0B5961" box="[159,236,1057,1083]" country="Philippines" name="Davao" pageId="91" pageNumber="92">Davao</collectingRegion>
/ Mindanao / Baker&quot;, &quot;
<emphasis id="B95A1B27621DFFB82C86B65AFD895961" box="[510,622,1057,1083]" italics="true" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82C86B65AFDBC5961" authorityName="Heller" authorityYear="1921" box="[510,603,1057,1083]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="hastatus">hastatus</taxonomicName>
!
</emphasis>
/ typus&quot; [red label], &quot;
<emphasis id="B95A1B27621DFFB82E1CB65AFC7B5961" box="[868,924,1057,1083]" italics="true" pageId="91" pageNumber="92">7243</emphasis>
&quot;, &quot;1920 /
<emphasis id="B95A1B27621DFFB8296FB65AFBC25961" box="[1047,1061,1057,1083]" italics="true" pageId="91" pageNumber="92">3</emphasis>
&quot; [yellow label]; &quot;
<emphasis id="B95A1B27621DFFB8298EB65AFAC95961" box="[1270,1326,1057,1083]" italics="true" pageId="91" pageNumber="92">1454</emphasis>
&quot;, &quot;Staatl. Museum für / Tierkunde, Dresden&quot;.
</paragraph>
<paragraph id="8B91C735621DFFB82DBFB613FDAD5991" blockId="91.[151,1437,804,2020]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DBFB613FDA059D8" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1940" box="[199,583,1128,1155]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatoides">
<emphasis id="B95A1B27621DFFB82DBFB613FDA059D8" bold="true" box="[199,583,1128,1155]" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82DBFB613FE3A59D9" bold="true" box="[199,477,1128,1155]" italics="true" pageId="91" pageNumber="92">Crossoglossa sellatoides</emphasis>
Jedlička
</emphasis>
</taxonomicName>
:
<emphasis id="B95A1B27621DFFB82F2DB612FD2559D9" bold="true" box="[597,706,1129,1155]" pageId="91" pageNumber="92">
<typeStatus id="54957997621DFFB82F2DB612FD2559D9" box="[597,706,1129,1155]" pageId="91" pageNumber="92" type="holotype">Holotype</typeStatus>
</emphasis>
(NMPC,
<figureCitation id="1315DBB0621DFFB82E57B612FC7F59DE" box="[815,920,1129,1156]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="91" pageNumber="92">Fig. 53D</figureCitation>
): female, body length =
<quantity id="4CD66AD0621DFFB829DEB612FB1A59D9" box="[1190,1277,1129,1156]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="91" pageNumber="92" unit="mm" value="7.5">7.5 mm</quantity>
, pin mounted, &quot;
<emphasis id="B95A1B27621DFFB82DDBB6F6FEE759FD" box="[163,256,1165,1191]" italics="true" pageId="91" pageNumber="92">KUARU</emphasis>
/
<collectingCountry id="F33987A5621DFFB82C6EB6F6FE7B59FD" box="[278,412,1165,1191]" name="Taiwan" pageId="91" pageNumber="92">FORMOSA</collectingCountry>
/
<emphasis id="B95A1B27621DFFB82CC9B6F6FDC959FD" box="[433,558,1165,1191]" italics="true" pageId="91" pageNumber="92">
<date id="FF90E1F5621DFFB82CC9B6F6FDC959FD" box="[433,558,1165,1191]" pageId="91" pageNumber="92" value="1937-06-15">15.VI.1937</date>
</emphasis>
/ COL. M. CHUJO&quot;, &quot;TYPUS&quot; [red label], &quot;
<emphasis id="B95A1B27621DFFB8295FB6F6FF785991" italics="true" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB8295FB6F6FB5A59FD" authority="Chaudoir, 1873" box="[1063,1213,1165,1191]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">Crossoglossa</taxonomicName>
/
<taxonomicName id="4C2EBCB6621DFFB829ABB6F6FAAC59FD" authorityName="Jedlicka" authorityYear="1940" box="[1235,1355,1165,1191]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatoides">sellatoides</taxonomicName>
/ sp. n. /
</emphasis>
DET. ING. JEDLIČKA&quot; [pink label].
</paragraph>
<paragraph id="8B91C735621DFFB82DBFB6AEFD655801" blockId="91.[151,1437,804,2020]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82DBFB6AEFD4059B5" authority="Louwerens" authorityName="Louwerens" authorityYear="1949" box="[199,679,1236,1263]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="variety" species="fasciata" variety="unicolor">
<emphasis id="B95A1B27621DFFB82DBFB6AEFD4059B5" bold="true" box="[199,679,1236,1263]" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82DBFB6AEFE9A59B4" bold="true" box="[199,381,1236,1263]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
var.
<emphasis id="B95A1B27621DFFB82CC1B6AFFDFC59B5" bold="true" box="[441,539,1236,1263]" italics="true" pageId="91" pageNumber="92">unicolor</emphasis>
Louwerens
</emphasis>
</taxonomicName>
:
<emphasis id="B95A1B27621DFFB82FCEB6AEFCC459B5" bold="true" box="[694,803,1237,1263]" pageId="91" pageNumber="92">
<typeStatus id="54957997621DFFB82FCEB6AEFCC459B5" box="[694,803,1237,1263]" pageId="91" pageNumber="92" type="holotype">Holotype</typeStatus>
</emphasis>
(NNML,
<figureCitation id="1315DBB0621DFFB82EECB6AEFC1C59AA" box="[916,1019,1237,1264]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="91" pageNumber="92">Fig. 53E</figureCitation>
): male, body length =
<quantity id="4CD66AD0621DFFB82980B6AEFAB759B5" box="[1272,1360,1237,1264]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="91" pageNumber="92" unit="mm" value="8.5">8.5 mm</quantity>
, board mounted, &quot;Bodja,
<emphasis id="B95A1B27621DFFB82C1CB682FE3E5849" box="[356,473,1273,1299]" italics="true" pageId="91" pageNumber="92">29-10-33 /</emphasis>
<collectingRegion id="49EA09D7621DFFB82C98B682FD955849" box="[480,626,1273,1299]" country="Indonesia" name="Jawa Tengah" pageId="91" pageNumber="92">Midden-Java</collectingRegion>
/ v. Doesburg.&quot;, &quot;Museum Leiden / ex. collection/ C.J. Louwerens / rec. 1979&quot;, &quot;
<emphasis id="B95A1B27621DFFB82D8CB765FDD8586D" box="[244,575,1309,1335]" italics="true" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82D8CB765FEA2586D" box="[244,325,1310,1335]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/
<taxonomicName id="4C2EBCB6621DFFB82C23B766FE16586D" authority="Chd." authorityName="Chd." box="[347,497,1309,1335]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">fasciata Chd.</taxonomicName>
/ var. /
</emphasis>
E.B. Britton det. / 194
<emphasis id="B95A1B27621DFFB82E45B766FCAC586D" box="[829,843,1309,1335]" italics="true" pageId="91" pageNumber="92">7</emphasis>
&quot;, &quot;
<emphasis id="B95A1B27621DFFB82E08B765FA8B586D" box="[880,1388,1309,1335]" italics="true" pageId="91" pageNumber="92">
Typus /
<taxonomicName id="4C2EBCB6621DFFB82EBFB765FB9F586D" box="[967,1144,1309,1335]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">Parena fasciata</taxonomicName>
/ Chd /
<taxonomicName id="4C2EBCB6621DFFB829AAB765FABA586D" authorityName="Louwerens" authorityYear="1949" box="[1234,1373,1309,1335]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="variety" species="fasciata" variety="unicolor">var unicolor</taxonomicName>
/
</emphasis>
det.
<emphasis id="B95A1B27621DFFB82DEFB739FF3C5801" box="[151,219,1346,1371]" italics="true" pageId="91" pageNumber="92">Louw.</emphasis>
/ C.J. Louwerens&quot;, &quot;type&quot; [red label].
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621DFFB82DBFB71EFABB5BC5" pageId="91" pageNumber="92" type="discussion">
<paragraph id="8B91C735621DFFB82DBFB71EFEEE589D" blockId="91.[151,1437,804,2020]" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82DBFB71EFDC25825" bold="true" box="[199,549,1381,1407]" pageId="91" pageNumber="92">Notes on types and synonyms</emphasis>
.
<emphasis id="B95A1B27621DFFB82F4BB71FFC7A5825" bold="true" box="[563,925,1380,1407]" pageId="91" pageNumber="92">
<taxonomicName id="4C2EBCB6621DFFB82F4BB71FFC7F5825" authority="Heller" authorityName="Heller" authorityYear="1921" box="[563,920,1380,1407]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B95A1B27621DFFB82F4BB71FFCA55825" bold="true" box="[563,834,1380,1407]" italics="true" pageId="91" pageNumber="92">Phloeodromus hastatus</emphasis>
Heller
</taxonomicName>
:
</emphasis>
The original description did not indicate how many types there are, but a serial number &quot;7243&quot; implied a single
<typeStatus id="54957997621DFFB82EFBB7F2FC0F58F9" box="[899,1000,1417,1443]" pageId="91" pageNumber="92" type="holotype">holotype</typeStatus>
, which is in accord with our examined specimen.
</paragraph>
<paragraph id="8B91C735621DFFB82DBFB7AAFB9158B1" blockId="91.[151,1437,804,2020]" box="[199,1142,1489,1515]" pageId="91" pageNumber="92">
For other names treated here, all
<typeStatus id="54957997621DFFB82F41B7AAFD4158B1" box="[569,678,1489,1515]" pageId="91" pageNumber="92" type="holotype">holotypes</typeStatus>
were original clearly fixed by monotypy.
</paragraph>
<paragraph id="8B91C735621DFFB82DBFB78EFABB5BC5" blockId="91.[151,1437,804,2020]" pageId="91" pageNumber="92">
All examined
<typeStatus id="54957997621DFFB82C1EB78EFE715B55" box="[358,406,1525,1551]" pageId="91" pageNumber="92">type</typeStatus>
materials of the above synonyms are very similar to each other except for the different patterns on their elytra. We studied specimens with different elytra pattern forms from various localities. Their identical male genitalia (
<figureCitation id="1315DBB0621DFFB82C15B446FE235B02" box="[365,452,1597,1624]" captionStart="FIGURE 54" captionStartId="93.[151,250,1150,1174]" captionTargetBox="[162,1422,193,1116]" captionTargetId="figure-19@93.[151,1436,181,1126]" captionTargetPageId="93" captionText="FIGURE 54. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Philippine, the &quot;hastata&quot; form [h]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959047" httpUri="https://zenodo.org/record/7959047/files/figure.png" pageId="91" pageNumber="92">Figs 54</figureCitation>
,
<figureCitation id="1315DBB0621DFFB82CA8B446FE0A5B02" box="[464,493,1597,1624]" captionStart="FIGURE 55" captionStartId="94.[151,250,1504,1528]" captionTargetBox="[163,1420,191,1453]" captionTargetId="figure-28@94.[151,1436,181,1481]" captionTargetPageId="94" captionText="FIGURE 55. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir) from different localities and in different elytra pattern forms, left-lateral (l) and dorsal (d) views: A Taiwan, the &quot;sellata&quot; form [s]. B Tonkin, the rhombic form [r]. C Java, the &quot;unicolor&quot; form [u]. D Papua New Guinea, the melanic form [m]. E Queensland, the &quot;fasciata&quot; form [f]. F Philippine, the &quot;sellata&quot; form [s]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959049" httpUri="https://zenodo.org/record/7959049/files/figure.png" pageId="91" pageNumber="92">55</figureCitation>
) suggest that they all belong to the same species. So, we herein treat
<taxonomicName id="4C2EBCB6621DFFB82982B446FED85B21" authority="Heller" authorityName="Heller" authorityYear="1921" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatus">
<emphasis id="B95A1B27621DFFB82982B446FF0A5B21" italics="true" pageId="91" pageNumber="92">Phloeodromus sellatus</emphasis>
Heller
</taxonomicName>
,
<taxonomicName id="4C2EBCB6621DFFB82C36B41AFD4C5B21" authority="Heller" authorityName="Heller" authorityYear="1921" box="[334,683,1633,1659]" class="Insecta" family="Pyralidae" genus="Phloeodromus" kingdom="Animalia" order="Lepidoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B95A1B27621DFFB82C36B41AFDBF5B21" box="[334,600,1633,1659]" italics="true" pageId="91" pageNumber="92">Phloeodromus hastatus</emphasis>
Heller
</taxonomicName>
,
<taxonomicName id="4C2EBCB6621DFFB82FC2B41AFBDB5B21" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1940" box="[698,1084,1633,1659]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sellatoides">
<emphasis id="B95A1B27621DFFB82FC2B41AFC355B21" box="[698,978,1633,1659]" italics="true" pageId="91" pageNumber="92">Crossoglossa sellatoides</emphasis>
Jedlička
</taxonomicName>
,
<taxonomicName id="4C2EBCB6621DFFB82932B419FEF15BC5" authority="Louwerens" authorityName="Louwerens" authorityYear="1949" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="variety" species="fasciata" variety="unicolor">
<emphasis id="B95A1B27621DFFB82932B419FB185B21" box="[1098,1279,1633,1659]" italics="true" pageId="91" pageNumber="92">Parena fasciata</emphasis>
var.
<emphasis id="B95A1B27621DFFB82845B41AFA7B5B21" box="[1341,1436,1633,1659]" italics="true" pageId="91" pageNumber="92">unicolor</emphasis>
Louwerens
</taxonomicName>
, and
<taxonomicName id="4C2EBCB6621DFFB82C29B4FDFD2F5BC5" authority="Kirschenhofer" authorityName="Kirschenhofer" authorityYear="2011" box="[337,712,1669,1695]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="sumatrana">
<emphasis id="B95A1B27621DFFB82C29B4FDFDC65BC5" box="[337,545,1670,1695]" italics="true" pageId="91" pageNumber="92">Parena sumatrana</emphasis>
Kirschenhofer
</taxonomicName>
all as junior synonyms of
<taxonomicName id="4C2EBCB6621DFFB82E89B4FEFABB5BC5" authority="Chaudoir." authorityName="Chaudoir." box="[1009,1372,1669,1695]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621DFFB82E89B4FEFB005BC5" box="[1009,1255,1669,1695]" italics="true" pageId="91" pageNumber="92">Crossoglossa fasciata</emphasis>
Chaudoir.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621DFFBF2DBFB4D2FCA85F3B" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92" type="materials_examined">
<paragraph id="8B91C735621DFFBF2DBFB4D2FCA85F3B" blockId="91.[151,1437,804,2020]" lastBlockId="92.[151,1437,150,1869]" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">
<emphasis id="B95A1B27621DFFB82DBFB4D2FDF95B99" bold="true" box="[199,542,1705,1731]" pageId="91" pageNumber="92">Non-type material examined</emphasis>
.
<materialsCitation id="3B46CD68621DFFB82F56B4D2FF205BB2" ID-GBIF-Occurrence="4115871502" collectorName="F. P. Dodd." country="Australia" location="Australia" pageId="91" pageNumber="92" specimenCount="1" specimenCount-male="1">
<emphasis id="B95A1B27621DFFB82F56B4D2FD795B99" bold="true" box="[558,670,1705,1731]" pageId="91" pageNumber="92">Australia</emphasis>
:
<specimenCount id="9D280CBC621DFFB82FD6B4D2FD1C5B99" box="[686,763,1705,1731]" pageId="91" pageNumber="92" type="male">1 male</specimenCount>
[f] (NHML), &quot;Townsville, Qld., 24.2.02,
<collectorName id="26DBA2E3621DFFB829AFB4D2FAB75B99" box="[1239,1360,1705,1731]" pageId="91" pageNumber="92">F.P. Dodd.</collectorName>
&quot; &lt;
<figureCitation id="1315DBB0621DFFB82808B4D2FF205BB2" captionStart="FIGURE 55" captionStartId="94.[151,250,1504,1528]" captionTargetBox="[163,1420,191,1453]" captionTargetId="figure-28@94.[151,1436,181,1481]" captionTargetPageId="94" captionText="FIGURE 55. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir) from different localities and in different elytra pattern forms, left-lateral (l) and dorsal (d) views: A Taiwan, the &quot;sellata&quot; form [s]. B Tonkin, the rhombic form [r]. C Java, the &quot;unicolor&quot; form [u]. D Papua New Guinea, the melanic form [m]. E Queensland, the &quot;fasciata&quot; form [f]. F Philippine, the &quot;sellata&quot; form [s]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959049" httpUri="https://zenodo.org/record/7959049/files/figure.png" pageId="91" pageNumber="92">Fig. 55E</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD68621DFFB82D9DB4B6FEC35A51" ID-GBIF-Occurrence="4115871403" collectingDate="1992-12-11" collectorName="W. Dressler &amp; P. Zborowski" country="Papua New Guinea" location="Papua New Guinea" pageId="91" pageNumber="92" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC621DFFB82D9DB4B6FEAE5BBD" box="[229,329,1741,1767]" pageId="91" pageNumber="92" type="female">1 female</specimenCount>
[f] (ANIC), &quot;12.40S 142.40E QLD, Batavia Downs Hs.,
<date id="FF90E1F5621DFFB82E92B4B6FB615BBD" box="[1002,1158,1741,1767]" pageId="91" pageNumber="92" value="1992-12-11">
<collectingDate id="EFD4181D621DFFB82E92B4B6FB615BBD" box="[1002,1158,1741,1767]" pageId="91" pageNumber="92" value="1992-12-11">11 Dec. 1992</collectingDate>
</date>
at light,
<collectorName id="26DBA2E3621DFFB8298AB4B6FA9D5BBD" box="[1266,1402,1741,1767]" pageId="91" pageNumber="92">W. Dressler</collectorName>
,
<collectorName id="26DBA2E3621DFFB828F1B4B6FEFF5A51" pageId="91" pageNumber="92">P. Zborowski</collectorName>
&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68621DFFB82C53B48AFD725A75" ID-GBIF-Occurrence="4115871374" collectingDate="1992-12-11" collectorName="W. Dressler &amp; P. Zborowski" country="Papua New Guinea" location="Papua New Guinea" pageId="91" pageNumber="92" specimenCount="1" specimenCount-female="1">
<emphasis id="B95A1B27621DFFB82C53B48AFDF45A51" bold="true" box="[299,531,1777,1803]" pageId="91" pageNumber="92">Papua New Guinea</emphasis>
:
<specimenCount id="9D280CBC621DFFB82F5BB48AFD625A51" box="[547,645,1777,1803]" pageId="91" pageNumber="92" type="female">1 female</specimenCount>
[f] (MNHN), &quot;Kiriwini Trobriand Is. iii.iv.v. 95. (
<collectorName id="26DBA2E3621DFFB829C5B48AFAD95A51" box="[1213,1342,1777,1803]" pageId="91" pageNumber="92">A.S. Meek</collectorName>
)&quot;, &quot;MUSEUM PARIS 1952, Coll. R. OBERTHUR&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68621DFFB82FD9B56EFE7D5A09" ID-GBIF-Occurrence="4115871386" collectorName="Papoa. N. Guinea" country="Papua New Guinea" location="Bololo Distr." pageId="91" pageNumber="92" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC621DFFB82FD9B56EFCE35A75" box="[673,772,1813,1839]" pageId="91" pageNumber="92" type="female">1 female</specimenCount>
[f] (NHML), &quot;
<collectorName id="26DBA2E3621DFFB82EC8B56EFB665A75" box="[944,1153,1813,1839]" pageId="91" pageNumber="92">Papoa. N. Guinea</collectorName>
,
<location id="8EF191EE621DFFB829F6B56EFAC25A75" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621DFFB829F6B56EFAC25A75" box="[1166,1317,1813,1839]" country="Papua New Guinea" name="Bololo Distr." pageId="91" pageNumber="92">Bololo Distr.</location>
, 12.xii.82. Baxch, BM 1983: 494&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68621DFFB82CDCB542FB875A0E" ID-GBIF-Occurrence="4115871452" box="[420,1120,1849,1876]" collectingDate="1937-08" collectorName="C. J. Louwerens &amp; G. Tompoe-Paloe" country="Papua New Guinea" location="Sulawesi" pageId="91" pageNumber="92" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D280CBC621DFFB82CDCB542FE095A09" box="[420,494,1849,1875]" pageId="91" pageNumber="92" type="male">1 male</specimenCount>
[m] &quot;Yanarba, Fgum Is. Meek. ii.95.&quot; &lt;
<figureCitation id="1315DBB0621DFFB82EDDB542FBEC5A09" box="[933,1035,1849,1876]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="91" pageNumber="92">Figs 53G</figureCitation>
,
<figureCitation id="1315DBB0621DFFB82960B542FBAB5A0E" box="[1048,1100,1849,1876]" captionStart="FIGURE 55" captionStartId="94.[151,250,1504,1528]" captionTargetBox="[163,1420,191,1453]" captionTargetId="figure-28@94.[151,1436,181,1481]" captionTargetPageId="94" captionText="FIGURE 55. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir) from different localities and in different elytra pattern forms, left-lateral (l) and dorsal (d) views: A Taiwan, the &quot;sellata&quot; form [s]. B Tonkin, the rhombic form [r]. C Java, the &quot;unicolor&quot; form [u]. D Papua New Guinea, the melanic form [m]. E Queensland, the &quot;fasciata&quot; form [f]. F Philippine, the &quot;sellata&quot; form [s]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959049" httpUri="https://zenodo.org/record/7959049/files/figure.png" pageId="91" pageNumber="92">55D</figureCitation>
&gt;.
</materialsCitation>
<materialsCitation id="3B46CD68621DFFB8291DB543FC9E5A2D" ID-GBIF-Occurrence="4115871359" collectingDate="1937-08" collectorName="C. J. Louwerens &amp; G. Tompoe-Paloe" country="Papua New Guinea" location="Sulawesi" pageId="91" pageNumber="92" specimenCount="1" specimenCount-male="1">
<emphasis id="B95A1B27621DFFB8291DB543FB2A5A08" bold="true" box="[1125,1229,1848,1874]" pageId="91" pageNumber="92">
<location id="8EF191EE621DFFB8291DB543FB2A5A08" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621DFFB8291DB543FB2A5A08" box="[1125,1229,1848,1874]" country="Papua New Guinea" name="Sulawesi" pageId="91" pageNumber="92">Sulawesi</location>
</emphasis>
: 8 ex [f] (NHML), &quot;
<collectorName id="26DBA2E3621DFFB82DD9B526FEB45A2D" box="[161,339,1885,1911]" pageId="91" pageNumber="92">C.J. Louwerens</collectorName>
, W-Celebes 2700',
<collectorName id="26DBA2E3621DFFB82F50B526FD095A2D" box="[552,750,1885,1911]" pageId="91" pageNumber="92">G. Tompoe-Paloe</collectorName>
,
<date id="FF90E1F5621DFFB82F8FB526FC8A5A2D" box="[759,877,1885,1911]" pageId="91" pageNumber="92" value="1937-08">
<collectingDate id="EFD4181D621DFFB82F8FB526FC8A5A2D" box="[759,877,1885,1911]" pageId="91" pageNumber="92" value="1937-08">VIII.1937</collectingDate>
</date>
&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68621DFFB82E07B526FE3E5AC1" ID-GBIF-Occurrence="4115871497" collectingDate="1937-08" collectorName="C. J. Louwerens &amp; G. Tompoe-Paloe" country="Papua New Guinea" elevation="4" location="Sulawesi" municipality="Timor" pageId="91" pageNumber="92" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<emphasis id="B95A1B27621DFFB82E07B526FC2C5A2D" bold="true" box="[895,971,1885,1911]" pageId="91" pageNumber="92">Timor</emphasis>
:
<specimenCount id="9D280CBC621DFFB82EA0B526FBDF5A2D" box="[984,1080,1885,1911]" pageId="91" pageNumber="92" type="female">1 female</specimenCount>
[u] (ZSM), &quot;
<collectingMunicipality id="6BF55D4F621DFFB829BEB526FAF75A2D" box="[1222,1296,1885,1911]" pageId="91" pageNumber="92">Timor</collectingMunicipality>
, 1911.24.IV, coll.
<collectorName id="26DBA2E3621DFFB82DB7B5FAFEFB5AC1" box="[207,284,1921,1947]" pageId="91" pageNumber="92">Haniel</collectorName>
,
<location id="8EF191EE621DFFB82C5FB5FAFE2A5AC1" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621DFFB82C5FB5FAFE2A5AC1" box="[295,461,1921,1947]" country="Papua New Guinea" municipality="Timor" name="central Semao" pageId="91" pageNumber="92">central Semao</location>
&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68621DFFB82CA7B5FAFCA65A9A" ID-GBIF-Occurrence="4115871347" collectingDate="1937-08" collectorName="C. J. Louwerens &amp; G. Tompoe-Paloe" country="Papua New Guinea" elevation="4" location="Sulawesi" municipality="Timor" pageId="91" pageNumber="92" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<emphasis id="B95A1B27621DFFB82CA7B5FAFDF05AC1" bold="true" box="[479,535,1921,1947]" pageId="91" pageNumber="92">Java</emphasis>
: 71 ex [29s, 27u, 10f,
<quantity id="4CD66AD0621DFFB82E69B5FAFCD05AC1" box="[785,823,1921,1947]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="91" pageNumber="92" unit="m" value="4.0">
<elevation id="00032006621DFFB82E69B5FAFCD05AC1" box="[785,823,1921,1947]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="91" pageNumber="92" unit="m" value="4.0">4m</elevation>
</quantity>
, 1h] (MNHN), &quot;Centr. Java, Nglirip.,
<collectorName id="26DBA2E3621DFFB829A6B5FAFA775AC1" box="[1246,1424,1921,1947]" pageId="91" pageNumber="92">Mme E. Walsh.</collectorName>
&quot;, &quot;Museum Paris, 1952, Coll. R. OBERthur&quot; &lt;
<figureCitation id="1315DBB0621DFFB82FF5B5DEFD175A9A" box="[653,752,1957,1984]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="91" pageNumber="92">Figs 53F</figureCitation>
,
<figureCitation id="1315DBB0621DFFB82F84B5DEFCCA5A9A" box="[764,813,1957,1984]" captionStart="FIGURE 55" captionStartId="94.[151,250,1504,1528]" captionTargetBox="[163,1420,191,1453]" captionTargetId="figure-28@94.[151,1436,181,1481]" captionTargetPageId="94" captionText="FIGURE 55. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir) from different localities and in different elytra pattern forms, left-lateral (l) and dorsal (d) views: A Taiwan, the &quot;sellata&quot; form [s]. B Tonkin, the rhombic form [r]. C Java, the &quot;unicolor&quot; form [u]. D Papua New Guinea, the melanic form [m]. E Queensland, the &quot;fasciata&quot; form [f]. F Philippine, the &quot;sellata&quot; form [s]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959049" httpUri="https://zenodo.org/record/7959049/files/figure.png" pageId="91" pageNumber="92">55C</figureCitation>
&gt;.
</materialsCitation>
<materialsCitation id="3B46CD68621DFFB82E30B5DEFB365AB9" ID-GBIF-Occurrence="4115871487" collectingDate="1937-08" collectorName="C. J. Louwerens &amp; G. Tompoe-Paloe" country="Papua New Guinea" elevation="4" location="Sulawesi" municipality="Timor" pageId="91" pageNumber="92" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
1 ex [u] (NHML), &quot;O.-Java; Tengger, Nongkodjadjar,
<quantity id="4CD66AD0621DFFB82DEFB5B2FF085AB9" box="[151,239,1993,2019]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="91" pageNumber="92" unit="m" value="1300.0">
<elevation id="00032006621DFFB82DEFB5B2FF085AB9" box="[151,239,1993,2019]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="91" pageNumber="92" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, Wegener&quot;, &quot;Gesch.12.1935, von Overbeck&quot;, &quot;
<collectorName id="26DBA2E3621DFFB82F83B5B2FBE35AB9" box="[763,1028,1993,2019]" pageId="91" pageNumber="92">F. Van Emden Bequest.</collectorName>
B.M. 1960-129.&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68621DFFBF29A1B5B3FD7C5DEB" ID-GBIF-Occurrence="4115871596" collectingDate="1937-08" collectorName="C. J. Louwerens &amp; G. Tompoe-Paloe" country="Papua New Guinea" elevation="4" lastPageId="92" lastPageNumber="93" location="Sulawesi" municipality="Timor" pageId="91" pageNumber="92" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<emphasis id="B95A1B27621DFFB829A1B5B3FAA65AB8" bold="true" box="[1241,1345,1992,2018]" pageId="91" pageNumber="92">Sumatra</emphasis>
:
<specimenCount id="9D280CBC621DFFB82828B5B2FA7B5AB9" box="[1360,1436,1993,2019]" pageId="91" pageNumber="92" type="male">1 male</specimenCount>
[s] (CRS), &quot;Sumatra,
<collectorName id="26DBA2E3621AFFBF2CFFB2ECFDC55DEB" box="[391,546,151,177]" pageId="92" pageNumber="93">M. Bukittingi</collectorName>
,
<date id="FF90E1F5621AFFBF2F55B2ECFD695DEB" box="[557,654,151,177]" pageId="92" pageNumber="93" value="1993-03">
<collectingDate id="EFD4181D621AFFBF2F55B2ECFD695DEB" box="[557,654,151,177]" pageId="92" pageNumber="93" value="1993-03">III.1993</collectingDate>
</date>
&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68621AFFBF2FD9B2ECFD165D8F" ID-GBIF-Occurrence="4115871376" collectingDate="1937-08" collectorName="C. J. Louwerens &amp; G. Tompoe-Paloe" country="Papua New Guinea" elevation="4" location="Sulawesi" municipality="Timor" pageId="92" pageNumber="93" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<emphasis id="B95A1B27621AFFBF2FD9B2ECFCBA5DEA" bold="true" box="[673,861,150,177]" pageId="92" pageNumber="93">The Philippines</emphasis>
: 1 ex [s] (SNSD), &quot;
<location id="8EF191EE621AFFBF294DB2ECFAF85DEB" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621AFFBF294DB2ECFAF85DEB" box="[1077,1311,151,177]" country="Philippines" name="Mt. Bhnahao Luzon" pageId="92" pageNumber="93">Mt. Bhnahao Luzon</location>
&quot;, &quot;1924 6&quot;, &quot;Pa 1454&quot;, &quot;Staatl. Museum für Tierkunde, Dresden&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68621AFFBF2F84B2C0FE505DA3" ID-GBIF-Occurrence="4115871339" collectingDate="2011-07" country="Philippines" county="Philippines" location="Mindoro" municipality="Haicon" pageId="92" pageNumber="93" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D280CBC621AFFBF2F84B2C0FCB95D8F" box="[764,862,187,213]" pageId="92" pageNumber="93" type="female">1 female</specimenCount>
[t] (CRS), &quot;
<collectingCounty id="62F0BFB9621AFFBF2E9EB2C0FB8F5D8F" box="[998,1128,187,213]" pageId="92" pageNumber="93">Philippines</collectingCounty>
S Luzon, M.
<collectingMunicipality id="6BF55D4F621AFFBF287DB2C0FABE5D8F" box="[1285,1369,187,213]" pageId="92" pageNumber="93">Haicon</collectingMunicipality>
,
<location id="8EF191EE621AFFBF281CB2C0FF2D5DA3" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621AFFBF281CB2C0FF2D5DA3" country="Philippines" county="Philippines" municipality="Haicon" name="Mindoro" pageId="92" pageNumber="93">Mindoro</location>
<date id="FF90E1F5621AFFBF2DAAB2A4FEDC5DA3" box="[210,315,223,249]" pageId="92" pageNumber="93" value="2011-07">
<collectingDate id="EFD4181D621AFFBF2DAAB2A4FEDC5DA3" box="[210,315,223,249]" pageId="92" pageNumber="93" value="2011-07">VII.2011</collectingDate>
</date>
&quot; &lt;
<figureCitation id="1315DBB0621AFFBF2C23B2A4FE505DA3" box="[347,439,222,249]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="92" pageNumber="93">Fig. 53I</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD68621AFFBF2CABB2A4FA955DA3" ID-GBIF-Occurrence="4115871481" box="[467,1394,222,249]" collectingDate="2014-08" collectorName="E. Luzon" country="Philippines" location="Dindin" municipality="Philippines" pageId="92" pageNumber="93" specimenCount="1" specimenCount-male="1" stateProvince="Isabela">
<specimenCount id="9D280CBC621AFFBF2CABB2A4FDF85DA3" box="[467,543,223,249]" pageId="92" pageNumber="93" type="male">1 male</specimenCount>
[r] (CRS), &quot;
<collectingMunicipality id="6BF55D4F621AFFBF2FD2B2A4FCCB5DA3" box="[682,812,223,249]" pageId="92" pageNumber="93">Philippines</collectingMunicipality>
-
<collectorName id="26DBA2E3621AFFBF2E3EB2A4FC575DA3" box="[838,944,223,249]" pageId="92" pageNumber="93">E. Luzon</collectorName>
, Isabela,
<location id="8EF191EE621AFFBF2964B2A4FB8A5DA3" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621AFFBF2964B2A4FB8A5DA3" box="[1052,1133,223,249]" country="Philippines" municipality="Philippines" name="Dindin" pageId="92" pageNumber="93" stateProvince="Isabela">Dindin</location>
,
<date id="FF90E1F5621AFFBF2900B2A4FB0C5DA3" box="[1144,1259,223,249]" pageId="92" pageNumber="93" value="2014-08">
<collectingDate id="EFD4181D621AFFBF2900B2A4FB0C5DA3" box="[1144,1259,223,249]" pageId="92" pageNumber="93" value="2014-08">VIII.2014</collectingDate>
</date>
&quot; &lt;
<figureCitation id="1315DBB0621AFFBF2873B2A4FA955DA3" box="[1291,1394,222,249]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="92" pageNumber="93">Fig. 53H</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD68621AFFBF28F6B2A4FC995C47" ID-GBIF-Occurrence="4115871572" collectingDate="2013-04" country="Philippines" location="Luzon" municipality="Philippines" pageId="92" pageNumber="93" specimenCount="1" specimenCount-male="1" stateProvince="Ifugao">
<specimenCount id="9D280CBC621AFFBF28F6B2A4FF2A5C47" pageId="92" pageNumber="93" type="male">1 male</specimenCount>
[h] (CRS), &quot;
<collectingMunicipality id="6BF55D4F621AFFBF2C24B378FE065C47" box="[348,481,259,285]" pageId="92" pageNumber="93">Philippines</collectingMunicipality>
, N
<location id="8EF191EE621AFFBF2F70B378FDB55C47" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621AFFBF2F70B378FDB55C47" box="[520,594,259,285]" country="Philippines" municipality="Philippines" name="Luzon" pageId="92" pageNumber="93" stateProvince="Ifugao">Luzon</location>
, Ifugao
<date id="FF90E1F5621AFFBF2FD7B378FCEC5C47" box="[687,779,259,285]" pageId="92" pageNumber="93" value="2013-04">
<collectingDate id="EFD4181D621AFFBF2FD7B378FCEC5C47" box="[687,779,259,285]" pageId="92" pageNumber="93" value="2013-04">IV.2013</collectingDate>
</date>
&quot; &lt;
<figureCitation id="1315DBB0621AFFBF2E53B378FC995C47" box="[811,894,258,285]" captionStart="FIGURE 54" captionStartId="93.[151,250,1150,1174]" captionTargetBox="[162,1422,193,1116]" captionTargetId="figure-19@93.[151,1436,181,1126]" captionTargetPageId="93" captionText="FIGURE 54. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir), right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Philippine, the &quot;hastata&quot; form [h]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959047" httpUri="https://zenodo.org/record/7959047/files/figure.png" pageId="92" pageNumber="93">Fig. 54</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD68621AFFBF2EE1B378FC1E5C3F" ID-GBIF-Occurrence="4115871506" collectingDate="2016-01" collectingDateMax="2016-02" collectingDateMin="2016-01" country="Vietnam" county="Philippines" location="Tanay" municipality="Luzon" pageId="92" pageNumber="93" specimenCount="1" specimenCount-male="1" stateProvince="Rizal">
<specimenCount id="9D280CBC621AFFBF2EE1B378FC025C47" box="[921,997,259,285]" pageId="92" pageNumber="93" type="male">1 male</specimenCount>
[s] (IZAS), &quot;
<collectingCounty id="62F0BFB9621AFFBF2902B378FAE75C47" box="[1146,1280,259,285]" pageId="92" pageNumber="93">Philippines</collectingCounty>
:
<collectingMunicipality id="6BF55D4F621AFFBF2873B378FAB25C47" box="[1291,1365,259,285]" pageId="92" pageNumber="93">Luzon</collectingMunicipality>
, Rizal prov.,
<location id="8EF191EE621AFFBF2DA7B35CFEC35C1B" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621AFFBF2DA7B35CFEC35C1B" box="[223,292,295,321]" country="Vietnam" county="Philippines" municipality="Luzon" name="Tanay" pageId="92" pageNumber="93" stateProvince="Rizal">Tanay</location>
,
<location id="8EF191EE621AFFBF2C49B35CFE2A5C1B" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621AFFBF2C49B35CFE2A5C1B" box="[305,461,295,321]" country="Vietnam" county="Philippines" municipality="Luzon" name="Sierra Madre." pageId="92" pageNumber="93" stateProvince="Rizal">Sierra Madre.</location>
<date id="FF90E1F5621AFFBF2CADB35CFDA45C1B" box="[469,579,295,321]" pageId="92" pageNumber="93" value="2016-02">
<collectingDate id="EFD4181D621AFFBF2CADB35CFDA45C1B" box="[469,579,295,321]" pageId="92" pageNumber="93" value="2016-02">Feb 2016</collectingDate>
</date>
. local collector&quot; &lt;
<figureCitation id="1315DBB0621AFFBF2E6BB35CFC815C1B" box="[787,870,295,321]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="92" pageNumber="93">Figs 8I</figureCitation>
,
<figureCitation id="1315DBB0621AFFBF2E0CB35DFC455C1B" box="[884,930,294,321]" captionStart="FIGURE 55" captionStartId="94.[151,250,1504,1528]" captionTargetBox="[163,1420,191,1453]" captionTargetId="figure-28@94.[151,1436,181,1481]" captionTargetPageId="94" captionText="FIGURE 55. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir) from different localities and in different elytra pattern forms, left-lateral (l) and dorsal (d) views: A Taiwan, the &quot;sellata&quot; form [s]. B Tonkin, the rhombic form [r]. C Java, the &quot;unicolor&quot; form [u]. D Papua New Guinea, the melanic form [m]. E Queensland, the &quot;fasciata&quot; form [f]. F Philippine, the &quot;sellata&quot; form [s]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959049" httpUri="https://zenodo.org/record/7959049/files/figure.png" pageId="92" pageNumber="93">55F</figureCitation>
&gt;. 3 ex [2s, 1f] (IZAS), &quot;Philippines: Luzon, Quirino prov., Maddela, Disimongal, Sierra Madre.
<date id="FF90E1F5621AFFBF2FAFB330FCDA5C3F" box="[727,829,331,357]" pageId="92" pageNumber="93" value="2016-01">
<collectingDate id="EFD4181D621AFFBF2FAFB330FCDA5C3F" box="[727,829,331,357]" pageId="92" pageNumber="93" value="2016-01">Jan 2016</collectingDate>
</date>
, local collector&quot;.
</materialsCitation>
<materialsCitation id="3B46CD68621AFFBF2E86B331FD855CD3" ID-GBIF-Occurrence="4115871482" collectingDate="2016-01" collectingDateMax="2016-02" collectingDateMin="2016-01" country="Vietnam" county="Philippines" location="Tanay" municipality="Luzon" pageId="92" pageNumber="93" specimenCount="1" specimenCount-male="1" stateProvince="Rizal">
<emphasis id="B95A1B27621AFFBF2E86B331FB835C3E" bold="true" box="[1022,1124,330,356]" pageId="92" pageNumber="93">Vietnam</emphasis>
:
<specimenCount id="9D280CBC621AFFBF2909B330FB5C5C3F" box="[1137,1211,331,357]" pageId="92" pageNumber="93" type="male">1 male</specimenCount>
[r] (MNHN), &quot;Hoa-Binh, Tonkin, A. de Cooman&quot; &lt;
<figureCitation id="1315DBB0621AFFBF2C85B314FD855CD3" box="[509,610,366,393]" captionStart="FIGURE 55" captionStartId="94.[151,250,1504,1528]" captionTargetBox="[163,1420,191,1453]" captionTargetId="figure-28@94.[151,1436,181,1481]" captionTargetPageId="94" captionText="FIGURE 55. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir) from different localities and in different elytra pattern forms, left-lateral (l) and dorsal (d) views: A Taiwan, the &quot;sellata&quot; form [s]. B Tonkin, the rhombic form [r]. C Java, the &quot;unicolor&quot; form [u]. D Papua New Guinea, the melanic form [m]. E Queensland, the &quot;fasciata&quot; form [f]. F Philippine, the &quot;sellata&quot; form [s]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959049" httpUri="https://zenodo.org/record/7959049/files/figure.png" pageId="92" pageNumber="93">Fig. 55B</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD68621AFFBF2F05B314FA7A5CD3" ID-GBIF-Occurrence="4115871499" box="[637,1437,367,393]" collectingDate="1997-04-08" collectingDateMax="1997-04-13" collectingDateMin="1997-04-08" collectorName="A. de Cooman &amp; Jiri Kolibac" country="Thailand" county="Tonkin" elevation="200" location="Thailand" municipality="Hoa-Binh" pageId="92" pageNumber="93" specimenCount="1" specimenCount-male="1" stateProvince="Satun">
<specimenCount id="9D280CBC621AFFBF2F05B314FD2E5CD3" box="[637,713,367,393]" pageId="92" pageNumber="93" type="male">1 male</specimenCount>
[s] (IZAS), &quot;
<collectingCounty id="62F0BFB9621AFFBF2E24B314FC555CD3" box="[860,946,367,393]" pageId="92" pageNumber="93">Tonkin</collectingCounty>
,
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, 1940.VIII, leg.
<collectorName id="26DBA2E3621AFFBF299FB314FA765CD3" box="[1255,1425,367,393]" pageId="92" pageNumber="93">A. de Cooman</collectorName>
&quot;.
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<emphasis id="B95A1B27621AFFBF2DEFB3E9FEE35CF6" bold="true" box="[151,260,402,428]" pageId="92" pageNumber="93">Thailand</emphasis>
: 1 ex [s] (NHMB), &quot;
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, S. Satun pr.
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, 6°45'N 100°09'E,
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,
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leg.&quot;
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.
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<emphasis id="B95A1B27621AFFBF2DA2B3CDFEC45C8A" bold="true" box="[218,291,438,464]" pageId="92" pageNumber="93">China</emphasis>
:
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[s] (SYUM), &quot;Hainan,
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,
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,
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<collectingDate id="EFD4181D621AFFBF2EE0B3CCFBF25C8B" box="[920,1045,439,465]" pageId="92" pageNumber="93" value="1981-07-03">1981.VII.3</collectingDate>
</date>
, Chen Zhenyue lgt.&quot;
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.
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[s] (IZAS), &quot;Taiwan, forestry test station&quot;.
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[s] (CCCC), &quot;
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,
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&quot;
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.
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[s] (CCCC), &quot;
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,
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,
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N&quot; &lt;
<figureCitation id="1315DBB0621AFFBF2941B384FB475F43" box="[1081,1184,511,537]" captionStart="FIGURE 10" captionStartId="16.[151,250,1909,1933]" captionTargetBox="[163,1429,1250,1879]" captionTargetId="figure-182@16.[152,1436,1241,1885]" captionTargetPageId="16" captionText="FIGURE 10. Parameres of male genitalia for Parena spp. A P. (Crossoglossa) cavipennis (Bates), Beijing. B P. (Crossoglossa) testacea (Chaudoir), Shanghai. C P. (Bothynoptera) dorsigera (Schaum), Zhejiang. D P. (Bothynoptera) taiwana Hua, Taiwan. E P. (Bothynoptera) tripunctata (Bates), Shaanxi. F P. (Parena) rubripicta Andrewes, Xizang. G P. (Parena) fasciata (Chaudoir), Taiwan. H P. (Parena) nigrolineata (Chaudoir), Taiwan. I P. (Parena) latecincta (Bates), Yunnan. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7958957" httpUri="https://zenodo.org/record/7958957/files/figure.png" pageId="92" pageNumber="93">Figs 10G</figureCitation>
,
<figureCitation id="1315DBB0621AFFBF29D6B384FB035F43" box="[1198,1252,511,537]" captionStart="FIGURE 11" captionStartId="18.[151,250,150,174]" captionTargetId="figure-19@17.[155,1432,181,1971]" captionTargetPageId="17" captionText="FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). AJ subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. KX subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Yγ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7963954" httpUri="https://zenodo.org/record/7963954/files/figure.png" pageId="92" pageNumber="93">11M</figureCitation>
,
<figureCitation id="1315DBB0621AFFBF2997B385FAC55F43" box="[1263,1314,510,537]" captionStart="FIGURE 55" captionStartId="94.[151,250,1504,1528]" captionTargetBox="[163,1420,191,1453]" captionTargetId="figure-28@94.[151,1436,181,1481]" captionTargetPageId="94" captionText="FIGURE 55. Median lobe of male genitalia for Parena (Parena) fasciata (Chaudoir) from different localities and in different elytra pattern forms, left-lateral (l) and dorsal (d) views: A Taiwan, the &quot;sellata&quot; form [s]. B Tonkin, the rhombic form [r]. C Java, the &quot;unicolor&quot; form [u]. D Papua New Guinea, the melanic form [m]. E Queensland, the &quot;fasciata&quot; form [f]. F Philippine, the &quot;sellata&quot; form [s]. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7959049" httpUri="https://zenodo.org/record/7959049/files/figure.png" pageId="92" pageNumber="93">55A</figureCitation>
</materialsCitation>
&gt;.
<materialsCitation id="3B46CD68621AFFBF2844B384FBD75F67" ID-GBIF-Occurrence="4115871587" collectingDate="2008-07-21" country="China" county="Taiwan" location="Lilongshan" municipality="Pingtung county" pageId="92" pageNumber="93" specimenCount="1" specimenCount-female="1">
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[s] (CCCC), &quot;
<collectingCounty id="62F0BFB9621AFFBF2C54B058FE635F67" box="[300,388,547,573]" pageId="92" pageNumber="93">Taiwan</collectingCounty>
,
<collectingMunicipality id="6BF55D4F621AFFBF2CF7B058FDA15F67" box="[399,582,547,573]" pageId="92" pageNumber="93">Pingtung county</collectingMunicipality>
,
<location id="8EF191EE621AFFBF2F28B058FD365F67" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621AFFBF2F28B058FD365F67" box="[592,721,547,573]" country="China" county="Taiwan" municipality="Pingtung county" name="Lilongshan" pageId="92" pageNumber="93">Lilongshan</location>
,
<date id="FF90E1F5621AFFBF2FA3B058FC805F67" box="[731,871,547,573]" pageId="92" pageNumber="93" value="2008-07-21">
<collectingDate id="EFD4181D621AFFBF2FA3B058FC805F67" box="[731,871,547,573]" pageId="92" pageNumber="93" value="2008-07-21">2008.VII.21</collectingDate>
</date>
, Chou Wen-I lgt.&quot;
</materialsCitation>
.
<materialsCitation id="3B46CD68621AFFBF2942B058FCAC5F3B" ID-GBIF-Occurrence="4115871357" collectingDate="1996-09-11" country="China" county="Taiwan" location="Kenting" municipality="Pingtung county" pageId="92" pageNumber="93" specimenCount="2" specimenCount-female="2">
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[s] (CCCC), &quot;
<collectingCounty id="62F0BFB9621AFFBF2838B058FA7F5F67" box="[1344,1432,547,573]" pageId="92" pageNumber="93">Taiwan</collectingCounty>
,
<collectingMunicipality id="6BF55D4F621AFFBF2DEFB03CFEB75F3B" box="[151,336,583,609]" pageId="92" pageNumber="93">Pingtung county</collectingMunicipality>
,
<location id="8EF191EE621AFFBF2C24B03CFE515F3B" LSID="urn:lsid:plazi:treatment:03877623621FFFBE2DEFB48CFE925AD0:8EF191EE621AFFBF2C24B03CFE515F3B" box="[348,438,583,609]" country="China" county="Taiwan" municipality="Pingtung county" name="Kenting" pageId="92" pageNumber="93">Kenting</location>
park,
<date id="FF90E1F5621AFFBF2C85B03CFD995F3B" box="[509,638,583,609]" pageId="92" pageNumber="93" value="1996-09-11">
<collectingDate id="EFD4181D621AFFBF2C85B03CFD995F3B" box="[509,638,583,609]" pageId="92" pageNumber="93" value="1996-09-11">1996.IX.11</collectingDate>
</date>
, Chou Wen-I lgt.&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621AFFBF2DBFB011FE945EB7" pageId="92" pageNumber="93" type="diagnosis">
<paragraph id="8B91C735621AFFBF2DBFB011FE0C5F97" blockId="92.[151,1437,150,1869]" pageId="92" pageNumber="93">
<emphasis id="B95A1B27621AFFBF2DBFB011FE815FDE" bold="true" box="[199,358,618,644]" pageId="92" pageNumber="93">Comparisons</emphasis>
. This species can be distinguished from all other Oriental-Australian species in subgenus
<taxonomicName id="4C2EBCB6621AFFBF2833B010FA7B5FDE" box="[1355,1436,619,644]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27621AFFBF2833B010FA7B5FDE" box="[1355,1436,619,644]" italics="true" pageId="92" pageNumber="93">Parena</emphasis>
</taxonomicName>
by the combination of the following two characters: (1) elytra yellow, with or without black patch; (2) postgenae with a pair of suborbital setae.
</paragraph>
<paragraph id="8B91C735621AFFBF2DBFB0ACFA7A5E07" blockId="92.[151,1437,150,1869]" pageId="92" pageNumber="93">
The &quot;unicolor&quot; form of
<taxonomicName id="4C2EBCB6621AFFBF2CA3B0ACFDB55FAB" box="[475,594,727,753]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621AFFBF2CA3B0ACFDB55FAB" box="[475,594,727,753]" italics="true" pageId="92" pageNumber="93">P. fasciata</emphasis>
</taxonomicName>
(
<figureCitation id="1315DBB0621AFFBF2F1BB0ACFCEE5FAB" box="[611,777,726,753]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="92" pageNumber="93">Figs 53E, 53F</figureCitation>
) is similar to some other uniformly brown species, especially those African species of the
<taxonomicName id="4C2EBCB6621AFFBF2F63B080FD4B5E4E" baseAuthorityName="Fairmaire" baseAuthorityYear="1899" box="[539,684,763,788]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="stigmatica">
<emphasis id="B95A1B27621AFFBF2F63B080FD4B5E4E" box="[539,684,763,788]" italics="true" pageId="92" pageNumber="93">P. stigmatica</emphasis>
</taxonomicName>
group, which also have the suborbital setae.
<taxonomicName id="4C2EBCB6621AFFBF29DEB080FAFC5E4F" box="[1190,1307,763,789]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621AFFBF29DEB080FAFC5E4F" box="[1190,1307,763,789]" italics="true" pageId="92" pageNumber="93">P. fasciata</emphasis>
</taxonomicName>
is different from these African species in having the antennae completely yellow, versus apical seven antennomeres black. We provide a key for all uniformly yellowish brown species belonging to the subgenus
<taxonomicName id="4C2EBCB6621AFFBF2947B138FB775E06" box="[1087,1168,835,860]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27621AFFBF2947B138FB775E06" box="[1087,1168,835,860]" italics="true" pageId="92" pageNumber="93">Parena</emphasis>
</taxonomicName>
at the end of this paper.
</paragraph>
<paragraph id="8B91C735621AFFBF2DBFB11CFE945EB7" blockId="92.[151,1437,150,1869]" pageId="92" pageNumber="93">
In
<taxonomicName id="4C2EBCB6621AFFBF2D9EB11CFEBC5EDB" box="[230,347,871,897]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621AFFBF2D9EB11CFEBC5EDB" box="[230,347,871,897]" italics="true" pageId="92" pageNumber="93">P. fasciata</emphasis>
</taxonomicName>
, the rhombic form (
<figureCitation id="1315DBB0621AFFBF2F47B11CFD425EDB" box="[575,677,870,897]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="92" pageNumber="93">Fig. 53H</figureCitation>
) is somewhat similar to
<taxonomicName id="4C2EBCB6621AFFBF2EC4B11CFB865EDA" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[956,1121,871,896]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="monostigma">
<emphasis id="B95A1B27621AFFBF2EC4B11CFB865EDA" box="[956,1121,871,896]" italics="true" pageId="92" pageNumber="93">P. monostigma</emphasis>
</taxonomicName>
, but differs in having elytra patch wider, extended laterally at least to interval 5, and the different position of first elytral discal pore. The melanic form (
<figureCitation id="1315DBB0621AFFBF2DA6B1D4FEA55E93" box="[222,322,942,969]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="92" pageNumber="93">Fig. 53G</figureCitation>
) is similar to the dark form of
<taxonomicName id="4C2EBCB6621AFFBF2FD9B1D4FCFB5E93" baseAuthorityName="Alluaud" baseAuthorityYear="1917" box="[673,796,943,969]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B95A1B27621AFFBF2FD9B1D4FCFB5E93" box="[673,796,943,969]" italics="true" pageId="92" pageNumber="93">P. africana</emphasis>
</taxonomicName>
, but differs in having suborbital setae and the elytra disc without depression.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621AFFBF2DBFB18DFD6E5B2F" pageId="92" pageNumber="93" type="description">
<paragraph id="8B91C735621AFFBF2DBFB18DFD6E5B2F" blockId="92.[151,1437,150,1869]" pageId="92" pageNumber="93">
<emphasis id="B95A1B27621AFFBF2DBFB18DFEB6594A" bold="true" box="[199,337,1014,1040]" pageId="92" pageNumber="93">Description</emphasis>
. Body length
<quantity id="4CD66AD0621AFFBF2C8EB18CFD65594B" box="[502,642,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" metricValueMax="8.9" metricValueMin="6.7" pageId="92" pageNumber="93" unit="mm" value="7.8" valueMax="8.9" valueMin="6.7">6.78.9 mm</quantity>
. Dorsum reddish yellow to yellowish brown, elytra with varied black patch, a single large patch behind middle in most specimens, without a black patch in some specimens; antennae, legs, and venter yellow. Postgenae with a pair of suborbital setae; antennae extended to elytral base; labrum quadrate, apex straight or very weakly concave; mentum with a pair of short median setae. Pronotum nearly cordate, PW/PL = 1.341.47, wider than head, PW/HW = 1.081.15; widest at anterior third, slightly rounded at anterior half, more or less sinuate before posterior angles; posterior angles widely obtuse. Elytra slightly dilated to apex, surface without or with very faint isodiametric microsculpture; striae very shallowly incised, with fine puncture rows; intervals very sparsely punctate; disc without depression at basal third. Males with biseriate adhesive setae on apical two-thirds of mesotarsomere 1 and almost full length of mesotarsomeres 2 and 3. Apex of abdominal sternite VII clearly emarginate in males (
<figureCitation id="1315DBB0621AFFBF2F6EB740FD81580F" box="[534,614,1339,1365]" captionStart="FIGURE 8" captionStartId="14.[151,250,1865,1889]" captionTargetBox="[198,1388,1017,1840]" captionTargetId="figure-59@14.[198,1388,1017,1841]" captionTargetPageId="14" captionText="FIGURE 8. Apex of sternite VII of Parena spp. AD females: A P. (Crossoglossa) sciakyi sp. n., holotype. B P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. C P. (Crossoglossa) mellea (Chaudoir), Philippine. D P. (Crossoglossa) cruralis Andrewes, Borneo. EJ males: E P. (Crossoglossa) cavipennis (Bates), Tianjin. F P. (Crossoglossa) mellea (Chaudoir), Philippine. G P. (Parena) monticola Shibata, Jiangxi. H P. (Parena) picipes sp. n., Borneo, paratype. I P. (Parena) fasciata (Chaudoir), Philippine. J P. (Parena) andrewesi Jedlička, Philippine. Scale bar = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958953" httpUri="https://zenodo.org/record/7958953/files/figure.png" pageId="92" pageNumber="93">Fig. 8I</figureCitation>
). Median lobe of aedeagus slender (AL/AW = 4.85.7), ventral margin gradually curved; apical third more or less bent to right side; apical lamella nearly ovate, LL subequal to LW, apex rounded, weakly bent upward in lateral view. Endophallus with primary sclerite small and linear, about one-seventh length of median lobe, with indistinct flared basal expansion; apical flagellum very short, ending near basal third of median lobe; apical sclerite widely V-shaped, well chitinized, basal core indistinct; basal sheath very small, dorsally surrounding apical half of primary sclerite; apical sheath simple, finely scaled; squamate sac not or partly divided, in similar size as basal sheath, right to squamate sheath. Gonocoxite II of ovipositor nearly quadrate, slightly wider than length, apex slightly concave, slightly projected near inner apical angle, with three ensiform setae on inner apical angle, two or three on outer apical angle.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE621AFFBE2DBFB405FE925AD0" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93" type="distribution">
<paragraph id="8B91C735621AFFBF2DBFB405FCA35BE7" blockId="92.[151,1437,150,1869]" pageId="92" pageNumber="93">
<emphasis id="B95A1B27621AFFBF2DBFB405FEBE5BC2" bold="true" box="[199,345,1662,1688]" pageId="92" pageNumber="93">Distribution</emphasis>
(
<figureCitation id="1315DBB0621AFFBF2C1FB404FE555BC3" box="[359,434,1663,1689]" captionStart="MAP 9" captionStartId="84.[152,211,1891,1915]" captionTargetBox="[151,1436,1041,1868]" captionTargetId="figure-773@84.[151,1436,1041,1868]" captionText="MAP 9. Distributions for species in the Parena (Parena) bicolor group." figureDoi="http://doi.org/10.5281/zenodo.7963679" httpUri="https://zenodo.org/record/7963679/files/figure.png" pageId="92" pageNumber="93">Map 9</figureCitation>
, blue).
<collectingCountry id="F33987A5621AFFBF2F7AB404FD8B5BC3" box="[514,620,1663,1689]" name="Australia" pageId="92" pageNumber="93">Australia</collectingCountry>
,
<collectingCountry id="F33987A5621AFFBF2F0DB404FCA85BC3" box="[629,847,1663,1689]" name="Papua New Guinea" pageId="92" pageNumber="93">Papua New Guinea</collectingCountry>
, Sulawesi, Maluku, Timor, Java, Sumatra, the
<collectingCountry id="F33987A5621AFFBF2828B404FF325BE7" name="Philippines" pageId="92" pageNumber="93">Philippines</collectingCountry>
,
<collectingCountry id="F33987A5621AFFBF2D98B4D8FEA55BE7" box="[224,322,1699,1725]" name="Vietnam" pageId="92" pageNumber="93">Vietnam</collectingCountry>
,
<collectingCountry id="F33987A5621AFFBF2C34B4D8FE555BE7" box="[332,434,1699,1725]" name="Thailand" pageId="92" pageNumber="93">Thailand</collectingCountry>
,
<collectingCountry id="F33987A5621AFFBF2CC6B4D8FDE65BE7" box="[446,513,1699,1725]" name="China" pageId="92" pageNumber="93">China</collectingCountry>
(
<collectingCountry id="F33987A5621AFFBF2F77B4D8FD815BE7" box="[527,614,1699,1725]" name="Taiwan" pageId="92" pageNumber="93">Taiwan</collectingCountry>
, Hainan, Zhejiang).
</paragraph>
<paragraph id="8B91C735621AFFBF2DBFB4BDFCA75A17" blockId="92.[151,1437,150,1869]" pageId="92" pageNumber="93">
<emphasis id="B95A1B27621AFFBF2DBFB4BDFEDF5BBA" bold="true" box="[199,312,1734,1760]" pageId="92" pageNumber="93">Variation</emphasis>
.
<taxonomicName id="4C2EBCB6621AFFBF2C3EB4BCFE5B5BBB" box="[326,444,1735,1761]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621AFFBF2C3EB4BCFE5B5BBB" box="[326,444,1735,1761]" italics="true" pageId="92" pageNumber="93">P. fasciata</emphasis>
</taxonomicName>
is widely distributed through Southeast Asia and the north part of Australasia. Its elytra color is highly varied, as might be expected with such a distribution. Generally, the following seven color forms can be recognized, but with intermediate specimens also seen. In the citations of examined materials above, we recorded the color form of each specimen by abbreviation in brackets.
</paragraph>
<caption id="DF5197BD621BFFBE2DEFB605FC7B59E3" ID-DOI="http://doi.org/10.5281/zenodo.7959047" ID-Zenodo-Dep="7959047" httpUri="https://zenodo.org/record/7959047/files/figure.png" pageId="93" pageNumber="94" startId="93.[151,250,1150,1174]" targetBox="[162,1422,193,1116]" targetPageId="93" targetType="figure">
<paragraph id="8B91C735621BFFBE2DEFB605FC7B59E3" blockId="93.[151,1437,1150,1210]" pageId="93" pageNumber="94">
<emphasis id="B95A1B27621BFFBE2DEFB605FEFF59CC" bold="true" box="[151,280,1150,1174]" pageId="93" pageNumber="94">FIGURE 54</emphasis>
. Median lobe of male genitalia for
<taxonomicName id="4C2EBCB6621BFFBE2F01B604FC0859CC" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[633,1007,1150,1174]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fasciata" subGenus="Parena">
<emphasis id="B95A1B27621BFFBE2F01B604FD2459CC" box="[633,707,1151,1174]" italics="true" pageId="93" pageNumber="94">Parena</emphasis>
(
<emphasis id="B95A1B27621BFFBE2FA8B604FCFD59CC" box="[720,794,1151,1174]" italics="true" pageId="93" pageNumber="94">Parena</emphasis>
)
<emphasis id="B95A1B27621BFFBE2E50B605FC9F59CC" box="[808,888,1150,1174]" italics="true" pageId="93" pageNumber="94">fasciata</emphasis>
(Chaudoir)
</taxonomicName>
, right-lateral (r), ventral (v), left-lateral (l), and dorsal (d) views, Philippine, the &quot;hastata&quot; form [h]. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B91C735621BFFBE2DEFB777FC3458C8" blockId="93.[151,1437,1292,1930]" pageId="93" pageNumber="94">
(1) The &quot;fasciata&quot; form [f] (
<figureCitation id="1315DBB0621BFFBE2CAAB777FDDD587D" box="[466,570,1292,1319]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="93" pageNumber="94">Fig. 53A</figureCitation>
): elytra with a transverse black patch.
<typeStatus id="54957997621BFFBE2E9AB776FBFD587D" box="[994,1050,1293,1319]" pageId="93" pageNumber="94">Type</typeStatus>
of
<taxonomicName id="4C2EBCB6621BFFBE2945B776FB56587C" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[1085,1201,1292,1318]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B95A1B27621BFFBE2945B776FB56587C" box="[1085,1201,1292,1318]" italics="true" pageId="93" pageNumber="94">P. fasciata</emphasis>
</taxonomicName>
belongs to this form. Most specimens from regions eastern to the Wallaces line are of the typical &quot;fasciata&quot; form, which has a very large black patch near elytral mid-length. A few specimens from Java and the
<collectingCountry id="F33987A5621BFFBE293AB72FFB275834" box="[1090,1216,1364,1390]" name="Philippines" pageId="93" pageNumber="94">Philippines</collectingCountry>
also have the &quot;fasciata&quot; form, but the patch is usually shorter than in typical specimens.
</paragraph>
<paragraph id="8B91C735621BFFBE2DEFB7E7FE0858A4" blockId="93.[151,1437,1292,1930]" pageId="93" pageNumber="94">
(2) The &quot;sellata&quot; form [s] (
<figureCitation id="1315DBB0621BFFBE2CB4B7E7FD9E58ED" box="[460,633,1436,1463]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="93" pageNumber="94">Figs 53B, 53D</figureCitation>
): elytra with a triangular black patch.
<typeStatus id="54957997621BFFBE2950B7E6FB8D58ED" box="[1064,1130,1437,1463]" pageId="93" pageNumber="94">Types</typeStatus>
of
<taxonomicName id="4C2EBCB6621BFFBE29EAB7E6FB1E58EC" baseAuthorityName="Heller" baseAuthorityYear="1921" box="[1170,1273,1436,1462]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="sellata">
<emphasis id="B95A1B27621BFFBE29EAB7E6FB1E58EC" box="[1170,1273,1436,1462]" italics="true" pageId="93" pageNumber="94">P. sellata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2EBCB6621BFFBE2870B7E6FA7B58EC" authorityName="Jedlicka" authorityYear="1940" box="[1288,1436,1436,1462]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="sellatoides">
<emphasis id="B95A1B27621BFFBE2870B7E6FA7B58EC" box="[1288,1436,1436,1462]" italics="true" pageId="93" pageNumber="94">P. sellatoides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2EBCB6621BFFBE2D8CB7BAFE605880" authorityName="Kirschenhofer" authorityYear="2011" box="[244,391,1473,1498]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="sumatrana">
<emphasis id="B95A1B27621BFFBE2D8CB7BAFE605880" box="[244,391,1473,1498]" italics="true" pageId="93" pageNumber="94">P. sumatrana</emphasis>
</taxonomicName>
all have this form. It is the most common form in regions west of Wallaces line, but not found in regions east of that line.
</paragraph>
<paragraph id="8B91C735621BFFBE2DEFB473FA9F5B1C" blockId="93.[151,1437,1292,1930]" pageId="93" pageNumber="94">
(3) The &quot;unicolor&quot; form [u] (
<figureCitation id="1315DBB0621BFFBE2CA7B473FD655B79" box="[479,642,1544,1571]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="93" pageNumber="94">Figs 53E, 53F</figureCitation>
): elytra unicolorous, without a black patch.
<typeStatus id="54957997621BFFBE2911B472FB465B79" box="[1129,1185,1545,1571]" pageId="93" pageNumber="94">Type</typeStatus>
of
<taxonomicName id="4C2EBCB6621BFFBE29BDB472FAD85B78" authorityName="Louwerens" authorityYear="1949" box="[1221,1343,1544,1570]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B95A1B27621BFFBE29BDB472FAD85B78" box="[1221,1343,1544,1570]" italics="true" pageId="93" pageNumber="94">P. unicolor</emphasis>
</taxonomicName>
belongs to this form. The &quot;unicolor&quot; form occurs in
<collectingRegion id="49EA09D7621BFFBE2FD6B456FD075B1D" box="[686,736,1581,1607]" country="Indonesia" name="Jawa Timur" pageId="93" pageNumber="94">Java</collectingRegion>
and Timor, and is relatively common in Javan populations.
</paragraph>
<paragraph id="8B91C735621BFFBE2DEFB42BFD6E5BD4" blockId="93.[151,1437,1292,1930]" pageId="93" pageNumber="94">
(4) The rhombic form [r] (
<figureCitation id="1315DBB0621BFFBE2CB0B42BFDD45B31" box="[456,563,1616,1643]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="93" pageNumber="94">Fig. 53H</figureCitation>
): elytra with a narrow rhombic black patch. This form occurs in
<collectingCountry id="F33987A5621BFFBE2875B42BFA885B30" box="[1293,1391,1616,1642]" name="Vietnam" pageId="93" pageNumber="94">Vietnam</collectingCountry>
, the
<collectingCountry id="F33987A5621BFFBE2DBEB40FFEA35BD4" box="[198,324,1652,1678]" name="Philippines" pageId="93" pageNumber="94">Philippines</collectingCountry>
and Java at a low frequency.
</paragraph>
<paragraph id="8B91C735621BFFBE2DEFB4E3FBD95B8C" blockId="93.[151,1437,1292,1930]" pageId="93" pageNumber="94">
(5) The &quot;hastata&quot; form [h] (
<figureCitation id="1315DBB0621BFFBE2CABB4E3FDDB5BE9" box="[467,572,1688,1715]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="93" pageNumber="94">Fig. 53C</figureCitation>
): elytra black patch with anterior margin notched.
<typeStatus id="54957997621BFFBE290BB4E2FB4C5BE9" box="[1139,1195,1689,1715]" pageId="93" pageNumber="94">Type</typeStatus>
of
<taxonomicName id="4C2EBCB6621BFFBE29A9B4E2FAD85BE8" baseAuthorityName="Heller" baseAuthorityYear="1921" box="[1233,1343,1688,1714]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="hastata">
<emphasis id="B95A1B27621BFFBE29A9B4E2FAD85BE8" box="[1233,1343,1688,1714]" italics="true" pageId="93" pageNumber="94">P. hastata</emphasis>
</taxonomicName>
has this form. The &quot;hastata&quot; form occurs in Java and the
<collectingCountry id="F33987A5621BFFBE2F9CB4C7FC855B8C" box="[740,866,1724,1750]" name="Philippines" pageId="93" pageNumber="94">Philippines</collectingCountry>
at a low frequency.
</paragraph>
<paragraph id="8B91C735621BFFBE2DEFB49BFEE45A19" blockId="93.[151,1437,1292,1930]" pageId="93" pageNumber="94">
(6) The melanic form [m] (
<figureCitation id="1315DBB0621BFFBE2CB6B49BFDD45BA0" box="[462,563,1760,1787]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="93" pageNumber="94">Fig. 53G</figureCitation>
): elytral black patch very large, with only a narrow apical region brown. This form was found in only
<specimenCount id="9D280CBC621BFFBE2CA0B57EFD945A44" box="[472,627,1796,1823]" pageId="93" pageNumber="94" type="generic">one specimen</specimenCount>
from
<collectingCountry id="F33987A5621BFFBE2FC1B57EFC745A44" box="[697,915,1796,1823]" name="Papua New Guinea" pageId="93" pageNumber="94">Papua New Guinea</collectingCountry>
and is very similar to another species,
<taxonomicName id="4C2EBCB6621BFFBE2836B57EFF1B5A18" baseAuthorityName="Alluaud" baseAuthorityYear="1917" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B95A1B27621BFFBE2836B57EFF1B5A18" italics="true" pageId="93" pageNumber="94">P. africana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B91C735621BFFBE2DEFB537FE925AD0" blockId="93.[151,1437,1292,1930]" pageId="93" pageNumber="94">
(7) The trident form [t] (
<figureCitation id="1315DBB0621BFFBE2CC8B537FDEA5A3D" box="[432,525,1868,1895]" captionStart="FIGURE 53" captionStartId="91.[151,250,150,174]" captionTargetId="figure-28@90.[177,1411,181,2003]" captionTargetPageId="90" captionText="FIGURE 53. Habitus of Parena (Parena) fasciata (Chaudoir). A Holotype of Parena fasciata (Chaudoir). B Holotype of Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)]. C Holotype of Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)]. D Holotype of Crossoglossa sellatoides Jedlička [= Parena fasciata (Chaudoir)]. E Holotype of Parena fasciata var. unicolor Louwerens [= Parena fasciata (Chaudoir)]. F Male from Java, the &quot;unicolor&quot; form [u]. G Male from Papua New Guinea, the melanic form [m]. H Male from Philippine, the rhombic form [r]. I Female from Philippine, the trident form [t]. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7963961" httpUri="https://zenodo.org/record/7963961/files/figure.png" pageId="93" pageNumber="94">Fig. 53I</figureCitation>
): elytra with trident black patch. This form was found in only
<specimenCount id="9D280CBC621BFFBE29BEB536FAB85A3C" box="[1222,1375,1868,1895]" pageId="93" pageNumber="94" type="generic">one specimen</specimenCount>
from the
<collectingCountry id="F33987A5621BFFBE2D97B50BFE965AD0" box="[239,369,1904,1930]" name="Philippines" pageId="93" pageNumber="94">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>