treatments-xml/data/03/86/A4/0386A43AEF59A2198B85FC10FE32FE3D.xml
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<mods:title id="5158073CDC8C1E2EC0072157DE3DCB3C">Review of selected species of Campsurus Eaton 1868 (Ephemeroptera: Polymitarcyidae), with description of eleven new species and a key to male imagos of the genus</mods:title>
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<taxonomicName id="4C2F6EAFEF59A2188B85FC10FD5EFC5B" ID-CoL="QF3K" authority="Molineri &amp; Emmerich" authorityName="Molineri &amp; Emmerich ." authorityYear="2010" box="[151,726,980,1006]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="46" pageNumber="347" phylum="Arthropoda" rank="species" species="amapaensis">
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<emphasis id="B95BC93EEF59A2188B85FC10FE3EFC5B" bold="true" box="[151,438,980,1006]" italics="true" pageId="46" pageNumber="347">Campsurus amapaensis</emphasis>
Molineri &amp; Emmerich
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(
<figureCitation id="131409A9EF59A2188B8DFC32FF77FBBA" box="[159,255,1014,1039]" captionStart="FIGURES 138 145" captionStartId="38.[151,264,1925,1948]" captionTargetBox="[283,1282,265,1875]" captionTargetId="figure@38.[277,1285,265,1880]" captionTargetPageId="38" captionText="FIGURES 138 145. C. segnis, holotype male imago: 138, genitalia, v. v.; 139, penes, v. v. Campsurus cotaxe sp. nov.: 140, genitalia, v. v.; 141, pedestal, v. v.; 142, penes, v. v.; 143, penes, l. v.; 144, female sternum VIII with detail of socket. C. amapaensis: 145, partial view of female sternum VIII with detail of socket." httpUri="https://zenodo.org/record/838472/files/figure.png" pageId="46" pageNumber="347">Fig. 145</figureCitation>
)
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<emphasis id="B95BC93EEF59A2188B85FBFEFE00FBFA" box="[151,392,1082,1103]" italics="true" pageId="46" pageNumber="347">Campsurus amapaensis</emphasis>
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: 276
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.
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:
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i and
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i from
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<location id="8EF043F7EF59A21889CBFBB2FCC9FB3A" LSID="urn:lsid:plazi:treatment:0386A43AEF59A2198B85FC10FE32FE3D:8EF043F7EF59A21889CBFBB2FCC9FB3A" box="[729,833,1142,1167]" country="Brazil" municipality="Material." name="Mineiros" pageId="46" pageNumber="347" stateProvince="Goias">Mineiros</location>
, riacho/ponte, L
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<collectorName id="26DA70FAEF59A2188AF2FB5FFDA3FB06" box="[480,555,1179,1203]" pageId="46" pageNumber="347">Santos</collectorName>
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si,
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si from
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si and
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,
<collectingMethod id="526E6D3BEF59A21889E1FAC2FC70FAAB" box="[755,1016,1286,1311]" pageId="46" pageNumber="347">Pennsylvania light trap</collectingMethod>
,
<collectorName id="26DA70FAEF59A2188F11FAC2FBC1FAAA" box="[1027,1097,1286,1311]" pageId="46" pageNumber="347">Fusari</collectorName>
&amp;
<collectorName id="26DA70FAEF59A2188F7FFAC2FB68FAAA" box="[1133,1248,1286,1311]" pageId="46" pageNumber="347">Fernandes</collectorName>
cols. (
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)
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;
<materialsCitation id="3B471F71EF59A2188E9CFAC3FB7EFAF6" ID-GBIF-Occurrence="1585352579" collectingDate="2012-02-24" collectionCode="CZNC" collectorName="E Raimundi" country="Brazil" latitude="-17.202583" location="Mineiros" longLatPrecision="1" longitude="-52.664" pageId="46" pageNumber="347" specimenCount="1" specimenCount-male="1" stateProvince="Goias">
<specimenCount id="9D29DEA5EF59A2188E9CFAC3FF25FAF6" pageId="46" pageNumber="347" type="male">1 ♂</specimenCount>
i,
<collectingRegion id="49EBDBCEEF59A2188BD4FAEEFE82FAF6" box="[198,266,1322,1347]" country="Brazil" name="Goias" pageId="46" pageNumber="347">Goiás</collectingRegion>
,
<location id="8EF043F7EF59A2188A0BFAEEFE09FAF6" LSID="urn:lsid:plazi:treatment:0386A43AEF59A2198B85FC10FE32FE3D:8EF043F7EF59A2188A0BFAEEFE09FAF6" box="[281,385,1322,1347]" country="Brazil" latitude="-17.202583" longLatPrecision="1" longitude="-52.664" name="Mineiros" pageId="46" pageNumber="347" stateProvince="Goias">Mineiros</location>
, L23,
<collectingDate id="EFD5CA04EF59A2188AC2FAEEFDCCFAF6" box="[464,580,1322,1347]" pageId="46" pageNumber="347" value="2012-02-24">
<date id="FF9133ECEF59A2188AC2FAEEFDCCFAF6" box="[464,580,1322,1347]" pageId="46" pageNumber="347" value="2012-02-24">24.ii.2012</date>
</collectingDate>
,
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/
<geoCoordinate id="EE1B73EBEF59A218881BFAEFFC33FAF6" box="[777,955,1322,1347]" direction="west" orientation="longitude" pageId="46" pageNumber="347" precision="1" value="-52.664">W 52°3950,4”</geoCoordinate>
,
<collectorName id="26DA70FAEF59A21888D9FAEFFBDDFAF6" box="[971,1109,1322,1347]" pageId="46" pageNumber="347">E Raimundi</collectorName>
col. (
<collectionCode id="ED3E8DE9EF59A2188F8FFAEFFB66FAF6" box="[1181,1262,1323,1347]" pageId="46" pageNumber="347">CZNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B471F71EF59A2188E17FAEFFBAAFAD2" ID-GBIF-Occurrence="1585352584" collectingDate="2011-06-01" collectorName="N Hamada" country="Brazil" latitude="-4.168333" location="Domingos Mourao" longLatPrecision="21" longitude="-41.581944" pageId="46" pageNumber="347" specimenCount="30" specimenCount-male="30" stateProvince="Piaui">
<specimenCount id="9D29DEA5EF59A2188E17FAEFFACAFAF6" box="[1285,1346,1322,1347]" pageId="46" pageNumber="347" type="male">30 ♂</specimenCount>
i,
<collectingRegion id="49EBDBCEEF59A2188E49FAEEFA1FFAF6" box="[1371,1431,1322,1347]" country="Brazil" name="Piaui" pageId="46" pageNumber="347">Piauí</collectingRegion>
,
<location id="8EF043F7EF59A2188B85FA8AFEFDFAD3" LSID="urn:lsid:plazi:treatment:0386A43AEF59A2198B85FC10FE32FE3D:8EF043F7EF59A2188B85FA8AFEFDFAD3" box="[151,373,1358,1383]" country="Brazil" latitude="-4.168333" longLatPrecision="21" longitude="-41.581944" name="Domingos Mourao" pageId="46" pageNumber="347" stateProvince="Piaui">Domingos Mourão</location>
,
<geoCoordinate id="EE1B73EBEF59A2188A94FA8BFD99FAD2" box="[390,529,1358,1383]" direction="south" orientation="latitude" pageId="46" pageNumber="347" precision="15" value="-4.168333">S 04°10'06&quot;</geoCoordinate>
/
<geoCoordinate id="EE1B73EBEF59A2188926FA8BFD53FAD2" box="[564,731,1358,1383]" direction="west" orientation="longitude" pageId="46" pageNumber="347" precision="15" value="-41.581944">W 041°34'55&quot;</geoCoordinate>
,
<collectingDate id="EFD5CA04EF59A21889FFFA8AFCD0FAD2" box="[749,856,1358,1383]" pageId="46" pageNumber="347" value="2011-06-01">
<date id="FF9133ECEF59A21889FFFA8AFCD0FAD2" box="[749,856,1358,1383]" pageId="46" pageNumber="347" value="2011-06-01">1.vi.2011</date>
</collectingDate>
,
<collectorName id="26DA70FAEF59A2188878FA8BFC60FAD2" box="[874,1000,1358,1383]" pageId="46" pageNumber="347">N Hamada</collectorName>
col.
</materialsCitation>
;
<materialsCitation id="3B471F71EF59A2188F26FA8BFF7CFA1A" ID-GBIF-Occurrence="1585352574" collectingDate="2012-02-26" collectionCode="CZNC" collectorName="E Raimundi" country="Brazil" county="Mineiros" latitude="-18.263334" location="Emas" longLatPrecision="21" longitude="-52.89278" municipality="Parque Nacional" pageId="46" pageNumber="347" specimenCount="1" specimenCount-female="1" stateProvince="Goias">
<specimenCount id="9D29DEA5EF59A2188F26FA8BFBEAFAD2" box="[1076,1122,1358,1383]" pageId="46" pageNumber="347" type="female">1 ♀</specimenCount>
i,
<collectingRegion id="49EBDBCEEF59A2188F92FA8AFB4CFAD2" box="[1152,1220,1358,1383]" country="Brazil" name="Goias" pageId="46" pageNumber="347">Goiás</collectingRegion>
,
<collectingCounty id="62F16DA0EF59A2188FC7FA8AFAB5FAD2" box="[1237,1341,1358,1383]" pageId="46" pageNumber="347">Mineiros</collectingCounty>
,
<collectingMunicipality id="6BF48F56EF59A2188E5CFA8BFF74FA3E" pageId="46" pageNumber="347">Parque Nacional</collectingMunicipality>
das
<location id="8EF043F7EF59A2188A29FAB7FEF1FA3F" LSID="urn:lsid:plazi:treatment:0386A43AEF59A2198B85FC10FE32FE3D:8EF043F7EF59A2188A29FAB7FEF1FA3F" box="[315,377,1395,1418]" country="Brazil" county="Mineiros" latitude="-18.263334" longLatPrecision="21" longitude="-52.89278" municipality="Parque Nacional" name="Emas" pageId="46" pageNumber="347" stateProvince="Goias">Emas</location>
(sede),
<geoCoordinate id="EE1B73EBEF59A2188ACCFAB7FDE2FA3E" box="[478,618,1394,1419]" direction="south" orientation="latitude" pageId="46" pageNumber="347" precision="15" value="-18.263334">S 18°15'48&quot;</geoCoordinate>
/
<geoCoordinate id="EE1B73EBEF59A2188999FAB7FCACFA3E" box="[651,804,1394,1419]" direction="west" orientation="longitude" pageId="46" pageNumber="347" precision="15" value="-52.89278">W 52°53'34&quot;</geoCoordinate>
,
<collectingDate id="EFD5CA04EF59A2188827FAB6FC21FA3E" box="[821,937,1394,1419]" pageId="46" pageNumber="347" value="2012-02-26">
<date id="FF9133ECEF59A2188827FAB6FC21FA3E" box="[821,937,1394,1419]" pageId="46" pageNumber="347" value="2012-02-26">26.ii.2012</date>
</collectingDate>
,
<collectingMethod id="526E6D3BEF59A21888A8FAB6FB41FA3F" box="[954,1225,1394,1419]" pageId="46" pageNumber="347">pennsylvania light trap</collectingMethod>
,
<collectorName id="26DA70FAEF59A2188FC8FAB7FAEEFA3E" box="[1242,1382,1394,1419]" pageId="46" pageNumber="347">E Raimundi</collectorName>
col. (
<collectionCode id="ED3E8DE9EF59A2188B8CFA53FF64FA1A" box="[158,236,1431,1455]" pageId="46" pageNumber="347">CZNC</collectionCode>
)
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</paragraph>
<paragraph id="8B90152CEF59A2188BD5FA7EFC91F833" blockId="46.[151,1437,1142,2034]" pageId="46" pageNumber="347">
Female imago. Length (mm): body, 12.513.2; fore wing, 11.713.0; hind wing, 5.25.5; caudal filaments lost. General coloration yellowish, shaded with gray. Head completely shaded with gray, except gena and venter. Thorax. Pronotum whitish translucent, shaded with gray medially and with black on medial line, lateral and posterior margins; propleura and sternum yellowish white, shaded gray; blackish line present between coxae. Mesonotum yellowish, extensively shaded with gray, with blackish medial line; mesopleura with blackish dash above coxa, furcasternum shaded with gray. Metanotum shaded completely with gray; pleura and sterna paler. Fore leg reddish gray, middle and hind legs whitish, shaded with gray on coxae. Wings. Wing membrane slightly tinged with yellow, veins C, Sc, R, MA and MP1 reddish, remaining veins yellowish; hind wing with yellowish veins. Abdomen with terga almost completely shaded with gray, except pale submedian circular dot and transversal dashes on segments IIVII; medial line of terga VIIX blackish; sterna whitish translucent, except sockets and anterior margin of tergum IX shaded with gray. Sternum VIII with anteromedian, small and single subcircular socket (
<figureCitation id="131409A9EF59A2188BE6F88DFED4F8D7" box="[244,348,1865,1890]" captionStart="FIGURES 138 145" captionStartId="38.[151,264,1925,1948]" captionTargetBox="[283,1282,265,1875]" captionTargetId="figure@38.[277,1285,265,1880]" captionTargetPageId="38" captionText="FIGURES 138 145. C. segnis, holotype male imago: 138, genitalia, v. v.; 139, penes, v. v. Campsurus cotaxe sp. nov.: 140, genitalia, v. v.; 141, pedestal, v. v.; 142, penes, v. v.; 143, penes, l. v.; 144, female sternum VIII with detail of socket. C. amapaensis: 145, partial view of female sternum VIII with detail of socket." httpUri="https://zenodo.org/record/838472/files/figure.png" pageId="46" pageNumber="347">Fig. 145</figureCitation>
); socket relatively deep medially. Sternum VIII almost completely covered with spine-like microtrichia directed rearward. Caudal filaments whitish.
</paragraph>
<paragraph id="8B90152CEF59A2188BD5F856FE8FF847" blockId="46.[151,1437,1142,2034]" pageId="46" pageNumber="347">Egg. Length 270285µm, width 210240µm, polar cap length 3045µm and width 150165µm. General coloration yellowish white with whitish large polar cap (0.7× maximum width of egg), and formed by 68 filaments.</paragraph>
</subSubSection>
<subSubSection id="C33546A7EF58A2198BD5FF53FB1CFF05" box="[199,1172,151,176]" pageId="47" pageNumber="348" type="distribution">
<paragraph id="8B90152CEF58A2198BD5FF53FB1CFF05" blockId="47.[151,1436,151,392]" box="[199,1172,151,176]" pageId="47" pageNumber="348">
<emphasis id="B95BC93EEF58A2198BD5FF53FED1FF05" bold="true" box="[199,345,151,176]" pageId="47" pageNumber="348">Distribution</emphasis>
(
<figureCitation id="131409A9EF58A2198A7AFF53FE43FF05" box="[360,459,151,176]" captionStart="FIGURES 178" captionStartId="47.[151,264,1951,1974]" captionTargetBox="[185,1402,447,1929]" captionTargetId="figure@47.[185,1402,447,1929]" captionTargetPageId="47" captionText="FIGURES 178. Distribution maps." httpUri="https://zenodo.org/record/838486/files/figure.png" pageId="47" pageNumber="348">Fig. 178</figureCitation>
).
<collectingCountry id="F33855BCEF58A2198ACCFF53FDAAFF05" box="[478,546,151,176]" name="Brazil" pageId="47" pageNumber="348">Brazil</collectingCountry>
(
<collectingRegion id="49EBDBCEEF58A2198923FF5CFD0CFF05" box="[561,644,152,176]" country="Brazil" name="Amapa" pageId="47" pageNumber="348">Amapá</collectingRegion>
orig. New records:
<collectingRegion id="49EBDBCEEF58A2198871FF53FC2FFF05" box="[867,935,151,176]" country="Brazil" name="Goias" pageId="47" pageNumber="348">Goiás</collectingRegion>
,
<collectingRegion id="49EBDBCEEF58A21988A0FF53FC64FF05" box="[946,1004,151,176]" country="Brazil" name="Piaui" pageId="47" pageNumber="348">Piauí</collectingRegion>
and
<collectingRegion id="49EBDBCEEF58A2198F30FF53FB00FF05" box="[1058,1160,151,176]" country="Brazil" name="Roraima" pageId="47" pageNumber="348">Roraima</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C33546A7EF58A2198BD5FF78FE32FE3D" pageId="47" pageNumber="348" type="discussion">
<paragraph id="8B90152CEF58A2198BD5FF78FE32FE3D" blockId="47.[151,1436,151,392]" pageId="47" pageNumber="348">
<emphasis id="B95BC93EEF58A2198BD5FF78FEC4FF60" bold="true" box="[199,332,188,213]" pageId="47" pageNumber="348">Discussion.</emphasis>
<bibRefCitation id="EFBE68DDEF58A2198A46FF78FD2AFF61" author="Molineri" box="[340,674,188,213]" pageId="47" pageNumber="348" refString="Molineri, C. &amp; Emmerich, D. (2010) New species and new stage descriptions of Campsurus major species-group (Polymitarcyidae, Campsurinae), with first report of silk-case construction in a mayfly nymphs. Aquatic Insects, 32, 265 - 280." type="journal article" year="2010">Molineri &amp; Emmerich (2010)</bibRefCitation>
described and diagnosed the species based only on males. We here give the first description of the female adult, but nymphs remain unknown. The female morphology of
<taxonomicName id="4C2F6EAFEF58A2198E90FF24FE93FEA9" authorityName="Molineri &amp; Emmerich ." authorityYear="2010" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="47" pageNumber="348" phylum="Arthropoda" rank="species" species="amapaensis">
<emphasis id="B95BC93EEF58A2198E90FF24FE93FEA9" italics="true" pageId="47" pageNumber="348">C. amapaensis</emphasis>
</taxonomicName>
coincides with that of the
<taxonomicName id="4C2F6EAFEF58A2198958FEC1FD05FEA9" authorityName="Needham &amp; Murphy" authorityYear="1924" box="[586,653,261,284]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="47" pageNumber="348" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B95BC93EEF58A2198958FEC1FD05FEA9" box="[586,653,261,284]" italics="true" pageId="47" pageNumber="348">major</emphasis>
</taxonomicName>
species group proposed by
<bibRefCitation id="EFBE68DDEF58A21988D8FEC0FA94FEA9" author="Molineri" box="[970,1308,260,285]" pageId="47" pageNumber="348" refString="Molineri, C. &amp; Emmerich, D. (2010) New species and new stage descriptions of Campsurus major species-group (Polymitarcyidae, Campsurinae), with first report of silk-case construction in a mayfly nymphs. Aquatic Insects, 32, 265 - 280." type="journal article" year="2010">Molineri &amp; Emmerich (2010)</bibRefCitation>
, but it may be separated from the other known females of this group (
<taxonomicName id="4C2F6EAFEF58A2198845FEECFB7AFEF5" authority="Esben-Petersen 1912" authorityName="Esben-Petersen" authorityYear="1912" box="[855,1266,295,320]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="47" pageNumber="348" phylum="Arthropoda" rank="species" species="argentinus">
<emphasis id="B95BC93EEF58A2198845FEECFC7CFEF5" box="[855,1012,296,320]" italics="true" pageId="47" pageNumber="348">C. argentinus</emphasis>
<bibRefCitation id="EFBE68DDEF58A2198F13FEE3FB7AFEF5" author="Esben-Petersen" box="[1025,1266,295,320]" pageId="47" pageNumber="348" refString="Esben-Petersen, T. (1912) New and little-known species of Ephemerida from Argentine (Neuropt.). Deutsche Entomologische Zeitschrift, 1912, 333 - 342." type="journal article" year="1912">Esben-Petersen 1912</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2F6EAFEF58A2198E21FEECFE4DFED1" authority="Needham &amp; Murphy 1924" authorityName="Needham &amp; Murphy" authorityYear="1924" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="47" pageNumber="348" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B95BC93EEF58A2198E21FEECFA14FEF5" box="[1331,1436,296,320]" italics="true" pageId="47" pageNumber="348">C. major</emphasis>
<bibRefCitation id="EFBE68DDEF58A2198B85FE88FE4DFED1" author="Needham" box="[151,453,332,357]" pageId="47" pageNumber="348" refString="Needham, J. G. &amp; Murphy, H. E. (1924) Neotropical mayflies. Bulletin of the Lloyd Library, 22, 1 - 79." type="journal article" year="1924">Needham &amp; Murphy 1924</bibRefCitation>
</taxonomicName>
) by the smaller size and darker coloration of
<taxonomicName id="4C2F6EAFEF58A21988DCFE89FBFDFED1" authorityName="Molineri &amp; Emmerich ." authorityYear="2010" box="[974,1141,333,356]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="47" pageNumber="348" phylum="Arthropoda" rank="species" species="amapaensis">
<emphasis id="B95BC93EEF58A21988DCFE89FBFDFED1" box="[974,1141,333,356]" italics="true" pageId="47" pageNumber="348">C. amapaensis</emphasis>
</taxonomicName>
and also by details of the female sockets (
<figureCitation id="131409A9EF58A2198A5EFEABFE26FE3D" box="[332,430,367,392]" captionStart="FIGURES 138 145" captionStartId="38.[151,264,1925,1948]" captionTargetBox="[283,1282,265,1875]" captionTargetId="figure@38.[277,1285,265,1880]" captionTargetPageId="38" captionText="FIGURES 138 145. C. segnis, holotype male imago: 138, genitalia, v. v.; 139, penes, v. v. Campsurus cotaxe sp. nov.: 140, genitalia, v. v.; 141, pedestal, v. v.; 142, penes, v. v.; 143, penes, l. v.; 144, female sternum VIII with detail of socket. C. amapaensis: 145, partial view of female sternum VIII with detail of socket." httpUri="https://zenodo.org/record/838472/files/figure.png" pageId="47" pageNumber="348">Fig. 145</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>