treatments-xml/data/03/86/87/038687B6697F8F7AFF1BF89AFC308007.xml

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<mods:title id="56B00550A2E7D126C1B4579A663BC6B3">A taxonomic revision of the Eucelatoria ferox species group (Diptera: Tachinidae)</mods:title>
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<paragraph id="8B9036A0697F8F78FF1BF89AFD858383" blockId="53.[151,650,1914,1978]" box="[151,551,1914,1941]" pageId="53" pageNumber="54">
<heading id="D0D881CC697F8F78FF1BF89AFD858383" bold="true" box="[151,551,1914,1941]" fontSize="11" level="2" pageId="53" pageNumber="54" reason="4">
<emphasis id="B95BEAB2697F8F78FF1BF89AFD858383" bold="true" box="[151,551,1914,1941]" pageId="53" pageNumber="54">
<taxonomicName id="4C2F4D23697F8F78FF1BF89AFE608383" ID-CoL="9J36F" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[151,450,1914,1941]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="ritavargasae" status="sp. nov.">
<emphasis id="B95BEAB2697F8F78FF1BF89AFE608383" bold="true" box="[151,450,1914,1941]" italics="true" pageId="53" pageNumber="54">Eucelatoria ritavargasae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857C9697F8F78FE46F89BFD858383" box="[458,551,1915,1941]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C335652B697F8F78FF1BF87FFD2883AC" box="[151,650,1950,1978]" pageId="53" pageNumber="54" type="description">
<paragraph id="8B9036A0697F8F78FF1BF87FFD2883AC" blockId="53.[151,650,1914,1978]" box="[151,650,1950,1978]" pageId="53" pageNumber="54">
(
<figureCitation id="13142A25697F8F78FF13F87FFF5883AF" box="[159,250,1951,1977]" captionStart="FIGURE 1" captionStartId="9.[151,250,1766,1790]" captionTargetBox="[164,1421,431,1729]" captionTargetId="figure-49@9.[153,1434,417,1743]" captionTargetPageId="9" captionText="FIGURE 1. CO1-based neighbor-joining tree of 12 species of the Eucelatoria ferox species group, with inset illustrations of mesonota. The outgroup is Eucelatoria armigera (Coquillett). Node labels indicate support values over 60. Scale bar = 2% genetic difference. The E. charapensis (Townsend) sequence with the axial break is of poor quality but is included to show unity of El Bigal specimens with the rest of the metapopulation. Mesonota are illustrated with setal annuli only for clarity. A. E. gustavogutierrezi sp. nov. B. E. kopis sp. nov. C. E. ritavargasae sp. nov. D. E. ferox (Townsend). E. E. huitepecensis sp. nov. F. E. auriceps (Aldrich). G. E. gladiatrix (Townsend). H. E. borealis sp. nov. I. E. yanayacu sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601406" httpUri="https://zenodo.org/record/6601406/files/figure.png" pageId="53" pageNumber="54">Figs 1C</figureCitation>
, 
<figureCitation id="13142A25697F8F78FE8AF87FFE8683AF" box="[262,292,1951,1977]" captionStart="FIGURES 22–25" captionStartId="67.[151,264,1943,1967]" captionTargetBox="[170,1416,181,1920]" captionTargetId="figure-15@67.[170,1416,181,1920]" captionTargetPageId="67" captionText="FIGURES 22–25. Female habitus photographs of Eucelatoria ferox group spp., dorsal view. 22. E. kopis sp. nov., holotype (ZLB_E.Ferox 00002) (CNC). 23. E. makhaira sp. nov., holotype (ZLB_E.Ferox 00344) (USNM). 24. E. ritavargasae sp. nov., Napo Province (ZLB_E.Ferox 00214) (USNM). 25. E. sica sp. nov., holotype (ZLB_E.Ferox 00391) (USNM)." figureDoi="http://doi.org/10.5281/zenodo.6601418" httpUri="https://zenodo.org/record/6601418/files/figure.png" pageId="53" pageNumber="54">24</figureCitation>
, 
<figureCitation id="13142A25697F8F78FEBCF87EFEEC83AF" box="[304,334,1950,1977]" captionStart="FIGURES 42–45" captionStartId="72.[151,264,1943,1967]" captionTargetBox="[172,1415,181,1920]" captionTargetId="figure-31@72.[172,1415,181,1920]" captionTargetPageId="72" captionText="FIGURES 42–45. Male habitus photographs of Eucelatoria ferox group spp., dorsal view. 42. E. hafelei sp. nov., Chapare Province (ZLB_E.Ferox 00386) (DMW). 43. E. kopis sp. nov., allotype (ZLB_E.Ferox 00003) (CNC). 44. E. makhaira sp. nov., allotype (ZLB_E.Ferox 00343) (USNM). 45. E. ritavargasae sp. nov., holotype (ZLB_E.Ferox 00004) (CNC)." figureDoi="http://doi.org/10.5281/zenodo.6601428" httpUri="https://zenodo.org/record/6601428/files/figure.png" pageId="53" pageNumber="54">45</figureCitation>
, 
<figureCitation id="13142A25697F8F78FED6F87FFEDA83AF" box="[346,376,1951,1977]" captionStart="FIGURES 68–69" captionStartId="84.[151,264,1943,1967]" captionTargetBox="[196,1391,181,1920]" captionTargetId="figure-31@84.[196,1391,181,1920]" captionTargetPageId="84" captionText="FIGURES 68–69. Female habitus photographs of Eucelatoria ferox group spp., lateral view. 68. E. makhaira sp. nov., holotype (ZLB_E.Ferox 00344) (USNM). 69. E. ritavargasae sp. nov., Napo Province (ZLB_E.Ferox 00214) (USNM)." figureDoi="http://doi.org/10.5281/zenodo.6601454" httpUri="https://zenodo.org/record/6601454/files/figure.png" pageId="53" pageNumber="54">69</figureCitation>
, 
<figureCitation id="13142A25697F8F78FE08F87FFE0183AF" box="[388,419,1951,1977]" captionStart="FIGURES 90–91" captionStartId="95.[151,264,1943,1967]" captionTargetBox="[196,1391,181,1920]" captionTargetId="figure-15@95.[196,1391,181,1920]" captionTargetPageId="95" captionText="FIGURES 90–91. Male habitus photographs of Eucelatoria ferox group spp., lateral view. 90. E. ritavargasae sp. nov., holotype (ZLB_E.Ferox 00004) (CNC). 91. E. tenebrionis sp. nov., Nova Teutonia (ZLB_E.Ferox 00268) (DMW)." figureDoi="http://doi.org/10.5281/zenodo.6601492" httpUri="https://zenodo.org/record/6601492/files/figure.png" pageId="53" pageNumber="54">90</figureCitation>
, 
<figureCitation id="13142A25697F8F78FE22F87FFDB083AF" box="[430,530,1950,1977]" captionStart="FIGURES 136–155" captionStartId="100.[151,264,1677,1701]" captionTargetBox="[277,1319,184,1650]" captionTargetId="figure-31@100.[264,1324,181,1653]" captionTargetPageId="100" captionText="FIGURES 136–155. Male terminalia of Eucelatoria ferox group spp., left lateral view, and caudal view of cerci and surstyli. Numbers are “ZLB_E.Ferox” specimen ID numbers. Specimen information for caudal views is identical to that for Figs 124– 139. 136. E. gustavogutierrezi sp. nov., lateral view, Guanacaste Province (00008) (CNC). 137. E. hafelei sp. nov., lateral view, Chapare Province (00386) (DMW). 138. E. ritavargasae sp. nov., lateral view, Narupa (00215) (USNM). 139. E. sica sp. nov., lateral view, Yanayacu (00213) (CNC). 140. E. aurata (Townsend), caudal view. 141. E. auriceps (Aldrich), caudal view. 142. E. borealis sp. nov., caudal view. 143. E. charapensis (Townsend), caudal view. 144. E. falcata sp. nov., caudal view. 145. E. fordlandia sp. nov., caudal view. 146. E. gladiatrix (Townsend), caudal view. 147. E. sabroskyi sp. nov., caudal view. 148. E. strigata (Wulp), caudal view. 149. E. texana (Reinhard), caudal view. 150. E. woodorum sp. nov., caudal view. 151. E. yanayacu sp. nov., caudal view. 152. E. gustavogutierrezi sp. nov., caudal view. 153. E. hafelei sp. nov., caudal view. 154. E. ritavargasae sp. nov., caudal view. 155. E. sica sp. nov., caudal view." figureDoi="http://doi.org/10.5281/zenodo.6601508" httpUri="https://zenodo.org/record/6601508/files/figure.png" pageId="53" pageNumber="54">138, 154</figureCitation>
, 
<figureCitation id="13142A25697F8F78FD92F87FFDE883AF" box="[542,586,1951,1977]" captionStart="FIGURES 156–173" captionStartId="101.[151,264,1755,1779]" captionTargetBox="[261,1320,195,1718]" captionTargetId="figure-15@101.[253,1335,181,1731]" captionTargetPageId="101" captionText="FIGURES 156–173. Male terminalia and fifth abdominal sternite of Eucelatoria ferox group spp. Specimen information for fifth sternites is identical to that for Figs 124–139. 156. Generic male terminalia, left lateral view (left) and caudal view (right), setae and membrane omitted.Abbreviations: aed, aedeagal apodeme; bas, basal lobe of surstylus; C, depth of postgonite anterior emargination; cer, cercus; epa, epandrium; hyp, hypandrium; pha, phallus; pos, postgonite; pre, pregonite; sur, surstylus. 157. E. aurata (Townsend). 158. E. auriceps (Aldrich). 159. E. borealis sp. nov. 160. E. charapensis (Townsend). 161. E. falcata sp. nov. 162. E. fordlandia sp. nov. 163. E. gladiatrix (Townsend). 164. E. sabroskyi sp. nov. 165. E. strigata (Wulp). 166. E. texana (Reinhard). 167. E. woodorum sp. nov. 168. E. yanayacu sp. nov. 169. E. gustavogutierrezi sp. nov. 170. E. hafelei sp. nov. 171. E. ritavargasae sp. nov. 172. E. sica sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601512" httpUri="https://zenodo.org/record/6601512/files/figure.png" pageId="53" pageNumber="54">171</figureCitation>
, 
<figureCitation id="13142A25697F8F78FDDAF87FFD2383AC" box="[598,641,1951,1978]" captionStart="FIGURES 174–179" captionStartId="103.[151,264,1856,1880]" captionTargetBox="[203,1374,189,1818]" captionTargetId="figure-14@103.[193,1393,181,1832]" captionTargetPageId="103" captionText="FIGURES 174–179. Distribution maps of Eucelatoria spp. 174. E. yanayacu sp. nov., E. gladiatrix (Townsend), E. falcata sp. nov., E. charapensis (Townsend), and E. fordlandia sp. nov. 175. E. strigata (Wulp) and E. aurata (Townsend). 176. E. hafelei sp. nov., E. gustavogutierrezi sp. nov., and E. ritavargasae sp. nov. 177. E. ferox (Townsend), E. huitepecensis sp. nov., and E. inclani sp. nov. 178. E. kopis sp. nov., E. sica sp. nov., E. makhaira sp. nov., and E. tenebrionis sp. nov. 179. E. rivalis (Reinhard), E. luctuosa (Wulp), E. woodorum sp. nov., E. crambivora sp. nov., and E. jorgecortesi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601514" httpUri="https://zenodo.org/record/6601514/files/figure.png" pageId="53" pageNumber="54">176</figureCitation>
)
</paragraph>
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<emphasis id="B95BEAB2697C8F7BFF1BFF76FEE784A6" bold="true" box="[151,325,150,176]" pageId="54" pageNumber="55">Type material.</emphasis>
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, labeled “DHJPAR0021970”, “Voucher: 
<collectorName id="26DA5376697C8F7BFC13FF77FB8B84A7" box="[927,1065,151,177]" pageId="54" pageNumber="55">D.H. Janzen</collectorName>
&amp; 
<collectorName id="26DA5376697C8F7BFBC1FF77FB4A84A6" box="[1101,1256,150,177]" pageId="54" pageNumber="55">W. Hallwachs</collectorName>
/ DB: http://janzen.sas.upen.edu/ 
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,/ 
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./ 07 -SRNP- 3758”, “ 
<typeStatus id="54948802697C8F7BFB07FF5AFA8384C2" box="[1163,1313,186,212]" pageId="54" pageNumber="55" type="holotype">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="4C2F4D23697C8F7BFABDFF5AFF1684EE" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
/ ritavargasae/ ZL Burington [red label]”, “LEGS AWAY/ FOR DNA [yellow label]”, “Janzen01”, “ 
<taxonomicName id="4C2F4D23697C8F7BFA9EFF3EFA3684EE" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" box="[1298,1428,222,248]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
/ Janzen11/ Vide/ Det ZL Burington 2015”, “ 
<specimenCount id="9D29FD29697C8F7BFDF2FEE2FD36850A" box="[638,660,258,284]" pageId="54" pageNumber="55" type="male">♂</specimenCount>
black/ scutellum [handwritten]”, “ZLB_E.Ferox 00004” (
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).
</materialsCitation>
</paragraph>
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<materialsCitation id="3B473CFD697C8F7BFF4BFEC7FCA68556" ID-GBIF-Occurrence="3878846381" box="[199,772,294,321]" location="Brazil" pageId="54" pageNumber="55" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Santa Catarina" typeStatus="paratype">
<typeStatus id="54948802697C8F7BFF4BFEC7FE958557" box="[199,311,295,321]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
, 
<specimenCount id="9D29FD29697C8F7BFEC8FEC6FECD8557" box="[324,367,294,321]" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
and 
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. 
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— 
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: 
<materialsCitation id="3B473CFD697C8F7BFC9CFEC6FAA3859F" ID-GBIF-Occurrence="3878846623" collectingDate="1961-12" collectionCode="DMW" collectorName="F. Plaumann" country="Brazil" location="Eucuador" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="9D29FD29697C8F7BFC9CFEC6FC9C8557" box="[784,830,294,321]" pageId="54" pageNumber="55" type="male">1 ♂</specimenCount>
, labeled “Nova Teutonia/ S.C.- 
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/ 
<date id="FF911060697C8F7BFAACFEC7FA318556" box="[1312,1427,294,321]" pageId="54" pageNumber="55" value="1961-12">
<collectingDate id="EFD5E988697C8F7BFAACFEC7FA318556" box="[1312,1427,294,321]" pageId="54" pageNumber="55" value="1961-12">Dec. 1961</collectingDate>
</date>
/ 
<collectorName id="26DA5376697C8F7BFF1BFEABFE8F8572" box="[151,301,330,357]" pageId="54" pageNumber="55">F. Plaumann</collectorName>
”, “ 
<typeStatus id="54948802697C8F7BFEC3FEAAFE7F8572" box="[335,477,330,356]" pageId="54" pageNumber="55" type="paratype">PARATYPE</typeStatus>
/ 
<taxonomicName id="4C2F4D23697C8F7BFE7CFEAAFDCD8572" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" box="[496,623,330,356]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
/ ritavargasae/ ZL Burington [yellow label]”, “ 
<typeStatus id="54948802697C8F7BFB03FEAAFABF8572" box="[1167,1309,330,356]" pageId="54" pageNumber="55" type="paratype">PARATYPE</typeStatus>
/ 
<taxonomicName id="4C2F4D23697C8F7BFABDFEAAFF16859E" baseAuthorityName="Sabrosky" baseAuthorityYear="1981" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Eucelatoria</taxonomicName>
/ ritavargasae/ ZL Burington [yellow label]”, “ZLB_E.Ferox 00267” (
<collectionCode id="ED3EAE65697C8F7BFC5AFE8EFB86859E" box="[982,1060,366,392]" pageId="54" pageNumber="55">DMW</collectionCode>
). 
<location id="8EF0607B697C8F7BFBB5FE8EFB06859E" LSID="urn:lsid:plazi:treatment:038687B6697F8F7AFF1BF89AFC308007:8EF0607B697C8F7BFBB5FE8EFB06859E" box="[1081,1188,366,392]" country="Brazil" name="Eucuador" pageId="54" pageNumber="55" stateProvince="Napo">Eucuador</location>
— 
<collectingRegion id="49EBF842697C8F7BFB4EFE8FFAA3859F" box="[1218,1281,367,393]" country="Ecuador" name="Napo" pageId="54" pageNumber="55">Napo</collectingRegion>
</materialsCitation>
: 
<materialsCitation id="3B473CFD697C8F7BFA82FE8EFDCF85E2" ID-GBIF-Occurrence="3878846490" collectingDate="2012-04" collectionCode="JOSC" country="Ecuador" elevation="1186" location="Narupa" longitude="-0.73125" municipality="Prov." pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="9D29FD29697C8F7BFA82FE8EFA9F859F" box="[1294,1341,366,393]" pageId="54" pageNumber="55" type="male">1 ♂</specimenCount>
, labeled “ 
<collectingCountry id="F3387630697C8F7BFF2DFE72FE9285BA" box="[161,304,402,428]" name="Ecuador" pageId="54" pageNumber="55">ECUADOR</collectingCountry>
: 
<collectingMunicipality id="6BF4ACDA697C8F7BFEB6FE72FED685BA" box="[314,372,402,428]" pageId="54" pageNumber="55">Prov.</collectingMunicipality>
<collectingRegion id="49EBF842697C8F7BFEFFFE72FE1285BA" box="[371,432,402,428]" country="Ecuador" name="Napo" pageId="54" pageNumber="55">Napo</collectingRegion>
/ 
<location id="8EF0607B697C8F7BFE4CFE73FDB785BB" LSID="urn:lsid:plazi:treatment:038687B6697F8F7AFF1BF89AFC308007:8EF0607B697C8F7BFE4CFE73FDB785BB" box="[448,533,403,429]" country="Ecuador" longitude="-0.73125" municipality="Prov." name="Narupa" pageId="54" pageNumber="55" stateProvince="Napo">Narupa</location>
, 
<quantity id="4CD79B45697C8F7BFDADFE72FDCF85BA" box="[545,621,402,428]" metricMagnitude="3" metricUnit="m" metricValue="1.186" pageId="54" pageNumber="55" unit="m" value="1186.0">
<elevation id="0002D193697C8F7BFDADFE72FDCF85BA" box="[545,621,402,428]" metricMagnitude="3" metricUnit="m" metricValue="1.186" pageId="54" pageNumber="55" unit="m" value="1186.0">1186m</elevation>
</quantity>
/ S 00°43′52.5″ W 077°46′25.3″/ REARED:/ 
<date id="FF911060697C8F7BFBE7FE72FB4685BA" box="[1131,1252,402,428]" pageId="54" pageNumber="55" value="2012-04">
<collectingDate id="EFD5E988697C8F7BFBE7FE72FB4685BA" box="[1131,1252,402,428]" pageId="54" pageNumber="55" value="2012-04">ABR 2012</collectingDate>
</date>
66666 [voucher YY66666]”, “Ecuador reared/ 
<taxonomicName id="4C2F4D23697C8F7BFE7FFE56FD1285C6" box="[499,688,438,465]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="undefined-8">Eucelatoria sp. 8</taxonomicName>
/ ZLBurington 2014”, “ 
<taxonomicName id="4C2F4D23697C8F7BFC37FE56FBDC85C6" box="[955,1150,438,465]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="undefined-8">Eucelatoria sp. 8</taxonomicName>
”, “ZLB_E.Ferox 00215” [genitalia in vial under specimen] (
<collectionCode id="ED3EAE65697C8F7BFD91FE3AFDC685E2" box="[541,612,474,500]" pageId="54" pageNumber="55">JOSC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B473CFD697C8F7BFDFBFE3AFB78862A" ID-GBIF-Occurrence="3878846533" collectingDate="2005-06" collectionCode="JOSC, MECN" country="Ecuador" elevation="2163" location="Napo Prov." longitude="-0.59833336" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="9D29FD29697C8F7BFDFBFE3AFD0385E3" box="[631,673,474,501]" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
, “ 
<collectingCountry id="F3387630697C8F7BFD34FE3AFCE585E2" box="[696,839,474,500]" name="Ecuador" pageId="54" pageNumber="55">ECUADOR</collectingCountry>
: 
<collectingRegion id="49EBF842697C8F7BFCDCFE3BFC6C85E2" box="[848,974,474,501]" country="Ecuador" name="Napo" pageId="54" pageNumber="55">Napo Prov.</collectingRegion>
/ Yanayacu Biological Station, 
<geoCoordinate id="EE1B5067697C8F7BFAA4FE3AFEB6860F" degrees="00" direction="west" minutes="35.9" orientation="longitude" pageId="54" pageNumber="55" precision="92" value="-0.59833336">S 00°35.9′ W 77°53.4</geoCoordinate>
, 
<quantity id="4CD79B45697C8F7BFEADFE1EFECC860E" box="[289,366,510,536]" metricMagnitude="3" metricUnit="m" metricValue="2.163" pageId="54" pageNumber="55" unit="m" value="2163.0">
<elevation id="0002D193697C8F7BFEADFE1EFECC860E" box="[289,366,510,536]" metricMagnitude="3" metricUnit="m" metricValue="2.163" pageId="54" pageNumber="55" unit="m" value="2163.0">2163m</elevation>
</quantity>
/ REARED [in red]/ 
<date id="FF911060697C8F7BFDD7FE1FFD6F860F" box="[603,717,510,537]" pageId="54" pageNumber="55" value="2005-06">
<collectingDate id="EFD5E988697C8F7BFDD7FE1FFD6F860F" box="[603,717,510,537]" pageId="54" pageNumber="55" value="2005-06">June 2005</collectingDate>
</date>
/ 4003”, “ 
<taxonomicName id="4C2F4D23697C8F7BFCB2FE1EFBA3860E" box="[830,1025,510,536]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="undefined-8">
<emphasis id="B95BEAB2697C8F7BFCB2FE1EFBA3860E" box="[830,1025,510,536]" italics="true" pageId="54" pageNumber="55">Eucelatoria sp. 8</emphasis>
</taxonomicName>
”, “ZLB_E.Ferox 00214” [puparium and genitalia respectively on card and in vial under specimen] (
<collectionCode id="ED3EAE65697C8F7BFCD1FDC2FC01862A" box="[861,931,546,572]" pageId="54" pageNumber="55">JOSC</collectionCode>
; to be deposited in 
<collectionCode id="ED3EAE65697C8F7BFBF7FDC2FB73862A" box="[1147,1233,546,572]" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" pageId="54" pageNumber="55">MECN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9036A0697C8F7BFF4BFDA6FCA886E6" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
<emphasis id="B95BEAB2697C8F7BFF4BFDA6FEFF8676" bold="true" box="[199,349,582,608]" pageId="54" pageNumber="55">Recognition.</emphasis>
This species is closest to 
<taxonomicName id="4C2F4D23697C8F7BFD01FDA7FD598676" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[653,763,582,608]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="hafelei" status="sp. nov.">
<emphasis id="B95BEAB2697C8F7BFD01FDA7FD598676" box="[653,763,582,608]" italics="true" pageId="54" pageNumber="55">E. hafelei</emphasis>
</taxonomicName>
<emphasis id="B95BEAB2697C8F7BFC88FDA6FCC28676" bold="true" box="[772,864,582,608]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="A26857C9697C8F7BFC88FDA6FCC28676" box="[772,864,582,608]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; in both species the palpi are dark, the presutural thoracic vittae are fused into two, and the scutellum bears visible tomentum only at the extreme apex. Males of 
<taxonomicName id="4C2F4D23697C8F7BFA08FD8BFE8786BE" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="ritavargasae" status="sp. nov.">
<emphasis id="B95BEAB2697C8F7BFA08FD8BFE8786BE" italics="true" pageId="54" pageNumber="55">E. ritavargasae</emphasis>
</taxonomicName>
<emphasis id="B95BEAB2697C8F7BFEA3FD6EFE2886BE" bold="true" box="[303,394,654,680]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="A26857C9697C8F7BFEA3FD6EFE2886BE" box="[303,394,654,680]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are distinguished from 
<taxonomicName id="4C2F4D23697C8F7BFD12FD6FFCA986BE" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[670,779,654,680]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="hafelei" status="sp. nov.">
<emphasis id="B95BEAB2697C8F7BFD12FD6FFCA986BE" box="[670,779,654,680]" italics="true" pageId="54" pageNumber="55">E. hafelei</emphasis>
</taxonomicName>
<emphasis id="B95BEAB2697C8F7BFC99FD6EFCD286BE" bold="true" box="[789,880,654,680]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="A26857C9697C8F7BFC99FD6EFCD286BE" box="[789,880,654,680]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the narrower, more diffuse dorsal abdominal bands, the more strongly dilated cercus apex in lateral view, the more widely separated basal lobes of the cercus, and the strongly angled, deeply emarginated postgonite.
</paragraph>
</subSubSection>
<subSubSection id="C335652B697C8F7BFF4BFD1AFEC18796" pageId="54" pageNumber="55" type="etymology">
<paragraph id="8B9036A0697C8F7BFF4BFD1AFEC18796" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">
<emphasis id="B95BEAB2697C8F7BFF4BFD1AFEEC8702" bold="true" box="[199,334,762,788]" pageId="54" pageNumber="55">Etymology.</emphasis>
By recommendation of Dr. Daniel Janzen, this species “is named in recognition of Dr. Rita Vargas Castillo for her full support of the new collaborations between Costa Rica’s Area de Conservación 
<collectingRegion id="49EBF842697C8F7BFB60FCFEFACF872E" box="[1260,1389,798,824]" country="Costa Rica" name="Guanacaste" pageId="54" pageNumber="55">Guanacaste</collectingRegion>
and the Universidad de 
<collectingCountry id="F3387630697C8F7BFEF8FCA2FE50874A" box="[372,498,834,860]" name="Costa Rica" pageId="54" pageNumber="55">Costa Rica</collectingCountry>
, and quite specifically, supporting ACG crustacean taxonomy and the ACG marine parataxonomists”.
</paragraph>
</subSubSection>
<subSubSection id="C335652B697C8F7AFF4BFC6AFB72860F" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="description">
<paragraph id="8B9036A0697C8F7BFF4BFC6AFB2687B3" blockId="54.[151,1437,150,2013]" box="[199,1156,906,933]" pageId="54" pageNumber="55">
<emphasis id="B95BEAB2697C8F7BFF4BFC6AFEFB87B2" bold="true" box="[199,345,906,932]" pageId="54" pageNumber="55">Description.</emphasis>
[Described from 
<specimenCount id="9D29FD29697C8F7BFD93FC6AFDEB87B3" box="[543,585,906,933]" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
and 
<specimenCount id="9D29FD29697C8F7BFDF3FC6AFD6187B3" box="[639,707,906,933]" pageId="54" pageNumber="55" type="male">3 ♂♂</specimenCount>
.] Length 6.0–7.0 mm (mean = 
<quantity id="4CD79B45697C8F7BFB91FC6AFBD587B3" box="[1053,1143,906,933]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="54" pageNumber="55" unit="mm" value="6.5">6.5 mm</quantity>
).
</paragraph>
<paragraph id="8B9036A0697C8F7BFF4BFC4EFB4D818A" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">Head. Parafacial, fronto-orbital plate, vertex, dorsal half of post-ocular plate, and vibrissal angle tomentum dense yellow, appearing gold. Genal dilation, postgena, ventral half of post-ocular plate, and occiput tomentum ash-gray. Pale occipital setae ash-gray, in female merging to yellow on upper half. Ocellar setae strong, two-thirds to subequal to length of posterior reclinate orbital seta, stronger in female. Gena with 3–4 setae, subvibrissal ridge with 1 seta. Facial ridge with setulae on lower one-third to one-half. Postocellar setae one-half to two-thirds length of ocellar setae. Paravertical seta one-half to three-fourths length of postocellar setae. Outer vertical seta undifferentiated in male, one-half length of inner vertical seta in female. Reclinate orbital setae 3. Frontal setae 4–7, less in female, last frontal seta level with postpedicel base to arista base. Eye with short setulae, each setula no more than three facets in length. Eye height to head height ratio 0.85–0.87 (mean = 0.85). Postpedicel length 0.38–0.45 (mean = 0.42) height of head. Facial ridge length 0.49–0.59 (mean = 0.54) height of head. Parafacial width 0.06–0.10 (mean = 0.08) lateral length of head. Pedicel 0.26–0.33 (mean = 0.32) length of postpedicel. Postpedicel 2.0–3.7 (mean = 2.6) width of parafacial in lateral view. Vertex 0.24–0.26 (mean = 0.25) width of head in dorsal view. Palpus dark brown to black, cylindrical in male, greatly dilated and flattened at apex in female; with small, short setae dorsoapically, less in female; several stout setae apicoventrally; 3–4 long, thin setae mediolaterally.</paragraph>
<paragraph id="8B9036A0697C8F7BFF4BFA46FAFC8312" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">Thorax. Dorsomedial length 1.3–1.4 times width of thorax. Lateral tomentum gray, merging to yellow on anepisternum. Dorsal tomentum yellow to gray, usually at least yellow-tinged gray. Presutural inner and outer vitta fused into subquadrate vitta, thus with only two vittae on presutural area, at most with a small triangular area of tomentum dividing inner and outer vitta them near notopleural suture; the two vittae at least partially fused near anterior margin, rarely completely fused, leaving at most a small medial triangle of tomentum near notopleural suture. Postsutural vittae fused into large, black vittae covering nearly the entirety of the postsutural area, with at most a small area of tomentum near supra-alar setae and along posterior margin. Inner thoracic vittae. Scutellar dorsal tomentum diffuse, in dorsal view apparently confined to apical area between subapical scutellar setae. Postpronotum with 3 setae. Presutural area with 2 supra-alar setae. Postsutural area with 3–4 dorsocentral setae. Fore tibia with 1–3 posterodorsal setae. Wing hyaline, at most lightly infuscated around costa and radial sector. Calypters tan to cream colored.</paragraph>
<paragraph id="8B9036A0697C8F7BFF4BF8EEFD728383" blockId="54.[151,1437,150,2013]" pageId="54" pageNumber="55">Abdomen. Cuticle and setulae entirely black. Dorsal tomentum bands gray to yellow-gray; one-fourth to onehalf length of T3–5, narrower in males. Ventral tomentum bands ash-gray; in males one-fourth to one-half length of T3–5; in female covering three-fourths to entire length of T3–5. T4 with 1 pair of median marginal setae and 3 or more pairs of widely spaced lateral marginal setae.</paragraph>
<paragraph id="8B9036A0697C8F7AFF4BF87EFB72859F" blockId="54.[151,1437,150,2013]" lastBlockId="55.[151,1437,151,1041]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
Male terminalia (
<figureCitation id="13142A25697C8F7BFE02F87EFD9283AF" box="[398,560,1950,1977]" captionStart="FIGURES 136–155" captionStartId="100.[151,264,1677,1701]" captionTargetBox="[277,1319,184,1650]" captionTargetId="figure-31@100.[264,1324,181,1653]" captionTargetPageId="100" captionText="FIGURES 136–155. Male terminalia of Eucelatoria ferox group spp., left lateral view, and caudal view of cerci and surstyli. Numbers are “ZLB_E.Ferox” specimen ID numbers. Specimen information for caudal views is identical to that for Figs 124– 139. 136. E. gustavogutierrezi sp. nov., lateral view, Guanacaste Province (00008) (CNC). 137. E. hafelei sp. nov., lateral view, Chapare Province (00386) (DMW). 138. E. ritavargasae sp. nov., lateral view, Narupa (00215) (USNM). 139. E. sica sp. nov., lateral view, Yanayacu (00213) (CNC). 140. E. aurata (Townsend), caudal view. 141. E. auriceps (Aldrich), caudal view. 142. E. borealis sp. nov., caudal view. 143. E. charapensis (Townsend), caudal view. 144. E. falcata sp. nov., caudal view. 145. E. fordlandia sp. nov., caudal view. 146. E. gladiatrix (Townsend), caudal view. 147. E. sabroskyi sp. nov., caudal view. 148. E. strigata (Wulp), caudal view. 149. E. texana (Reinhard), caudal view. 150. E. woodorum sp. nov., caudal view. 151. E. yanayacu sp. nov., caudal view. 152. E. gustavogutierrezi sp. nov., caudal view. 153. E. hafelei sp. nov., caudal view. 154. E. ritavargasae sp. nov., caudal view. 155. E. sica sp. nov., caudal view." figureDoi="http://doi.org/10.5281/zenodo.6601508" httpUri="https://zenodo.org/record/6601508/files/figure.png" pageId="54" pageNumber="55">Figs 138, 154</figureCitation>
, 
<figureCitation id="13142A25697C8F7BFDB3F87EFDCE83AE" box="[575,620,1950,1976]" captionStart="FIGURES 156–173" captionStartId="101.[151,264,1755,1779]" captionTargetBox="[261,1320,195,1718]" captionTargetId="figure-15@101.[253,1335,181,1731]" captionTargetPageId="101" captionText="FIGURES 156–173. Male terminalia and fifth abdominal sternite of Eucelatoria ferox group spp. Specimen information for fifth sternites is identical to that for Figs 124–139. 156. Generic male terminalia, left lateral view (left) and caudal view (right), setae and membrane omitted.Abbreviations: aed, aedeagal apodeme; bas, basal lobe of surstylus; C, depth of postgonite anterior emargination; cer, cercus; epa, epandrium; hyp, hypandrium; pha, phallus; pos, postgonite; pre, pregonite; sur, surstylus. 157. E. aurata (Townsend). 158. E. auriceps (Aldrich). 159. E. borealis sp. nov. 160. E. charapensis (Townsend). 161. E. falcata sp. nov. 162. E. fordlandia sp. nov. 163. E. gladiatrix (Townsend). 164. E. sabroskyi sp. nov. 165. E. strigata (Wulp). 166. E. texana (Reinhard). 167. E. woodorum sp. nov. 168. E. yanayacu sp. nov. 169. E. gustavogutierrezi sp. nov. 170. E. hafelei sp. nov. 171. E. ritavargasae sp. nov. 172. E. sica sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601512" httpUri="https://zenodo.org/record/6601512/files/figure.png" pageId="54" pageNumber="55">171</figureCitation>
). Sternite 5 basal plate with median teeth separated 0.7 width of basal notch; apical lobes 1.5 times length of basal plate. Postgonite right-angled on posterior margin, slightly dilated at base, then tapered to long digitiform apex; anterior emargination depth two times width of postgonite at mid length. Surstylus paddle-shaped, length more than two and one-half times width; gradually curved on posterior and anterior margins; apex blunt; basal lobe subquadrate with distinct marginal notch. Cercus in lateral view dilated at base, narrowed to mid length, then dilated and slightly upturned as subquadrate apex; in caudal view curved from basal lobe, only slightly tapered to apex, subparallel along nearly entire length, apex blunt. Upper lobe subquadrate, nearly digitiform, with margins broadly rounded; 0.25 length of cercus. Median section 0.5 length of cercus. Apical cleft narrow, 0.25 length of cercus. Syncercus apex width in caudal view 0.4 width of syncercus base.
</paragraph>
<paragraph id="8B9036A0697D8F7AFF4BFE73FB72860F" blockId="55.[151,1437,151,1041]" pageId="55" pageNumber="56">Female terminalia. Piercer generally nearly to base of T4, in lateral view ribbon-like, sharply bent near base, then straight to slightly bent apex; in posterior view wide, parallel sided on basal third, then gradually tapered to apex. Aculeate lobe at least 2 times height of segment seven base. End tergite not visible. Cercus with 3–4 setae; ventral elongation length subequal to width of cercus main body. Postgenital plate with 10 setae.</paragraph>
</subSubSection>
<subSubSection id="C335652B697D8F7AFF4BFDC3FC6486E4" pageId="55" pageNumber="56" type="biology_ecology">
<paragraph id="8B9036A0697D8F7AFF4BFDC3FB1286BF" blockId="55.[151,1437,151,1041]" pageId="55" pageNumber="56">
<emphasis id="B95BEAB2697D8F7AFF4BFDC3FEB3862B" bold="true" box="[199,273,547,573]" pageId="55" pageNumber="56">Hosts.</emphasis>
Two individuals (“Janzen11”) have been reared from an 
<taxonomicName id="4C2F4D23697D8F7AFC13FDC3FB7D8628" box="[927,1247,547,574]" class="Insecta" family="Tortricidae" genus="Olethreutes" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95BEAB2697D8F7AFC13FDC3FB83862B" box="[927,1057,547,573]" italics="true" pageId="55" pageNumber="56">Olethreutes</emphasis>
sp. (Tortricidae)
</taxonomicName>
feeding on 
<taxonomicName id="4C2F4D23697D8F7AFAE5FDC3FE528674" authority="Benth." authorityName="Benth." class="Magnoliopsida" family="Fabaceae" genus="Inga" kingdom="Plantae" order="Fabales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="oerstediana">
<emphasis id="B95BEAB2697D8F7AFAE5FDC3FEB98677" italics="true" pageId="55" pageNumber="56">Inga oerstediana</emphasis>
Benth. (Fabaceae)
</taxonomicName>
in 
<collectingCountry id="F3387630697D8F7AFD98FDA7FD2C8677" box="[532,654,583,609]" name="Costa Rica" pageId="55" pageNumber="56">Costa Rica</collectingCountry>
(
<bibRefCitation id="EFBE4B51697D8F7AFD12FDA7FC698677" author="Janzen, D. H. &amp; Hallwachs, W." box="[670,971,583,609]" pageId="55" pageNumber="56" refId="ref67919" refString="Janzen, D. H. &amp; Hallwachs, W. (2008) Event-based database of caterpillars, their food plants, and their parasitoids in Area de Conservacion Guanacaste, Northwestern Costa Rica. Available from: http: // janzen. sas. upenn. edu (accessed 11 February 2015)" type="url" year="2008">Janzen &amp; Hallwachs 2008</bibRefCitation>
). One individual has been reared from an unknown 
<taxonomicName id="4C2F4D23697D8F7AFE8BFD8BFE238693" box="[263,385,619,645]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="family">Tortricidae</taxonomicName>
feeding on an unknown species of 
<taxonomicName id="4C2F4D23697D8F7AFC94FD8BFC638693" box="[792,961,619,645]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
, and another has been reared from an unknown “ 
<taxonomicName id="4C2F4D23697D8F7AFF78FD6FFEC186BF" box="[244,355,655,681]" class="Insecta" family="Pyralidae" kingdom="Animalia" order="Lepidoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="family">Pyralidae</taxonomicName>
” on 
<taxonomicName id="4C2F4D23697D8F7AFE1BFD6FFE4786BF" box="[407,485,655,681]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="genus">
<emphasis id="B95BEAB2697D8F7AFE1BFD6FFE4786BF" box="[407,485,655,681]" italics="true" pageId="55" pageNumber="56">Croton</emphasis>
</taxonomicName>
L. sp. (Euphorbiaecae), both in 
<collectingCountry id="F3387630697D8F7AFCC3FD6FFC0E86BF" box="[847,940,655,681]" name="Ecuador" pageId="55" pageNumber="56">Ecuador</collectingCountry>
(
<bibRefCitation id="EFBE4B51697D8F7AFC37FD6FFB0786BF" author="Stireman, J. O. III &amp; Dyer, L. A. &amp; Greeney, H. F." box="[955,1189,655,681]" pageId="55" pageNumber="56" pagination="367 - 384" refId="ref68830" refString="Stireman, J. O. III, Dyer, L. A. &amp; Greeney, H. F. (2017) Specialised generalists? Food web structure of a tropical tachinid-caterpillar community. Insect Conservation and Diversity, 10 (5), 367 - 384. https: // doi. org / 10.1111 / icad. 12238" type="journal article" year="2017">
Stireman 
<emphasis id="B95BEAB2697D8F7AFBA5FD6FFBC386BF" box="[1065,1121,655,681]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9036A0697D8F7AFF4BFD52FC6486E4" blockId="55.[151,1437,151,1041]" pageId="55" pageNumber="56">
<emphasis id="B95BEAB2697D8F7AFF4BFD52FD7A86DB" bold="true" box="[199,728,690,717]" pageId="55" pageNumber="56">Geographic extent and seasonal occurrence.</emphasis>
Individuals of 
<taxonomicName id="4C2F4D23697D8F7AFC0AFD53FB9686DA" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[902,1076,691,716]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="ritavargasae" status="sp. nov.">
<emphasis id="B95BEAB2697D8F7AFC0AFD53FB9686DA" box="[902,1076,691,716]" italics="true" pageId="55" pageNumber="56">E. ritavargasae</emphasis>
</taxonomicName>
<emphasis id="B95BEAB2697D8F7AFBB0FD53FB3586DB" bold="true" box="[1084,1175,691,717]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A26857C9697D8F7AFBB0FD53FB3586DB" box="[1084,1175,691,717]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have been collected in tropical and subtropical moist forest at 
<quantity id="4CD79B45697D8F7AFDC2FD36FD7986E7" box="[590,731,726,753]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="55" pageNumber="56" unit="m" value="850.0" valueMax="1200.0" valueMin="500.0">500–1200 m</quantity>
elevation (
<figureCitation id="13142A25697D8F7AFCD4FD37FC1886E4" box="[856,954,727,754]" captionStart="FIGURES 174–179" captionStartId="103.[151,264,1856,1880]" captionTargetBox="[203,1374,189,1818]" captionTargetId="figure-14@103.[193,1393,181,1832]" captionTargetPageId="103" captionText="FIGURES 174–179. Distribution maps of Eucelatoria spp. 174. E. yanayacu sp. nov., E. gladiatrix (Townsend), E. falcata sp. nov., E. charapensis (Townsend), and E. fordlandia sp. nov. 175. E. strigata (Wulp) and E. aurata (Townsend). 176. E. hafelei sp. nov., E. gustavogutierrezi sp. nov., and E. ritavargasae sp. nov. 177. E. ferox (Townsend), E. huitepecensis sp. nov., and E. inclani sp. nov. 178. E. kopis sp. nov., E. sica sp. nov., E. makhaira sp. nov., and E. tenebrionis sp. nov. 179. E. rivalis (Reinhard), E. luctuosa (Wulp), E. woodorum sp. nov., E. crambivora sp. nov., and E. jorgecortesi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601514" httpUri="https://zenodo.org/record/6601514/files/figure.png" pageId="55" pageNumber="56">Fig. 176</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C335652B697D8F7AFF4BFD1AFC308007" pageId="55" pageNumber="56" type="discussion">
<paragraph id="8B9036A0697D8F7AFF4BFD1AFC308007" blockId="55.[151,1437,151,1041]" pageId="55" pageNumber="56">
<emphasis id="B95BEAB2697D8F7AFF4BFD1AFEEE8702" bold="true" box="[199,332,762,788]" pageId="55" pageNumber="56">Discussion.</emphasis>
This species includes “ 
<taxonomicName id="4C2F4D23697D8F7AFDC1FD1BFC848703" authority="Janzen" authorityName="Janzen" box="[589,806,763,789]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">Eucelatoria Janzen</taxonomicName>
11” in the ACG caterpillar database (
<bibRefCitation id="EFBE4B51697D8F7AFB3BFD1BFF76872F" author="Janzen, D. H. &amp; Hallwachs, W." pageId="55" pageNumber="56" refId="ref67919" refString="Janzen, D. H. &amp; Hallwachs, W. (2008) Event-based database of caterpillars, their food plants, and their parasitoids in Area de Conservacion Guanacaste, Northwestern Costa Rica. Available from: http: // janzen. sas. upenn. edu (accessed 11 February 2015)" type="url" year="2008">Janzen &amp; Hallwachs 2008</bibRefCitation>
). The characteristic confinement of dorsal scutellar tomentum to the extreme apex and two presutural thoracic vittae will separate both males and females from all known 
<taxonomicName id="4C2F4D23697D8F7AFCBCFCA3FC2A874B" box="[816,904,835,861]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="ferox">
<emphasis id="B95BEAB2697D8F7AFCBCFCA3FC2A874B" box="[816,904,835,861]" italics="true" pageId="55" pageNumber="56">E. ferox</emphasis>
</taxonomicName>
group species except 
<taxonomicName id="4C2F4D23697D8F7AFBF2FCA3FB4A874B" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[1150,1256,835,861]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="hafelei" status="sp. nov.">
<emphasis id="B95BEAB2697D8F7AFBF2FCA3FB4A874B" box="[1150,1256,835,861]" italics="true" pageId="55" pageNumber="56">E. hafelei</emphasis>
</taxonomicName>
<emphasis id="B95BEAB2697D8F7AFB63FCA3FAEA874B" bold="true" box="[1263,1352,835,861]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A26857C9697D8F7AFB63FCA3FAEA874B" box="[1263,1352,835,861]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Generally, the widths of the abdominal bands are adequate to separate the two species in males; it is unknown if females are similar in this regard. The single male from Nova Teutonia is included here despite the wide geographic distance, pending more material. Genetic data suggest some population structure between Ecuadorian and 
<collectingRegion id="49EBF842697D8F7AFB54FC4FFABA87DF" box="[1240,1304,943,969]" country="New Zealand" name="West Coast" pageId="55" pageNumber="56">Costa</collectingRegion>
Rican individuals, but this is only from a few sequences (
<figureCitation id="13142A25697D8F7AFD2DFC33FD4087FB" box="[673,738,979,1005]" captionStart="FIGURE 1" captionStartId="9.[151,250,1766,1790]" captionTargetBox="[164,1421,431,1729]" captionTargetId="figure-49@9.[153,1434,417,1743]" captionTargetPageId="9" captionText="FIGURE 1. CO1-based neighbor-joining tree of 12 species of the Eucelatoria ferox species group, with inset illustrations of mesonota. The outgroup is Eucelatoria armigera (Coquillett). Node labels indicate support values over 60. Scale bar = 2% genetic difference. The E. charapensis (Townsend) sequence with the axial break is of poor quality but is included to show unity of El Bigal specimens with the rest of the metapopulation. Mesonota are illustrated with setal annuli only for clarity. A. E. gustavogutierrezi sp. nov. B. E. kopis sp. nov. C. E. ritavargasae sp. nov. D. E. ferox (Townsend). E. E. huitepecensis sp. nov. F. E. auriceps (Aldrich). G. E. gladiatrix (Townsend). H. E. borealis sp. nov. I. E. yanayacu sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601406" httpUri="https://zenodo.org/record/6601406/files/figure.png" pageId="55" pageNumber="56">Fig. 1</figureCitation>
). The phylogenetic analysis strongly supports a close relationship between 
<taxonomicName id="4C2F4D23697D8F7AFEBDFC17FE298007" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[305,395,1015,1041]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="kopis" status="sp. nov.">
<emphasis id="B95BEAB2697D8F7AFEBDFC17FE298007" box="[305,395,1015,1041]" italics="true" pageId="55" pageNumber="56">E. kopis</emphasis>
</taxonomicName>
<emphasis id="B95BEAB2697D8F7AFE1EFC17FE4E8007" bold="true" box="[402,492,1015,1041]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A26857C9697D8F7AFE1EFC17FE4E8007" box="[402,492,1015,1041]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C2F4D23697D8F7AFDAEFC17FD6D8006" authority="Burington, 2022" authorityName="Burington" authorityYear="2022" box="[546,719,1015,1040]" class="Insecta" family="Tachinidae" genus="Eucelatoria" kingdom="Animalia" order="Diptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="ritavargasae" status="sp. nov.">
<emphasis id="B95BEAB2697D8F7AFDAEFC17FD6D8006" box="[546,719,1015,1040]" italics="true" pageId="55" pageNumber="56">E. ritavargasae</emphasis>
</taxonomicName>
<emphasis id="B95BEAB2697D8F7AFD5AFC17FC928007" bold="true" box="[726,816,1015,1041]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A26857C9697D8F7AFD5AFC17FC928007" box="[726,816,1015,1041]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13142A25697D8F7AFCB3FC17FC238007" box="[831,897,1015,1041]" captionStart="FIGURE 1" captionStartId="9.[151,250,1766,1790]" captionTargetBox="[164,1421,431,1729]" captionTargetId="figure-49@9.[153,1434,417,1743]" captionTargetPageId="9" captionText="FIGURE 1. CO1-based neighbor-joining tree of 12 species of the Eucelatoria ferox species group, with inset illustrations of mesonota. The outgroup is Eucelatoria armigera (Coquillett). Node labels indicate support values over 60. Scale bar = 2% genetic difference. The E. charapensis (Townsend) sequence with the axial break is of poor quality but is included to show unity of El Bigal specimens with the rest of the metapopulation. Mesonota are illustrated with setal annuli only for clarity. A. E. gustavogutierrezi sp. nov. B. E. kopis sp. nov. C. E. ritavargasae sp. nov. D. E. ferox (Townsend). E. E. huitepecensis sp. nov. F. E. auriceps (Aldrich). G. E. gladiatrix (Townsend). H. E. borealis sp. nov. I. E. yanayacu sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6601406" httpUri="https://zenodo.org/record/6601406/files/figure.png" pageId="55" pageNumber="56">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>