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<document id="D5508DD35741EFD52074F9BFA7C57610" ID-DOI="10.5281/zenodo.195647" ID-GBIF-Dataset="5dbc17e2-43e6-4ca4-9921-56eb43a9aec3" ID-ISSN="1175-5326" ID-Zenodo-Dep="195647" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460231312424" checkinUser="plazi" docAuthor="Zaragoza, Juan A." docDate="2010" docId="0386879EFFA1B864FF5F766B9CECFADF" docLanguage="en" docName="zt02491p060.pdf" docOrigin="Zootaxa 2491" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Microcreagrella caeca subsp. caeca Simon 1883" docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="FFBFFFE6FFABB86FFFC8743C9C0FFA23" masterDocTitle="Arcanobisium, a remarkable new genus, representing a new subfamily with a relictual distribution from eastern Spain (Arachnida: Pseudoscorpiones: Syarinidae)" masterLastPageNumber="60" masterPageNumber="41" pageNumber="51" updateTime="1698239000673" updateUser="plazi">
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<mods:title id="F1090ABAEA253773F5A62068258A8F52">Arcanobisium, a remarkable new genus, representing a new subfamily with a relictual distribution from eastern Spain (Arachnida: Pseudoscorpiones: Syarinidae)</mods:title>
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<mods:namePart id="A6E1123E77C1E282013CCD813AFDC39B">Zaragoza, Juan A.</mods:namePart>
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<mods:date id="C3679C5C35A3A413C4781F93A453BCBF">2010</mods:date>
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<treatment id="0386879EFFA1B864FF5F766B9CECFADF" ID-DOI="http://doi.org/10.5281/zenodo.6208437" ID-GBIF-Taxon="119387343" ID-Zenodo-Dep="6208437" LSID="urn:lsid:plazi:treatment:0386879EFFA1B864FF5F766B9CECFADF" httpUri="http://treatment.plazi.org/id/0386879EFFA1B864FF5F766B9CECFADF" lastPageId="11" lastPageNumber="51" pageId="10" pageNumber="51">
<subSubSection id="C3356503FFA1B865FF5F766B9EAAF852" box="[151,677,599,625]" pageId="10" pageNumber="51" type="nomenclature">
<paragraph id="8B903688FFA1B865FF5F766B9EAAF852" blockId="10.[151,677,599,625]" box="[151,677,599,625]" pageId="10" pageNumber="51">
<heading id="D0D881E4FFA1B865FF5F766B9EAAF852" bold="true" box="[151,677,599,625]" fontSize="11" level="1" pageId="10" pageNumber="51" reason="1">
<taxonomicName id="4C2F4D0BFFA1B865FF5F766B9EAAF852" ID-CoL="5JP2W" authority="Simon, 1883" authorityName="Simon" authorityYear="1883" box="[151,677,599,625]" class="Arachnida" family="Syarinidae" genus="Microcreagrella" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="51" phylum="Arthropoda" rank="subSpecies" species="caeca" subSpecies="caeca">
<emphasis id="B95BEA9AFFA1B865FF5F766B9EAAF852" bold="true" box="[151,677,599,625]" pageId="10" pageNumber="51">
<emphasis id="B95BEA9AFFA1B865FF5F766B9DFEF852" bold="true" box="[151,497,599,625]" italics="true" pageId="10" pageNumber="51">Microcreagrella caeca caeca</emphasis>
(
<bibRefCitation id="EFBE4B79FFA1B865FDC9766B9E92F852" author="Simon" box="[513,669,599,625]" pageId="10" pageNumber="60" refString="Simon, E. (1883) Materiaux pour servir a la faune arachnologique des iles de l'Ocean Atlantique (Acores, Madere, Salvages, Canaries, Cap Vert, Sainte-Helene et Bermudes). Annales de la Societe Entomologique de France, (6) 3, 259 - 314." type="journal article" year="1883">Simon, 1883</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8B903688FFA1B865FF5F76A19E4BF897" blockId="10.[151,775,669,756]" box="[151,580,669,692]" pageId="10" pageNumber="51">
<treatmentCitationGroup id="AB3F11A6FFA1B865FF5F76A19E4BF897" box="[151,580,669,692]" pageId="10" pageNumber="51">
<taxonomicName id="4C2F4D0BFFA1B865FF5F76A19E07F897" authority="Simon 1883: 279" authorityName="Simon" authorityPageNumber="279" authorityYear="1883" box="[151,520,669,692]" class="Arachnida" family="Cheliferidae" genus="Obisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="caecum">
<emphasis id="B95BEA9AFFA1B865FF5F76A19D46F897" box="[151,329,669,692]" italics="true" pageId="10" pageNumber="51">Obisium caecum</emphasis>
<treatmentCitation id="0A8E1099FFA1B865FE9876A19E07F897" author="Simon" box="[336,520,669,692]" page="279" pageId="10" pageNumber="51" year="1883">
<bibRefCitation id="EFBE4B79FFA1B865FE9876A19DDAF897" author="Simon" box="[336,469,669,692]" pageId="10" pageNumber="60" refString="Simon, E. (1883) Materiaux pour servir a la faune arachnologique des iles de l'Ocean Atlantique (Acores, Madere, Salvages, Canaries, Cap Vert, Sainte-Helene et Bermudes). Annales de la Societe Entomologique de France, (6) 3, 259 - 314." type="journal article" year="1883">Simon 1883</bibRefCitation>
: 279
</treatmentCitation>
</taxonomicName>
280.
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</paragraph>
<paragraph id="8B903688FFA1B865FF5F76819E9CF8F7" blockId="10.[151,775,669,756]" box="[151,659,701,724]" pageId="10" pageNumber="51">
<treatmentCitationGroup id="AB3F11A6FFA1B865FF5F76819E9CF8F7" box="[151,659,701,724]" pageId="10" pageNumber="51">
<taxonomicName id="4C2F4D0BFFA1B865FF5F76819DD9F8F7" authority="Simon" authorityName="Simon" box="[151,470,701,724]" class="Arachnida" family="Neobisiidae" genus="Microcreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B95BEA9AFFA1B865FF5F76819D7CF8F7" box="[151,371,701,724]" italics="true" pageId="10" pageNumber="51">Microcreagris caeca</emphasis>
(Simon)
</taxonomicName>
:
<treatmentCitation id="0A8E1099FFA1B865FE2876819E81F8F7" author="Beier" box="[480,654,701,724]" page="157" pageId="10" pageNumber="51" year="1932">
<bibRefCitation id="EFBE4B79FFA1B865FE2876819E56F8F7" author="Beier" box="[480,601,701,724]" pageId="10" pageNumber="58" refString="Beier, M. (1932) Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich, 57, i - xx, 1 - 258." type="journal article" year="1932">Beier 1932</bibRefCitation>
: 157
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3356503FFA1B865FF5F76E19F08F8D7" box="[151,775,733,756]" pageId="10" pageNumber="51" type="description">
<paragraph id="8B903688FFA1B865FF5F76E19F08F8D7" blockId="10.[151,775,669,756]" box="[151,775,733,756]" pageId="10" pageNumber="51">
<treatmentCitationGroup id="AB3F11A6FFA1B865FF5F76E19F08F8D7" box="[151,775,733,756]" pageId="10" pageNumber="51">
<taxonomicName id="4C2F4D0BFFA1B865FF5F76E19DE4F8D7" ID-CoL="6RG62" authority="Simon" authorityName="Simon" box="[151,491,733,756]" class="Arachnida" family="Syarinidae" genus="Microcreagrella" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B95BEA9AFFA1B865FF5F76E19D86F8D7" box="[151,393,733,756]" italics="true" pageId="10" pageNumber="51">Microcreagrella caeca</emphasis>
(Simon)
</taxonomicName>
:
<treatmentCitation id="0A8E1099FFA1B865FE3C76E19E9CF8D7" author="Beier" box="[500,659,733,756]" page="72" pageId="10" pageNumber="51" year="1961">
<bibRefCitation id="EFBE4B79FFA1B865FE3C76E19E61F8D7" author="Beier" box="[500,622,733,756]" pageId="10" pageNumber="58" refString="Beier, M. (1961) Pseudoscorpione von den Azoren und Madeira. Boletim do Museu Municipal do Funchal, 14, 67 - 74." type="journal article" year="1961">Beier 1961</bibRefCitation>
: 72
</treatmentCitation>
74, fig. 2.
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</paragraph>
</subSubSection>
<subSubSection id="C3356503FFA1B865FF5F771E98B6FC51" pageId="10" pageNumber="51" type="materials_examined">
<paragraph id="8B903688FFA1B865FF5F771E9DF9F941" blockId="10.[151,1437,802,2023]" pageId="10" pageNumber="51">
<emphasis id="B95BEA9AFFA1B865FF5F771E9D94F91F" bold="true" box="[151,411,802,828]" pageId="10" pageNumber="51">Material examined.</emphasis>
<collectingCountry id="F3387618FFA1B865FE6C771E9E1DF91F" box="[420,530,802,828]" name="Portugal" pageId="10" pageNumber="51">Portugal</collectingCountry>
, Setúbal, Sesimbra, Lapa do Vento,
<date id="FF911048FFA1B865FC14771E989EF91F" box="[988,1169,802,828]" pageId="10" pageNumber="51" value="2007-04-11">11 April 2007</date>
:
<specimenCount id="9D29FD01FFA1B865FB56771E9900F91F" box="[1182,1295,802,828]" pageId="10" pageNumber="51" type="female">1 female</specimenCount>
, leg. Pedro Cardoso, deposited in
<collectionCode id="ED3EAE4DFFA1B865FE5577749DFDF941" box="[413,498,840,866]" pageId="10" pageNumber="51">DEUA</collectionCode>
.
</paragraph>
<paragraph id="8B903688FFA1B865FF0E775198F3F9D4" blockId="10.[151,1437,802,2023]" pageId="10" pageNumber="51">
<emphasis id="B95BEA9AFFA1B865FF0E77519E2EF9A4" bold="true" box="[198,545,877,903]" pageId="10" pageNumber="51">Supplementary description.</emphasis>
Carapace, chelicera and pedipalps brownish-red, legs pale and opisthosoma desclerotised. Opisthosoma elongate, all tergites and sternites wide and easy discernible. Tergites IXI chaetotaxy: 8: 9: 9: 9: 8: 8: 8: 8: 8: 9: 7, opisthosoma ventrally crushed, anal cone with 2+2 setae. Stigmata III and IV each with 4 setae. Pleural membrane laterally striate but also medially weakly granulate.
</paragraph>
<paragraph id="8B903688FFA1B865FF0E703E9F32FE61" blockId="10.[151,1437,802,2023]" pageId="10" pageNumber="51">
Body length
<quantity id="4CD79B6DFFA1B865FE94703E9DC7FE3F" box="[348,456,1026,1052]" metricMagnitude="-3" metricUnit="m" metricValue="2.14" pageId="10" pageNumber="51" unit="mm" value="2.14">2.14 mm</quantity>
. Carapace elongate, 1.37× (0.70/
<quantity id="4CD79B6DFFA1B865FC8F703E9FBDFE3F" box="[839,946,1026,1052]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="10" pageNumber="51" unit="mm" value="0.51">0.51 mm</quantity>
), without eyes or eye-spots, with 24 setae, formula 4: 6: 8: 6. Epistome small and weakly prominent.
</paragraph>
<paragraph id="8B903688FFA1B865FF0E70719D4CFEF4" blockId="10.[151,1437,802,2023]" pageId="10" pageNumber="51">
Chelicera with 5 setae on palm, galea long (
<quantity id="4CD79B6DFFA1B865FD0670719F34FE44" box="[718,827,1101,1127]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="51" unit="mm" value="0.06">0.06 mm</quantity>
) and simple. Hand 2.00× (0.38/
<quantity id="4CD79B6DFFA1B865FB7C7071992DFE44" box="[1204,1314,1101,1127]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="10" pageNumber="51" unit="mm" value="0.19">0.19 mm</quantity>
), movable finger
<quantity id="4CD79B6DFFA1B865FF20704E9D5CFEAF" box="[232,339,1138,1164]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="10" pageNumber="51" unit="mm" value="0.26">0.26 mm</quantity>
long. Movable finger with subgaleal seta situated 0.73× from base, dental row ending far basal; fixed finger with 15 medium or small teeth, movable finger with 9 large teeth and 4 weak basal protuberances.
</paragraph>
<paragraph id="8B903688FFA1B865FF0E70DE9DCFFC21" blockId="10.[151,1437,802,2023]" pageId="10" pageNumber="51">
Apex of pedipalpal coxa triangular and long, with 2 distal setae. Trichobothrium
<emphasis id="B95BEA9AFFA1B865FBB170DF9880FEDF" box="[1145,1167,1251,1276]" italics="true" pageId="10" pageNumber="51">ib</emphasis>
dorsal, in basal half of hand,
<emphasis id="B95BEA9AFFA1B865FF1771349CF4FF02" box="[223,251,1288,1313]" italics="true" pageId="10" pageNumber="51">eb</emphasis>
on antiaxial face of hand and far basal,
<emphasis id="B95BEA9AFFA1B865FD1171349EE6FF02" box="[729,745,1288,1313]" italics="true" pageId="10" pageNumber="51">it</emphasis>
level with
<emphasis id="B95BEA9AFFA1B865FCA771349F8BFF02" box="[879,900,1288,1313]" italics="true" pageId="10" pageNumber="51">et</emphasis>
and both close to apex of finger,
<emphasis id="B95BEA9AFFA1B865FADC7134993AFF02" box="[1300,1333,1288,1313]" italics="true" pageId="10" pageNumber="51">est</emphasis>
distal to
<emphasis id="B95BEA9AFFA1B865FF5F71119CBCFF65" box="[151,179,1325,1350]" italics="true" pageId="10" pageNumber="51">ist</emphasis>
,
<emphasis id="B95BEA9AFFA1B865FF0B71119CE9FF65" box="[195,230,1325,1350]" italics="true" pageId="10" pageNumber="51">isb</emphasis>
at base of finger,
<emphasis id="B95BEA9AFFA1B865FE7671119DEAFF65" box="[446,485,1325,1350]" italics="true" pageId="10" pageNumber="51">esb</emphasis>
on antiaxial distal portion of finger. Trichobothrium
<emphasis id="B95BEA9AFFA1B865FBA871119867FF65" box="[1120,1128,1325,1350]" italics="true" pageId="10" pageNumber="51">t</emphasis>
shorter, thicker and more acuminate than others on the movable finger, not flattened or lanceolate, and bent backward;
<emphasis id="B95BEA9AFFA1B865FB14716F98FFFF4F" box="[1244,1264,1363,1388]" italics="true" pageId="10" pageNumber="51">st</emphasis>
basal, close to
<emphasis id="B95BEA9AFFA1B865FF5F71449C90FFB2" box="[151,159,1400,1425]" italics="true" pageId="10" pageNumber="51">t</emphasis>
, distance
<emphasis id="B95BEA9AFFA1B865FED171449D44FFB2" box="[281,331,1400,1426]" italics="true" pageId="10" pageNumber="51">b -sb</emphasis>
1.49× longer than
<emphasis id="B95BEA9AFFA1B865FDF971449E66FFB2" box="[561,617,1400,1426]" italics="true" pageId="10" pageNumber="51">sb -st</emphasis>
,
<emphasis id="B95BEA9AFFA1B865FDB171449E9BFFB2" box="[633,660,1400,1425]" italics="true" pageId="10" pageNumber="51">sb</emphasis>
not forming group with
<emphasis id="B95BEA9AFFA1B865FC7471449FEDFFB2" box="[956,994,1400,1426]" italics="true" pageId="10" pageNumber="51">st -t</emphasis>
. A row of 4 antiaxial feather-shaped setae distal of trichobothrium
<emphasis id="B95BEA9AFFA1B865FDC871A19E2CFF95" box="[512,547,1437,1462]" italics="true" pageId="10" pageNumber="51">isb</emphasis>
. Pedipalps: trochanter 0.39/0.18 (2.14×), femur 0.63/0.21 (3.00×), patella 0.56/0.28 (2.04×) and its pedicel 0.19 long, chela with pedicel 1.05/0.37 (2.82×), hand with pedicel 0.54/0.37 (1.48×), finger 0.55 long.
</paragraph>
<paragraph id="8B903688FFA1B865FF0E723198B6FC51" blockId="10.[151,1437,802,2023]" pageId="10" pageNumber="51">Junction between femur-patella of leg IV perpendicular, patella 1.41× longer than femur. Subterminal seta with 56 apical rami and one internal small medial tooth. Femur+patella 0.53/0.20 (2.70×), tibia 0.48/0.10 (4.80×) TS 0.71, metatarsus 0.15/0.07 (2.16×) TS 0.30, tarsus 0.26/0.005 (4.91×) TS 0.36.</paragraph>
</subSubSection>
<subSubSection id="C3356503FFA1B864FF0E72419CECFADF" lastPageId="11" lastPageNumber="52" pageId="10" pageNumber="51" type="discussion">
<paragraph id="8B903688FFA1B865FF0E7241989BFD0F" blockId="10.[151,1437,802,2023]" pageId="10" pageNumber="51">
<emphasis id="B95BEA9AFFA1B865FF0E72419D30FCB4" bold="true" box="[198,319,1661,1687]" pageId="10" pageNumber="51">Remarks.</emphasis>
Using the key provided by
<bibRefCitation id="EFBE4B79FFA1B865FD4D72419F13FCB4" author="Beier" box="[645,796,1661,1687]" pageId="10" pageNumber="58" refString="Beier, M. (1963) Ordnung Pseudoscorpionidea (Afterskorpione). Bestimmungsbucher zur Bodenfauna Europas. Vol. 1. Akademie-Verlag. Berlin, 313 pp." type="book" year="1963">Beier (1963)</bibRefCitation>
, this specimen is identified by having the chelal finger as long as the hand with pedicel as
<taxonomicName id="4C2F4D0BFFA1B865FD9B729F9FB2FC9F" box="[595,957,1699,1724]" class="Arachnida" family="Syarinidae" genus="Microcreagrella" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="51" phylum="Arthropoda" rank="subSpecies" species="caeca" subSpecies="caeca">
<emphasis id="B95BEA9AFFA1B865FD9B729F9FB2FC9F" box="[595,957,1699,1724]" italics="true" pageId="10" pageNumber="51">Microcreagrella caeca caeca</emphasis>
</taxonomicName>
, known from the Azores and Canarias Islands. However, the chela and hand are distinctly more slender than in the nominate subspecies; this variability should be verified with more specimens from new localities. The tarsi of each leg are not basally enlarged in this specimen, unlike the sample from Coimbra studied by
<bibRefCitation id="EFBE4B79FFA1B865FC1C732E989EFD0F" author="Mahnert" box="[980,1169,1810,1836]" pageId="10" pageNumber="59" refString="Mahnert, V. (1980) Pseudoskorpione (Arachnida) aus Hohlen Italiens, mit Bemerkungen zur Gattung Pseudoblothrus. Grotte d'Italia, (4 a) 8, 21 - 38." type="journal article" year="1980">Mahnert (1980)</bibRefCitation>
.
</paragraph>
<paragraph id="8B903688FFA1B864FF0E73049CECFADF" blockId="10.[151,1437,802,2023]" lastBlockId="11.[151,1435,152,252]" lastPageId="11" lastPageNumber="52" pageId="10" pageNumber="51">
The presence of
<taxonomicName id="4C2F4D0BFFA1B865FE4773049E94FD72" box="[399,667,1848,1873]" class="Arachnida" family="Syarinidae" genus="Microcreagrella" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B95BEA9AFFA1B865FE4773049E94FD72" box="[399,667,1848,1873]" italics="true" pageId="10" pageNumber="51">Microcreagrella caeca</emphasis>
</taxonomicName>
in mainland
<collectingCountry id="F3387618FFA1B865FCF373049FAEFD71" box="[827,929,1848,1874]" name="Portugal" pageId="10" pageNumber="51">Portugal</collectingCountry>
, at the Botanical Garden of Coimbra, was reported by
<bibRefCitation id="EFBE4B79FFA1B865FEE073619DE8FD54" author="Mahnert" box="[296,487,1885,1911]" pageId="10" pageNumber="59" refString="Mahnert, V. (1980) Pseudoskorpione (Arachnida) aus Hohlen Italiens, mit Bemerkungen zur Gattung Pseudoblothrus. Grotte d'Italia, (4 a) 8, 21 - 38." type="journal article" year="1980">Mahnert (1980)</bibRefCitation>
, hence it is possible that its presence there was due to the importation of plants from one of the Macaronesian islands (V. Mahnert,
<emphasis id="B95BEA9AFFA1B865FCDA73BF9F52FDBF" box="[786,861,1923,1948]" italics="true" pageId="10" pageNumber="51">in litt.</emphasis>
). However, the specimen described above was found in a cave
<quantity id="4CD79B6DFFA1B865FE9C73949D90FDE1" box="[340,415,1960,1986]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="51" unit="km" value="30.0">30 km</quantity>
from Lisbon and more than
<quantity id="4CD79B6DFFA1B865FD3973949F45FDE1" box="[753,842,1960,1986]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="51" unit="km" value="200.0">200 km</quantity>
from Coimbra, 7 meters deep in a semi-dark zone and under a stone; the outside vegetation was xerophilic and no imported plants were present (Pedro Cardoso,
<emphasis id="B95BEA9AFFA0B864FF5F74A49CD3FA92" box="[151,220,152,177]" italics="true" pageId="11" pageNumber="52">in litt.</emphasis>
). Thus it would appear that this species has a wider distribution than previously supposed, being present in continental Europe and the Macaronesian islands, as is the case for
<taxonomicName id="4C2F4D0BFFA0B864FC1574819CD0FADF" authority="Ellingsen, 1910" authorityName="Ellingsen" authorityYear="1910" class="Arachnida" family="Syarinidae" genus="Microcreagrina" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="B95BEA9AFFA0B864FC157481991CFAF5" box="[989,1299,189,214]" italics="true" pageId="11" pageNumber="52">Microcreagrina hispanica</emphasis>
(Ellingsen, 1910)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>