treatments-xml/data/03/86/87/038687B2FFFC0633C4EE80E9FC0DF9C7.xml
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<document id="3E1A8F80E330D87D8D33D70A47855DD0" ID-DOI="10.5281/zenodo.5328783" ID-ISSN="0374-1036" ID-Zenodo-Dep="5328783" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630159881273" checkinUser="felipe" docAuthor="Fikáček, Martin, Barclay, Maxwell V. L., Perkins, Philip D. &amp; Sw, London" docDate="2011" docId="038687B2FFFC0633C4EE80E9FC0DF9C7" docLanguage="en" docName="ActaEntMusNatPra.2011.51.2.477-504.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 51 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Epimetopus mendeli Fikáček &amp; Barclay &amp; Perkins &amp; Sw 2011, sp. nov." docType="treatment" docVersion="5" lastPageNumber="487" masterDocId="FFBFFFCAFFFF0639C5558428FFABFFF4" masterDocTitle="Two new species of the Epimetopus mendeli species group and notes on its adult and larval morphology (Coleoptera: Hydrophiloidea: Epimetopidae)" masterLastPageNumber="504" masterPageNumber="477" pageNumber="480" updateTime="1699169192168" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C14B0E3FE0F956508D4C4E6B2CEFE3A8">Two new species of the Epimetopus mendeli species group and notes on its adult and larval morphology (Coleoptera: Hydrophiloidea: Epimetopidae)</mods:title>
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<mods:namePart id="487CDBE7F6DD8C1C8F431400F65D6111">Fikáček, Martin</mods:namePart>
<mods:affiliation id="14A520F70E7120FF35712AD0899A7490">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="026867977155744E6B3A6B20DAB55BC6">Barclay, Maxwell V. L.</mods:namePart>
<mods:affiliation id="8B6A1C2DB6C872A03160E651FE20AF5A">Department of Entomology, The Natural History Museum, London SW 7 5 BD, United Kingdom; e-mail: m. barclay @ nhm. ac. uk</mods:affiliation>
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<mods:namePart id="CCAD8C9B1C02649AACCEFA5E80552803">Perkins, Philip D.</mods:namePart>
<mods:affiliation id="CC9B06008FC4D984226CE192F2277BC7">Department of Entomology, Museum of Comparative Zoology, Harvard University, Cambridge, MA 02138, USA; e-mail: perkins @ oeb. harvard. edu</mods:affiliation>
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<mods:namePart id="51EDB6225900D9C27278F9966DEDD1E6">Sw, London</mods:namePart>
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<mods:title id="7A2DD28EE59C23DE5003A935D89D876E">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="E761D7A5750E141F317746B9AB02DADC">2011</mods:date>
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<taxonomicName id="4C2F4D27FFFC063AC4EE80E9FD30FB2D" ID-CoL="6G5XR" authority="Fikáček &amp; Barclay &amp; Perkins &amp; Sw, 2011" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[443,667,1216,1241]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="480" phylum="Arthropoda" rank="species" species="mendeli" status="sp. nov.">
<emphasis id="B95BEAB6FFFC063AC4EE80E9FD30FB2D" bold="true" box="[443,667,1216,1241]" italics="true" pageId="3" pageNumber="480">Epimetopus mendeli</emphasis>
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locality.
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,
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, Kosñipata Valley, Cock of the Rock Lodge,
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a.s.l.
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<emphasis id="B95BEAB6FFFC063AC524811BFF53FAB2" bold="true" box="[113,248,1330,1350]" pageId="3" pageNumber="480">Type material.</emphasis>
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):
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<collectorName id="26DA5372FFFC063AC73D8165FD62FA95" box="[616,713,1357,1377]" pageId="3" pageNumber="480">Mendel, H.</collectorName>
&amp;
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//
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{E} / 2009-126 / under stones/ by torrent
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.
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(
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,
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<collectionCode id="ED3EAE61FFFC063AC7A08140FC8EFA8F" box="[757,805,1384,1403]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="480" type="Museum">MCZ</collectionCode>
,
<collectionCode id="ED3EAE61FFFC063AC6658140FCDDFA8F" box="[816,886,1384,1403]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="480" type="Museum">NHMW</collectionCode>
,
<collectionCode id="ED3EAE61FFFC063AC6D58140FC7EFA8F" box="[896,981,1384,1403]" pageId="3" pageNumber="480">UNMSM</collectionCode>
): same label data as the holotype
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;
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<specimenCount id="9D29FD2DFFFC063AC47081ABFECAFA62" box="[293,353,1411,1430]" pageId="3" pageNumber="480" type="generic">9 spec.</specimenCount>
(
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,
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,
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):
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,
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/
<collectingCounty id="62F14E28FFFC063AC62781ABFBAFFA62" box="[882,1028,1411,1430]" pageId="3" pageNumber="480">Kosñipata Valley</collectingCounty>
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<collectingMunicipality id="6BF4ACDEFFFC063AC14181ABFBEAFA62" box="[1044,1089,1411,1430]" pageId="3" pageNumber="480">Cock</collectingMunicipality>
of the
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/
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</materialsCitation>
<materialsCitation id="3B473CF9FFFC063AC4F381B6FE6BFA3F" collectingDate="2009-11-20" collectingDateMax="2009-12-15" collectingDateMin="2009-11-20" collectionCode="BMNH" collectorName="Mendel &amp; Barclay, M. V. L." country="Peru" elevation="1500" latitude="-13.056056" pageId="3" pageNumber="480" specimenCount="1" stateProvince="Cusco" typeStatus="paratype">
;
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<collectingDate id="EFD5E98CFFFC063AC76F81B5FD72FA44" box="[570,729,1437,1456]" pageId="3" pageNumber="480" value="2009-11-20" valueMax="2009-12-15" valueMin="2009-11-20">20.xi.15.xii.2009</collectingDate>
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,
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/
<collectorName id="26DA5372FFFC063AC66181B5FCDCFA44" box="[820,887,1437,1456]" pageId="3" pageNumber="480">Mendel</collectorName>
, H. &amp;
<collectorName id="26DA5372FFFC063AC6E381B5FBE9FA45" box="[950,1090,1437,1457]" pageId="3" pageNumber="480">Barclay, M.V.L.</collectorName>
//
<collectionCode id="ED3EAE61FFFC063AC5D18190FF60FA3F" box="[132,203,1464,1483]" pageId="3" pageNumber="480">BMNH</collectionCode>
{E} / 2009-126 / light sheet
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;
<materialsCitation id="3B473CF9FFFC063AC49F8190FC4AFA12" collectingDate="2000-10-25" collectionCode="KSEM" country="Peru" county="Panliacolla Lodge" elevation="400" latitude="-12.656111" location="Alto" longLatPrecision="21" longitude="-71.23194" municipality="Monk Saki Trail" pageId="3" pageNumber="480" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
1 J (
<collectionCode id="ED3EAE61FFFC063AC4AD8190FD92FA3F" box="[504,569,1464,1483]" pageId="3" pageNumber="480">KSEM</collectionCode>
):
<collectingCountry id="F3387634FFFC063AC71E8190FD22FA3F" box="[587,649,1464,1483]" name="Peru" pageId="3" pageNumber="480">PERU</collectingCountry>
:
<collectingRegion id="49EBF846FFFC063AC7C78190FCB9FA3F" box="[658,786,1464,1483]" country="Peru" name="Madre de Dios" pageId="3" pageNumber="480">Madre de Dios</collectingRegion>
/
<collectingCounty id="62F14E28FFFC063AC6738190FC69FA3F" box="[806,962,1464,1483]" pageId="3" pageNumber="480">Panliacolla Lodge</collectingCounty>
[= Pantiacolla Lodge],
<collectingMunicipality id="6BF4ACDEFFFC063AC5EF81FBFEEDFA12" box="[186,326,1491,1510]" pageId="3" pageNumber="480">Monk Saki Trail</collectingMunicipality>
/
<location id="8EF0607FFFFC063AC40381FBFED7FA12" LSID="urn:lsid:plazi:treatment:038687B2FFFC0633C4EE80E9FC0DF9C7:8EF0607FFFFC063AC40381FBFED7FA12" box="[342,380,1491,1510]" country="Peru" county="Panliacolla Lodge" latitude="-12.656111" longLatPrecision="21" longitude="-71.23194" municipality="Monk Saki Trail" name="Alto" pageId="3" pageNumber="480" stateProvince="Madre de Dios">Alto</location>
<collectingRegion id="49EBF846FFFC063AC4D481FBFDABFA12" box="[385,512,1491,1510]" country="Peru" name="Madre de Dios" pageId="3" pageNumber="480">Madre de Dios</collectingRegion>
<location id="8EF0607FFFFC063AC75381FBFD9CFA12" LSID="urn:lsid:plazi:treatment:038687B2FFFC0633C4EE80E9FC0DF9C7:8EF0607FFFFC063AC75381FBFD9CFA12" box="[518,567,1491,1510]" country="Peru" county="Panliacolla Lodge" latitude="-12.656111" longLatPrecision="21" longitude="-71.23194" municipality="Monk Saki Trail" name="River" pageId="3" pageNumber="480" stateProvince="Madre de Dios">River</location>
,
<quantity id="4CD79B41FFFC063AC71581FBFDDAFA12" box="[576,625,1491,1510]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="480" unit="m" value="400.0">
<elevation id="0002D197FFFC063AC71581FBFDDAFA12" box="[576,625,1491,1510]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="480" unit="m" value="400.0">400m</elevation>
</quantity>
/
<geoCoordinate id="EE1B5063FFFC063AC7D481FBFD4FFA12" box="[641,740,1490,1510]" degrees="12" direction="south" minutes="39" orientation="latitude" pageId="3" pageNumber="480" precision="15" seconds="22" value="-12.656111">12°3922″S</geoCoordinate>
,
<geoCoordinate id="EE1B5063FFFC063AC7BB81FBFCF3FA12" box="[750,856,1490,1510]" degrees="71" direction="west" minutes="13" orientation="longitude" pageId="3" pageNumber="480" precision="15" seconds="55" value="-71.23194">71°1355″W</geoCoordinate>
/
<date id="FF911064FFFC063AC63C81FBFC4AFA12" box="[873,993,1491,1510]" pageId="3" pageNumber="480" value="2000-10-25">
<collectingDate id="EFD5E98CFFFC063AC63C81FBFC4AFA12" box="[873,993,1491,1510]" pageId="3" pageNumber="480" value="2000-10-25">25 OCT 2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B473CF9FFFC063AC6BF81FBFE12F9F4" country="Peru" location="Brooks" pageId="3" pageNumber="480" specimenCount="1" typeStatus="paratype">
R.
<location id="8EF0607FFFFC063AC15681FBFBEAFA12" LSID="urn:lsid:plazi:treatment:038687B2FFFC0633C4EE80E9FC0DF9C7:8EF0607FFFFC063AC15681FBFBEAFA12" box="[1027,1089,1491,1510]" country="Peru" name="Brooks" pageId="3" pageNumber="480">Brooks</location>
/
<collectingCountry id="F3387634FFFC063AC52B81C6FF12F9F5" box="[126,185,1518,1537]" name="Peru" pageId="3" pageNumber="480">PERU</collectingCountry>
1B00 098A ex: at black light
</materialsCitation>
;
<materialsCitation id="3B473CF9FFFC063AC49781C6FF13F9EF" collectingDate="1979-01-28" collectionCode="USNM" collectorName="W. E. Steiner'" country="Peru" location="Quince Mil" pageId="3" pageNumber="480" specimenCount="3" specimenCount-female="3" stateProvince="Cusco" typeStatus="paratype">
4 JJ
<specimenCount id="9D29FD2DFFFC063AC4AC81C6FD81F9F4" box="[505,554,1511,1537]" pageId="3" pageNumber="480" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="ED3EAE61FFFC063AC76081C5FDD3F9F4" box="[565,632,1517,1536]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="480" type="Museum">USNM</collectionCode>
):
<collectingCountry id="F3387634FFFC063AC7DE81C6FD62F9F5" box="[651,713,1518,1537]" name="Peru" pageId="3" pageNumber="480">PERU</collectingCountry>
:
<collectingRegion id="49EBF846FFFC063AC78481C6FCA3F9F5" box="[721,776,1518,1537]" country="Peru" name="Cusco" pageId="3" pageNumber="480">Cuzco</collectingRegion>
/
<location id="8EF0607FFFFC063AC64F81C5FCD6F9F4" LSID="urn:lsid:plazi:treatment:038687B2FFFC0633C4EE80E9FC0DF9C7:8EF0607FFFFC063AC64F81C5FCD6F9F4" box="[794,893,1517,1536]" country="Peru" name="Quince Mil" pageId="3" pageNumber="480" stateProvince="Cusco">Quince Mil</location>
/
<date id="FF911064FFFC063AC6C581C6FC54F9F5" box="[912,1023,1518,1537]" pageId="3" pageNumber="480" value="1979-01-28">
<collectingDate id="EFD5E98CFFFC063AC6C581C6FC54F9F5" box="[912,1023,1518,1537]" pageId="3" pageNumber="480" value="1979-01-28">28 Jan. 1979</collectingDate>
</date>
/
<collectorName id="26DA5372FFFC063AC14581C6FF13F9EF" pageId="3" pageNumber="480">W. E. Steiner</collectorName>
</materialsCitation>
;
<materialsCitation id="3B473CF9FFFC063AC5948220FEDFF9C2" collectionCode="USNM" country="Peru" location="Fikacek" pageId="3" pageNumber="480" specimenCount="1" specimenCount-female="1" stateProvince="Cusco" typeStatus="paratype">
<specimenCount id="9D29FD2DFFFC063AC5948220FF4BF9EF" box="[193,224,1538,1563]" pageId="3" pageNumber="480" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED3EAE61FFFC063AC5BE8220FE85F9EF" box="[235,302,1544,1563]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="480" type="Museum">USNM</collectionCode>
): same label data plus larvae from the egg case / carried by this specimen / used for description by /
<location id="8EF0607FFFFC063AC5EB820BFEAAF9C2" LSID="urn:lsid:plazi:treatment:038687B2FFFC0633C4EE80E9FC0DF9C7:8EF0607FFFFC063AC5EB820BFEAAF9C2" box="[190,257,1571,1590]" country="Peru" name="Fikacek" pageId="3" pageNumber="480" stateProvince="Cusco">Fikáček</location>
et al. (2011)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9036A4FFFC063AC5248215FF17F99F" blockId="3.[113,1090,1261,1643]" pageId="3" pageNumber="480">
<emphasis id="B95BEAB6FFFC063AC5248215FE15F9A5" bold="true" box="[113,446,1597,1617]" pageId="3" pageNumber="480">Additional adult material examined.</emphasis>
<specimenCount id="9D29FD2DFFFC063AC4968216FE4FF9A4" box="[451,484,1591,1617]" pageId="3" pageNumber="480" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED3EAE61FFFC063AC4BA8215FD99F9A4" box="[495,562,1597,1616]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="480" type="Museum">USNM</collectionCode>
):
<collectingCountry id="F3387634FFFC063AC7118216FD29F9A5" box="[580,642,1598,1617]" name="Peru" pageId="3" pageNumber="480">PERU</collectingCountry>
:
<collectingRegion id="49EBF846FFFC063AC7DF8216FD6AF9A5" box="[650,705,1598,1617]" country="Peru" name="Cusco" pageId="3" pageNumber="480">Cuzco</collectingRegion>
/
<quantity id="4CD79B41FFFC063AC7878215FCA9F9A4" box="[722,770,1597,1616]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="480" unit="km" value="25.0">25km</quantity>
SW Pilcopata /
<date id="FF911064FFFC063AC6DA8215FBABF9A5" box="[911,1024,1597,1617]" pageId="3" pageNumber="480" value="1979-02-16">16 Feb. 1979</date>
/ W. E. Steiner.
</paragraph>
</subSubSection>
<caption id="DF50662CFFFB063DC5C2820AFC51F99E" ID-DOI="http://doi.org/10.5281/zenodo.5328787" ID-Zenodo-Dep="5328787" httpUri="https://zenodo.org/record/5328787/files/figure.png" pageId="4" pageNumber="481" startId="4.[151,190,1570,1589]" targetBox="[153,1096,241,1475]" targetPageId="4">
<paragraph id="8B9036A4FFFB063DC5C2820AFC51F99E" blockId="4.[151,1126,1570,1643]" pageId="4" pageNumber="481">
Figs. 15. General habitus and aedeagi of
<taxonomicName id="4C2F4D27FFFB063DC758820BFDD9F9C1" authorityName="Lacordaire" authorityYear="1854" box="[525,626,1571,1589]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEAB6FFFB063DC758820BFDD9F9C1" box="[525,626,1571,1589]" italics="true" pageId="4" pageNumber="481">Epimetopus</emphasis>
</taxonomicName>
. 12
<taxonomicName id="4C2F4D27FFFB063DC7ED820BFCBEF9C1" authority="Fikáček &amp; Barclay &amp; Perkins &amp; Sw, 2011" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[696,789,1571,1589]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="481" phylum="Arthropoda" rank="species" species="mendeli" status="sp. nov.">
<emphasis id="B95BEAB6FFFB063DC7ED820BFCBEF9C1" box="[696,789,1571,1589]" italics="true" pageId="4" pageNumber="481">E. mendeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857CDFFFB063DC649820AFCC8F9C1" box="[796,867,1570,1589]" pageId="4" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
, general habitus (body length 3.4 mm): 1 dorsal view; 2 lateral view. 35 aedeagus of the holotype: 3
<taxonomicName id="4C2F4D27FFFB063DC6798216FC2DF9A4" authority="Fikáček &amp; Barclay &amp; Perkins &amp; Sw, 2011" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[812,902,1598,1616]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="481" phylum="Arthropoda" rank="species" species="mendeli" status="sp. nov.">
<emphasis id="B95BEAB6FFFB063DC6798216FC2DF9A4" box="[812,902,1598,1616]" italics="true" pageId="4" pageNumber="481">E. mendeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857CDFFFB063DC6DF8215FC65F9A4" box="[906,974,1597,1616]" pageId="4" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
, 4
<taxonomicName id="4C2F4D27FFFB063DC6A68216FBCDF9A4" authority="Fikáček &amp; Barclay &amp; Perkins &amp; Sw, 2011" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[1011,1126,1598,1616]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="481" phylum="Arthropoda" rank="species" species="flavicaptus" status="sp. nov.">
<emphasis id="B95BEAB6FFFB063DC6A68216FBCDF9A4" box="[1011,1126,1598,1616]" italics="true" pageId="4" pageNumber="481">E. flavicaptus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857CDFFFB063DC5C28270FF77F99F" box="[151,220,1624,1643]" pageId="4" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
, 5
<taxonomicName id="4C2F4D27FFFB063DC4568270FD93F99E" authority="Balfour-Browne, 1949" authorityName="Balfour-Browne" authorityYear="1949" box="[259,568,1623,1642]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="481" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="B95BEAB6FFFB063DC4568270FEC4F99E" box="[259,367,1624,1642]" italics="true" pageId="4" pageNumber="481">E. angulatus</emphasis>
<bibRefCitation id="EFBE4B55FFFB063DC420827FFD93F99E" author="BALFOUR-BROWNE J." box="[373,568,1623,1642]" pageId="4" pageNumber="481" pagination="12 - 18" refId="ref12091" refString="BALFOUR-BROWNE J. 1949: Notes on the subfamily Epimetopinae (Col., Palpicornia) with the description of a new genus and of three new species. Proceedings of the Royal Entomological Society of London, Series B 18: 12 - 18." type="journal article" year="1949">Balfour-Browne, 1949</bibRefCitation>
</taxonomicName>
. Abbreviations:
<emphasis id="B95BEAB6FFFB063DC7968270FD75F99E" box="[707,734,1624,1642]" italics="true" pageId="4" pageNumber="481">lap</emphasis>
lateral process of median lobe.
</paragraph>
</caption>
<caption id="DF50662CFFFA063CC52481EAFBB7F9D1" ID-DOI="http://doi.org/10.5281/zenodo.5328789" ID-Zenodo-Dep="5328789" httpUri="https://zenodo.org/record/5328789/files/figure.png" pageId="5" pageNumber="482" startId="5.[113,152,1474,1493]" targetBox="[113,1087,220,1423]" targetPageId="5">
<paragraph id="8B9036A4FFFA063CC52481EAFBB7F9D1" blockId="5.[113,1089,1474,1574]" pageId="5" pageNumber="482">
Figs. 610. First instar larva of
<taxonomicName id="4C2F4D27FFFA063CC42B81EBFD87FA21" authority="Fikáček &amp; Barclay &amp; Perkins &amp; Sw, 2011" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[382,556,1475,1493]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="482" phylum="Arthropoda" rank="species" species="mendeli" status="sp. nov.">
<emphasis id="B95BEAB6FFFA063CC42B81EBFD87FA21" box="[382,556,1475,1493]" italics="true" pageId="5" pageNumber="482">Epimetopus mendeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857CDFFFA063CC76481EBFDDFFA22" box="[561,628,1475,1494]" pageId="5" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
6 general habitus in ventrolateral view; 7 detail of frontoclypeus; 8 head capsule in dorsal view; 9 abdominal apex in dorsal view; 10 abdominal apex in dorsolateral view.Abbreviations:
<emphasis id="B95BEAB6FFFA063CC41A81D1FE24F9FF" box="[335,399,1529,1547]" italics="true" pageId="5" pageNumber="482">abd79</emphasis>
abdominal segments 79;
<emphasis id="B95BEAB6FFFA063CC72A81D1FD01F9FF" box="[639,682,1529,1547]" italics="true" pageId="5" pageNumber="482">app8</emphasis>
lateral projection of the abdominal segment 8;
<emphasis id="B95BEAB6FFFA063CC524823BFF37F9D1" box="[113,156,1555,1573]" italics="true" pageId="5" pageNumber="482">app9</emphasis>
lateral projection of the abdominal segment 9;
<emphasis id="B95BEAB6FFFA063CC715823BFDE0F9D1" box="[576,587,1555,1573]" italics="true" pageId="5" pageNumber="482">fl</emphasis>
frontal line;
<emphasis id="B95BEAB6FFFA063CC79F823BFD76F9D1" box="[714,733,1555,1573]" italics="true" pageId="5" pageNumber="482">ur</emphasis>
urogomphus;
<emphasis id="B95BEAB6FFFA063CC63D823BFCD7F9D1" box="[872,892,1555,1573]" italics="true" pageId="5" pageNumber="482">vp</emphasis>
ventral papillae.
</paragraph>
</caption>
<caption id="DF50662CFFF9063FC5CD8035FBA5FB74" ID-DOI="http://doi.org/10.5281/zenodo.5328793" ID-Zenodo-Dep="5328793" httpUri="https://zenodo.org/record/5328793/files/figure.png" pageId="6" pageNumber="483" startId="6.[152,191,1053,1072]" targetBox="[173,1103,221,1015]" targetPageId="6">
<paragraph id="8B9036A4FFF9063FC5CD8035FBA5FB74" blockId="6.[151,1127,1053,1152]" pageId="6" pageNumber="483">
Figs. 1113. Morphological features of
<taxonomicName id="4C2F4D27FFF9063FC48A8036FDE8FBC4" authorityName="Lacordaire" authorityYear="1854" box="[479,579,1054,1072]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEAB6FFF9063FC48A8036FDE8FBC4" box="[479,579,1054,1072]" italics="true" pageId="6" pageNumber="483">Epimetopus</emphasis>
</taxonomicName>
. 1112 head in lateral view:11
<taxonomicName id="4C2F4D27FFF9063FC6318036FC16FBC4" authority="Fikáček &amp; Barclay &amp; Perkins &amp; Sw, 2011" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[868,957,1054,1072]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="mendeli" status="sp. nov.">
<emphasis id="B95BEAB6FFF9063FC6318036FC16FBC4" box="[868,957,1054,1072]" italics="true" pageId="6" pageNumber="483">E. mendeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857CDFFF9063FC6948035FBAFFBC4" box="[961,1028,1053,1072]" pageId="6" pageNumber="483" rank="species">sp.nov.</taxonomicNameLabel>
; 12
<taxonomicName id="4C2F4D27FFF9063FC1648036FEE0FBBE" authority="(Sharp, 1875)" baseAuthorityName="Sharp" baseAuthorityYear="1875" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="trogoides">
<emphasis id="B95BEAB6FFF9063FC1648036FF64FBBF" italics="true" pageId="6" pageNumber="483">E.trogoides</emphasis>
(Sharp, 1875)
</taxonomicName>
. 13 hind wing of
<taxonomicName id="4C2F4D27FFF9063FC4B88011FDE3FBBF" authority="Fikáček &amp; Barclay &amp; Perkins &amp; Sw, 2011" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[493,584,1081,1099]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="483" phylum="Arthropoda" rank="species" species="mendeli" status="sp. nov.">
<emphasis id="B95BEAB6FFF9063FC4B88011FDE3FBBF" box="[493,584,1081,1099]" italics="true" pageId="6" pageNumber="483">E. mendeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857CDFFF9063FC7188010FD24FBBF" box="[589,655,1080,1099]" pageId="6" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations:
<emphasis id="B95BEAB6FFF9063FC6408011FC84FBBF" box="[789,815,1081,1099]" italics="true" pageId="6" pageNumber="483">AA</emphasis>
anterior branch of anal vein;
<emphasis id="B95BEAB6FFF9063FC16D8011FBCDFBBF" box="[1080,1126,1081,1099]" italics="true" pageId="6" pageNumber="483">anins</emphasis>
antennal insertion;
<emphasis id="B95BEAB6FFF9063FC41E807BFEDAFB91" box="[331,369,1107,1125]" italics="true" pageId="6" pageNumber="483">CuA</emphasis>
anterior branch of cubital vein;
<emphasis id="B95BEAB6FFF9063FC7C0807BFD18FB91" box="[661,691,1107,1125]" italics="true" pageId="6" pageNumber="483">MP</emphasis>
posterior branch of media;
<emphasis id="B95BEAB6FFF9063FC6E4807BFC6FFB91" box="[945,964,1107,1125]" italics="true" pageId="6" pageNumber="483">r4</emphasis>
transverse radial vein;
<emphasis id="B95BEAB6FFF9063FC5928046FF4AFB74" box="[199,225,1134,1152]" italics="true" pageId="6" pageNumber="483">RA</emphasis>
anterior branch of radius;
<emphasis id="B95BEAB6FFF9063FC4818046FE45FB74" box="[468,494,1134,1152]" italics="true" pageId="6" pageNumber="483">RP</emphasis>
posterior branch of radius;
<emphasis id="B95BEAB6FFF9063FC7BF8046FCA0FB74" box="[746,779,1134,1152]" italics="true" pageId="6" pageNumber="483">ScA</emphasis>
anterior branch of subcosta.
</paragraph>
</caption>
<subSubSection id="C335652FFFF90633C5C28087FF50FBC0" lastPageId="10" lastPageNumber="487" pageId="6" pageNumber="483" type="description">
<paragraph id="8B9036A4FFF9063FC5C28087FD7FFA7C" blockId="6.[151,1127,1199,1576]" pageId="6" pageNumber="483">
<emphasis id="B95BEAB6FFF9063FC5C28087FE89FB33" bold="true" box="[151,290,1199,1223]" pageId="6" pageNumber="483">Description.</emphasis>
Body widely elongate (
<figureCitation id="13142A21FFF9063FC74C8098FDD6FB3C" box="[537,637,1200,1224]" captionStart="Figs" captionStartId="4.[151,190,1570,1589]" captionTargetBox="[153,1096,241,1475]" captionTargetId="figure-10@4.[151,1096,215,1485]" captionTargetPageId="4" captionText="Figs. 15. General habitus and aedeagi of Epimetopus. 12 E. mendeli sp. nov., general habitus (body length 3.4 mm): 1 dorsal view; 2 lateral view. 35 aedeagus of the holotype: 3 E. mendeli sp. nov., 4 E. flavicaptus sp. nov., 5 E. angulatus Balfour-Browne, 1949. Abbreviations: lap lateral process of median lobe." figureDoi="http://doi.org/10.5281/zenodo.5328787" httpUri="https://zenodo.org/record/5328787/files/figure.png" pageId="6" pageNumber="483">Figs. 12</figureCitation>
), widest at posterior third of elytra, moderately convex in lateral view. Dorsal surface piceous brown, elytra with a pale reddish half-moonshaped spot at ca. posterior 0.4 of intervals 15. Ventral surface piceous black, epipleura turning reddish brown posteriorly; antennae reddish with brown antennal club, maxillary palpi reddish with palpomere 4 brown; trochanters and bases of femora dark brown to brown, distal portions of femora and entire tibiae and tarsi reddish. Body length
<quantity id="4CD79B41FFF9063FC6208178FC5CFA93" box="[885,1015,1360,1384]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.5" metricValueMin="2.3" pageId="6" pageNumber="483" unit="mm" value="2.9" valueMax="3.5" valueMin="2.3">2.33.5 mm</quantity>
(
<typeStatus id="54948806FFF9063FC1528178FBCDFA9C" box="[1031,1126,1360,1384]" pageId="6" pageNumber="483" type="holotype">holotype</typeStatus>
<quantity id="4CD79B41FFF9063FC5C28158FF47FA7C" box="[151,236,1392,1416]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="483" unit="mm" value="3.1">3.1 mm</quantity>
), body width
<quantity id="4CD79B41FFF9063FC42B8158FE55FA73" box="[382,510,1392,1416]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="6" pageNumber="483" unit="mm" value="1.55" valueMax="1.8" valueMin="1.3">1.31.8 mm</quantity>
(
<typeStatus id="54948806FFF9063FC7598158FDC0FA7C" box="[524,619,1392,1416]" pageId="6" pageNumber="483" type="holotype">holotype</typeStatus>
<quantity id="4CD79B41FFF9063FC7278158FD63FA7C" box="[626,712,1392,1416]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="483" unit="mm" value="1.5">1.5 mm</quantity>
).
</paragraph>
<paragraph id="8B9036A4FFF9063EC5E081B9FCDCFD87" blockId="6.[151,1127,1199,1576]" lastBlockId="7.[113,1089,219,1587]" lastPageId="7" lastPageNumber="484" pageId="6" pageNumber="483">
<emphasis id="B95BEAB6FFF9063FC5E081B9FF44FA5C" box="[181,239,1425,1448]" italics="true" pageId="6" pageNumber="483">Head</emphasis>
. Clypeus with uniform rather dense granulation, clypeus feebly concave, without marginal rim, lacking systematic punctures. Frons with granulation similar to that of clypeus, lacking systematic punctures at inner margin of eyes. Eyes large, consisting of rather large ommatidia; elongate oval in dorsal view, incompletely divided into dorsal and ventral portion in anterior 0.7 by a wide canthus; ventral portion large, slightly larger than dorsal portion, delimited posteriorly by short but distinct postocular bridge. Gular sutures widely joined in anterior 0.7, widely divergent in posterior 0.3. Labrum large, exposed, lacking granulation and systematic punctures. Maxillary palpi moderately long, palpomeres 2 and 4 subequal in length, each ca. 2.5× as long as palpomere 3; palpomere 4 strongly asymmetrical, nearly straight on outer margin, widened in basal 0.4 on inner margin; cardo and stipes lacking systematic punctures. Mentum ca. as long as wide, flat, bearing fine punctation; labial palpus short, basal palpomere the smallest, palpomere 2 ca. half as long as palpomere 3, palpomere 3 lacking spiniform sensillae, bearing only a few pore-like sensillae apically. Antenna with 9 antennomeres; scape slightly longer than antennomeres 29 combined, widest at distal margin; pedicel small, bulbous; antennomeres 36 very short, subequal in length, antennomere 6 (cupula) not differentiated from antennomeres
<quantity id="4CD79B41FFF8063EC7D78633FD6CFDC7" box="[642,711,539,563]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="7" pageNumber="484" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">35 in</quantity>
size and shape; antennomeres 79 forming large pubescent club, first club antennomere the shortest and narrowest, 9
<superScript id="7C5A9BECFFF8063EC6878612FC75FDBD" attach="left" box="[978,990,570,585]" fontSize="6" pageId="7" pageNumber="484">th</superScript>
antennomere the longest and widest, ca. as long as antennomeres 78 combined.
</paragraph>
<paragraph id="8B9036A4FFF8063EC5C58655FC5EFBE7" blockId="7.[113,1089,219,1587]" pageId="7" pageNumber="484">
<emphasis id="B95BEAB6FFF8063EC5C58655FEAFFD60" box="[144,260,637,660]" italics="true" pageId="7" pageNumber="484">Prothorax.</emphasis>
Pronotum slightly wider than long; anterior lobe large, covering large basal portion of head up to anterior margin of eyes, bearing a deep distinct groove at anterior margin mesally; lateral lobes large, directed anterolaterad, nearly rectangular in shape. Pronotal sculpture consisting of four elevated ridges; two median ones close to each other, slightly sinuate in shape and joining each other anteriorly, high and distinct especially in their anterior third, defining a median groove; lateral ones weak and rather indistinct. Entire surface of pronotum except for medioanterior margin and anterior third of median ridges covered by moderately dense granulation similar to that of clypeus. Prosternum ca. as long as half of procoxal cavity, flat, bearing a reticulate microsculpture on the whole surface with a few isolated granules at midwidth; prosternal process strongly narrowing between procoxae, but widened again posteriorly, joining the hypomeral process posteriorly (hence, procoxal cavities closed posteriorly). Hypomeron divided by a ridge into the lateral narrow granulate portion and large mesal portion bearing the reticulate microsculpture and scattered granules.
</paragraph>
<paragraph id="8B9036A4FFF8063EC5C58035FD1BF9C7" blockId="7.[113,1089,219,1587]" pageId="7" pageNumber="484">
<emphasis id="B95BEAB6FFF8063EC5C58035FEBDFBC0" box="[144,278,1053,1076]" italics="true" pageId="7" pageNumber="484">Mesothorax.</emphasis>
Scutellar shield slightly longer than wide, rounded posteriorly. Elytra narrowest subanteriorly, widest in posterior 0.3. Each elytron with 10 longitudinal punctuate striae, punctures large, close to each other, not connected by a longitudinal tubercles; scutellar stria absent. Each elytron elevated on sutural interval and bearing three additional high elevated ridges on intervals 3, 5 and 7; ridge on interval 5 interrupted in anterior 0.2, ridges on intervals 3 and 7 complete, ridge on interval 7 reaching large humeral bulge; all ridges reaching subapically. Remaining intervals (i.e. those not elevated to ridges) bearing irregular series of scattered granules; systematic punctures absent from all elytral intervals. Elytral interval 11 with additional highly elevated ridge forming a false elytral margin in dorsal view. Epipleuron not divided into pubescent inner and bare outer portions, wide throughout, reaching elytral apex. Mesoventrite distinctly divided from mesanepisterna, bearing highly elevated transverse ridge on mesoventral process. Anapleural sutures nearly straight, joining anteriorly; mesanepisterna joining mesally. Anterior collar of mesothorax well defined, widest mesally. Mesepimeron large, subtriangular. Whole ventral surface of mesothorax except for anterior portion of mesoventrite and anterior collar bearing reticulate microsculpture, posterior portions of mesoventrite, mesanepisterna and whole mesepimera with scattered granules. Mesoventral process short, loosely contacting metaventral process.
</paragraph>
<paragraph id="8B9036A4FFF70630C5E084F5FEFAFE07" blockId="8.[151,1127,220,340]" lastBlockId="9.[113,1089,219,691]" lastPageId="9" lastPageNumber="486" pageId="8" pageNumber="485">
<emphasis id="B95BEAB6FFF70631C5E084F5FE91FF00" box="[181,314,221,244]" italics="true" pageId="8" pageNumber="485">Metathorax.</emphasis>
Ventral portion of metathorax slightly longer than mesoventrite, bearing reticulate sculpture and scattered granules over its entire surface. Metaventral process long and rather wide, projecting far between mesocoxae; anterolateral rim of metaventrite distinct, reaching submesally. Katepisternum very wide sublaterally, divided by very distinct katepisternal suture, bearing large deeply bifurcate metacoxal process. Metanepisternum ca. 7.5× as long as wide, lacking anterior transverse ridge strongly widened and projecting mesad anteriorly (nearly contacting mesocoxal cavity), bent mesad posteriorly and delimiting the outer wall of metacoxal cavity. Posteroventral portion of metepimeron widely exposed sublaterally. Metacoxal cavities not reaching lateral portion of metathorax. Hind wing well developed, with rather large distal area lacking any veins; RP preserved nearly from base, basal cell incompletely delimited due to interrupted AA
<subScript id="17AB34E1FFF60630C7968583FD70FE4E" attach="right" box="[707,731,427,442]" fontSize="6" pageId="9" pageNumber="486">3+4</subScript>
, wedge cell well developed, very small, AA
<subScript id="17AB34E1FFF60630C58B85E3FF4DFE2E" attach="left" box="[222,230,459,474]" fontSize="6" pageId="9" pageNumber="486">3</subScript>
well developed, bifurcation of MP
<subScript id="17AB34E1FFF60630C70E85E3FDC8FE2E" attach="left" box="[603,611,459,474]" fontSize="6" pageId="9" pageNumber="486">4</subScript>
and CuA
<subScript id="17AB34E1FFF60630C79C85E3FD7AFE2E" attach="left" box="[713,721,459,474]" fontSize="6" pageId="9" pageNumber="486">2</subScript>
preserved as weak remnants only; jugal lobe preserved.
</paragraph>
<caption id="DF50662CFFF70631C5C281F9FDFFF9C1" ID-DOI="http://doi.org/10.5281/zenodo.5328795" ID-Zenodo-Dep="5328795" httpUri="https://zenodo.org/record/5328795/files/figure.png" pageId="8" pageNumber="485" startId="8.[151,190,1489,1508]" targetBox="[162,1116,379,1464]" targetPageId="8">
<paragraph id="8B9036A4FFF70631C5C281F9FDFFF9C1" blockId="8.[151,1128,1489,1589]" pageId="8" pageNumber="485">
Figs. 1419.External morphology of
<taxonomicName id="4C2F4D27FFF70631C48481FAFDD7FA10" authority="Fikáček &amp; Barclay &amp; Perkins &amp; Sw, 2011" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[465,636,1490,1508]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="485" phylum="Arthropoda" rank="species" species="mendeli" status="sp. nov.">
<emphasis id="B95BEAB6FFF70631C48481FAFDD7FA10" box="[465,636,1490,1508]" italics="true" pageId="8" pageNumber="485">Epimetopus mendeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857CDFFF70631C7D481FAFD69FA11" box="[641,706,1490,1509]" pageId="8" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
14 head, ventral view; 15 detail of mouthparts; 16 antenna; 17 detail of prosternum; 18 prothorax, ventral view; 19 microsculpture of prothoracic hypomeron. Abbreviations:
<emphasis id="B95BEAB6FFF70631C44F8220FE92F9EE" box="[282,313,1544,1562]" italics="true" pageId="8" pageNumber="485">cup</emphasis>
cupula;
<emphasis id="B95BEAB6FFF70631C4DB8220FE19F9EE" box="[398,434,1544,1562]" italics="true" pageId="8" pageNumber="485">guls</emphasis>
gular sutures;
<emphasis id="B95BEAB6FFF70631C76E8220FDC8F9EE" box="[571,611,1544,1562]" italics="true" pageId="8" pageNumber="485">hpps</emphasis>
hypomeral process;
<emphasis id="B95BEAB6FFF70631C6758220FC94F9EE" box="[800,831,1544,1562]" italics="true" pageId="8" pageNumber="485">hyp</emphasis>
hypomeron;
<emphasis id="B95BEAB6FFF70631C6E98220FC49F9EE" box="[956,994,1544,1562]" italics="true" pageId="8" pageNumber="485">labp</emphasis>
labial palpus;
<emphasis id="B95BEAB6FFF70631C5C2820AFF1DF9C0" box="[151,182,1570,1588]" italics="true" pageId="8" pageNumber="485">ped</emphasis>
pedicel;
<emphasis id="B95BEAB6FFF70631C441820AFE99F9C0" box="[276,306,1570,1588]" italics="true" pageId="8" pageNumber="485">pps</emphasis>
prosternal process;
<emphasis id="B95BEAB6FFF70631C4B8820AFE54F9C0" box="[493,511,1570,1588]" italics="true" pageId="8" pageNumber="485">sc</emphasis>
scapus.
</paragraph>
</caption>
<paragraph id="8B9036A4FFF60630C5C585D5FC77FD47" blockId="9.[113,1089,219,691]" pageId="9" pageNumber="486">
<emphasis id="B95BEAB6FFF60630C5C585D5FF5EFDE0" box="[144,245,509,532]" italics="true" pageId="9" pageNumber="486">Abdomen</emphasis>
with 5 visible ventrites. Ventrite 1 with large metacoxal grooves covering nearly its whole surface. Ventrites 25 flat, lacking pubescence; ventrites 23 and anterior portion of ventrite 4 with fine punctation (i.e. having shagreened appearance when examined under the binocular at 90× magnification), posterior portion of ventrite 4 and whole ventrite 5 shiny, without punctation; lateral margins of ventrites 24 and posterior margin of ventrite 5 with fine submarginal ridge. Posteromedian portion of ventrite 5 without emargination.
</paragraph>
<caption id="DF50662CFFF60630C52481FBFD76F9C2" ID-DOI="http://doi.org/10.5281/zenodo.5367795" ID-Zenodo-Dep="5367795" httpUri="https://zenodo.org/record/5367795/files/figure.png" pageId="9" pageNumber="486" startId="9.[113,152,1491,1510]" targetBox="[113,1084,728,1466]" targetPageId="9">
<paragraph id="8B9036A4FFF60630C52481FBFD76F9C2" blockId="9.[113,1090,1491,1591]" pageId="9" pageNumber="486">
Figs. 2025. External morphology of
<taxonomicName id="4C2F4D27FFF60630C4E081FCFDCFFA12" authority="Fikáček &amp; Barclay &amp; Perkins &amp; Sw, 2011" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[437,612,1492,1510]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="mendeli" status="sp. nov.">
<emphasis id="B95BEAB6FFF60630C4E081FCFDCFFA12" box="[437,612,1492,1510]" italics="true" pageId="9" pageNumber="486">Epimetopus mendeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857CDFFF60630C73C81FCFD07FA13" box="[617,684,1492,1511]" pageId="9" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
20 mesothorax, ventral view; 21 abdomen; 22 middle leg; 23 metathorax, ventral view; 24 detail of pronotal granulation; 25 detail of middle tarsus. Abbreviations:
<emphasis id="B95BEAB6FFF60630C5AD8222FEB5F9E8" box="[248,286,1546,1564]" italics="true" pageId="9" pageNumber="486">aes3</emphasis>
metanepisternum;
<emphasis id="B95BEAB6FFF60630C48C8222FE57F9E8" box="[473,508,1546,1564]" italics="true" pageId="9" pageNumber="486">em2</emphasis>
mesepimeron;
<emphasis id="B95BEAB6FFF60630C7C28222FD11F9E8" box="[663,698,1546,1564]" italics="true" pageId="9" pageNumber="486">em3</emphasis>
metepimeron;
<emphasis id="B95BEAB6FFF60630C6078222FCD3F9E8" box="[850,888,1546,1564]" italics="true" pageId="9" pageNumber="486">keps</emphasis>
katepisternal suture;
<emphasis id="B95BEAB6FFF60630C524820CFF39F9C2" box="[113,146,1572,1590]" italics="true" pageId="9" pageNumber="486">msv</emphasis>
mesoventrite;
<emphasis id="B95BEAB6FFF60630C474820CFEEBF9C2" box="[289,320,1572,1590]" italics="true" pageId="9" pageNumber="486">mtv</emphasis>
metaventrite;
<emphasis id="B95BEAB6FFF60630C49E820CFE74F9C2" box="[459,479,1572,1590]" italics="true" pageId="9" pageNumber="486">v2</emphasis>
second abdominal ventrite.
</paragraph>
</caption>
<paragraph id="8B9036A4FFF50633C5E084F5FCA1FD80" blockId="10.[151,1127,220,1588]" pageId="10" pageNumber="487">
<emphasis id="B95BEAB6FFF50633C5E084F5FF45FF00" box="[181,238,221,244]" italics="true" pageId="10" pageNumber="487">Legs.</emphasis>
Profemora widest in basal third, covered with large tubercle-like projections on ventral face, bearing dense pubescence in basal third dorsally; mesofemora apparently widened at midlength, bearing stout decumbent sparsely arranged setae on ventral surface; metafemora long and slender, only indistinctly widened in distal third, ventral surface with same setation as mesofemora. All femora without tibial grooves. Pro- and mesotibiae distinctly longer than femora, metatibiae ca. as long as metafemora; all tibiae narrowest basally, slightly widened subbasally and then conical throughout in meso- and metatibiae, protibiae distinctly widened ca. at midlength; all tibiae bearing series of equidistant series of stout setae, those on inner face of metatibiae longer and thinner than remaining ones. Tibial spurs short, only slightly longer than other spines on tibial apices. Tarsi with five tarsomeres, tarsomere 1 shortest, rather indistinct in ventral view, tarsomeres 24 subequal in length, ca. as long as tarsomere 5 when combined; each tarsomere with few stout erect setae ventrally and few short decumbent setae on dorsal face. Claws moderately large, semicircular.
</paragraph>
<paragraph id="8B9036A4FFF50633C5E08655FEC3FCA0" blockId="10.[151,1127,220,1588]" pageId="10" pageNumber="487">
<emphasis id="B95BEAB6FFF50633C5E08655FEF9FD60" box="[181,338,637,660]" italics="true" pageId="10" pageNumber="487">Male genitalia</emphasis>
(
<figureCitation id="13142A21FFF50633C4358654FE00FD60" box="[352,427,636,660]" captionStart="Figs" captionStartId="4.[151,190,1570,1589]" captionTargetBox="[153,1096,241,1475]" captionTargetId="figure-10@4.[151,1096,215,1485]" captionTargetPageId="4" captionText="Figs. 15. General habitus and aedeagi of Epimetopus. 12 E. mendeli sp. nov., general habitus (body length 3.4 mm): 1 dorsal view; 2 lateral view. 35 aedeagus of the holotype: 3 E. mendeli sp. nov., 4 E. flavicaptus sp. nov., 5 E. angulatus Balfour-Browne, 1949. Abbreviations: lap lateral process of median lobe." figureDoi="http://doi.org/10.5281/zenodo.5328787" httpUri="https://zenodo.org/record/5328787/files/figure.png" pageId="10" pageNumber="487">Figs. 3</figureCitation>
,
<figureCitation id="13142A21FFF50633C4E28653FE55FD67" box="[439,510,635,659]" captionStart="Figs" captionStartId="11.[113,152,1521,1540]" captionTargetBox="[233,968,504,1497]" captionTargetId="figure-17@11.[229,973,502,1504]" captionTargetPageId="11" captionText="Figs. 2629. Details of the aedeagal morphology. 2627 Epimetopus mendeli sp. nov., paratype from the type locality: 26 ventral view; 27 dorsal view. 2829 E. flavicaptus sp. nov., holotype:28 ventral view; 29 dorsal view. Abbreviations: dl dorsal lobe; vl ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.5367797" httpUri="https://zenodo.org/record/5367797/files/figure.png" pageId="10" pageNumber="487">2627</figureCitation>
). Aedeagus
<quantity id="4CD79B41FFF50633C72B8654FD64FD67" box="[638,719,636,660]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="487" unit="mm" value="0.9">0.9 mm</quantity>
long. Phallobase symmetrical, widely rounded basally, slightly shorter than parameres. Paramere wide throughout, continuously arcuate on outer margin, apex widely rounded. Median lobe narrower than paramere; basal portion narrow, forming ca. 0.4 of length of median lobe, projecting into small lateral and large median projections ca. at midlength; apical portion bilobate, bearing two sets of spines subapically. Sternite 9 with semicircular median portion. Sternite 8 without median anteriaddirected projection.
</paragraph>
<paragraph id="8B9036A4FFF50633C5C28773FF50FBC0" blockId="10.[151,1127,220,1588]" pageId="10" pageNumber="487">
<emphasis id="B95BEAB6FFF50633C5C28773FEA1FC87" bold="true" box="[151,266,859,883]" pageId="10" pageNumber="487">Variation.</emphasis>
The type material varies particularly in size, which seems to be connected to sex (all dissected male specimens are smaller and less robust than female specimens (recognized by the presence of an egg case and/or projecting cerci). The shape of the lateral margin of lateral lobes of the pronotum varies from straight to slightly concave or shallowly excised. The pale spot on the elytra is rather indistinct in several
<typeStatus id="54948806FFF50633C7BB87F4FCF3FC00" box="[750,856,988,1012]" pageId="10" pageNumber="487" type="paratype">paratypes</typeStatus>
. All specimens examined are constant in the characters mentioned in the differential diagnosis, including those of male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C335652FFFF50633C5C28014FC50FB47" pageId="10" pageNumber="487" type="etymology">
<paragraph id="8B9036A4FFF50633C5C28014FC50FB47" blockId="10.[151,1127,220,1588]" pageId="10" pageNumber="487">
<emphasis id="B95BEAB6FFF50633C5C28014FEB3FBA0" bold="true" box="[151,280,1084,1108]" pageId="10" pageNumber="487">Etymology.</emphasis>
The species is dedicated to one of its collectors, Howard Mendel, recently retired Head of Collections at the Natural History Museum, London. His persistence in climbing down the ravine to extend the
<typeStatus id="54948806FFF50633C4C38054FE6FFB60" box="[406,452,1148,1172]" pageId="10" pageNumber="487">type</typeStatus>
series of this new species mirrors his persistence in fighting for the best interests of the Entomology Collections throughout his successful career.
</paragraph>
</subSubSection>
<subSubSection id="C335652FFFF50633C5C28093FD4FFA07" pageId="10" pageNumber="487" type="biology_ecology">
<paragraph id="8B9036A4FFF50633C5C28093FD4FFA07" blockId="10.[151,1127,220,1588]" pageId="10" pageNumber="487">
<emphasis id="B95BEAB6FFF50633C5C28093FF58FB27" bold="true" box="[151,243,1211,1235]" pageId="10" pageNumber="487">Biology.</emphasis>
The
<typeStatus id="54948806FFF50633C47B8094FE20FB20" box="[302,395,1212,1236]" pageId="10" pageNumber="487" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D29FD2DFFF50633C4908094FDE4FB20" box="[453,591,1212,1236]" pageId="10" pageNumber="487" type="generic" typeStatus="paratypes">18 paratypes</specimenCount>
of this species were collected under loose rocks and rock litter lying on or partly buried in wet gravel substrate, close to the edge of a very fast flowing mountain stream. Males and females (generally with egg cases) were found in groups under stones. The stream flows through good quality cloud forest, but can be accessed by climbing down a ravine beneath a road bridge close to the entrance to Cock of the Rock Lodge. The habitat is not species-rich, although
<taxonomicName id="4C2F4D27FFF50633C7CA8175FBCCFA80" authority="(Horn, 1938)" baseAuthorityName="Horn" baseAuthorityYear="1938" box="[671,1127,1372,1396]" class="Insecta" family="Carabidae" genus="Oxycheila" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="487" phylum="Arthropoda" rank="species" species="pseudonigroaenea">
<emphasis id="B95BEAB6FFF50633C7CA8175FC7EFA80" box="[671,981,1373,1396]" italics="true" pageId="10" pageNumber="487">Oxycheila pseudonigroaenea</emphasis>
(Horn, 1938)
</taxonomicName>
(
<taxonomicName id="4C2F4D27FFF50633C5CA8154FEBAFA60" authorityName="Latreille" authorityYear="1802" box="[159,273,1404,1428]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="487" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
:
<taxonomicName id="4C2F4D27FFF50633C4498154FE0DFA60" authorityName="Latreille" authorityYear="1802" box="[284,422,1404,1428]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="487" phylum="Arthropoda" rank="subFamily" subFamily="Cicindelinae">Cicindelinae</taxonomicName>
, det. By F. Cassola) and some smaller
<taxonomicName id="4C2F4D27FFF50633C61B8154FC11FA60" authorityName="Latreille" authorityYear="1802" box="[846,954,1404,1428]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="487" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
were abundant, and the Torrent Duck
<taxonomicName id="4C2F4D27FFF50633C4D081B5FD4FFA40" authority="Gould, 1842" authorityName="Gould" authorityYear="1842" box="[389,740,1436,1460]" class="Aves" family="Anatidae" genus="Merganetta" kingdom="Animalia" order="Anseriformes" pageId="10" pageNumber="487" phylum="Chordata" rank="species" species="armata">
<emphasis id="B95BEAB6FFF50633C4D081B5FDFEFA40" box="[389,597,1437,1460]" italics="true" pageId="10" pageNumber="487">Merganetta armata</emphasis>
Gould, 1842
</taxonomicName>
(
<taxonomicName id="4C2F4D27FFF50633C7A681B4FC86FA40" authorityName="Linnaeus" authorityYear="1758" box="[755,813,1436,1460]" class="Aves" kingdom="Animalia" pageId="10" pageNumber="487" phylum="Chordata" rank="class">Aves</taxonomicName>
:
<taxonomicName id="4C2F4D27FFF50633C66281B4FC37FA40" box="[823,924,1436,1460]" class="Aves" family="Anatidae" kingdom="Animalia" order="Anseriformes" pageId="10" pageNumber="487" phylum="Chordata" rank="family">Anatidae</taxonomicName>
), a predator in fast flowing Andean streams, was present. An additional
<specimenCount id="9D29FD2DFFF50633C7EB8194FC9DFA20" box="[702,822,1468,1492]" pageId="10" pageNumber="487" type="generic" typeStatus="paratypes">9 paratypes</specimenCount>
of
<taxonomicName id="4C2F4D27FFF50633C6028195FC6CFA20" authorityName="Fikáček &amp; Barclay &amp; Perkins &amp; Sw" authorityYear="2011" box="[855,967,1469,1492]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="487" phylum="Arthropoda" rank="species" species="mendeli">
<emphasis id="B95BEAB6FFF50633C6028195FC6CFA20" box="[855,967,1469,1492]" italics="true" pageId="10" pageNumber="487">E. mendeli</emphasis>
</taxonomicName>
were collected at MV light within a few hundred meters of the stream.
</paragraph>
</subSubSection>
<subSubSection id="C335652FFFF50633C5C281D3FC0DF9C7" pageId="10" pageNumber="487" type="distribution">
<paragraph id="8B9036A4FFF50633C5C281D3FC0DF9C7" blockId="10.[151,1127,220,1588]" pageId="10" pageNumber="487">
<emphasis id="B95BEAB6FFF50633C5C281D3FE82F9E7" bold="true" box="[151,297,1531,1555]" pageId="10" pageNumber="487">Distribution.</emphasis>
The species is known from three rather closely situated localities on the eastern slope of the Andes Mts. in the
<collectingRegion id="49EBF846FFF50633C4888234FD8BF9C0" box="[477,544,1564,1588]" country="Peru" name="Cusco" pageId="10" pageNumber="487">Cusco</collectingRegion>
and
<collectingRegion id="49EBF846FFF50633C7018234FD59F9C0" box="[596,754,1564,1588]" country="Peru" name="Madre de Dios" pageId="10" pageNumber="487">Madre de Dios</collectingRegion>
provinces,
<collectingCountry id="F3387634FFF50633C63B8234FC09F9C7" box="[878,930,1564,1587]" name="Peru" pageId="10" pageNumber="487">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>