545 lines
65 KiB
XML
545 lines
65 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5522461" ID-GBIF-Dataset="95f9938e-946f-495a-bd6c-71fa002305bc" ID-GBIF-Taxon="158950212" ID-Zenodo-Dep="5522461" approvalRequired="1" approvalRequired_for_treatments="1" checkinTime="1564744755810" checkinUser="jeremy" docAuthor="Kosterin, O. E." docDate="2015" docId="038687D6FFE73361890A0764BC10A55E" docLanguage="en" docName="Kosterin2015ABBYY.pdf" docOrigin="IDF-Report 81" docStyle="DocumentStyle{}" docTitle="Indolestes gracilis subsp. expressior Kosterin 2015" docType="treatment" docVersion="1" lastPageId="8" lastPageNumber="9" masterDocId="FFBFFFAEFFE333698D050531BF6CAC74" masterDocTitle="Taxonomical notes on Indolestes Fraser, 1922 (Lestidae, Zygoptera). 1. Indolestes gracilis expressior ssp. nov. from eastern Cambodia" masterLastPageNumber="11" masterPageNumber="1" pageId="4" pageNumber="5" updateTime="1651598569305" updateUser="jonas">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomical notes on Indolestes Fraser, 1922 (Lestidae, Zygoptera). 1. Indolestes gracilis expressior ssp. nov. from eastern Cambodia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kosterin, O. E.</mods:namePart>
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<mods:affiliation>Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev Ave. 10, Novosibirsk, 630090, Russia; Novosibirsk State University, Pirogova Str. 2, Novosibirsk, 630090, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">kosterin@bionet.nsc.ru</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>IDF-Report</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-04-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>81</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>11</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.3358796</mods:identifier>
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<mods:identifier type="GBIF-Dataset">95f9938e-946f-495a-bd6c-71fa002305bc</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3358796</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5522461" ID-GBIF-Taxon="158950212" ID-Zenodo-Dep="5522461" LSID="urn:lsid:plazi:treatment:038687D6FFE73361890A0764BC10A55E" httpUri="http://treatment.plazi.org/id/038687D6FFE73361890A0764BC10A55E" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
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<subSubSection box="[1039,1688,597,630]" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph blockId="4.[697,1973,597,2659]" box="[1039,1688,597,630]" pageId="4" pageNumber="5">
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<heading box="[1039,1688,597,630]" centered="true" fontSize="10" level="4" pageId="4" pageNumber="5" reason="1">
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<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" baseAuthorityName="Kosterin" baseAuthorityYear="2015" box="[1039,1688,597,630]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="gracilis" status="ssp. nov." subSpecies="expressior">
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Indolestes
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<emphasis box="[1225,1349,597,630]" italics="true" pageId="4" pageNumber="5">gracilis</emphasis>
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<emphasis bold="true" box="[1356,1534,597,630]" pageId="4" pageNumber="5">expressior</emphasis>
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<emphasis bold="true" box="[1542,1603,597,630]" pageId="4" pageNumber="5">ssp</emphasis>
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. nov.
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection box="[1277,1449,650,679]" pageId="4" pageNumber="5" type="description">
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||
<paragraph blockId="4.[697,1973,597,2659]" box="[1277,1449,650,679]" pageId="4" pageNumber="5">
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||
<figureCitation box="[1277,1401,650,679]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4</figureCitation>
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a-h
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="materials_examined">
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<paragraph blockId="4.[697,1973,597,2659]" box="[697,948,745,776]" pageId="4" pageNumber="5">
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<emphasis bold="true" box="[697,826,745,776]" pageId="4" pageNumber="5">Material</emphasis>
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studied
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</paragraph>
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<paragraph blockId="4.[697,1973,597,2659]" pageId="4" pageNumber="5">
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<materialsCitation ID-GBIF-Occurrence="2302281265" collectingDate="2014-06-09" collectionCode="RMNH" country="Cambodia" elevation="502" latitude="12.565278" location="river upstream Buu Sraa Waterfall, the left bank at the bridge" longitude="107.41917" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Mondulkiri Province" typeStatus="holotype">
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<typeStatus box="[700,833,794,823]" pageId="4" pageNumber="5">Holotype</typeStatus>
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<specimenCount box="[845,871,794,823]" pageId="4" pageNumber="5" type="male">
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<emphasis bold="true" box="[845,871,794,823]" pageId="4" pageNumber="5">♂</emphasis>
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</specimenCount>
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,
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<collectingCountry box="[886,1042,794,823]" name="Cambodia" pageId="4" pageNumber="5" value="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion box="[1058,1350,794,823]" pageId="4" pageNumber="5">Mondulkiri Province</collectingRegion>
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, the
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<location LSID="urn:lsid:plazi:treatment:038687D6FFE73361890A0764BC10A55E:8EF0601BFFE7336D888B062BBB61AF15" country="Cambodia" latitude="12.565278" longLatPrecision="21" longitude="107.41917" name="river upstream Buu Sraa Waterfall, the left bank at the bridge" pageId="4" pageNumber="5" stateProvince="Mondulkiri Province">river upstream Buu Sraa Waterfall, the left bank at the bridge</location>
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,
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<geoCoordinate box="[1052,1239,836,865]" degrees="12" direction="north" minutes="33" orientation="latitude" pageId="4" pageNumber="5" precision="15" seconds="55" value="12.565278">12°33'55'' N</geoCoordinate>
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<geoCoordinate box="[1248,1449,836,865]" degrees="107" direction="east" minutes="25" orientation="longitude" pageId="4" pageNumber="5" precision="15" seconds="09" value="107.41917">107°25'09'' E</geoCoordinate>
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,
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<elevation box="[1464,1561,836,865]" metricMagnitude="2" metricUnit="m" metricValue="5.02" pageId="4" pageNumber="5" unit="m" value="502.0">
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<quantity box="[1464,1561,836,865]" metricMagnitude="2" metricUnit="m" metricValue="5.02" pageId="4" pageNumber="5" unit="m" value="502.0">502 m</quantity>
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</elevation>
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a.s.l.,
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<collectingDate box="[1653,1802,836,865]" pageId="4" pageNumber="5" value="2014-06-09">
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<date box="[1653,1802,836,865]" pageId="4" pageNumber="5" value="2014-06-09">9 VI 2014</date>
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</collectingDate>
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; deposited in Naturalis Biodiversity Centre, Leiden, the Netherlands (
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<collectionCode box="[1560,1666,890,919]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="4" pageNumber="5">RMNH</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="etymology">
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<paragraph blockId="4.[697,1973,597,2659]" box="[697,860,986,1015]" pageId="4" pageNumber="5">Etymology</paragraph>
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<paragraph blockId="4.[697,1973,597,2659]" pageId="4" pageNumber="5">
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<emphasis box="[699,883,1035,1062]" italics="true" pageId="4" pageNumber="5">Expressions</emphasis>
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a Latin
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<emphasis box="[995,1128,1033,1062]" italics="true" pageId="4" pageNumber="5">adjective</emphasis>
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in gradus comparativus and genus masculinus, meaning ‘more expressed', referring to a more expressed differentiation of the apical part of the cercus than in other subspecies.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
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<paragraph blockId="4.[697,1973,597,2659]" box="[697,925,1223,1252]" pageId="4" pageNumber="5">
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Male (
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<figureCitation box="[791,915,1223,1252]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4</figureCitation>
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)
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</paragraph>
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<paragraph blockId="4.[697,1973,597,2659]" pageId="4" pageNumber="5">
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Head. Labium bluish in central part, yellowish at margins. Labrum, mandible bases, genae dull bluish (
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<figureCitation box="[972,1152,1320,1349]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4a, b</figureCitation>
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). Anteclypeus of the same colour with a small indistinct dark spot at centre, postclypeus mostly dark-bronze with anterior part dull bluish forming a central projection of this colour. Frons, vertex and occiput dark-bronze, there is a pair of greenish spots just below lateral ocelli and another one lateroposteriorly of the former. Antennae blackish-brown (
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<figureCitation box="[1316,1464,1512,1541]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4b</figureCitation>
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).
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</paragraph>
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<paragraph blockId="4.[697,1973,597,2659]" pageId="4" pageNumber="5">
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Thorax. Prothorax dull bluish with dorsal suture slightly darker brown and a pair of two indistinct bronze dorso-lateral patches (
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<figureCitation box="[1368,1514,1629,1658]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4c</figureCitation>
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). Posterior lobe slightly raised with smooth margins, light-brown, darker dorsally (
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<figureCitation box="[1481,1665,1677,1706]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4a, c</figureCitation>
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). Mesostigmal plate broadly-triangular, with a transversal lens-shaped central hollow deepening to its lateral ends, and deep middorsal groove. Central hollow and adjacent anterior margin brown, the rest dark-bronze but with brownish lightenings at sides of middorsal groove (
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<figureCitation box="[830,983,1868,1897]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4c</figureCitation>
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). Synthorax ground colour changes from olivaceus at mesepisternum to dull blue on sides (
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<figureCitation box="[1141,1283,1917,1946]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4a</figureCitation>
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). There is a broad middorsal dark-bronze band but dorsal ridge narrowly brown.
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<emphasis box="[1214,1301,1964,1993]" italics="true" pageId="4" pageNumber="5">There</emphasis>
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are elongate black spots at top of humeral and the first lateral sutures and a slight trace of a brownish stripe below mesopleural suture, with a small brown spot with indistinct margins in its dorsal part (
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<figureCitation box="[1806,1956,2060,2089]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4d</figureCitation>
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). Sclerites at wing bases blue. Coxae and trochanters bluish, rest legs brownish, with outer sides of femora, ventral ribs of tibia, tips of tarsi, spines and hooks blackish (
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<figureCitation box="[707,851,2204,2233]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4a</figureCitation>
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).
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</paragraph>
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<paragraph blockId="4.[697,1973,597,2659]" pageId="4" pageNumber="5">
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<emphasis box="[698,791,2273,2302]" italics="true" pageId="4" pageNumber="5">Wings</emphasis>
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hyaline, major veins dark brown, minor ones black. Discoidal cell very narrow, its dorsal side is ca 0.35 as long as ventral side in fw and ca half as long on hw postnodals 10 on fw, 9 on hw. Pterostigmata of folded fore and hind
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<emphasis box="[1734,1822,2368,2397]" italics="true" pageId="4" pageNumber="5">wings</emphasis>
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disposed exactly near each other. They are ca 2.2 as long as high, accompanied with two cells below, dark-brown, bordering longitudinal veins somewhat swollen and darkened (
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<figureCitation box="[707,853,2512,2541]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4e</figureCitation>
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).
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</paragraph>
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<paragraph blockId="4.[697,1973,597,2659]" lastBlockId="6.[697,1973,596,916]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
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Abdomen. Ground colour dull blue at S1-S3 (
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<figureCitation box="[1385,1528,2581,2610]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4a</figureCitation>
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) changing to brownish at S5- S8 (partly shown in
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<figureCitation box="[988,1125,2628,2657]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">Figure 4f</figureCitation>
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) but
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<emphasis box="[1203,1283,2628,2657]" italics="true" pageId="4" pageNumber="5">again</emphasis>
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greyish blue at S9-S10 (
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<figureCitation box="[1647,1832,2628,2657]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="4" pageNumber="5">
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Figure
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<quantity box="[1752,1793,2628,2657]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="4" pageNumber="5" unit="g" value="4.0">4g</quantity>
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, h
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</figureCitation>
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). Tergites 1-7 with solid dark-bronze to blackish dorsal
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<emphasis box="[1363,1469,596,623]" italics="true" pageId="6" pageNumber="7">stripes</emphasis>
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.That on S2 shaped as squid body with a tail directed anteriorly. Those on S2-S7 are constricted anteriorly, before tergite anterior margins, marked with black rings, and rather indistinctly expanding to tergite ventral margins posteriorly (
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<figureCitation box="[1142,1329,740,767]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="6" pageNumber="7">Figure 4a, f</figureCitation>
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). S8 dark with indistinct lateral brownish patches in its anterior half (
|
||
<figureCitation box="[1100,1239,787,814]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="6" pageNumber="7">Figure 4f</figureCitation>
|
||
). S9 blackish with a pair of indistinct greyish-blue spots at posterior margin, nearly fused to each other, occupying half of its length. S10 greyish blue above changing through reddish-brown to blackish below (
|
||
<figureCitation box="[1791,1938,883,910]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="6" pageNumber="7">Figure 4h</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="5" pageNumber="6" startId="5.[581,676,2501,2530]" targetBox="[578,1856,590,2456]" targetPageId="5">
|
||
<paragraph blockId="5.[580,1853,2501,2673]" pageId="5" pageNumber="6">
|
||
Figure 4. Details of
|
||
<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" baseAuthorityName="Kosterin" baseAuthorityYear="2015" box="[887,1333,2501,2532]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="gracilis" status="ssp. nov." subSpecies="expressior">
|
||
Indolestes
|
||
<emphasis box="[1054,1163,2501,2530]" italics="true" pageId="5" pageNumber="6">gracilis</emphasis>
|
||
expressior
|
||
</taxonomicName>
|
||
♂ ssp. nov.
|
||
<emphasis bold="true" box="[1523,1542,2501,2530]" pageId="5" pageNumber="6">-</emphasis>
|
||
a, head, thorax
|
||
<emphasis bold="true" box="[1797,1851,2501,2530]" pageId="5" pageNumber="6">and</emphasis>
|
||
abdominal segments
|
||
<emphasis bold="true" box="[909,939,2548,2577]" pageId="5" pageNumber="6">1-</emphasis>
|
||
2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d,
|
||
<emphasis bold="true" box="[803,854,2597,2626]" pageId="5" pageNumber="6">top</emphasis>
|
||
of synthorax, lateral view;
|
||
<emphasis bold="true" box="[1254,1278,2597,2626]" pageId="5" pageNumber="6">e,</emphasis>
|
||
pterostigmata; f, S6
|
||
<emphasis bold="true" box="[1581,1591,2597,2626]" pageId="5" pageNumber="6">-</emphasis>
|
||
S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[697,1974,979,1682]" pageId="6" pageNumber="7">
|
||
Cerci about twice as long as S10. In dorsal view (
|
||
<figureCitation box="[1406,1550,979,1006]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="6" pageNumber="7">Figure 4h</figureCitation>
|
||
), their outer outline (bearing sparse robust spines) smoothly curving towards each other to distinct apical portions occupying about 1/3 of the cercus projection to the central
|
||
<emphasis box="[1581,1643,1075,1102]" italics="true" pageId="6" pageNumber="7">axis</emphasis>
|
||
. Apical portions dropshaped, rounded proximally, bluntly pointed distally, directed caudally but very slightly diverging. Inner outline of cerci elaborate: narrowing at ca 1/5 of their length, then with a ventro-adaxial ledge ending with a long process, then slightly broadening again; cercus apical portion forms a prominent rounded backward ‘heel'. In lateral view (
|
||
<figureCitation box="[784,932,1315,1342]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="6" pageNumber="7">
|
||
Figure
|
||
<quantity box="[891,932,1315,1342]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="6" pageNumber="7" unit="g" value="4.0">4g</quantity>
|
||
</figureCitation>
|
||
), cercus dorsal outline nearly straight at basal 2/3 then bends down. Cercus ventral outline narrowing at basal 1/4, ventro-adaxial ledge with a subbasal blunt spine (not seen in dorsal view) and apical process; cercus apical portion hoof- or pen-like, with a rounded base and attenuated apex, occupies 1/3 of cercus length. Cercus dorsal side greyish blue,apical portion blackish-brown,rest brown (
|
||
<figureCitation box="[1782,1956,1506,1533]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="6" pageNumber="7">
|
||
Figure
|
||
<quantity box="[1882,1925,1506,1533]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="6" pageNumber="7" unit="g" value="4.0">4g</quantity>
|
||
,h
|
||
</figureCitation>
|
||
). Paraprocts light-brown, thrice as short as
|
||
<emphasis box="[1329,1405,1554,1581]" italics="true" pageId="6" pageNumber="7">cerci</emphasis>
|
||
, rounded in dorsal
|
||
<emphasis box="[1704,1776,1554,1581]" italics="true" pageId="6" pageNumber="7">view</emphasis>
|
||
, trapezoid in lateral
|
||
<emphasis box="[796,867,1602,1629]" italics="true" pageId="6" pageNumber="7">view</emphasis>
|
||
, their dorsolateral side deeply concave with semicircular ridges occupying the concavities (
|
||
<figureCitation box="[940,1124,1649,1676]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="6" pageNumber="7">
|
||
Figure
|
||
<quantity box="[1046,1089,1649,1676]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="6" pageNumber="7" unit="g" value="4.0">4g</quantity>
|
||
, h
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="6.[700,1902,1746,1779]" box="[700,1902,1750,1777]" pageId="6" pageNumber="7">Measurements [mm] - hw 18; abd (without apps) 29; total length (with apps) 38.</paragraph>
|
||
<paragraph blockId="6.[697,956,1842,1869]" box="[697,956,1842,1869]" pageId="6" pageNumber="7">Female unknown</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
|
||
<paragraph blockId="6.[697,1974,1960,2664]" box="[697,1223,1961,1992]" pageId="6" pageNumber="7">
|
||
<heading box="[697,1223,1961,1992]" fontSize="10" level="2" pageId="6" pageNumber="7" reason="1">
|
||
<emphasis bold="true" box="[697,1025,1961,1992]" pageId="6" pageNumber="7">Differential diagnosis</emphasis>
|
||
and remarks
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="6.[697,1974,1960,2664]" lastBlockId="7.[579,1853,596,771]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
The shape of the cerci of the new subspecies (
|
||
<figureCitation box="[1406,1552,2012,2039]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="6" pageNumber="7">Figure 4h</figureCitation>
|
||
)
|
||
<emphasis box="[1573,1592,2012,2039]" italics="true" pageId="6" pageNumber="7">is</emphasis>
|
||
close to that of
|
||
<taxonomicName baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1841,1974,2010,2039]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis box="[1841,1974,2010,2039]" italics="true" pageId="6" pageNumber="7">I. gracilis</emphasis>
|
||
</taxonomicName>
|
||
spp. (
|
||
<figureCitation box="[779,898,2060,2087]" captionStart="Figure 1" captionStartId="1.[581,674,2430,2457]" captionTargetBox="[588,1832,1694,2358]" captionTargetPageId="1" captionText="Figure 1. Indolestes cf taxa as illustrated in literature - anal appendages, dorsal. - a, I. cyaneus after Fraser (1933: fig. 34, as Ceylonolestes cyanea); - b-d, I. gracilis gracilis: b, after Asahina (1 976: figs. 10, as Indolestes gracilis); c, the type of Lestes gracilis from Hagen's collection (after Asahina 1976: fig. 13, as Indolestes gracilis); d, after Fraser (1933: fig. 28, as Ceylonolestes gracilis). - e, I. gracilis davenporti, after Fraser (1933: fig. 31, as Ceylonolestes davenporti). Not to scale." httpUri="https://zenodo.org/record/3358798/files/figure.png" pageId="6" pageNumber="7">Figure 1</figureCitation>
|
||
b-e) and
|
||
<taxonomicName authorityName="Ris" authorityYear="1916" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1037,1230,2058,2087]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="peregrinus">
|
||
<emphasis box="[1037,1230,2058,2087]" italics="true" pageId="6" pageNumber="7">I. peregrinus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[1240,1364,2060,2087]" captionStart="Figure 2" captionStartId="2.[700,793,2551,2578]" captionTargetBox="[718,1910,1874,2498]" captionTargetPageId="2" captionText="Figure 2. Indolestes peregrinus ♂ as illustrated in literature - anal appendages lateral (a), dorsal (b+c). — a,b, after Ris (1916: fig. 2, as Indolestes gracilis perigrinus); c, after Asahina (1976: figs 5, 7). Not to scale ." httpUri="https://zenodo.org/record/3358800/files/figure.png" pageId="6" pageNumber="7">Figure 2</figureCitation>
|
||
). However, the apical portion of the cerci
|
||
<emphasis box="[700,719,2104,2131]" italics="true" pageId="6" pageNumber="7">is</emphasis>
|
||
curiously inflated basally in dorsal view, forming a prominent ‘heel' protruding medio-anteriorly. So the cerci resemble rather legs and feet of a man in the act of brass swimming than the arms and hands of a man in the act of
|
||
<emphasis box="[1565,1659,2204,2231]" italics="true" pageId="6" pageNumber="7">diving</emphasis>
|
||
, as in the mentioned species. None of so far described taxa of
|
||
<taxonomicName authorityName="Fraser" authorityYear="1922" box="[1323,1479,2250,2279]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Indolestes</taxonomicName>
|
||
displays this shape of the cercus apical part, modification of which in the new subspecies looks most pronounced as compared to the related taxa. In other subspecies of
|
||
<emphasis box="[1528,1878,2346,2375]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1528,1664,2346,2375]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gracilis">I. gracilis</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Ris" authorityYear="1916" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1683,1878,2346,2375]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="peregrinus">I. peregrinus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1862" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cyaneus">
|
||
<emphasis box="[1958,1972,2346,2375]" italics="true" pageId="6" pageNumber="7">I.</emphasis>
|
||
<emphasis bold="true" box="[699,822,2393,2422]" pageId="6" pageNumber="7">cyaneus</emphasis>
|
||
</taxonomicName>
|
||
, the inner outline of the cercus at the apical portion base in dorsal view just turns caudad without a concavity before the apical part to form the ‘heel'. Drawings of the appendages of
|
||
<taxonomicName baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1002,1133,2489,2518]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis box="[1002,1133,2489,2518]" italics="true" pageId="6" pageNumber="7">I. gracilis</emphasis>
|
||
</taxonomicName>
|
||
spp and
|
||
<taxonomicName authorityName="Ris" authorityYear="1916" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1272,1462,2489,2518]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="peregrinus">
|
||
<emphasis box="[1272,1462,2489,2518]" italics="true" pageId="6" pageNumber="7">I. peregrinus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<bibRefCitation author="Ris, F." box="[1548,1694,2491,2518]" journalOrPublisher="Supplementa Entomologica" pageId="6" pageNumber="7" pagination="1 - 81" part="5" refId="ref4481" refString="Ris, F. 1916. H. Sauter's Formosa-Ausbeute. Odonata. Supplementa Entomologica 5: 1 - 81." title="H. Sauter's Formosa-Ausbeute. Odonata" type="journal article" year="1916">Ris (1916)</bibRefCitation>
|
||
, Fraser (1933) and Asahina (1976) are reproduced in
|
||
<figureCitation box="[1221,1356,2540,2567]" captionStart="Figure 1" captionStartId="1.[581,674,2430,2457]" captionTargetBox="[588,1832,1694,2358]" captionTargetPageId="1" captionText="Figure 1. Indolestes cf taxa as illustrated in literature - anal appendages, dorsal. - a, I. cyaneus after Fraser (1933: fig. 34, as Ceylonolestes cyanea); - b-d, I. gracilis gracilis: b, after Asahina (1 976: figs. 10, as Indolestes gracilis); c, the type of Lestes gracilis from Hagen's collection (after Asahina 1976: fig. 13, as Indolestes gracilis); d, after Fraser (1933: fig. 28, as Ceylonolestes gracilis). - e, I. gracilis davenporti, after Fraser (1933: fig. 31, as Ceylonolestes davenporti). Not to scale." httpUri="https://zenodo.org/record/3358798/files/figure.png" pageId="6" pageNumber="7">Figures 1</figureCitation>
|
||
b-e + 2a-c.
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1862" box="[1534,1823,2538,2567]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="2" phylum="Arthropoda" rank="species" species="cyaneus">
|
||
Indolestes
|
||
<emphasis bold="true" box="[1700,1823,2538,2567]" pageId="6" pageNumber="7">cyaneus</emphasis>
|
||
</taxonomicName>
|
||
has more robust cerci with a less attenuated and inflated apical part (
|
||
<figureCitation box="[1602,1744,2587,2614]" captionStart="Figure 1" captionStartId="1.[581,674,2430,2457]" captionTargetBox="[588,1832,1694,2358]" captionTargetPageId="1" captionText="Figure 1. Indolestes cf taxa as illustrated in literature - anal appendages, dorsal. - a, I. cyaneus after Fraser (1933: fig. 34, as Ceylonolestes cyanea); - b-d, I. gracilis gracilis: b, after Asahina (1 976: figs. 10, as Indolestes gracilis); c, the type of Lestes gracilis from Hagen's collection (after Asahina 1976: fig. 13, as Indolestes gracilis); d, after Fraser (1933: fig. 28, as Ceylonolestes gracilis). - e, I. gracilis davenporti, after Fraser (1933: fig. 31, as Ceylonolestes davenporti). Not to scale." httpUri="https://zenodo.org/record/3358798/files/figure.png" pageId="6" pageNumber="7">Figure 1a</figureCitation>
|
||
) (
|
||
<bibRefCitation author="Fraser, F. C." box="[1776,1956,2587,2614]" journalOrPublisher="Taylor and Francis, London" pageId="6" pageNumber="7" refId="ref4053" refString="Fraser, F. C. 1933. The fauna of British India, including Ceylon and Burma. Odonata. Vol. I. Taylor and Francis, London." title="The fauna of British India, including Ceylon and Burma. Odonata. Vol. I." type="book" year="1933">Fraser 1933</bibRefCitation>
|
||
). The coloration of the cerci, with distinctly darker apical portion,
|
||
<emphasis box="[1671,1690,2635,2662]" italics="true" pageId="6" pageNumber="7">is</emphasis>
|
||
as in
|
||
<taxonomicName authorityName="Ris" authorityYear="1916" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1783,1973,2633,2662]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="peregrinus">
|
||
<emphasis box="[1783,1973,2633,2662]" italics="true" pageId="6" pageNumber="7">
|
||
<botanicCollectionCode box="[1783,1798,2633,2662]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[1783,1794,2633,2662]" italics="true" pageId="6" pageNumber="7">I</emphasis>
|
||
.
|
||
</botanicCollectionCode>
|
||
peregrinus
|
||
</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Asahina S" box="[588,791,600,627]" journalOrPublisher="Kontyu" pageId="7" pageNumber="8" pagination="1 - 12" part="44" refId="ref3844" refString="Asahina S. 1976. Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces. Kontyu 44: 1 - 12." title="Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces" type="journal article" year="1976">Asahina 1976</bibRefCitation>
|
||
), while in
|
||
<taxonomicName authority="gracilis" authorityName="gracilis (Lieftinck" authorityYear="1940" box="[936,1177,596,629]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracilis" subSpecies="gracilis">
|
||
<emphasis box="[936,1063,598,627]" italics="true" pageId="7" pageNumber="8">
|
||
<botanicCollectionCode box="[936,950,598,627]" pageId="7" pageNumber="8">I.</botanicCollectionCode>
|
||
gracilis
|
||
</emphasis>
|
||
gracilis
|
||
</taxonomicName>
|
||
they are almost dark throughout (
|
||
<bibRefCitation author="Fraser, F. C." box="[1670,1845,600,627]" journalOrPublisher="Taylor and Francis, London" pageId="7" pageNumber="8" refId="ref4053" refString="Fraser, F. C. 1933. The fauna of British India, including Ceylon and Burma. Odonata. Vol. I. Taylor and Francis, London." title="The fauna of British India, including Ceylon and Burma. Odonata. Vol. I." type="book" year="1933">Fraser 1933</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Asahina S" box="[579,798,644,671]" journalOrPublisher="Kontyu" pageId="7" pageNumber="8" pagination="1 - 12" part="44" refId="ref3844" refString="Asahina S. 1976. Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces. Kontyu 44: 1 - 12." title="Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces" type="journal article" year="1976">Asahina 1976</bibRefCitation>
|
||
). However this character may vary with age, individually and geographically. The paraprocts in
|
||
<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" baseAuthorityName="Kosterin" baseAuthorityYear="2015" box="[1051,1359,694,723]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="gracilis" status="ssp. nov." subSpecies="expressior">
|
||
/.
|
||
<emphasis box="[1077,1359,694,723]" italics="true" pageId="7" pageNumber="8">gracilis expressior</emphasis>
|
||
</taxonomicName>
|
||
ssp. nov. (
|
||
<figureCitation box="[1520,1668,696,723]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="7" pageNumber="8">Figure 4h</figureCitation>
|
||
) are bluntly rounded in dorsal view, as should be in
|
||
<emphasis box="[1181,1316,742,771]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1181,1312,742,771]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="9" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<botanicCollectionCode box="[1181,1196,742,771]" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[1181,1192,742,771]" italics="true" pageId="7" pageNumber="8">I</emphasis>
|
||
.
|
||
</botanicCollectionCode>
|
||
gracilis
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="7.[578,1854,813,1323]" pageId="7" pageNumber="8">
|
||
The body coloration and pattern, with straight margins of the synthorax median black band and the solid abdominal black markings not divided into anterior/posterior or left/
|
||
<emphasis box="[642,711,913,940]" italics="true" pageId="7" pageNumber="8">right</emphasis>
|
||
parts,
|
||
<emphasis box="[811,830,913,940]" italics="true" pageId="7" pageNumber="8">is</emphasis>
|
||
similar to those in
|
||
<taxonomicName baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1115,1246,911,940]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis box="[1115,1246,911,940]" italics="true" pageId="7" pageNumber="8">I. gracilis</emphasis>
|
||
</taxonomicName>
|
||
and contrasted to
|
||
<taxonomicName authorityName="Ris" authorityYear="1916" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1528,1724,911,940]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="peregrinus">
|
||
<emphasis box="[1528,1724,911,940]" italics="true" pageId="7" pageNumber="8">I. peregrinus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Asahina S" journalOrPublisher="Kontyu" pageId="7" pageNumber="8" pagination="1 - 12" part="44" refId="ref3844" refString="Asahina S. 1976. Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces. Kontyu 44: 1 - 12." title="Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces" type="journal article" year="1976">Asahina 1976</bibRefCitation>
|
||
). Moreover, reduction of the humeral spots is similar to the nominotypical subspecies
|
||
<taxonomicName authorityName="gracilis (Lieftinck" authorityYear="1940" box="[756,919,1004,1037]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracilis" subSpecies="gracilis">
|
||
<emphasis box="[756,771,1007,1036]" italics="true" pageId="7" pageNumber="8">I.</emphasis>
|
||
g. gracilis
|
||
</taxonomicName>
|
||
from Sri Lanka.
|
||
<bibRefCitation author="Lieftinck, M. A." box="[1155,1381,1009,1036]" journalOrPublisher="Ceylon Journal of Sciences (B" pageId="7" pageNumber="8" pagination="79 - 117" part="22" refId="ref4265" refString="Lieftinck, M. A. 1940. On some Odonata collected in Ceylon, with descriptions of new species and larvae. Ceylon Journal of Sciences (B) 22: 79 - 117." title="On some Odonata collected in Ceylon, with descriptions of new species and larvae" type="journal article" year="1940">Lieftinck (1940)</bibRefCitation>
|
||
pointed that in that subspecies, these spots are variable from complete absence to 3-4 isolated spots or even fused into irregular fascia (as in
|
||
<taxonomicName authorityName="Fraser" authorityYear="1930" box="[971,1202,1102,1131]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="9" phylum="Arthropoda" rank="species" species="davenporti" subSpecies="davenporti">
|
||
<emphasis box="[971,986,1102,1131]" italics="true" pageId="7" pageNumber="8">I.</emphasis>
|
||
g. davenporti
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Selys" authorityYear="1891" baseAuthorityName="Selys" baseAuthorityYear="1891" box="[1278,1446,1102,1131]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="birmanus">
|
||
<emphasis box="[1278,1293,1102,1131]" italics="true" pageId="7" pageNumber="8">I.</emphasis>
|
||
birmanus
|
||
</taxonomicName>
|
||
<emphasis box="[1446,1461,1102,1131]" italics="true" pageId="7" pageNumber="8">).</emphasis>
|
||
The small spot below the mesopleural suture in the
|
||
<typeStatus box="[993,1128,1152,1179]" pageId="7" pageNumber="8">holotype</typeStatus>
|
||
of
|
||
<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" baseAuthorityName="Kosterin" baseAuthorityYear="2015" box="[1185,1495,1150,1179]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="gracilis" status="ssp. nov." subSpecies="expressior">
|
||
<emphasis box="[1185,1495,1150,1179]" italics="true" pageId="7" pageNumber="8">
|
||
<emphasis bold="true" box="[1185,1196,1150,1179]" italics="true" pageId="7" pageNumber="8">I</emphasis>
|
||
. gracilis expressior
|
||
</emphasis>
|
||
</taxonomicName>
|
||
ssp. nov. (
|
||
<figureCitation box="[1658,1808,1152,1179]" captionStart="Figure 4" captionStartId="5.[581,676,2501,2530]" captionTargetBox="[578,1856,590,2456]" captionTargetPageId="5" captionText="Figure 4. Details of Indolestes gracilis expression ssp. nov. - a, head, thorax and abdominal segments 1-2; b, head, dorsal view; c, prothorax and synthorax dorsum, dorsal view; d, top of synthorax, lateral view; e, pterostigmata; f, S6-S10 of abdomen; g, anal appendages, lateral view; h, the same, dorsal view. Not to scale." httpUri="https://zenodo.org/record/3358804/files/figure.png" pageId="7" pageNumber="8">Figure 4d</figureCitation>
|
||
)
|
||
<emphasis box="[1833,1851,1152,1179]" italics="true" pageId="7" pageNumber="8">is</emphasis>
|
||
brown and diffuse at margins, as it often happens with a variable, environmentally induced melanisation in some damselflies, e.g. in northern populations of
|
||
<taxonomicName authority="Charpentier, 1840" authorityName="Charpentier" authorityYear="1840" class="Insecta" family="Coenagrionidae" genus="Enallagma" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cyathigerum">
|
||
<emphasis bold="true" box="[1696,1854,1246,1275]" pageId="7" pageNumber="8">Enallagma</emphasis>
|
||
cyathigerum Charpentier,
|
||
<quantity box="[977,1049,1296,1323]" metricMagnitude="1" metricUnit="m" metricValue="4.6736" pageId="7" pageNumber="8" unit="in" value="1840.0">1840</quantity>
|
||
</taxonomicName>
|
||
in Eurasia (
|
||
<bibRefCitation author="Kosterin, O. E. & V. V. Zaika" box="[1223,1554,1296,1323]" journalOrPublisher="International Journal of Odonatology" pageId="7" pageNumber="8" pagination="277 - 327" part="13" refId="ref4234" refString="Kosterin, O. E. & V. V. Zaika. 2010. Odonata of Tuva, Russia. International Journal of Odonatology 13: 277 - 327." title="Odonata of Tuva, Russia" type="journal article" year="2010">
|
||
Kosterin & Zaika
|
||
<date box="[1477,1554,1296,1323]" pageId="7" pageNumber="8" value="2010">2010</date>
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="7.[578,1853,1364,1779]" pageId="7" pageNumber="8">
|
||
<emphasis box="[578,659,1364,1391]" italics="true" pageId="7" pageNumber="8">Yokoi</emphasis>
|
||
& Souphanthong (
|
||
<date box="[942,1024,1364,1391]" pageId="7" pageNumber="8" value="2014">2014</date>
|
||
: fig. 4) provided a drawing of the appendages of a male of «
|
||
<taxonomicName authorityName="Fraser" authorityYear="1922" box="[638,798,1412,1446]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">lndolestes</taxonomicName>
|
||
sp. 3» (
|
||
<figureCitation box="[903,1028,1417,1444]" captionStart="Figure 5" captionStartId="7.[581,675,2586,2613]" captionTargetBox="[824,1638,1930,2496]" captionTargetPageId="7" captionText="Figure 5. Male anal appendages, dorsal. - Indolestes?gracilis expressior ♂ ssp. nov. after Yokoi & Southpanthong (2014: fig. 4, as Indolestes sp. 3)." httpUri="https://zenodo.org/record/3358806/files/figure.png" pageId="7" pageNumber="8">Figure 5</figureCitation>
|
||
) from Paksong, Bolaven Plateau, Champasak Province, southern Laos. Their shape
|
||
<emphasis box="[1013,1032,1464,1491]" italics="true" pageId="7" pageNumber="8">is</emphasis>
|
||
very similar to
|
||
<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1278,1588,1462,1491]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="9" phylum="Arthropoda" rank="species" species="gracilis" status="ssp. nov." subSpecies="expressior">
|
||
<emphasis box="[1278,1588,1462,1491]" italics="true" pageId="7" pageNumber="8">
|
||
I.
|
||
<emphasis bold="true" box="[1304,1588,1462,1491]" italics="true" pageId="7" pageNumber="8">gracilis expressior</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
ssp. nov., so that specimen most probably represents the same subspecies. Unlike the here described
|
||
<typeStatus box="[756,898,1560,1587]" pageId="7" pageNumber="8">holotype</typeStatus>
|
||
, the photo of the general habitus of this specimen shows a complete humeral
|
||
<emphasis box="[866,949,1604,1631]" italics="true" pageId="7" pageNumber="8">stripe</emphasis>
|
||
(Ibid.: pl. 1). However, the great variation of the humeral pattern is common in
|
||
<taxonomicName authorityName="Fraser" authorityYear="1951" box="[932,1173,1656,1683]" class="Insecta" family="Lestidae" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subFamily" subFamily="Sympecmatinae">Sympecmatinae</taxonomicName>
|
||
and observed in
|
||
<taxonomicName authority="gracilis (Lieftinck, 1940)" authorityName="gracilis (Lieftinck" authorityYear="1940" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracilis" subSpecies="gracilis">
|
||
<emphasis box="[1444,1694,1654,1683]" italics="true" pageId="7" pageNumber="8">I. gracilis gracilis</emphasis>
|
||
(
|
||
<bibRefCitation author="Lieftinck, M. A." journalOrPublisher="Ceylon Journal of Sciences (B" pageId="7" pageNumber="8" pagination="79 - 117" part="22" refId="ref4265" refString="Lieftinck, M. A. 1940. On some Odonata collected in Ceylon, with descriptions of new species and larvae. Ceylon Journal of Sciences (B) 22: 79 - 117." title="On some Odonata collected in Ceylon, with descriptions of new species and larvae" type="journal article" year="1940">Lieftinck, 1940</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, hence
|
||
<emphasis box="[784,803,1704,1731]" italics="true" pageId="7" pageNumber="8">is</emphasis>
|
||
expected in
|
||
<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" baseAuthorityName="Lieftinck" baseAuthorityYear="1940" box="[998,1434,1702,1731]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="3" phylum="Arthropoda" rank="species" species="gracilis" status="ssp. nov." subSpecies="expressior">
|
||
Indolestes
|
||
<emphasis box="[1159,1434,1702,1731]" italics="true" pageId="7" pageNumber="8">gracilis expressior</emphasis>
|
||
</taxonomicName>
|
||
as well, perhaps depending on environmental conditions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption httpUri="https://zenodo.org/record/3358806/files/figure.png" pageId="7" pageNumber="8" startId="7.[581,675,2586,2613]" subCaptionStartIDs="7.[1136,1177,2641,2668]" subCaptionStarts="fig. 4" targetBox="[824,1638,1930,2496]" targetPageId="7">
|
||
<paragraph blockId="7.[580,1852,2584,2668]" pageId="7" pageNumber="8">
|
||
Figure
|
||
<emphasis bold="true" box="[687,712,2586,2613]" pageId="7" pageNumber="8">5.</emphasis>
|
||
Male anal appendages, dorsal.
|
||
<emphasis bold="true" box="[1199,1218,2586,2613]" pageId="7" pageNumber="8">-</emphasis>
|
||
<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" box="[1228,1682,2586,2613]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="3" phylum="Arthropoda" rank="species" species="gracilis" status="ssp. nov." subSpecies="expressior">
|
||
Indolestes
|
||
<emphasis box="[1395,1516,2586,2613]" italics="true" pageId="7" pageNumber="8">?gracilis</emphasis>
|
||
expressior
|
||
</taxonomicName>
|
||
♂
|
||
<emphasis bold="true" box="[1726,1783,2586,2613]" pageId="7" pageNumber="8">ssp.</emphasis>
|
||
nov. after Yokoi
|
||
<emphasis bold="true" box="[749,771,2641,2668]" pageId="7" pageNumber="8">&</emphasis>
|
||
Southpanthong
|
||
<emphasis bold="true" box="[1028,1039,2641,2668]" pageId="7" pageNumber="8">(</emphasis>
|
||
2014:
|
||
<emphasis bold="true" box="[1136,1183,2641,2668]" pageId="7" pageNumber="8">fig.</emphasis>
|
||
4,
|
||
<emphasis bold="true" box="[1230,1429,2639,2668]" pageId="7" pageNumber="8">
|
||
as
|
||
<taxonomicName authorityName="Fraser" authorityYear="1922" box="[1272,1429,2639,2668]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Indolestes</taxonomicName>
|
||
</emphasis>
|
||
sp. 3).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection box="[697,1921,596,671]" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph blockId="8.[697,1921,596,677]" box="[697,877,596,623]" pageId="8" pageNumber="9">Distribution</paragraph>
|
||
<paragraph blockId="8.[697,1921,596,677]" box="[697,1921,644,671]" pageId="8" pageNumber="9">
|
||
The subspecies
|
||
<emphasis box="[939,963,644,671]" italics="true" pageId="8" pageNumber="9">is</emphasis>
|
||
known from eastern Cambodia and (tentatively) southern Laos.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="9" type="biology_ecology">
|
||
<paragraph blockId="8.[697,1973,739,1156]" box="[697,808,739,766]" pageId="8" pageNumber="9">Habitat</paragraph>
|
||
<paragraph blockId="8.[697,1973,739,1156]" pageId="8" pageNumber="9">
|
||
The
|
||
<typeStatus box="[762,898,787,814]" pageId="8" pageNumber="9">holotype</typeStatus>
|
||
was startled from a bush branch in a secondary growth at the left bank of the large river which form the well-known Buu Sraa Waterfall. There were also some shallow stagnant pond and pools nearby. This place was at
|
||
<quantity box="[1589,1685,883,910]" metricMagnitude="2" metricUnit="m" metricValue="5.02" pageId="8" pageNumber="9" unit="m" value="502.0">502 m</quantity>
|
||
a.s.l. (as to Google Earth) and was surrounded by countryside but close to the evergreen forest in the river valley downstream of that place. The Laotian male presumably of this new subspecies was collected at
|
||
<quantity box="[1120,1235,1027,1054]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="8" pageNumber="9" unit="m" value="1310.0">1310 m</quantity>
|
||
a.s.l. (Yokoi & Souphanthong 2014).The nominotypical Sri Lankan subspecies inhabits mountains, with localities between
|
||
<date pageId="8" pageNumber="9" value="1800" valueMax="2500">
|
||
1800 and
|
||
<quantity box="[699,773,1127,1154]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="9" unit="m" value="2500.0">2500</quantity>
|
||
</date>
|
||
m a.s.l. (
|
||
<bibRefCitation author="Bedjanic, M. & Coniff, K. & N. van der Poorten & A. Salamun" box="[905,1310,1127,1154]" journalOrPublisher="Pensoft, Sofia" pageId="8" pageNumber="9" refId="ref3871" refString="Bedjanic, M., K. Coniff, K., N. van der Poorten N. & A. Salamun. 2014. Dragonfly Fauna of Sri Lanka. Distribution and biology, with threat status of its endemics. Pensoft, Sofia" title="Dragonfly Fauna of Sri Lanka. Distribution and biology, with threat status of its endemics" type="book" year="2014">Bedjanic et al. 2014: 76-77</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="9" type="discussion">
|
||
<paragraph blockId="8.[696,880,1216,1249]" box="[696,880,1216,1249]" pageId="8" pageNumber="9">
|
||
<heading bold="true" box="[696,880,1216,1249]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="0">
|
||
<emphasis bold="true" box="[696,880,1216,1249]" pageId="8" pageNumber="9">Discussion</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="8.[697,1971,1287,1608]" pageId="8" pageNumber="9">
|
||
<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" box="[699,837,1290,1319]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" sensu="lato" species="gracilis" status="sensu lato">
|
||
<emphasis box="[699,837,1290,1319]" italics="true" pageId="8" pageNumber="9">I. gracilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[849,1006,1292,1319]" pageId="8" pageNumber="9" sensu="lato">sensu lato</taxonomicNameLabel>
|
||
ranges from Sri Lanka through western (Western Ghats) and southern (Tamil Nadu State) India (
|
||
<bibRefCitation author="Ris, F." box="[1224,1355,1339,1366]" journalOrPublisher="Supplementa Entomologica" pageId="8" pageNumber="9" pagination="1 - 81" part="5" refId="ref4481" refString="Ris, F. 1916. H. Sauter's Formosa-Ausbeute. Odonata. Supplementa Entomologica 5: 1 - 81." title="H. Sauter's Formosa-Ausbeute. Odonata" type="journal article" year="1916">Ris 1916</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Fraser, F. C." box="[1374,1552,1339,1366]" journalOrPublisher="Taylor and Francis, London" pageId="8" pageNumber="9" refId="ref4053" refString="Fraser, F. C. 1933. The fauna of British India, including Ceylon and Burma. Odonata. Vol. I. Taylor and Francis, London." title="The fauna of British India, including Ceylon and Burma. Odonata. Vol. I." type="book" year="1933">Fraser 1933</bibRefCitation>
|
||
), while
|
||
<taxonomicName authorityName="Ris" authorityYear="1916" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1670,1862,1337,1366]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="peregrinus">
|
||
<emphasis box="[1670,1862,1337,1366]" italics="true" pageId="8" pageNumber="9">I. peregrinus</emphasis>
|
||
</taxonomicName>
|
||
ranges from S China to Korea and Japan (
|
||
<bibRefCitation author="Asahina S" box="[1242,1460,1384,1411]" journalOrPublisher="Kontyu" pageId="8" pageNumber="9" pagination="1 - 12" part="44" refId="ref3844" refString="Asahina S. 1976. Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces. Kontyu 44: 1 - 12." title="Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces" type="journal article" year="1976">Asahina 1976</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Wilson, K. D. P" box="[1476,1673,1384,1411]" journalOrPublisher="The IUCN Red List of Threatened Species. Version 2014.3" pageId="8" pageNumber="9" publicationUrl="www.iucnredlist.org" refId="ref4556" refString="Wilson, K. D. P 2009. Indolestes peregrinus. The IUCN Red List of Threatened Species. Version 2014.3. <www. iucnredlist. org>. Downloaded on 26 II 2015." title="Indolestes peregrinus" type="book" year="2009">Wilson 2009</bibRefCitation>
|
||
). East Cambodia
|
||
<emphasis italics="true" pageId="8" pageNumber="9">is situated</emphasis>
|
||
far to the east from the range of the former and far to the south from the range of the latter. Hence
|
||
<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" baseAuthorityName="Kosterin" baseAuthorityYear="2015" box="[1002,1306,1481,1510]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="gracilis" status="ssp. nov." subSpecies="expressior">
|
||
<emphasis box="[1002,1306,1481,1510]" italics="true" pageId="8" pageNumber="9">
|
||
I. gracilis
|
||
<emphasis bold="true" box="[1147,1306,1481,1510]" italics="true" pageId="8" pageNumber="9">expressior</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
subsp. nov seems to represent the hitherto unknown south-eastern, Indochinese subspecies of this species, characterised by a more elaborated shape of the cerci with a more modified apical part.
|
||
</paragraph>
|
||
<paragraph blockId="8.[697,1973,1643,2346]" pageId="8" pageNumber="9">
|
||
No
|
||
<taxonomicName authorityName="Fraser" authorityYear="1922" box="[759,918,1646,1675]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Indolestes</taxonomicName>
|
||
spp. have been reported for Vietnam (
|
||
<bibRefCitation author="Do M. C. & T. T. H. Dang" box="[1547,1823,1648,1675]" journalOrPublisher="Vietnam National University Publisher, Hanoi" pageId="8" pageNumber="9" refId="ref3917" refString="Do M. C. & T. T. H. Dang 2006. Checklist of Dragonflies from Vietnam. Vietnam National University Publisher, Hanoi." title="Checklist of Dragonflies from Vietnam" type="book" year="2006">Do & Dang 2006</bibRefCitation>
|
||
). Yokoi & Souphanthong (2014) listed three not identified
|
||
<taxonomicName authorityName="Fraser" authorityYear="1922" box="[1431,1587,1693,1722]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[1431,1587,1693,1722]" pageId="8" pageNumber="9">Indolestes</emphasis>
|
||
</taxonomicName>
|
||
spp. from Laos, with their
|
||
<emphasis box="[699,714,1742,1771]" italics="true" pageId="8" pageNumber="9">«</emphasis>
|
||
<emphasis bold="true" box="[714,873,1742,1771]" pageId="8" pageNumber="9">lndolestes</emphasis>
|
||
sp3» most probably representing
|
||
<taxonomicName authority="Kosterin, 2015" authorityName="Kosterin" authorityYear="2015" baseAuthorityName="Kosterin" baseAuthorityYear="2015" box="[1422,1735,1742,1771]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="gracilis" status="ssp. nov." subSpecies="expressior">
|
||
<emphasis box="[1422,1735,1742,1771]" italics="true" pageId="8" pageNumber="9">I. gracilis expressior</emphasis>
|
||
</taxonomicName>
|
||
ssp. nov. Since subspecies is an entity of intraspecies variation, specifically geographical variation, it
|
||
<emphasis box="[700,719,1836,1863]" italics="true" pageId="8" pageNumber="9">is</emphasis>
|
||
undesirable to describe a subspecies by one or two specimens. However, I am quite
|
||
<emphasis box="[699,852,1883,1910]" italics="true" pageId="8" pageNumber="9">convinced</emphasis>
|
||
in existence of this Indochinese taxon because of the unique apical part of the cercus in an area so remote from other subspecies. Of course, further specimens are needed to reveal the variation of the new species and its range and to finally prove its distinctness. Quite likely, it may appear bona species. Note, however, that this was the first and only
|
||
<taxonomicName authorityName="Fraser" authorityYear="1922" box="[954,1110,2077,2106]" class="Insecta" family="Lestidae" genus="Indolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Indolestes</taxonomicName>
|
||
specimen obtained on my five 2-3 week long expeditions to Cambodia in 2010-2014, that
|
||
<emphasis box="[1181,1200,2123,2150]" italics="true" pageId="8" pageNumber="9">is</emphasis>
|
||
they are very rare. To postpone the description until a consider-able collection accumulates from Cambodia would mean for a long time to operate in discussions of the fauna of Cambodia with an unnamed taxon, among so many others of Indochinese
|
||
<taxonomicName authorityName="Fabricius" authorityYear="1793" box="[1187,1319,2266,2293]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
(e.g. Yokoi & Souphanthong 2014), that
|
||
<emphasis box="[1952,1970,2266,2293]" italics="true" pageId="8" pageNumber="9">is</emphasis>
|
||
inconvenient.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |