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<document id="0F7BEF46ACE8FF3D936B87A148CC55BC" ID-CLB-Dataset="25496" ID-DOI="10.11646/zootaxa.4668.1.10" ID-GBIF-Dataset="3b51725f-e873-48d2-918c-bb926e54a7b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3448971" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568897291135" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="FC7187BEA467001992F269840ABBFEDC" docLanguage="en" docName="zootaxa.4668.1.10.pdf" docOrigin="Zootaxa 4668 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marcepania princesa Jałoszyński 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="143" masterDocId="0048FFC6A466001A92656A550958FF9E" masterDocTitle="The first Philippine species of Marcepania (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="144" masterPageNumber="141" pageNumber="142" updateTime="1698755374918" updateUser="ExternalLinkService">
<mods:mods id="5C2A12B3B0A7147D227F14771FFA3520" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B1BF47A304CBBE74A1F592F436509CE0">
<mods:title id="8D7C8A5A17D2EE7FDC3870A4C9B9BC1B">The first Philippine species of Marcepania (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="7231F9003FDBFD44A99F72DF3DE0E74B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="93BBA1E38687FD6BC415748A44707F7A">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="F25C469AC65F3AE3574AF4A4EED19BEA">text</mods:typeOfResource>
<mods:relatedItem id="89BB3F3F606DF917FAA410C25166ECD5" type="host">
<mods:titleInfo id="6A2548365E06896D83F50FB6E3A52F74">
<mods:title id="EEE273DA93AA2014076229C231B5C1C8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D1BBF1892BCA51BE59E8500EBE6023B1">
<mods:date id="37C2FD04241700949A1F25B360A1EFCD">2019</mods:date>
<mods:detail id="7CB6D9943350290D404AB0B5C755E4E4" type="pubDate">
<mods:number id="13A51466A0A8B18D9BE1E59FD4DA2EE2">2019-09-09</mods:number>
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<mods:number id="309F9324B357AA54A5FA6C4DDC351605">4668</mods:number>
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<mods:number id="808B674AD0120020B33A37A5918F93DB">1</mods:number>
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<mods:classification id="1EC838BF430BAF690F797D7555EA0144">journal article</mods:classification>
<mods:identifier id="4D15B66FBECE419CE0A0F3B0B47D60B5" type="CLB-Dataset">25496</mods:identifier>
<mods:identifier id="1F9DCF03EE34D8D9E738370213FC1819" type="DOI">10.11646/zootaxa.4668.1.10</mods:identifier>
<mods:identifier id="632C60522EFDA2D891799035E4056541" type="GBIF-Dataset">3b51725f-e873-48d2-918c-bb926e54a7b7</mods:identifier>
<mods:identifier id="1816274453B1D37B1C8657C1D1743669" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B837877918EDAE89CCC187008EB34EDE" type="Zenodo-Dep">3448971</mods:identifier>
</mods:mods>
<treatment id="FC7187BEA467001992F269840ABBFEDC" ID-GBIF-Taxon="159405168" LSID="urn:lsid:plazi:treatment:FC7187BEA467001992F269840ABBFEDC" httpUri="http://treatment.plazi.org/id/FC7187BEA467001992F269840ABBFEDC" lastPageId="3" lastPageNumber="143" pageId="1" pageNumber="142">
<subSubSection id="3CC26523A467001B92F269840B3CFC75" box="[151,612,977,1004]" pageId="1" pageNumber="142" type="nomenclature">
<paragraph id="746736A8A467001B92F269840B3CFC75" blockId="1.[151,1437,977,2011]" box="[151,612,977,1004]" pageId="1" pageNumber="142">
<heading id="2F2F81C4A467001B92F269840B3CFC75" bold="true" box="[151,612,977,1004]" fontSize="11" level="1" pageId="1" pageNumber="142" reason="1">
<emphasis id="46ACEABAA467001B92F269840B3CFC75" bold="true" box="[151,612,977,1004]" pageId="1" pageNumber="142">
<taxonomicName id="B3D84D2BA467001B92F2698408C0FC75" ID-CoL="8Y67Q" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,408,977,1003]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="princesa" status="sp. nov.">
<emphasis id="46ACEABAA467001B92F2698408C0FC75" bold="true" box="[151,408,977,1003]" italics="true" pageId="1" pageNumber="142">Marcepania princesa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5D9F57C1A467001B93FA698708B9FC72" box="[415,481,978,1004]" pageId="1" pageNumber="142" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="ECE32A2DA467001B939569840B02FC75" box="[496,602,977,1003]" captionStart="FIGURES 1–4" captionStartId="2.[151,258,1863,1885]" captionTargetBox="[160,1428,265,1843]" captionTargetId="figure@2.[160,1428,265,1843]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus (1) and aedeagus in ventral view (2–3) of Marcepania princesa sp. n.; distribution of Marcepania species (4)." figureDoi="http://doi.org/10.5281/zenodo.3448973" httpUri="https://zenodo.org/record/3448973/files/figure.png" pageId="1" pageNumber="142">Figs 1–4</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3CC26523A467001B92F269A20BCDFBC9" pageId="1" pageNumber="142" type="materials_examined">
<paragraph id="746736A8A467001B92F269A20BCDFBC9" blockId="1.[151,1437,977,2011]" pageId="1" pageNumber="142">
<emphasis id="46ACEABAA467001B92F269A20866FB91" bold="true" box="[151,318,1015,1039]" pageId="1" pageNumber="142">Type material.</emphasis>
<materialsCitation id="C4B03CF5A467001B932669A20BCDFBC9" ID-GBIF-Occurrence="2416711684" collectingDate="2001-07-15" collectingDateMax="2018-10" collectingDateMin="2001-07-15" collectionCode="NHMW" collectorName="P. Princesa &amp; Freitag &amp; P. Jaloszynski" country="Philippines" latitude="10.162666" location="Maratarpi" longLatPrecision="1" longitude="118.83234" pageId="1" pageNumber="142" specimenCount="1" stateProvince="PALAWAN" typeStatus="holotype">
<emphasis id="46ACEABAA467001B932669A20A5BFB91" bold="true" box="[323,771,1015,1039]" pageId="1" pageNumber="142">
<typeStatus id="AB63880AA467001B932669A208F7FB91" box="[323,431,1015,1039]" pageId="1" pageNumber="142" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="0CCF7638A467001B93DC69A20B3DFB91" box="[441,613,1015,1039]" name="Philippines" pageId="1" pageNumber="142">PHILIPPINES</collectingCountry>
,
<collectingRegion id="B61CF84AA467001B901569A20BA6FB91" box="[624,766,1015,1039]" country="Philippines" name="Palawan" pageId="1" pageNumber="142">PALAWAN</collectingRegion>
:
</emphasis>
♁, two labels: &quot;PHIL.: Palawan,
<collectorName id="D92D537EA467001B960769A20D8FFB91" box="[1122,1239,1015,1039]" pageId="1" pageNumber="142">P. Princesa</collectorName>
/ SSW
<location id="71076073A467001B974369A20CD7FB91" LSID="urn:lsid:plazi:treatment:FC7187BEA467001992F269840ABBFEDC:71076073A467001B974369A20CD7FB91" box="[1318,1423,1015,1039]" country="Philippines" latitude="10.162666" longLatPrecision="1" longitude="118.83234" name="Maratarpi" pageId="1" pageNumber="142" stateProvince="PALAWAN">Maratarpi</location>
/
<geoCoordinate id="11EC506FA467001B92F26E4E0846FBAD" box="[151,286,1051,1075]" degrees="10" direction="north" minutes="09.760" orientation="latitude" pageId="1" pageNumber="142" precision="1" value="10.162666">10°09.760'N</geoCoordinate>
<geoCoordinate id="11EC506FA467001B93466E4E08ECFBAD" box="[291,436,1051,1075]" degrees="118" direction="east" minutes="49.940" orientation="longitude" pageId="1" pageNumber="142" precision="1" value="118.83234">118°49.940'E</geoCoordinate>
/
<date id="00661068A467001B93A06E4E0B6AFBAD" box="[453,562,1051,1075]" pageId="1" pageNumber="142" value="2001-07-15">
<collectingDate id="1022E980A467001B93A06E4E0B6AFBAD" box="[453,562,1051,1075]" pageId="1" pageNumber="142" value="2001-07-15">15.7.2001</collectingDate>
</date>
, leg.
<collectorName id="D92D537EA467001B900D6E4E0BECFBAD" box="[616,692,1051,1075]" pageId="1" pageNumber="142">Freitag</collectorName>
(
<date id="00661068A467001B90A46E4E0BB3FBAD" box="[705,747,1051,1075]" pageId="1" pageNumber="142" value="2018-10">
<collectingDate id="1022E980A467001B90A46E4E0BB3FBAD" box="[705,747,1051,1075]" pageId="1" pageNumber="142" value="2018-10">x18</collectingDate>
</date>
)&quot; [white, printed], &quot;
<taxonomicName id="B3D84D2BA467001B91D86E4E0D39FBAC" authorityName="Jaloszynski" authorityYear="2013" box="[957,1121,1051,1074]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="46ACEABAA467001B91D86E4E0D39FBAC" box="[957,1121,1051,1074]" italics="true" pageId="1" pageNumber="142">MARCEPANIA</emphasis>
</taxonomicName>
/
<taxonomicName id="B3D84D2BA467001B96166E490D97FBAD" authorityName="Jałoszyński" authorityYear="2019" box="[1139,1231,1052,1075]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="princesa">
<emphasis id="46ACEABAA467001B96166E490D97FBAD" box="[1139,1231,1052,1075]" italics="true" pageId="1" pageNumber="142">princesa</emphasis>
</taxonomicName>
m. /
<collectorName id="D92D537EA467001B97656E4E0CC1FBAD" box="[1280,1433,1051,1075]" pageId="1" pageNumber="142">P. Jałoszyński</collectorName>
, 2019 / HOLOTYPUS&quot; [red, printed] (
<collectionCode id="12C9AE6DA467001B90496E6A0BD2FBC9" box="[556,650,1087,1111]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="142" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3CC26523A467001B92A26E360BB5FB01" pageId="1" pageNumber="142" type="diagnosis">
<paragraph id="746736A8A467001B92A26E360BB5FB01" blockId="1.[151,1437,977,2011]" pageId="1" pageNumber="142">
<emphasis id="46ACEABAA467001B92A26E360864FBE5" bold="true" box="[199,316,1123,1147]" pageId="1" pageNumber="142">Diagnosis.</emphasis>
Body very slender, EI&gt; 1.50; aedeagus elongate, pear-shaped, with a darkly sclerotized pair of slender median components in subapical region of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="3CC26523A467001B92A26EFE08CCF80D" pageId="1" pageNumber="142" type="description">
<paragraph id="746736A8A467001B92A26EFE0990FB78" blockId="1.[151,1437,977,2011]" pageId="1" pageNumber="142">
<emphasis id="46ACEABAA467001B92A26EFE080AFB5D" bold="true" box="[199,338,1195,1219]" pageId="1" pageNumber="142">Description.</emphasis>
Body of male (
<figureCitation id="ECE32A2DA467001B93916EFE0B69FB5D" box="[500,561,1195,1219]" captionStart="FIGURES 1–4" captionStartId="2.[151,258,1863,1885]" captionTargetBox="[160,1428,265,1843]" captionTargetId="figure@2.[160,1428,265,1843]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus (1) and aedeagus in ventral view (2–3) of Marcepania princesa sp. n.; distribution of Marcepania species (4)." figureDoi="http://doi.org/10.5281/zenodo.3448973" httpUri="https://zenodo.org/record/3448973/files/figure.png" pageId="1" pageNumber="142">Fig. 1</figureCitation>
) elongate and slender, flattened, brown, covered with yellowish vestiture; BL
<quantity id="B3209B4DA467001B97086EFE099CFB78" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="142" unit="mm" value="0.81">0.81 mm</quantity>
.
</paragraph>
<paragraph id="746736A8A467001B92A26EA60AD6FA20" blockId="1.[151,1437,977,2011]" pageId="1" pageNumber="142">
Head broadest at large, strongly convex, bean-shaped and coarsely faceted eyes, HL
<quantity id="B3209B4DA467001B96356EA60DE9FA94" box="[1104,1201,1267,1291]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="142" unit="mm" value="0.1">0.10 mm</quantity>
, HW 0.18; vertex and posterior part of frons confluent and weakly convex; anterior portion of frons flattened; supraantennal tubercles barely marked; frons between eyes and vertex nearly impunctate, frons on sides with fine but dense, inconspicuous punctures; setae sparse and short, suberect. Antennae short, AnL
<quantity id="B3209B4DA467001B90946F0A0A0FFAE8" box="[753,855,1375,1399]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="142" unit="mm" value="0.3">0.30 mm</quantity>
; antennomeres I–II strongly elongate; III indistinctly transverse, IV–VI each about as long as broad; VII–VIII each slightly transverse; IX and X each distinctly transverse, XI only slightly longer than broad, with evenly and broadly rounded apex.
</paragraph>
<paragraph id="746736A8A467001B92A26F9E08D2F9D1" blockId="1.[151,1437,977,2011]" pageId="1" pageNumber="142">
Pronotum nearly semi-oval, broadest slightly in front of middle; PL
<quantity id="B3209B4DA467001B91C56F9E0D5AFA7C" box="[928,1026,1483,1507]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="142" unit="mm" value="0.24">0.24 mm</quantity>
, PW
<quantity id="B3209B4DA467001B965E6F9E0DC6FA7C" box="[1083,1182,1483,1507]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="142" unit="mm" value="0.28">0.28 mm</quantity>
. Anterior margin nearly straight; lateral margins weakly and nearly evenly rounded; hind corners nearly right-angled; posterior margin weakly arcuate; lateral ante-basal impressions shallow but distinct. Punctures on disc fine and shallow, inconspicuous; vestiture sparse, short, suberect.
</paragraph>
<paragraph id="746736A8A467001B92A26C0E0B0BF941" blockId="1.[151,1437,977,2011]" pageId="1" pageNumber="142">
Elytra together oval, broadest near anterior fourth; EL
<quantity id="B3209B4DA467001B917D6C0E0A23F9EC" box="[792,891,1627,1651]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="142" unit="mm" value="0.48">0.48 mm</quantity>
, EW
<quantity id="B3209B4DA467001B91DD6C0E0D43F9EC" box="[952,1051,1627,1651]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="142" unit="mm" value="0.31">0.31 mm</quantity>
, EI 1.52; apices broadly, separately rounded; punctures more distinct than those on pronotum but still shallow and inconspicuous, those on anterior half separated by spaces shorter than diameters of punctures, punctures reducing in diameter and depth toward sides and apices; setae short, sparse, only slightly suberect.
</paragraph>
<paragraph id="746736A8A467001B92A26CBE0B40F89D" blockId="1.[151,1437,977,2011]" box="[199,536,1771,1795]" pageId="1" pageNumber="142">Legs short, robust; unmodified.</paragraph>
<paragraph id="746736A8A467001B92A26D5A0B0FF8F0" blockId="1.[151,1437,977,2011]" pageId="1" pageNumber="142">
Aedeagus (
<figureCitation id="ECE32A2DA467001B93256D5A08F9F8B9" box="[320,417,1807,1831]" captionStart="FIGURES 1–4" captionStartId="2.[151,258,1863,1885]" captionTargetBox="[160,1428,265,1843]" captionTargetId="figure@2.[160,1428,265,1843]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus (1) and aedeagus in ventral view (2–3) of Marcepania princesa sp. n.; distribution of Marcepania species (4)." figureDoi="http://doi.org/10.5281/zenodo.3448973" httpUri="https://zenodo.org/record/3448973/files/figure.png" pageId="1" pageNumber="142">Figs 2–3</figureCitation>
) pear-shaped, broadest in sub-basal region, apical margin broadly and weakly rounded; endophallus with a pair of slender median sclerites in subapical region; parameres slender, exceeding apex of median lobe, each with one apical seta; AeL
<quantity id="B3209B4DA467001B93866D020B0BF8F0" box="[483,595,1879,1903]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="142" unit="mm" value="0.125">0.125 mm</quantity>
.
</paragraph>
<paragraph id="746736A8A467001B92A26D2E08CCF80D" blockId="1.[151,1437,977,2011]" box="[199,404,1915,1939]" pageId="1" pageNumber="142">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="3CC26523A467001B92A26DCA0BBEF829" box="[199,742,1951,1975]" pageId="1" pageNumber="142" type="distribution">
<paragraph id="746736A8A467001B92A26DCA0BBEF829" blockId="1.[151,1437,977,2011]" box="[199,742,1951,1975]" pageId="1" pageNumber="142">
<emphasis id="46ACEABAA467001B92A26DCA0802F829" bold="true" box="[199,346,1951,1975]" pageId="1" pageNumber="142">Distribution.</emphasis>
<collectingCountry id="0CCF7638A467001B93056DCA0883F829" box="[352,475,1951,1975]" name="Philippines" pageId="1" pageNumber="142">Philippines</collectingCountry>
, central
<collectingRegion id="B61CF84AA467001B90516DCA0BD6F829" box="[564,654,1951,1975]" country="Philippines" name="Palawan" pageId="1" pageNumber="142">Palawan</collectingRegion>
(
<figureCitation id="ECE32A2DA467001B90F86DCA0B81F829" box="[669,729,1951,1975]" captionStart="FIGURES 1–4" captionStartId="2.[151,258,1863,1885]" captionTargetBox="[160,1428,265,1843]" captionTargetId="figure@2.[160,1428,265,1843]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus (1) and aedeagus in ventral view (2–3) of Marcepania princesa sp. n.; distribution of Marcepania species (4)." figureDoi="http://doi.org/10.5281/zenodo.3448973" httpUri="https://zenodo.org/record/3448973/files/figure.png" pageId="1" pageNumber="142">Fig 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3CC26523A467001892A26D960980FF64" lastPageId="2" lastPageNumber="143" pageId="1" pageNumber="142" type="etymology">
<paragraph id="746736A8A467001892A26D960980FF64" blockId="1.[151,1437,977,2011]" lastBlockId="2.[151,1436,154,250]" lastPageId="2" lastPageNumber="143" pageId="1" pageNumber="142">
<emphasis id="46ACEABAA467001B92A26D960810F845" bold="true" box="[199,328,1987,2011]" pageId="1" pageNumber="142">Etymology.</emphasis>
Locotypical, after
<collectingRegion id="B61CF84AA467001B90706D960BAAF845" box="[533,754,1987,2011]" country="Philippines" name="Puerto Princesa" pageId="1" pageNumber="142">Puerto Princesa City</collectingRegion>
(used as a noun in apposition). Coincidentally, &quot;Princessa&quot; is a Polish chocolate bar of the same etymology as the Philippine city (except that not dedicated to any particular princess), which combines well with the generic name, as
<taxonomicName id="B3D84D2BA464001890F06AEB0A4EFF4B" authorityName="Jaloszynski" authorityYear="2013" box="[661,790,190,213]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="46ACEABAA464001890F06AEB0A4EFF4B" box="[661,790,190,213]" italics="true" pageId="2" pageNumber="143">Marcepania</emphasis>
</taxonomicName>
has been derived from the Polish word 'marcepan', i.e., mar- zipan.
</paragraph>
</subSubSection>
<caption id="20A76620A464001892F26D12085DF81C" ID-DOI="http://doi.org/10.5281/zenodo.3448973" ID-Zenodo-Dep="3448973" httpUri="https://zenodo.org/record/3448973/files/figure.png" pageId="2" pageNumber="143" startId="2.[151,258,1863,1885]" targetBox="[160,1428,265,1843]" targetPageId="2">
<paragraph id="746736A8A464001892F26D12085DF81C" blockId="2.[151,1436,1863,1922]" pageId="2" pageNumber="143">
<emphasis id="46ACEABAA464001892F26D12086CF8C2" bold="true" box="[151,308,1863,1885]" pageId="2" pageNumber="143">FIGURES 1–4.</emphasis>
Dorsal habitus (1) and aedeagus in ventral view (2–3) of
<taxonomicName id="B3D84D2BA464001891106D1D0D1DF8C3" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[885,1093,1864,1885]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="princesa" status="sp. nov.">
<emphasis id="46ACEABAA464001891106D1D0D1DF8C3" box="[885,1093,1864,1885]" italics="true" pageId="2" pageNumber="143">Marcepania princesa</emphasis>
</taxonomicName>
<emphasis id="46ACEABAA464001896286D1D0DDEF8C3" bold="true" box="[1101,1158,1864,1885]" pageId="2" pageNumber="143">
<taxonomicNameLabel id="5D9F57C1A464001896286D1D0DDEF8C3" box="[1101,1158,1864,1885]" pageId="2" pageNumber="143" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; distribution of
<taxonomicName id="B3D84D2BA464001897426D1D0CC4F8C3" authorityName="Jaloszynski" authorityYear="2013" box="[1319,1436,1864,1885]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="46ACEABAA464001897426D1D0CC4F8C3" box="[1319,1436,1864,1885]" italics="true" pageId="2" pageNumber="143">Marcepania</emphasis>
</taxonomicName>
species (4).
</paragraph>
</caption>
<subSubSection id="3CC26523A464001992A26DE70ABBFEDC" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143" type="discussion">
<paragraph id="746736A8A464001992A26DE70ABBFEDC" blockId="2.[151,1437,1970,2030]" lastBlockId="3.[151,1437,154,322]" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143">
<emphasis id="46ACEABAA464001892A26DE7086EF854" bold="true" box="[199,310,1970,1994]" pageId="2" pageNumber="143">Remarks.</emphasis>
Species of
<taxonomicName id="B3D84D2BA464001893D46DE60B6AF854" authorityName="Jaloszynski" authorityYear="2013" box="[433,562,1971,1994]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="46ACEABAA464001893D46DE60B6AF854" box="[433,562,1971,1994]" italics="true" pageId="2" pageNumber="143">Marcepania</emphasis>
</taxonomicName>
are morphologically very uniform, including male genital structures. They can be distinguished by differences in proportions of body parts, the general shape of the aedeagus, and, to a lesser extent, by weakly sclerotized and often poorly visible endophallic structures.
<taxonomicName id="B3D84D2BA465001991006ACF0D11FF2F" authorityName="Jałoszyński" authorityYear="2019" box="[869,1097,154,177]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="princesa">
<emphasis id="46ACEABAA465001991006ACF0D11FF2F" box="[869,1097,154,177]" italics="true" pageId="3" pageNumber="144">Marcepania princesa</emphasis>
</taxonomicName>
resembles most its closest geographical relative,
<taxonomicName id="B3D84D2BA465001993396AEB0883FF48" box="[348,475,190,214]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="46ACEABAA465001993396AEB0883FF48" box="[348,475,190,214]" italics="true" pageId="3" pageNumber="144">M. elongata</emphasis>
</taxonomicName>
, known to occur in the northern region of Borneo. These are the only such slender, strongly elongate species with the elytral indices exceeding 1.50.
<taxonomicName id="B3D84D2BA465001990916AB70A80FF67" authorityName="Jałoszyński" authorityYear="2019" box="[756,984,226,249]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="princesa">
<emphasis id="46ACEABAA465001990916AB70A80FF67" box="[756,984,226,249]" italics="true" pageId="3" pageNumber="144">Marcepania princesa</emphasis>
</taxonomicName>
is distinctly smaller than
<taxonomicName id="B3D84D2BA4650019968E6AB70C33FF64" box="[1259,1387,226,250]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="46ACEABAA4650019968E6AB70C33FF64" box="[1259,1387,226,250]" italics="true" pageId="3" pageNumber="144">M. elongata</emphasis>
</taxonomicName>
(BL
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vs.
<quantity id="B3209B4DA465001993406B5008DFFE83" box="[293,391,261,285]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="144" unit="mm" value="0.95">0.95 mm</quantity>
, respectively), and its aedeagus is much more elongate, ~ 1.7 × as long as broad in ventral view, whereas in
<taxonomicName id="B3D84D2BA465001993746B7F08C9FEDC" box="[273,401,298,322]" class="Insecta" family="Staphylinidae" genus="Marcepania" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="46ACEABAA465001993746B7F08C9FEDC" box="[273,401,298,322]" italics="true" pageId="3" pageNumber="144">M. elongata</emphasis>
</taxonomicName>
it is conspicuously stout, only ~ 1.3 × as long as broad.
</paragraph>
</subSubSection>
</treatment>
</document>
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