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<document id="2AB2799AF564986E02D4E92660B967AE" ID-DOI="10.5281/zenodo.189132" ID-GBIF-Dataset="86143b99-7b3c-492d-9494-d645fc9c647b" ID-ISSN="1175-5326" ID-Zenodo-Dep="189132" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461099427313" checkinUser="plazi" docAuthor="Hutchinson, Mark N., Doughty, Paul &amp; Oliver, Paul M." docDate="2009" docId="038687D23B6A755165990B383ED614FC" docLanguage="en" docName="zt02167p046.pdf" docOrigin="Zootaxa 2167" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Diplodactylus calcicolus Hutchinson, Doughty &amp; Oliver, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="39" masterDocId="FFBFFFAA3B66755F650E0E353D7A115B" masterDocTitle="Taxonomic revision of the stone geckos (Squamata: Diplodactylidae: Diplodactylus) of southern Australia" masterLastPageNumber="46" masterPageNumber="25" pageNumber="37" updateTime="1698590674586" updateUser="plazi">
<mods:mods id="0E6CCD458BB837340D5218E287048106" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="50C24D7683D1AC59DD1A3444457911BC">
<mods:title id="24C409AFC3829A043766665F1376BA88">Taxonomic revision of the stone geckos (Squamata: Diplodactylidae: Diplodactylus) of southern Australia</mods:title>
</mods:titleInfo>
<mods:name id="03468A97347C423AAA8FC8936549B8CD" type="personal">
<mods:role id="8E093D7D7502BF619A27D397C236B976">
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</mods:role>
<mods:namePart id="212D67C10DD2A1C0ECF595C4915E9628">Hutchinson, Mark N.</mods:namePart>
</mods:name>
<mods:name id="CD7ECF6A1AF3B550E2538D8BBAB39D6F" type="personal">
<mods:role id="0E10360C01BE76AE7E70AAB8C85EFAD3">
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</mods:role>
<mods:namePart id="AFDD5F27A35D72A3CF39449FF406EABC">Doughty, Paul</mods:namePart>
</mods:name>
<mods:name id="7063FAF9035E033EFF50D31579FDC188" type="personal">
<mods:role id="ECD8703F89355FCE9D66EA66245208BA">
<mods:roleTerm id="BE51C0F87CD89A7EDEDF83C728F70300">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="68A26D6BCC79D0F5F5ACBDC68918B5EF">Oliver, Paul M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="DDCF66768895C7C6EDA363E65A7EB4B6">text</mods:typeOfResource>
<mods:relatedItem id="0DF1E69CAE5AF94B1F3997ECCA84A1D6" type="host">
<mods:titleInfo id="1730CD4C751EA73A1BD04C5860F6784A">
<mods:title id="B4B961E770E772507745EDD6BA69A484">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BEDDB67A613437CEF960CFA58AF4B646">
<mods:date id="E9CD3C43708878B89157AD3D3B62FC37">2009</mods:date>
<mods:detail id="73867E6EFE745B09DC91D99FDC28AAAE" type="volume">
<mods:number id="1CF2619DCB0A694E313F83387AD04728">2167</mods:number>
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<mods:start id="BA677D54213B86E4F3BC3D76DAA6D29F">25</mods:start>
<mods:end id="53016F69E2879BC04D88D538506951E0">46</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F4DF841F40235240BAE5372128061847">journal article</mods:classification>
<mods:identifier id="0A02FD69F9A66C5FCA69CDC99566AFC8" type="DOI">10.5281/zenodo.189132</mods:identifier>
<mods:identifier id="44F1B84BBEC7FD0E33370264FBBA82D4" type="GBIF-Dataset">86143b99-7b3c-492d-9494-d645fc9c647b</mods:identifier>
<mods:identifier id="B67AD4E8F1570E14DA8A8A354DA179D3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E2FC896DDD8AC0DD065EBF94617BD2DB" type="Zenodo-Dep">189132</mods:identifier>
</mods:mods>
<treatment id="038687D23B6A755165990B383ED614FC" ID-DOI="http://doi.org/10.5281/zenodo.6221900" ID-GBIF-Taxon="119621288" ID-Zenodo-Dep="6221900" LSID="urn:lsid:plazi:treatment:038687D23B6A755165990B383ED614FC" httpUri="http://treatment.plazi.org/id/038687D23B6A755165990B383ED614FC" lastPageId="14" lastPageNumber="39" pageId="12" pageNumber="37">
<subSubSection id="C335654F3B6A755365990B383C031411" pageId="12" pageNumber="37" type="nomenclature">
<paragraph id="8B9036C43B6A755365990B383F5B147C" blockId="12.[151,545,1293,1392]" box="[151,545,1293,1319]" pageId="12" pageNumber="37">
<heading id="D0D881A83B6A755365990B383F5B147C" bold="true" box="[151,545,1293,1319]" fontSize="11" level="1" pageId="12" pageNumber="37" reason="1">
<emphasis id="B95BEAD63B6A755365990B383F5B147C" bold="true" box="[151,545,1293,1319]" pageId="12" pageNumber="37">
<taxonomicName id="4C2F4D473B6A755365990B383CC6147C" box="[151,444,1293,1319]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="37" phylum="Chordata" rank="species" species="calcicolus" status="sp. nov.">
<emphasis id="B95BEAD63B6A755365990B383CC6147C" bold="true" box="[151,444,1293,1319]" italics="true" pageId="12" pageNumber="37">Diplodactylus calcicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857AD3B6A755364CA0B383F5B147C" box="[452,545,1293,1319]" pageId="12" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9036C43B6A755365990B053C031411" blockId="12.[151,545,1293,1392]" box="[151,377,1328,1354]" pageId="12" pageNumber="37">South Coast Gecko</paragraph>
</subSubSection>
<subSubSection id="C335654F3B6A755365990B633F0E14EC" pageId="12" pageNumber="37" type="description">
<paragraph id="8B9036C43B6A755365990B633C24142B" blockId="12.[151,545,1293,1392]" box="[151,350,1366,1392]" pageId="12" pageNumber="37">
<figureCitation id="13142A413B6A755365990B633D8D142B" box="[151,247,1366,1392]" captionStart="FIGURE 10" captionStartId="13.[151,255,873,897]" captionText="FIGURE 10. Holotype of Diplodactylus calcicolus sp. nov, SAMA R 25343." pageId="12" pageNumber="37">Figs. 10</figureCitation>
,
<figureCitation id="13142A413B6A7553640A0B633C58142B" box="[260,290,1366,1392]" captionStart="FIGURE 11" captionStartId="13.[151,257,1535,1559]" captionTargetBox="[295,1238,970,1518]" captionTargetId="figure@13.[295,1304,948,1519]" captionTargetPageId="13" captionText="FIGURE 11. Variation in colour pattern in Diplodactylus calcicolus sp. nov. From left to right: western specimens (WAM R 140935, WAM R 156947, WAM R 156943), Nullarbor specimens (SAM R 59582, SAM R 59524), and eastern specimens (SAM R 40554, SAM R 58537, SAM R 36644)." httpUri="https://zenodo.org/record/189139/files/figure.png" pageId="12" pageNumber="37">11</figureCitation>
,
<figureCitation id="13142A413B6A755364210B633C31142B" box="[303,331,1366,1392]" captionStart="FIGURE 14" captionStartId="16.[151,257,1878,1902]" captionTargetBox="[210,1388,208,1840]" captionTargetId="figure@16.[194,1400,194,1851]" captionTargetPageId="16" captionText="FIGURE 14. Species of the Diplodactylus vittatus complex in life. A, D. vittatus, Warrumbungle Range, NSW, (M. Hutchinson); B, D. furcosus, Whyalla, South Australia (T. Peters, SA Museum); C, D. g. granariensis, Newman Rock, WA (B. Maryan); D, D. g. rex, Wildara Pinnacle, WA (B. Maryan); E, D. calcicolus sp. nov., vicinity of the WA-SA border (G. Harold); F, D. wiru sp. nov. Lake Ifould, SA (P. Oliver)." httpUri="https://zenodo.org/record/189141/files/figure.png" pageId="12" pageNumber="37">14</figureCitation>
E
</paragraph>
<paragraph id="8B9036C43B6A755365990B953F0E14EC" blockId="12.[151,628,1440,1463]" box="[151,628,1440,1463]" pageId="12" pageNumber="37">
<taxonomicName id="4C2F4D473B6A755365990B953C5714EC" box="[151,301,1440,1463]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="B95BEAD63B6A755365990B953C5714EC" box="[151,301,1440,1463]" italics="true" pageId="12" pageNumber="37">Diplodactylus</emphasis>
</taxonomicName>
'southern',
<bibRefCitation id="EFBE4B353B6A755364AB0B953F0E14EC" author="Oliver" box="[421,628,1440,1463]" pageId="12" pageNumber="45" refString="Oliver, P., Hugall, A., Adams, M., Cooper, S. J. B. &amp; Hutchinson, M. (2007 a) Genetic elucidation of ancient and cryptic diversity in a group of Australian geckos: the Diplodactylus vittatus complex. Molecular Phylogenetics and Evolution, 44, 77 - 88." type="journal article" year="2007" yearSuffix="a">
Oliver
<emphasis id="B95BEAD63B6A755364FE0B953F6514EC" box="[496,543,1440,1463]" italics="true" pageId="12" pageNumber="37">et al</emphasis>
., 2007a
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C335654F3B6A755265910BD7394510C1" lastPageId="13" lastPageNumber="38" pageId="12" pageNumber="37" type="materials_examined">
<paragraph id="8B9036C43B6A755365910BD73DCD1711" blockId="12.[151,1437,1506,2035]" pageId="12" pageNumber="37">
<emphasis id="B95BEAD63B6A755365910BD73C6214A7" bold="true" box="[159,280,1506,1532]" pageId="12" pageNumber="37">
<typeStatus id="549488663B6A755365910BD73C6E14A7" box="[159,276,1506,1532]" pageId="12" pageNumber="37" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED3EAE013B6A7553642E0BD73C0F14A7" LSID="urn:lsid:biocol.org:col:34982" box="[288,373,1506,1532]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="12" pageNumber="37">SAMA</collectionCode>
R25343, adult male, from Hambidge Conservation Park, Eyre Peninsula, SA (
<geoCoordinate id="EE1B50033B6A755360160BD73DD31778" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="15" value="-33.444447">33º 26' 40&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A755365B8083C3C1B1778" box="[182,353,1545,1571]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="15" value="136.04167">136º 02' 30&quot; E</geoCoordinate>
), collected by the Mammal Club of the SA Field Naturalists’ Society on
<date id="FF9110043B6A755361B6083C38271778" box="[1208,1373,1545,1571]" pageId="12" pageNumber="37" value="1984-04-23">23 April 1984</date>
(
<figureCitation id="13142A413B6A75536062083C3DDD1711" captionStart="FIGURE 8" captionStartId="11.[151,255,966,990]" captionText="FIGURE 8. Holotype of Diplodactylus granariensis rex, WAM R 97288." pageId="12" pageNumber="37">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8B9036C43B6A755365CB086338571603" blockId="12.[151,1437,1506,2035]" pageId="12" pageNumber="37">
<emphasis id="B95BEAD63B6A755365CB08633C30172B" bold="true" box="[197,330,1622,1648]" pageId="12" pageNumber="37">
<typeStatus id="549488663B6A755365CB08633C3F172B" box="[197,325,1622,1648]" pageId="12" pageNumber="37" type="paratype">Paratypes</typeStatus>
.
</emphasis>
South
<collectingCountry id="F33876543B6A7553649108633F71172B" box="[415,523,1622,1648]" name="Australia" pageId="12" pageNumber="37">Australia</collectingCountry>
(
<collectionCode id="ED3EAE013B6A7553671508633F08172B" LSID="urn:lsid:biocol.org:col:34982" box="[539,626,1622,1648]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="12" pageNumber="37">SAMA</collectionCode>
specimens): R24763 (M) Lighthouse near Stenhouse Bay (
<geoCoordinate id="EE1B50033B6A7553603C086338EC172B" box="[1330,1430,1622,1648]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-35.283333">35º17' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A7553659908483C6817CC" box="[151,274,1661,1687]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="136.95">136º 57' E</geoCoordinate>
); R25382 (M) Ifould Campsite No. 2, 62 Km N Colona H/S (
<geoCoordinate id="EE1B50033B6A755366EB0848392A17CC" box="[997,1104,1661,1687]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-30.933332">30º 56' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A75536153084839A217CC" box="[1117,1240,1661,1687]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="132.08333">132º 05' E</geoCoordinate>
); R26257 (F)
<quantity id="4CD79B213B6A7553607008483DC717E5" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="37" unit="km" value="10.0">10 km</quantity>
NE Border Village (
<geoCoordinate id="EE1B50033B6A755364B908913F2D17E5" box="[439,599,1700,1726]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="15" value="-31.61389">31º 36' 50&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A7553676808913E6217E5" box="[614,792,1700,1726]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="15" value="129.10556">129º 06' 20&quot; E</geoCoordinate>
); R26322 (F)
<quantity id="4CD79B213B6A755366CC0891397417E5" box="[962,1038,1700,1726]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="12" pageNumber="37" unit="km" value="48.0">48 km</quantity>
SE Koonalda Stn (
<geoCoordinate id="EE1B50033B6A755361F8089138EC17E5" box="[1270,1430,1700,1726]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="15" value="-31.51111">31º 30' 40&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A7553659908FF3C3F17BF" box="[151,325,1738,1764]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="15" value="130.35834">130º 21' 30&quot; E</geoCoordinate>
); R26495 (F)
<quantity id="4CD79B213B6A755364E308FF3F4D17BF" box="[493,567,1738,1764]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="37" unit="km" value="11.0">11 km</quantity>
NE Border Village (
<geoCoordinate id="EE1B50033B6A7553662308FF3EB017BF" box="[813,970,1738,1764]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="15" value="-31.591667">31º 35' 30&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A755366D608FF39FC17BF" box="[984,1158,1738,1764]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="15" value="129.12222">129º 07' 20&quot; E</geoCoordinate>
); R26496 (M), R26497 (M)
<quantity id="4CD79B213B6A755365C208C43C6C1650" box="[204,278,1777,1803]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="37" unit="km" value="21.0">21 km</quantity>
NE Border Village (
<geoCoordinate id="EE1B50033B6A7553670508C43FD21650" box="[523,680,1777,1803]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="15" value="-31.5">31º 30' 00&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A755367BB08C43E191650" box="[693,867,1777,1803]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="15" value="129.14722">129º 08' 50&quot; E</geoCoordinate>
); R36643 (F) Venus Bay (
<geoCoordinate id="EE1B50033B6A7553619608C4387E1650" box="[1176,1284,1777,1803]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-33.233334">33º 14' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A7553601F08C438F71650" box="[1297,1421,1777,1803]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="134.66667">134º 40' E</geoCoordinate>
); R39517 (M)
<quantity id="4CD79B213B6A75536421092D3C111669" box="[303,363,1816,1842]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="37" unit="km" value="5.0">5 km</quantity>
S of Monarto South (
<geoCoordinate id="EE1B50033B6A75536769092D3E7B1669" box="[615,769,1816,1842]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="15" value="-35.176113">35º 10' 34&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A75536600092D3EC21669" box="[782,952,1816,1842]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="15" value="139.12334">139º 07' 24&quot; E</geoCoordinate>
); R44119 (F) Mt Younghusband, St Peter Island (
<geoCoordinate id="EE1B50033B6A755365E0090B3C231603" box="[238,345,1854,1880]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-32.25">32º 15' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A75536468090B3C9B1603" box="[358,481,1854,1880]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="133.61667">133º 37' E</geoCoordinate>
); R52488 (F) 28.3 km SSE Port Lincoln (
<geoCoordinate id="EE1B50033B6A755366C5090B391C1603" box="[971,1126,1854,1880]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="15" value="-34.973057">34º 58' 23&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A7553617D090B38651603" box="[1139,1311,1854,1880]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="15" value="135.95222">135º 57' 08&quot; E</geoCoordinate>
).
</paragraph>
<paragraph id="8B9036C43B6A755265CB0950394510C1" blockId="12.[151,1437,1506,2035]" lastBlockId="13.[151,1437,152,410]" lastPageId="13" lastPageNumber="38" pageId="12" pageNumber="37">
Western
<collectingCountry id="F33876543B6A7553642309503CE31624" box="[301,409,1893,1919]" name="Australia" pageId="12" pageNumber="37">Australia</collectingCountry>
(
<collectionCode id="ED3EAE013B6A755364A709503C891624" LSID="urn:lsid:biocol.org:col:34981" box="[425,499,1893,1919]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="37">WAM</collectionCode>
specimens): R24602 (F) Eucla (
<geoCoordinate id="EE1B50033B6A7553667809503E991624" box="[886,995,1893,1919]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-31.716667">31º 43' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A755366FF0950390A1624" box="[1009,1136,1893,1919]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="128.9">128º 54' E</geoCoordinate>
); R39921 (M)
<quantity id="4CD79B213B6A7553602F095038021624" box="[1313,1400,1893,1919]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="12" pageNumber="37" unit="mi" value="2.0">2 miles</quantity>
W Greenshield Soak (
<geoCoordinate id="EE1B50033B6A7553648F09B93C8816FD" box="[385,498,1932,1958]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-33.5">33º 30' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A7553670E09B93FF816FD" box="[512,642,1932,1958]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="118.816666">118º 49' E</geoCoordinate>
); R56866 (F) Eyre (
<geoCoordinate id="EE1B50033B6A7553667409B93E9616FD" box="[890,1004,1932,1958]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-32.25">32º 15' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A755366F409B9390616FD" box="[1018,1148,1932,1958]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="126.3">126º 18' E</geoCoordinate>
); R91602 (M)
<quantity id="4CD79B213B6A7553603A09B938FB16FD" box="[1332,1409,1932,1958]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="37" unit="km" value="16.0">16 km</quantity>
E Toolinna Rockhole (
<geoCoordinate id="EE1B50033B6A7553649C09873F7F1697" box="[402,517,1970,1996]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-32.666668">32º 40' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A7553671A09873FE21697" box="[532,664,1970,1996]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="125.11667">125º 07' E</geoCoordinate>
); R91627, R91628 and R91630 (all M)
<quantity id="4CD79B213B6A75536184098739AA1697" box="[1162,1232,1970,1996]" metricMagnitude="4" metricUnit="m" metricValue="5.3" pageId="12" pageNumber="37" unit="km" value="53.0">53km</quantity>
WNW Toolinna Rockhole (
<geoCoordinate id="EE1B50033B6A7553641909EC3CFF16A8" box="[279,389,2009,2035]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-32.666668">32º 40' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A7553649D09EC3F6B16A8" box="[403,529,2009,2035]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="125.11667">125º 07' E</geoCoordinate>
); R93273 (F) and R93275 (F)
<quantity id="4CD79B213B6A7553667209EC3EB216A8" box="[892,968,2009,2035]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="37" unit="km" value="32.0">32 km</quantity>
NW Clyde Hill (
<geoCoordinate id="EE1B50033B6A7553619A09EC387B16A8" box="[1172,1281,2009,2035]" direction="south" orientation="latitude" pageId="12" pageNumber="37" precision="925" value="-33.2">33º 12' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6A7553600109EC38F416A8" box="[1295,1422,2009,2035]" direction="east" orientation="longitude" pageId="12" pageNumber="37" precision="925" value="122.71667">122º 43' E</geoCoordinate>
); R114154 (F)
<quantity id="4CD79B213B6B7552643A0EAD3C0A11E9" box="[308,368,152,178]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="38" unit="km" value="4.0">4 km</quantity>
N Ravensthorpe (
<geoCoordinate id="EE1B50033B6B7552674A0EAD3FD511E9" box="[580,687,152,178]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="925" value="-33.533333">33º 32' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B755267B50EAD3E4D11E9" box="[699,823,152,178]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="925" value="120.05">120º 03' E</geoCoordinate>
); R116051 (F)
<geoCoordinate id="EE1B50033B6B755266E40EAD395211E9" box="[1002,1064,152,178]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="5555" value="3.5">3.5 E</geoCoordinate>
Calyerup Rocks (
<geoCoordinate id="EE1B50033B6B755261F50EAD38EC11E9" box="[1275,1430,152,178]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="15" value="-33.890556">33º 53' 26&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B755265990E8B3C381183" box="[151,322,190,216]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="15" value="119.12556">119º 07' 32&quot; E</geoCoordinate>
); R133572 (F) Saint Ronans Nature Reserve (
<geoCoordinate id="EE1B50033B6B755266510E8B3E811183" box="[863,1019,190,216]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="15" value="-31.875">31º 52' 30&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B755261060E8B39C91183" box="[1032,1203,190,216]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="15" value="116.602776">116º 36' 10&quot; E</geoCoordinate>
); R
<date id="FF9110043B6B755261D30E8B384F1183" box="[1245,1333,190,216]" pageId="13" pageNumber="38" value="32-09-14">140932</date>
(M) and R
<date id="FF9110043B6B755265A20ED03C7211A4" box="[172,264,229,255]" pageId="13" pageNumber="38" value="41-09-14">140941</date>
(M) 27.5 km SSE Peak Eleanora (
<geoCoordinate id="EE1B50033B6B755267B40ED03E5411A4" box="[698,814,229,255]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="925" value="-33.166668">33º 10' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B755266330ED03EB911A4" box="[829,963,229,255]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="925" value="121.26667">121º 16' E</geoCoordinate>
); R143815 (M) Lake Magenta Nature Reserve (
<geoCoordinate id="EE1B50033B6B755264080F393CD9107D" box="[262,419,268,294]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="15" value="-33.494724">33º 29' 41&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B755264BE0F393F26107D" box="[432,604,268,294]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="15" value="119.04806">119º 02' 53&quot; E</geoCoordinate>
); R144224 (M) Bandalup Hill (
<geoCoordinate id="EE1B50033B6B755266DC0F393912107D" box="[978,1128,268,294]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="15" value="-33.66611">33º 39' 58&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B7552617A0F393861107D" box="[1140,1307,268,294]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="15" value="120.398605">120º 23' 55&quot;E</geoCoordinate>
); R149069 (F) Scaddan area (
<geoCoordinate id="EE1B50033B6B755264600F073F791017" box="[366,515,306,332]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="15" value="-33.386112">33º 23' 10&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B7552671E0F073FCF1017" box="[528,693,306,332]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="15" value="121.58278">121º 34' 58&quot;E</geoCoordinate>
); R
<date id="FF9110043B6B755267D10F073E4D1017" box="[735,823,306,332]" pageId="13" pageNumber="38" value="09-12-15">151209</date>
(M) Salmon Gums area (
<geoCoordinate id="EE1B50033B6B7552616C0F0739851017" box="[1122,1279,306,332]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="15" value="-32.809998">32º 48' 36&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B755260020F073DD71028" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="15" value="121.39">121º 23' 24&quot; E</geoCoordinate>
); R154175 (M) near Kundip (
<geoCoordinate id="EE1B50033B6B7552671C0F6C3FCB1028" box="[530,689,345,371]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="15" value="-33.68778">33º 41' 16&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B755267B10F6C3E151028" box="[703,879,345,371]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="15" value="120.19778">120º 11' 52&quot; E</geoCoordinate>
); R156209 (F) Kundip (
<geoCoordinate id="EE1B50033B6B755261810F6C38541028" box="[1167,1326,345,371]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="15" value="-33.68528">33º 41' 07&quot; S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B755260330F6C3DA610C1" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="15" value="120.19722">120º 11' 50&quot; E</geoCoordinate>
); R156943 (M) and R156947 (M) Forrestania area (
<geoCoordinate id="EE1B50033B6B755266310FB53ED010C1" box="[831,938,384,410]" direction="south" orientation="latitude" pageId="13" pageNumber="38" precision="925" value="-32.4">32º 24' S</geoCoordinate>
,
<geoCoordinate id="EE1B50033B6B755266B90FB5394B10C1" box="[951,1073,384,410]" direction="east" orientation="longitude" pageId="13" pageNumber="38" precision="925" value="119.683334">119º 41' E</geoCoordinate>
).
</paragraph>
</subSubSection>
<caption id="DF50664C3B6B755265990D5C3EB712DA" box="[151,973,873,897]" pageId="13" pageNumber="38">
<paragraph id="8B9036C43B6B755265990D5C3EB712DA" blockId="13.[151,973,873,897]" box="[151,973,873,897]" pageId="13" pageNumber="38">
<emphasis id="B95BEAD63B6B755265990D5C3C5C12DA" bold="true" box="[151,294,873,897]" pageId="13" pageNumber="38">FIGURE 10.</emphasis>
Holotype of
<taxonomicName id="4C2F4D473B6B755264BA0D5C3FC612DB" box="[436,700,873,896]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="38" phylum="Chordata" rank="species" species="calcicolus" status="sp. nov.">
<emphasis id="B95BEAD63B6B755264BA0D5C3FC612DB" box="[436,700,873,896]" italics="true" pageId="13" pageNumber="38">Diplodactylus calcicolus</emphasis>
</taxonomicName>
<emphasis id="B95BEAD63B6B755267CD0D5C3E6B12DA" bold="true" box="[707,785,873,897]" pageId="13" pageNumber="38">
<taxonomicNameLabel id="A26857AD3B6B755267CD0D5C3E6B12DA" box="[707,785,873,897]" pageId="13" pageNumber="38" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
, SAMA R25343.
</paragraph>
</caption>
<caption id="DF50664C3B6B755265990BCA3F851707" httpUri="https://zenodo.org/record/189139/files/figure.png" pageId="13" pageNumber="38" targetBox="[295,1238,970,1518]" targetPageId="13">
<paragraph id="8B9036C43B6B755265990BCA3F851707" blockId="13.[151,1436,1535,1628]" pageId="13" pageNumber="38">
<emphasis id="B95BEAD63B6B755265990BCA3C50174C" bold="true" box="[151,298,1535,1559]" pageId="13" pageNumber="38">FIGURE 11.</emphasis>
Variation in colour pattern in
<taxonomicName id="4C2F4D473B6B7552677308353EEB174C" box="[637,913,1536,1559]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="38" phylum="Chordata" rank="species" species="calcicolus" status="sp. nov.">
<emphasis id="B95BEAD63B6B7552677308353EEB174C" box="[637,913,1536,1559]" italics="true" pageId="13" pageNumber="38">Diplodactylus calcicolus</emphasis>
</taxonomicName>
<emphasis id="B95BEAD63B6B755266970BCA3E88174C" bold="true" box="[921,1010,1535,1559]" pageId="13" pageNumber="38">
<taxonomicNameLabel id="A26857AD3B6B755266970BCA3E88174C" box="[921,1010,1535,1559]" pageId="13" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
From left to right: western specimens (WAM R140935, WAM R156947, WAM R156943), Nullarbor specimens (SAM R59582, SAM R59524), and eastern specimens (SAM R40554, SAM R58537, SAM R36644).
</paragraph>
</caption>
<subSubSection id="C335654F3B6B755265CB08BF3D9E1642" pageId="13" pageNumber="38" type="diagnosis">
<paragraph id="8B9036C43B6B755265CB08BF3D9E1642" blockId="13.[151,1436,1674,2010]" pageId="13" pageNumber="38">
<emphasis id="B95BEAD63B6B755265CB08BF3C3217FF" bold="true" box="[197,328,1674,1700]" pageId="13" pageNumber="38">Diagnosis.</emphasis>
A moderate-sized, longer tailed member of the
<taxonomicName id="4C2F4D473B6B7552668308BE397617FF" box="[909,1036,1675,1700]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="38" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="B95BEAD63B6B7552668308BE397617FF" box="[909,1036,1675,1700]" italics="true" pageId="13" pageNumber="38">D. vittatus</emphasis>
</taxonomicName>
complex, typically with a highly broken and spotted pattern, but showing considerable within-population variation. Flanks marked with spots of various sizes, and background colour of dorsum usually variegated by a mixture of lighter and darker scales.
</paragraph>
</subSubSection>
<subSubSection id="C335654F3B6B755165C809103F3E1317" lastPageId="14" lastPageNumber="39" pageId="13" pageNumber="38" type="description">
<paragraph id="8B9036C43B6B755265C8091039CC1681" blockId="13.[151,1436,1674,2010]" pageId="13" pageNumber="38">
<emphasis id="B95BEAD63B6B755265C809103C2D1664" bold="true" box="[198,343,1829,1855]" pageId="13" pageNumber="38">Description</emphasis>
(
<figureCitation id="13142A413B6B7552646909103CB91664" box="[359,451,1829,1855]" captionStart="FIGURE 10" captionStartId="13.[151,255,873,897]" captionText="FIGURE 10. Holotype of Diplodactylus calcicolus sp. nov, SAMA R 25343." pageId="13" pageNumber="38">Figs 10</figureCitation>
,
<figureCitation id="13142A413B6B755264DE09103C941664" box="[464,494,1829,1855]" captionStart="FIGURE 11" captionStartId="13.[151,257,1535,1559]" captionTargetBox="[295,1238,970,1518]" captionTargetId="figure@13.[295,1304,948,1519]" captionTargetPageId="13" captionText="FIGURE 11. Variation in colour pattern in Diplodactylus calcicolus sp. nov. From left to right: western specimens (WAM R 140935, WAM R 156947, WAM R 156943), Nullarbor specimens (SAM R 59582, SAM R 59524), and eastern specimens (SAM R 40554, SAM R 58537, SAM R 36644)." httpUri="https://zenodo.org/record/189139/files/figure.png" pageId="13" pageNumber="38">11</figureCitation>
,
<figureCitation id="13142A413B6B755264F209103F611664" box="[508,539,1829,1855]" captionStart="FIGURE 14" captionStartId="16.[151,257,1878,1902]" captionTargetBox="[210,1388,208,1840]" captionTargetId="figure@16.[194,1400,194,1851]" captionTargetPageId="16" captionText="FIGURE 14. Species of the Diplodactylus vittatus complex in life. A, D. vittatus, Warrumbungle Range, NSW, (M. Hutchinson); B, D. furcosus, Whyalla, South Australia (T. Peters, SA Museum); C, D. g. granariensis, Newman Rock, WA (B. Maryan); D, D. g. rex, Wildara Pinnacle, WA (B. Maryan); E, D. calcicolus sp. nov., vicinity of the WA-SA border (G. Harold); F, D. wiru sp. nov. Lake Ifould, SA (P. Oliver)." httpUri="https://zenodo.org/record/189141/files/figure.png" pageId="13" pageNumber="38">14</figureCitation>
E). SVL 31.0–58.0 mm (mean 48.9, n=34); tailL 24.5–38.0 mm, mean 32.4, (56.6–72.3% SVL, mean 64.5) (n=18). Supralabials 10–13 (mean 11.5, n=17), first and second supralabial generally subequal or the second slightly higher than the first. Infralabials 8–13 (mean 10.4, n=17). Apex of rostral scale with median division descending to about the mid-height of the scale. Supranasals usually in contact; a single internarial present in 5 of 34 examined. 2–5 postnasals (mean 3.1, n=34).
</paragraph>
<paragraph id="8B9036C43B68755165CB0EAD3840137C" blockId="14.[151,1437,152,1447]" pageId="14" pageNumber="39">The most variable in colour and pattern of the group. Dorsal colour pale to very dark brown, grey, or greybrown. Beige to pale grey vertebral stripe most often broken into a disconnected series of irregularly-shaped dorsal blotches, but within population variation can include individuals with a more continuous chain of blotches, or a continuous dorsal stripe with deeply to moderately scalloped margins. Vertebral stripe usually bifurcates on the nape with well-defined lateral margins but irregularly defined inner margins, so that the crown of the head tends to be paler laterally but with a darker, weakly defined central patch of background colour. Blackish paravertebral margins of vertebral stripe variable in contrast, but even if prominent, often discontinuous due to random interruptions by lighter scales. Upper lateral zone sometimes patterned with irregularly distributed circular spots; some specimens with very large spots, similar in size to the vertebral series of blotches. Apart from such spots, the upper lateral surfaces have a ‘messy’ texturing of lighter and darker scales overlying the medium-toned background colour. Ventral surface white to greyish white.</paragraph>
<paragraph id="8B9036C43B68755165C80C073F3E1317" blockId="14.[151,1437,152,1447]" box="[198,580,562,588]" pageId="14" pageNumber="39">
<emphasis id="B95BEAD63B68755165C80C073C291317" bold="true" box="[198,339,562,588]" pageId="14" pageNumber="39">Karyotype.</emphasis>
2n=38 (
<bibRefCitation id="EFBE4B353B68755164BB0C073F4D1317" author="King" box="[437,567,562,588]" pageId="14" pageNumber="45" refString="King, M. (1977) Chromosomal and morphometric variation in the gecko Diplodactylus vittatus (Gray). Australian Journal of Zoology, 25, 42 - 57." type="journal article" year="1977">King 1977</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C335654F3B68755165C80C623DB613B9" pageId="14" pageNumber="39" type="distribution">
<paragraph id="8B9036C43B68755165C80C623DB613B9" blockId="14.[151,1437,152,1447]" pageId="14" pageNumber="39">
<emphasis id="B95BEAD63B68755165C80C623C1D132A" bold="true" box="[198,359,599,625]" pageId="14" pageNumber="39">Distribution.</emphasis>
Distributed along the dry temperate to arid southern coast and hinterland of
<collectingCountry id="F33876543B68755161E60C6D382C1329" box="[1256,1366,600,626]" name="Australia" pageId="14" pageNumber="39">Australia</collectingCountry>
, from near Perth, WA, south to Albany (absent from southwest corner) and along the Great Australian Bight to the Eyre Peninsula, with outliers on the southern York Peninsula and the mouth of the Murray River (
<figureCitation id="13142A413B68755160130C97381213E7" box="[1309,1384,674,700]" captionStart="FIGURE 1" captionStartId="2.[151,256,1270,1294]" captionTargetBox="[269,1335,477,1269]" captionTargetId="figure@2.[226,1360,438,1303]" captionTargetPageId="2" captionText="FIGURE 1. Map of southern Australia showing the distribution of the members of the Diplodactylus vittatus species complex described or re-described in the present study. Insets shows details of species distributions in overlap areas in Western Australia and South Australia (See Figs 15 and 16). Diplodactylus vittatus, square; D. furcosus, black cross, D. g. granariensis, circle; D. g. rex, circle with central dot, D. calcicolus sp. nov., grey cross, D. wiru sp. nov., triangle." httpUri="https://zenodo.org/record/189133/files/figure.png" pageId="14" pageNumber="39">Figs 1</figureCitation>
,
<figureCitation id="13142A413B68755160790C9738ED13E7" box="[1399,1431,674,700]" captionStart="FIGURE 15" captionStartId="18.[151,256,859,883]" captionTargetBox="[384,1197,189,822]" captionTargetId="figure@18.[377,1210,177,835]" captionTargetPageId="18" captionText="FIGURE 15. Detail of distribution of D. vittatus complex in Western Australia. Note significant range overlap of D. granariensis and D. calcicolus sp. nov. in the southwest. Symbols as in Fig. 1." httpUri="https://zenodo.org/record/189142/files/figure.png" pageId="14" pageNumber="39">15</figureCitation>
,
<figureCitation id="13142A413B68755165990CFD3DCF13B9" box="[151,181,712,738]" captionStart="FIGURE 16" captionStartId="18.[151,258,1592,1616]" captionTargetBox="[309,1241,963,1592]" captionTargetId="figure@18.[302,1285,931,1605]" captionTargetPageId="18" captionText="FIGURE 16. Detail of distribution of D. vittatus complex in South Australia. Note significant range overlap of D. furcosus and D. calcicolus sp. nov. on Eyre Peninsula, and the two regions (arrowed) where three species are sympatric. Symbols as in Fig. 1." httpUri="https://zenodo.org/record/189143/files/figure.png" pageId="14" pageNumber="39">16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C335654F3B68755165C80CD83E2F146C" pageId="14" pageNumber="39" type="etymology">
<paragraph id="8B9036C43B68755165C80CD83F1C1209" blockId="14.[151,1437,152,1447]" pageId="14" pageNumber="39">
<emphasis id="B95BEAD63B68755165C80CD83C29125C" bold="true" box="[198,339,749,775]" pageId="14" pageNumber="39">Etymology.</emphasis>
The specific name has been constructed from Latin roots '
<emphasis id="B95BEAD63B687551610E0CD83943125D" box="[1024,1081,749,774]" italics="true" pageId="14" pageNumber="39">calci</emphasis>
' (lime) and '
<emphasis id="B95BEAD63B68755161C20CD8387F125D" box="[1228,1285,749,774]" italics="true" pageId="14" pageNumber="39">cola'</emphasis>
(inhabitant), referring to the dominant limestone geology of the coast and hinterland of the Great Australian Bight, the central area of the species’ distribution.
</paragraph>
<paragraph id="8B9036C43B68755165CB0D683E2F146C" blockId="14.[151,1437,152,1447]" pageId="14" pageNumber="39">
<emphasis id="B95BEAD63B68755165CB0D683CF1122C" bold="true" box="[197,395,861,887]" pageId="14" pageNumber="39">Similar Species.</emphasis>
This is the most variable species within the complex, and it overlaps geographically with all other taxa treated here except
<taxonomicName id="4C2F4D473B68755167100DB73FFE12C0" box="[542,644,898,923]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="subSpecies" species="granariensis" subSpecies="rex">
<emphasis id="B95BEAD63B68755167100DB73FFE12C0" box="[542,644,898,923]" italics="true" pageId="14" pageNumber="39">D. g. rex</emphasis>
</taxonomicName>
. Within most populations of
<taxonomicName id="4C2F4D473B68755166D30DB7390212C0" box="[989,1144,898,923]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="calcicolus">
<emphasis id="B95BEAD63B68755166D30DB7390212C0" box="[989,1144,898,923]" italics="true" pageId="14" pageNumber="39">D. calcicolus</emphasis>
</taxonomicName>
a minority of specimens have a continuous vertebral stripe combined with weakly marked flanks. Such individuals are difficult to distinguish from sympatric individuals of
<taxonomicName id="4C2F4D473B687551679E0DF83E6312BD" box="[656,793,973,998]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="furcosus">
<emphasis id="B95BEAD63B687551679E0DF83E6312BD" box="[656,793,973,998]" italics="true" pageId="14" pageNumber="39">D. furcosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F4D473B68755166270DF8397E12BD" box="[809,1028,973,998]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="subSpecies" species="granariensis" subSpecies="granariensis">
<emphasis id="B95BEAD63B68755166270DF8397E12BD" box="[809,1028,973,998]" italics="true" pageId="14" pageNumber="39">D. g. granariensis</emphasis>
</taxonomicName>
, or
<taxonomicName id="4C2F4D473B687551613A0DF839C812BD" box="[1076,1202,973,998]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="B95BEAD63B687551613A0DF839C812BD" box="[1076,1202,973,998]" italics="true" pageId="14" pageNumber="39">D. vittatus</emphasis>
</taxonomicName>
. The stripe is more irregular than in either of the other species, the flanks have intermixed darker and lighter scales rather than being uniformly coloured and the occipital markings are usually more poorly defined (
<figureCitation id="13142A413B687551619E0A2D39A21569" box="[1168,1240,1048,1074]" captionStart="FIGURE 9" captionStartId="11.[151,255,1826,1850]" captionTargetBox="[217,1369,1034,1802]" captionTargetId="figure@11.[217,1369,1034,1803]" captionTargetPageId="11" captionText="FIGURE 9. Variation in colour pattern in Diplodactylus granariensis rex" httpUri="https://zenodo.org/record/189138/files/figure.png" pageId="14" pageNumber="39">Fig. 9</figureCitation>
). The longer tail of
<taxonomicName id="4C2F4D473B68755165B70A083C27150D" box="[185,349,1085,1110]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="calcicolus" status="sp. nov.">
<emphasis id="B95BEAD63B68755165B70A083C27150D" box="[185,349,1085,1110]" italics="true" pageId="14" pageNumber="39">D. calcicolus</emphasis>
</taxonomicName>
<emphasis id="B95BEAD63B687551646B0A083CB2150C" bold="true" box="[357,456,1085,1111]" pageId="14" pageNumber="39">
<taxonomicNameLabel id="A26857AD3B687551646B0A083CB2150C" box="[357,456,1085,1111]" pageId="14" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
further distinguishes it from
<taxonomicName id="4C2F4D473B687551663B0A083ECD150D" box="[821,951,1085,1110]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="B95BEAD63B687551663B0A083ECD150D" box="[821,951,1085,1110]" italics="true" pageId="14" pageNumber="39">D. vittatus</emphasis>
</taxonomicName>
in the small overlap zone on the lower Murray River. In the southwest of Western
<collectingCountry id="F33876543B68755167940A573E731527" box="[666,777,1122,1148]" name="Australia" pageId="14" pageNumber="39">Australia</collectingCountry>
, the diffuse spotted markings of
<taxonomicName id="4C2F4D473B68755161810A5638151527" box="[1167,1391,1123,1148]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="polyophthalmus">
<emphasis id="B95BEAD63B68755161810A5638151527" box="[1167,1391,1123,1148]" italics="true" pageId="14" pageNumber="39">D. polyophthalmus</emphasis>
</taxonomicName>
are unlike the more solidly delineated and irrregular spots of
<taxonomicName id="4C2F4D473B68755166440ABD3E9D15FA" box="[842,999,1160,1185]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="calcicolus">
<emphasis id="B95BEAD63B68755166440ABD3E9D15FA" box="[842,999,1160,1185]" italics="true" pageId="14" pageNumber="39">D. calcicolus</emphasis>
</taxonomicName>
. At present, distinguishing some far western
<taxonomicName id="4C2F4D473B68755165F70A983CEE159D" box="[249,404,1197,1222]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="calcicolus" status="sp. nov.">
<emphasis id="B95BEAD63B68755165F70A983CEE159D" box="[249,404,1197,1222]" italics="true" pageId="14" pageNumber="39">D. calcicolus</emphasis>
</taxonomicName>
<emphasis id="B95BEAD63B68755164940A983C8D159C" bold="true" box="[410,503,1197,1223]" pageId="14" pageNumber="39">
<taxonomicNameLabel id="A26857AD3B68755164940A983C8D159C" box="[410,503,1197,1223]" pageId="14" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from geographically proximate individuals of
<taxonomicName id="4C2F4D473B68755161140A9839ED159D" box="[1050,1175,1197,1222]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="B95BEAD63B68755161140A9839ED159D" box="[1050,1175,1197,1222]" italics="true" pageId="14" pageNumber="39">D. ornatus</emphasis>
</taxonomicName>
can be difficult. More detailed genetic comparisons among Perth area
<taxonomicName id="4C2F4D473B68755167E20AE63EE015B7" box="[748,922,1235,1260]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="genus">
<emphasis id="B95BEAD63B68755167E20AE63EE015B7" box="[748,922,1235,1260]" italics="true" pageId="14" pageNumber="39">Diplodactylus</emphasis>
</taxonomicName>
populations, focusing on the distinctions between
<taxonomicName id="4C2F4D473B687551640F0ACD3CE6144A" box="[257,412,1272,1297]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="calcicolus" status="sp. nov.">
<emphasis id="B95BEAD63B687551640F0ACD3CE6144A" box="[257,412,1272,1297]" italics="true" pageId="14" pageNumber="39">D. calcicolus</emphasis>
</taxonomicName>
<emphasis id="B95BEAD63B68755164AA0AC23F78144A" bold="true" box="[420,514,1271,1297]" pageId="14" pageNumber="39">
<taxonomicNameLabel id="A26857AD3B68755164AA0AC23F78144A" box="[420,514,1271,1297]" pageId="14" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C2F4D473B68755167330ACD3FC1144A" box="[573,699,1272,1297]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="39" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="B95BEAD63B68755167330ACD3FC1144A" box="[573,699,1272,1297]" italics="true" pageId="14" pageNumber="39">D. ornatus</emphasis>
</taxonomicName>
, will be necessary before the unambiguous genetic distinction can be matched by equally clear morphological characters..
</paragraph>
</subSubSection>
<subSubSection id="C335654F3B68755165CB0B773ED614FC" pageId="14" pageNumber="39" type="discussion">
<paragraph id="8B9036C43B68755165CB0B773ED614FC" blockId="14.[151,1437,152,1447]" pageId="14" pageNumber="39">
<emphasis id="B95BEAD63B68755165CB0B773C3A1407" bold="true" box="[197,320,1346,1372]" pageId="14" pageNumber="39">Remarks.</emphasis>
This species mostly occurs in cool dry habitats, typically on limestone or calcareous coastal sands.
<bibRefCitation id="EFBE4B353B68755165E90B5D3CA314D9" box="[231,473,1384,1410]" pageId="14" pageNumber="45" refString="Oliver, P., Hugall, A., Adams, M., Cooper, S. J. B. &amp; Hutchinson, M. (2007 a) Genetic elucidation of ancient and cryptic diversity in a group of Australian geckos: the Diplodactylus vittatus complex. Molecular Phylogenetics and Evolution, 44, 77 - 88." type="journal article">
Oliver
<emphasis id="B95BEAD63B68755164340B5D3C0C14DA" box="[314,374,1384,1409]" italics="true" pageId="14" pageNumber="39">et al.</emphasis>
(2007a)
</bibRefCitation>
noted that this species comprises three genetic lineages that do not correspond to any obvious patterns of variation in the characters discussed above.
</paragraph>
</subSubSection>
</treatment>
</document>
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