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<document id="5E49D0176C7445D037ED7D01ECECE454" ID="10.11646/zootaxa.4254.1.10SLASH10789" ID-CLB-Dataset="33243" ID-DOI="10.11646/zootaxa.4254.1.10" ID-GBIF-Dataset="c5b23ffc-6b96-4686-aba1-3ecdb21f618b" ID-ISSN="1175-5326" ID-Zenodo-Dep="545752" ID-ZooBank="FD092AA7-4407-41A4-A402-8E887EC54DC6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1491985124231" checkinUser="plazi" docAuthor="Nascimento, Dariane Amorim Do, Mendonça, Maria Thayane Da Silva &amp; Fernandes, Jose Antonio Marin" docDate="2017" docId="755387CAFFB4BA6F6CFA04D2FD00297E" docLanguage="en" docName="zootaxa.4254.1.10SLASH10789.pdf" docOrigin="Zootaxa 4254 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Edessa rugulata Nascimento, Mendonça &amp; Fernandes, 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="143" masterDocId="896AFFB2FFB3BA686C6D0445FFBD2A01" masterDocTitle="Description of a new group of species of Edessa (Hemiptera: Pentatomidae: Edessinae)" masterLastPageNumber="150" masterPageNumber="136" pageNumber="143" updateTime="1698438871090" updateUser="ExternalLinkService">
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<mods:title id="F87A5F051EAB7DDC252DCFFB31A1BE18">Description of a new group of species of Edessa (Hemiptera: Pentatomidae: Edessinae)</mods:title>
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<mods:namePart id="D5B51F6FE2CB0205E6EF99DABED2DD94">Nascimento, Dariane Amorim Do</mods:namePart>
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<mods:namePart id="F7684C3CCFB1B613281FAA1ADF8D7053">Mendonça, Maria Thayane Da Silva</mods:namePart>
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<mods:namePart id="8AAC09EFE75F036F486665BE8C9F57E2">Fernandes, Jose Antonio Marin</mods:namePart>
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<mods:title id="54DC63DA913DADF7BE522FBFA9C2AE2F">Zootaxa</mods:title>
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<mods:date id="9D050999F3DBB953295EA69F06B6A2FD">2017</mods:date>
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<treatment id="755387CAFFB4BA6F6CFA04D2FD00297E" ID-DOI="http://doi.org/10.5281/zenodo.6000621" ID-GBIF-Taxon="128347668" ID-Zenodo-Dep="6000621" LSID="urn:lsid:plazi:treatment:755387CAFFB4BA6F6CFA04D2FD00297E" httpUri="http://treatment.plazi.org/id/755387CAFFB4BA6F6CFA04D2FD00297E" lastPageNumber="143" pageId="7" pageNumber="143">
<subSubSection id="B5E06557FFB4BA6F6CFA04D2FE272AD2" pageId="7" pageNumber="143" type="nomenclature">
<paragraph id="FD4536DCFFB4BA6F6CFA04D2FE192AB3" blockId="7.[151,420,151,211]" box="[151,420,151,178]" pageId="7" pageNumber="143">
<heading id="A60D81B0FFB4BA6F6CFA04D2FE192AB3" bold="true" box="[151,420,151,178]" fontSize="11" level="1" pageId="7" pageNumber="143" reason="1">
<emphasis id="CF8EEACEFFB4BA6F6CFA04D2FE192AB3" bold="true" box="[151,420,151,178]" pageId="7" pageNumber="143">
<taxonomicName id="3AFA4D5FFFB4BA6F6CFA04D2FEE62AB0" box="[151,347,151,177]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="143" phylum="Arthropoda" rank="species" species="rugulata" status="sp. nov.">
<emphasis id="CF8EEACEFFB4BA6F6CFA04D2FEE62AB0" bold="true" box="[151,347,151,177]" italics="true" pageId="7" pageNumber="143">Edessa rugulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4BD57B5FFB4BA6F6D0F04DDFE192AB3" box="[354,420,152,178]" pageId="7" pageNumber="143" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FD4536DCFFB4BA6F6CFA04FFFE272AD2" blockId="7.[151,420,151,211]" box="[151,410,186,211]" pageId="7" pageNumber="143">
(
<figureCitation id="65C12A59FFB4BA6F6CF204FFFF5A2AD2" box="[159,231,186,211]" captionStart="FIGURE 4" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[230,1378,230,1829]" captionTargetId="figure@8.[200,1387,193,1938]" captionTargetPageId="8" captionText="FIGURE 4. A D. Edessa rugulata sp. n.: A C, pygophore in dorsal, ventral and posterior view, respectively; D, metasternal process. Scale 1 mm." httpUri="https://zenodo.org/record/545756/files/figure.png" pageId="7" pageNumber="143">Figs 4</figureCitation>
,
<figureCitation id="65C12A59FFB4BA6F6C9E04FEFEFD2AD2" box="[243,320,187,211]" captionStart="FIGURE 7" captionStartId="11.[151,250,1959,1981]" captionTargetBox="[218,1327,208,1924]" captionTargetId="figure@11.[200,1387,193,1938]" captionTargetPageId="11" captionText="FIGURE 7. A D. Dorsal and ventral view: A B, Edessa rugulata sp. n.; C D, Edessa eucnema sp. n.. Scale 5 mm." httpUri="https://zenodo.org/record/545759/files/figure.png" pageId="7" pageNumber="143">7 AB</figureCitation>
,
<figureCitation id="65C12A59FFB4BA6F6D2604FEFECA2AD3" box="[331,375,187,211]" captionStart="FIGURE 8" captionStartId="12.[151,250,1959,1981]" captionTargetBox="[212,1372,254,1875]" captionTargetId="figure@12.[200,1387,193,1938]" captionTargetPageId="12" captionText="FIGURE 8. A E. Peritreme: A, Edessa viridula sp. n.; B, Edessa exigusternata sp. n.; C, Edessa angusticlada sp. n.; D, Edessa rugulata sp. n.; E, Edessa eucnema sp. n.. Scale 1 mm." httpUri="https://zenodo.org/record/545760/files/figure.png" pageId="7" pageNumber="143">8 D</figureCitation>
,
<figureCitation id="65C12A59FFB4BA6F6DEE04FEFE2C2AD2" box="[387,401,187,211]" captionStart="FIGURE 9" captionStartId="13.[151,250,1948,1970]" captionTargetBox="[232,1255,303,1814]" captionTargetId="figure@13.[176,1410,193,2007]" captionTargetPageId="13" captionText="FIGURE 9. Distribution of the species of the viridula group." httpUri="https://zenodo.org/record/545761/files/figure.png" pageId="7" pageNumber="143">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B5E06557FFB4BA6F6CFA0547FE5D2B62" pageId="7" pageNumber="143" type="etymology">
<paragraph id="FD4536DCFFB4BA6F6CFA0547FB842B1B" blockId="7.[151,1437,258,895]" box="[151,1081,258,283]" pageId="7" pageNumber="143">
<emphasis id="CF8EEACEFFB4BA6F6CFA0547FEA32B1A" bold="true" box="[151,286,258,283]" pageId="7" pageNumber="143">Etymology.</emphasis>
Reference to the short peritreme of the species. Rugula—small crease.
</paragraph>
<paragraph id="FD4536DCFFB4BA6F6CAA0563FE5D2B62" blockId="7.[151,1437,258,895]" pageId="7" pageNumber="143">
<materialsCitation id="4D923C81FFB4BA6F6CAA0563FE5D2B62" ID-GBIF-Occurrence="1455870632" collectingDate="1954-12-10" collectionCode="USNM" collectorName="Luis E." country="Bolivia" latitude="-17.006283" location="Chapare" longLatPrecision="1" longitude="-65.64128" municipality="Cristal Mayu" pageId="7" pageNumber="143" specimenCount="1" stateProvince="Cochabamba" typeStatus="holotype">
<typeStatus id="2241887EFFB4BA6F6CAA0563FE922B3E" box="[199,303,294,319]" pageId="7" pageNumber="143" type="holotype">Holotype</typeStatus>
male.
<emphasis id="CF8EEACEFFB4BA6F6D110563FD162B3E" bold="true" box="[380,683,294,319]" pageId="7" pageNumber="143">
<collectingCountry id="85ED764CFFB4BA6F6D110563FE4A2B3E" box="[380,503,294,319]" name="Bolivia" pageId="7" pageNumber="143">BOLIVIA</collectingCountry>
,
<collectingRegion id="3F3EF83EFFB4BA6F6E690563FD1A2B3E" box="[516,679,294,319]" country="Bolivia" name="Cochabamba" pageId="7" pageNumber="143">Cochabamba</collectingRegion>
:
</emphasis>
<collectingMunicipality id="1D21ACA6FFB4BA6F6EDE0563FCF42B3E" box="[691,841,294,319]" pageId="7" pageNumber="143">Cristal Mayu</collectingMunicipality>
,
<location id="F8256007FFB4BA6F6F380563FC082B3E" LSID="urn:lsid:plazi:treatment:755387CAFFB4BA6F6CFA04D2FD00297E:F8256007FFB4BA6F6F380563FC082B3E" box="[853,949,294,319]" country="Bolivia" latitude="-17.006283" longLatPrecision="1" longitude="-65.64128" municipality="Cristal Mayu" name="Chapare" pageId="7" pageNumber="143" stateProvince="Cochabamba">Chapare</location>
,
<collectingDate id="9900E9F4FFB4BA6F6FAC0563FBE72B3E" box="[961,1114,294,319]" pageId="7" pageNumber="143" value="1954-12-10">
<date id="8944101CFFB4BA6F6FAC0563FBE72B3E" box="[961,1114,294,319]" pageId="7" pageNumber="143" value="1954-12-10">10XII1954</date>
</collectingDate>
,
<collectorName id="500F530AFFB4BA6F680B0563FB052B3F" box="[1126,1208,294,319]" pageId="7" pageNumber="143">Luis E.</collectorName>
Peña (
<geoCoordinate id="98CE501BFFB4BA6F69680563FF1A2B62" direction="south" orientation="latitude" pageId="7" pageNumber="143" precision="1" value="-17.006283">17°0022.62” S</geoCoordinate>
<geoCoordinate id="98CE501BFFB4BA6F6CC0050FFEDB2B62" box="[173,358,330,355]" direction="west" orientation="longitude" pageId="7" pageNumber="143" precision="1" value="-65.64128">65°3828.62” W</geoCoordinate>
) (
<collectionCode id="9BEBAE19FFB4BA6F6D11050EFE6E2B62" box="[380,467,331,355]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="143">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B5E06557FFB4BA6F6CAA052BFEAB2912" pageId="7" pageNumber="143" type="description">
<paragraph id="FD4536DCFFB4BA6F6CAA052BFD8B2BCE" blockId="7.[151,1437,258,895]" pageId="7" pageNumber="143">
<emphasis id="CF8EEACEFFB4BA6F6CAA052BFEC02B86" bold="true" box="[199,381,366,391]" pageId="7" pageNumber="143">Measurements.</emphasis>
Head length (1.4); head width (2.5); pronotum length (2.5); pronotum width (6.0); scutellum length (4.9); scutellum width (3.6); length of antennal segments (I: 0.7; II: 1.0; III: 1.0; IV: 1.5; V missing); total length (11.6); abdominal width (6.6).
</paragraph>
<paragraph id="FD4536DCFFB4BA6F6CAA059FFC8128A6" blockId="7.[151,1437,258,895]" pageId="7" pageNumber="143">
<emphasis id="CF8EEACEFFB4BA6F6CAA059FFE862BF2" bold="true" box="[199,315,474,499]" pageId="7" pageNumber="143">Diagnose.</emphasis>
Pronotum rugose. Arms of the anterior bifurcation of the metasternal process clavate (
<figureCitation id="65C12A59FFB4BA6F6945059FFA322BF2" box="[1320,1423,474,499]" captionStart="FIGURE 4" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[230,1378,230,1829]" captionTargetId="figure@8.[200,1387,193,1938]" captionTargetPageId="8" captionText="FIGURE 4. A D. Edessa rugulata sp. n.: A C, pygophore in dorsal, ventral and posterior view, respectively; D, metasternal process. Scale 1 mm." httpUri="https://zenodo.org/record/545756/files/figure.png" pageId="7" pageNumber="143">Fig. 4 D</figureCitation>
); excavation short receiving part of the fourth rostral segment. Peritreme reaching 1/3 of the distance between ostiole of the scent gland and lateral margin of the body (
<figureCitation id="65C12A59FFB4BA6F6E9A0667FCDF283A" box="[759,866,546,571]" captionStart="FIGURE 8" captionStartId="12.[151,250,1959,1981]" captionTargetBox="[212,1372,254,1875]" captionTargetId="figure@12.[200,1387,193,1938]" captionTargetPageId="12" captionText="FIGURE 8. A E. Peritreme: A, Edessa viridula sp. n.; B, Edessa exigusternata sp. n.; C, Edessa angusticlada sp. n.; D, Edessa rugulata sp. n.; E, Edessa eucnema sp. n.. Scale 1 mm." httpUri="https://zenodo.org/record/545760/files/figure.png" pageId="7" pageNumber="143">Fig. 8 D</figureCitation>
). Evaporatorium with a long and raised crest in mesopleuron delimited by sulci; contiguous margin of meso and metapleuron also raised; metapleuron with two distinct raised crests one short medial and the other long distal. Spiracles concolorous with ventral surface. Trichobothria in yellowish calli. Antennae and legs spotless.
</paragraph>
<paragraph id="FD4536DCFFB4BA6F6CAA06F7FEAB2912" blockId="7.[151,1437,258,895]" pageId="7" pageNumber="143">
<emphasis id="CF8EEACEFFB4BA6F6CAA06F7FEB928CA" bold="true" box="[199,260,690,715]" pageId="7" pageNumber="143">Male</emphasis>
(
<figureCitation id="65C12A59FFB4BA6F6D7906F7FE2628CA" box="[276,411,690,715]" captionStart="FIGURE 4" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[230,1378,230,1829]" captionTargetId="figure@8.[200,1387,193,1938]" captionTargetPageId="8" captionText="FIGURE 4. A D. Edessa rugulata sp. n.: A C, pygophore in dorsal, ventral and posterior view, respectively; D, metasternal process. Scale 1 mm." httpUri="https://zenodo.org/record/545756/files/figure.png" pageId="7" pageNumber="143">Figs 4 AC</figureCitation>
). Superior process of the genital cup reduced to a black line. Ventral part of the paramere with one barely distinct and rounded lateral lobe, base barely wider than distal part; lobes not distinctly projected posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="B5E06557FFB4BA6F6CAA075BFC1B295B" pageId="7" pageNumber="143" type="discussion">
<paragraph id="FD4536DCFFB4BA6F6CAA075BFC1B295B" blockId="7.[151,1437,258,895]" pageId="7" pageNumber="143">
<emphasis id="CF8EEACEFFB4BA6F6CAA075BFEF22936" bold="true" box="[199,335,798,823]" pageId="7" pageNumber="143">Comments.</emphasis>
Female unknown. This species is very similar to the others has the shortest peritreme ruga in the group and is the only one with claviform arms of metasternal process.
</paragraph>
</subSubSection>
<subSubSection id="B5E06557FFB4BA6F6CAA0723FD00297E" box="[199,701,870,895]" pageId="7" pageNumber="143" type="distribution">
<paragraph id="FD4536DCFFB4BA6F6CAA0723FD00297E" blockId="7.[151,1437,258,895]" box="[199,701,870,895]" pageId="7" pageNumber="143">
<emphasis id="CF8EEACEFFB4BA6F6CAA0723FEE4297E" bold="true" box="[199,345,870,895]" pageId="7" pageNumber="143">Distribution</emphasis>
(
<figureCitation id="65C12A59FFB4BA6F6D050723FE10297E" box="[360,429,870,895]" captionStart="FIGURE 9" captionStartId="13.[151,250,1948,1970]" captionTargetBox="[232,1255,303,1814]" captionTargetId="figure@13.[176,1410,193,2007]" captionTargetPageId="13" captionText="FIGURE 9. Distribution of the species of the viridula group." httpUri="https://zenodo.org/record/545761/files/figure.png" pageId="7" pageNumber="143">Fig. 9</figureCitation>
).
<collectingCountry id="85ED764CFFB4BA6F6DAF0723FDA4297E" box="[450,537,870,895]" name="Bolivia" pageId="7" pageNumber="143">Bolivia</collectingCountry>
:
<collectingRegion id="3F3EF83EFFB4BA6F6E4E0723FD04297E" box="[547,697,870,895]" country="Bolivia" name="Cochabamba" pageId="7" pageNumber="143">Cochabamba</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>