1026 lines
146 KiB
XML
1026 lines
146 KiB
XML
<document id="ECBA703427A86A0B442735BECCCEE57A" ID-DOI="10.5852/ejt.2011.3" ID-GBIF-Dataset="112f950a-a2bb-42bc-bece-1a8a75ce9b02" ID-ISSN="2118-9773" ID-Zenodo-Dep="3711435" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1584284816757" checkinUser="plazi" docAuthor="Kok, Philippe J. R. & Rivas, Gilson A." docDate="2011" docId="327B87E66162FFCBFDB4447DFC5CFCEB" docLanguage="en" docName="ejt-2011-3.pdf.imf" docOrigin="European Journal of Taxonomy 3" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Anadia mcdiarmidi Kok & Rivas 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="CE42FF9E6160FFC5FFC04156FFBAFFA4" masterDocTitle="A new species of Anadia (Reptilia, Squamata) from the Venezuelan ' Lost World', northern South America" masterLastPageNumber="18" masterPageNumber="1" pageNumber="3" updateTime="1738173478122" updateUser="ExternalLinkService">
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<mods:mods id="433279B189178259F83C2483116F9EC9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="57398F5936D91B30A5E2BCD86B88AA22">
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<mods:title id="096B94125D3C07535F1E0ED59C62B40F">A new species of Anadia (Reptilia, Squamata) from the Venezuelan ' Lost World', northern South America</mods:title>
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<mods:name id="1E882CC6FB168032AE7D82AE63DEE804" type="personal">
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<mods:roleTerm id="D462107A60BFC3F5D9058BC1A781B9FB">Author</mods:roleTerm>
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<mods:namePart id="CDA727DDDB2D671238DB7366D60F1751">Kok, Philippe J. R.</mods:namePart>
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<mods:affiliation id="1DE06D57CA0208AED0967EF3FE2450CC">Philippe.</mods:affiliation>
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<mods:nameIdentifier id="EE21EAF565249E3C1235641415C99E94" type="email">Philippe.Kok@vub.ac.be</mods:nameIdentifier>
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<mods:roleTerm id="90683840EBE33DF999322A369B1F8E9D">Author</mods:roleTerm>
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<mods:namePart id="CA043B393F3CC7D2E443F018F8F9C71E">Rivas, Gilson A.</mods:namePart>
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<mods:typeOfResource id="D22778EAEB3F9CA4508AE2BF60B5786A">text</mods:typeOfResource>
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<mods:title id="71661BC97BBC80B81D1514A56136358E">European Journal of Taxonomy</mods:title>
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<mods:part id="2D045D3E2A63DC8B0F20286A3584BAB7">
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<mods:date id="03A3F51E584DA407CC3EBF8183803AFF">2011</mods:date>
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<mods:number id="C78B38F33D1E85D072E413A2421CD246">2011-09-29</mods:number>
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<mods:number id="3291D55310B11929E8792EAF9302266D">3</mods:number>
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<mods:classification id="ECB90C23A6BBF771A3FC76363A81D27C">journal article</mods:classification>
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<mods:identifier id="8BEECF43689F122726D89C55B462092C" type="DOI">10.5852/ejt.2011.3</mods:identifier>
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<mods:identifier id="21AB63789A8FF3087CAEAC7806C4ADDB" type="ISSN">2118-9773</mods:identifier>
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<treatment id="327B87E66162FFCBFDB4447DFC5CFCEB" ID-DOI="http://doi.org/10.5281/zenodo.3717557" ID-GBIF-Taxon="162514291" ID-Zenodo-Dep="3717557" LSID="urn:lsid:zoobank.org:act:7BB5317B-BE36-41A4-B34E-8A66EBDB0FCC" httpUri="http://treatment.plazi.org/id/327B87E66162FFCBFDB4447DFC5CFCEB" lastPageId="14" lastPageNumber="15" pageId="2" pageNumber="3" scope_class="Squamata">
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<subSubSection id="F2C8657B6162FFC7FDB4447DFC7AFAE2" box="[628,960,1323,1350]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="BA6D36F06162FFC7FDB4447DFC7AFAE2" blockId="2.[628,960,1323,1389]" box="[628,960,1323,1350]" pageId="2" pageNumber="3">
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<heading id="E125819C6162FFC7FDB4447DFC7AFAE2" box="[628,960,1323,1350]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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<taxonomicName id="7DD24D736162FFC7FDB4447DFCE4FAE2" authority="Kok & Rivas, 2011" authorityName="Kok & Rivas" authorityYear="2011" box="[628,862,1323,1350]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="mcdiarmidi" status="sp. nov.">
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<emphasis id="88A6EAE26162FFC7FDB4447DFCE4FAE2" bold="true" box="[628,862,1323,1350]" italics="true" pageId="2" pageNumber="3">Anadia mcdiarmidi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="939557996162FFC7FCA5447AFC7AFAE2" box="[869,960,1324,1350]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="F2C8657B6162FFC7FD774405FCC7FAC9" box="[695,893,1363,1389]" pageId="2" pageNumber="3" type="description">
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<paragraph id="BA6D36F06162FFC7FD774405FCC7FAC9" blockId="2.[628,960,1323,1389]" box="[695,893,1363,1389]" pageId="2" pageNumber="3">
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<figureCitation id="22E92A756162FFC7FD774405FCA3FAC9" box="[695,793,1363,1389]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="3.[189,232,1895,1921]" captionStartId-1="4.[189,232,1965,1991]" captionStartId-2="5.[189,232,1930,1956]" captionStartId-3="6.[189,232,1000,1026]" captionStartId-4="6.[189,232,1824,1850]" captionStartId-5="8.[189,232,1930,1956]" captionStartId-6="9.[189,232,1930,1956]" captionTargetBox-0="[207,1380,265,1855]" captionTargetBox-1="[189,1372,1072,1925]" captionTargetBox-2="[189,1399,467,1890]" captionTargetBox-3="[189,1399,1085,1787]" captionTargetBox-4="[189,1399,1085,1787]" captionTargetBox-5="[248,1389,501,1830]" captionTargetBox-6="[219,1380,874,1814]" captionTargetId-0="figure@3.[207,1380,265,1855]" captionTargetId-1="figure@4.[189,1398,1072,1925]" captionTargetId-2="figure@5.[189,1399,467,1890]" captionTargetId-3="figure@6.[189,1399,1099,1787]" captionTargetId-4="figure@6.[189,1399,1099,1787]" captionTargetId-5="figure@8.[189,1398,462,1889]" captionTargetId-6="figure@9.[207,1382,855,1890]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="Fig. 1. Anadia mcdiarmidi sp. nov. A. IRSNB 2677, ♂ holotype in life. B. IRSNB 2675, juvenile male paratype in life. C. Ventral view of the anaesthetized holotype (grid squares = 5 mm). D. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok." captionText-1="Fig. 2. Anadia mcdiarmidi sp. nov. Ventral, dorsal, and lateral views of the head of the holotype (IRSNB 2677). Scale lines = 5 mm. Photographs by Philippe J.R. Kok." captionText-2="Fig. 3. Anadia mcdiarmidi sp. nov. Subpelvic region of the preserved ♂ holotype (IRSNB 2677), showing preanal scales and femoral pores, and trunk of the preserved holotype in lateral view (head to left), showing scalation at midbody. Scale lines = 5 mm. Photographs by Philippe J.R. Kok." captionText-3="Fig. 4. Anadia mcdiarmidi sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok." captionText-4="Fig. 5. Anadia mcdiarmidi sp. nov. The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok." captionText-5="Fig. 6. Anadia mcdiarmidi sp. nov. Intrapopulation variation in dorsal (above) and ventral (below) colour pattern in preserved specimens. From left to right: IRSNB 2677, IRSNB 2674, IRSNB 2676, IRSNB 2675. Photographs by Philippe J.R. Kok." captionText-6="Fig. 7. Anadia mcdiarmidi sp. nov. Intrapopulation variation in the condition of the parietals and the interparietal (frontoparietals in grey). Top, from left to right: IRSNB 2677, IRSNB 2676; bottom, from left to right: IRSNB 2675, IRSNB 2674. Drawings not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.3711437" figureDoi-1="http://doi.org/10.5281/zenodo.3711439" figureDoi-2="http://doi.org/10.5281/zenodo.3711441" figureDoi-3="http://doi.org/10.5281/zenodo.14766542" figureDoi-4="http://doi.org/10.5281/zenodo.3711443" figureDoi-5="http://doi.org/10.5281/zenodo.3711447" figureDoi-6="http://doi.org/10.5281/zenodo.3711449" httpUri-0="https://zenodo.org/record/3711437/files/figure.png" httpUri-1="https://zenodo.org/record/3711439/files/figure.png" httpUri-2="https://zenodo.org/record/3711441/files/figure.png" httpUri-3="https://zenodo.org/record/14766542/files/figure.png" httpUri-4="https://zenodo.org/record/3711443/files/figure.png" httpUri-5="https://zenodo.org/record/3711447/files/figure.png" httpUri-6="https://zenodo.org/record/3711449/files/figure.png" pageId="2" pageNumber="3">Figs 1-7</figureCitation>
|
||
,
|
||
<tableCitation id="F750034B6162FFC7FCE64405FCC7FAC9" box="[806,893,1363,1389]" captionStart="Table 1" captionStartId="13.[189,258,822,848]" captionTargetId="graphics@13.[191,1397,948,2017]" captionTargetPageId="13" captionText="Table 1. Sex, measurements (in mm) and scale counts of the type series of Anadia mcdiarmidi sp. nov. 1 = tail incomplete or obviously regenerated, 2 = data corresponding to right/left side, other abbreviations are defined in the text." httpUri="http://table.plazi.org/id/EEAD6678616DFFC8FF7D4260FE79FC33" pageId="2" pageNumber="3" tableUuid="EEAD6678616DFFC8FF7D4260FE79FC33">Table 1</tableCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F2C8657B6162FFC7FF7D44CEFB70F983" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="BA6D36F06162FFC7FF7D44CEFC75FA16" blockId="2.[189,1226,1432,1576]" box="[189,975,1432,1459]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="9AC211DE6162FFC7FF7D44CEFC75FA16" box="[189,975,1432,1459]" pageId="2" pageNumber="3">
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<taxonomicName id="7DD24D736162FFC7FF7D44CEFEC9FA16" authorityName="Gorzula" authorityYear="1992" box="[189,371,1432,1458]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="breweri">
|
||
<emphasis id="88A6EAE26162FFC7FF7D44CEFEC9FA16" box="[189,371,1432,1458]" italics="true" pageId="2" pageNumber="3">Anadia breweri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="88A6EAE26162FFC7FE4444CFFD90FA16" box="[388,554,1432,1458]" italics="true" pageId="2" pageNumber="3">nomen nudum</emphasis>
|
||
) –
|
||
<treatmentCitation id="3B7310E16162FFC7FD9144CEFC8AFA16" author="Gorzula S." box="[593,816,1432,1458]" page="276" pageId="2" pageNumber="3" year="1992">
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||
<bibRefCitation id="DE434B016162FFC7FD9144CEFC8AFA16" author="Gorzula S." box="[593,816,1432,1458]" editor="Huber O" firstAuthor="Gorzula" journalOrPublisher="Oscar Todtmann Editores, Caracas." pageId="2" pageNumber="3" pagination="304 - 310" part="280" refId="ref7629" refString="Gorzula S. 1992. La herpetofauna del macizo del Chimanta. In: Huber O (ed.), El Macizo del Chimanta: 267 - 280, 304 - 310. Oscar Todtmann Editores, Caracas." title="La herpetofauna del macizo del Chimanta" type="journal article" volumeTitle="El Macizo del Chimanta: 267" year="1992">Gorzula 1992: 276</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs 168-171
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F06162FFC7FF7D44E9FC3EFA7D" blockId="2.[189,1226,1432,1576]" box="[189,900,1471,1498]" pageId="2" pageNumber="3">
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||
<treatmentCitationGroup id="9AC211DE6162FFC7FF7D44E9FC3EFA7D" box="[189,900,1471,1498]" pageId="2" pageNumber="3">
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<taxonomicName id="7DD24D736162FFC7FF7D44E9FE38FA7E" authorityName="Gray" authorityYear="1845" box="[189,386,1471,1498]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="88A6EAE26162FFC7FF7D44E9FEA8FA7D" box="[189,274,1471,1497]" italics="true" pageId="2" pageNumber="3">Anadia</emphasis>
|
||
species a
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="3B7310E16162FFC7FE5F44E9FC97FA7E" author="Gorzula S. & Senaris J. C." box="[415,813,1471,1498]" page="114" pageId="2" pageNumber="3" year="1999">
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<bibRefCitation id="DE434B016162FFC7FE5F44E9FC97FA7E" author="Gorzula S. & Senaris J. C." box="[415,813,1471,1498]" firstAuthor="Gorzula" journalOrPublisher="Scientia Guaianae" pageId="2" pageNumber="3" pagination="1 - 269" part="8" refId="ref7669" refString="Gorzula S. & Senaris J. C. 1999 " 1998 ". Contribution to the herpetofauna of the Venezuelan Guayana: I. A data base. Scientia Guaianae 8: 1 - 269." title="Contribution to the herpetofauna of the Venezuelan Guayana: I. A data base" type="journal article" year="1999">Gorzula & Señaris 1999: 114-115</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 88
|
||
</treatmentCitationGroup>
|
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</paragraph>
|
||
<paragraph id="BA6D36F06162FFC7FF7D44B0FCB0F9A5" blockId="2.[189,1226,1432,1576]" box="[189,778,1510,1537]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="9AC211DE6162FFC7FF7D44B0FCB0F9A5" box="[189,778,1510,1537]" pageId="2" pageNumber="3">
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<taxonomicName id="7DD24D736162FFC7FF7D44B0FEEEF9A5" authorityName="Gray" authorityYear="1845" box="[189,340,1510,1537]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="88A6EAE26162FFC7FF7D44B0FEA8F9A4" box="[189,274,1510,1536]" italics="true" pageId="2" pageNumber="3">Anadia</emphasis>
|
||
sp. A
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="3B7310E16162FFC7FEB144B0FCB0F9A5" author="McDiarmid R. W. & Donnelly M. A." box="[369,778,1510,1537]" page="514" pageId="2" pageNumber="3" year="2005">
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<bibRefCitation id="DE434B016162FFC7FEB144B0FCB0F9A5" author="McDiarmid R. W. & Donnelly M. A." box="[369,778,1510,1537]" editor="Donnelly M. A. & Crother B. I. & Guyer C. & Wake M. H. & White M. E." firstAuthor="McDiarmid" journalOrPublisher="University of Chicago Press, Chicago, Illinois" pageId="2" pageNumber="3" pagination="461 - 560" refId="ref7922" refString="McDiarmid R. W. & Donnelly M. A. 2005. The herpetofauna of the Guayana Highlands: amphibians and reptiles of the Lost World. In: Donnelly M. A., Crother B. I., Guyer C., Wake M. H. & White M. E. (eds.) Ecology and Evolution in the Tropics: A Herpetological Perspective: 461 - 560. University of Chicago Press, Chicago, Illinois." title="The herpetofauna of the Guayana Highlands: amphibians and reptiles of the Lost World" type="book chapter" volumeTitle="Ecology and Evolution in the Tropics: A Herpetological Perspective" year="2005">McDiarmid & Donnelly 2005: 514</bibRefCitation>
|
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</treatmentCitation>
|
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</treatmentCitationGroup>
|
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</paragraph>
|
||
<paragraph id="BA6D36F06162FFC7FF7D475BFB70F983" blockId="2.[189,1226,1432,1576]" box="[189,1226,1549,1576]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="9AC211DE6162FFC7FF7D475BFB70F983" box="[189,1226,1549,1576]" pageId="2" pageNumber="3">
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<taxonomicName id="7DD24D736162FFC7FF7D475BFEC9F983" authorityName="Gorzula" authorityYear="1992" box="[189,371,1549,1575]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="breweri">
|
||
<emphasis id="88A6EAE26162FFC7FF7D475BFEC9F983" box="[189,371,1549,1575]" italics="true" pageId="2" pageNumber="3">Anadia breweri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="88A6EAE26162FFC7FE444758FD90F983" box="[388,554,1549,1575]" italics="true" pageId="2" pageNumber="3">nomen nudum</emphasis>
|
||
) –
|
||
<treatmentCitation id="3B7310E16162FFC7FD91475BFC5BF98C" author="Brewer-Carias C. & Audy M." box="[593,993,1549,1576]" page="210" pageId="2" pageNumber="3" year="2010">
|
||
<bibRefCitation id="DE434B016162FFC7FD91475BFC5BF98C" author="Brewer-Carias C. & Audy M." box="[593,993,1549,1576]" firstAuthor="Brewer-Carias" journalOrPublisher="Caracas" pageId="2" pageNumber="3" refId="ref7483" refString="Brewer-Carias C. & Audy M. 2010. Entranas del Mundo Perdido. Carlos Capriles de Altolitho C. A., Caracas." title="Entranas del Mundo Perdido. Carlos Capriles de Altolitho C. A" type="book" year="2010">Brewer-Carías & Audy 2010: 210</bibRefCitation>
|
||
</treatmentCitation>
|
||
, unnumbered figure
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F2C8657B6162FFC7FF7D4705FACDF84F" pageId="2" pageNumber="3" type="materials_examined">
|
||
<paragraph id="BA6D36F06162FFC7FF7D4705FA88F91F" blockId="2.[189,1399,1619,1724]" pageId="2" pageNumber="3">
|
||
<materialsCitation id="0ABA3CAD6162FFC7FF7D4705FA88F91F" ID-GBIF-Occurrence="2575931620" collectingDate="2011-05-03" collectionCode="R" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2201" latitude="5.1858335" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.29333" pageId="2" pageNumber="3" specimenCode="IRSNB 2677" specimenCount="1" specimenCount-adult="1" specimenCount-male="1" stateProvince="Bolivar State" typeStatus="holotype">
|
||
<heading id="E125819C6162FFC7FF7D4705FE95F9C9" bold="true" box="[189,303,1619,1645]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="88A6EAE26162FFC7FF7D4705FE95F9C9" bold="true" box="[189,303,1619,1645]" pageId="2" pageNumber="3">
|
||
<typeStatus id="656988526162FFC7FF7D4705FE95F9C9" box="[189,303,1619,1645]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
<specimenCode id="EA749E8B6162FFC7FF7D472CFEEDF930" box="[189,343,1658,1684]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="3">IRSNB 2677</specimenCode>
|
||
(field number PK 3565),
|
||
<specimenCount id="ACD4FD796162FFC7FD48472CFD42F930" box="[648,760,1658,1684]" pageId="2" pageNumber="3">1 adult ♂</specimenCount>
|
||
collected by
|
||
<collectorName id="172753266162FFC7FC58472CFBD6F930" box="[920,1132,1658,1684]" pageId="2" pageNumber="3">Philippe J.R. Kok</collectorName>
|
||
,
|
||
<date id="CE6C10306162FFC7FBBA472CFAB8F931" box="[1146,1282,1658,1685]" pageId="2" pageNumber="3" value="2011-05-03">
|
||
<collectingDate id="DE28E9D86162FFC7FBBA472CFAB8F931" box="[1146,1282,1658,1685]" pageId="2" pageNumber="3" value="2011-05-03">03-05-2011</collectingDate>
|
||
</date>
|
||
at 11h39,
|
||
<location id="BF0D602B6162FFC7FF7D47F7FE50F91F" LSID="urn:lsid:plazi:treatment:327B87E66162FFCBFDB4447DFC5CFCEB:BF0D602B6162FFC7FF7D47F7FE50F91F" box="[189,490,1697,1723]" country="Venezuela" latitude="5.1858335" longLatPrecision="21" longitude="-62.29333" name="summit of Abakapa-tepui" pageId="2" pageNumber="3" stateProvince="Bolivar State">summit of Abakapá-tepui</location>
|
||
,
|
||
<collectingRegion id="7816F8126162FFC7FE3647F7FD29F91F" box="[502,659,1697,1723]" country="Venezuela" name="Bolivar" pageId="2" pageNumber="3">Bolívar State</collectingRegion>
|
||
,
|
||
<collectingCountry id="C2C576606162FFC7FD5E47F7FCA0F91F" box="[670,794,1697,1723]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
|
||
,
|
||
<geoCoordinate id="DFE650376162FFC7FCE547F7FC17F918" box="[805,941,1697,1724]" degrees="05" direction="north" minutes="11" orientation="latitude" pageId="2" pageNumber="3" precision="15" seconds="09" value="5.1858335">05°11'09''N</geoCoordinate>
|
||
<geoCoordinate id="DFE650376162FFC7FC7447F7FBE8F91F" box="[948,1106,1697,1723]" degrees="062" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="36" value="-62.29333">062°17'36''W</geoCoordinate>
|
||
,
|
||
<quantity id="7D2A9B156162FFC7FB9E47F7FB0DF91F" box="[1118,1207,1697,1723]" metricMagnitude="3" metricUnit="m" metricValue="2.2009999999999996" pageId="2" pageNumber="3" unit="m" value="2201.0">
|
||
<elevation id="31FFD1C36162FFC7FB9E47F7FB0DF91F" box="[1118,1207,1697,1723]" metricMagnitude="3" metricUnit="m" metricValue="2.2009999999999996" pageId="2" pageNumber="3" unit="m" value="2201.0">2201 m</elevation>
|
||
</quantity>
|
||
elevation.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F06162FFC7FF7D47B0FACDF84F" blockId="2.[189,1399,1766,2027]" pageId="2" pageNumber="3">
|
||
<materialsCitation id="0ABA3CAD6162FFC7FF7D47B0FACCF8EB" ID-GBIF-Occurrence="2575931621" collectingDate="2011-05-06" collectionCode="IRSNB, PK" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2242" latitude="5.1805553" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.296944" pageId="2" pageNumber="3" specimenCode="IRSNB 2674" specimenCount="2" specimenCount-adult="1" specimenCount-male="1" stateProvince="Bolivar State" typeStatus="paratopotypes">
|
||
<heading id="E125819C6162FFC7FF7D47B0FEC8F8A4" bold="true" box="[189,370,1766,1792]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="88A6EAE26162FFC7FF7D47B0FEC8F8A4" bold="true" box="[189,370,1766,1792]" pageId="2" pageNumber="3">
|
||
<typeStatus id="656988526162FFC7FF7D47B0FEC8F8A4" box="[189,370,1766,1792]" pageId="2" pageNumber="3">Paratopotypes</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
(n=3)
|
||
<specimenCount id="ACD4FD796162FFC7FEC7465BFEE0F883" box="[263,346,1805,1831]" pageId="2" pageNumber="3" type="adult">1 adult</specimenCount>
|
||
<specimenCount id="ACD4FD796162FFC7FEA34658FEC3F88C" box="[355,377,1806,1832]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
|
||
(
|
||
<specimenCode id="EA749E8B6162FFC7FE4A465BFD93F883" box="[394,553,1805,1831]" pageId="2" pageNumber="3">IRSNB 2674</specimenCode>
|
||
, field number PK 3578) collected by
|
||
<collectorName id="172753266162FFC7FC2C465BFB78F883" box="[1004,1218,1805,1832]" pageId="2" pageNumber="3">Philippe J.R. Kok</collectorName>
|
||
,
|
||
<date id="CE6C10306162FFC7FB10465BFAE2F88C" box="[1232,1368,1805,1832]" pageId="2" pageNumber="3" value="2011-05-06">
|
||
<collectingDate id="DE28E9D86162FFC7FB10465BFAE2F88C" box="[1232,1368,1805,1832]" pageId="2" pageNumber="3" value="2011-05-06">06-05-2011</collectingDate>
|
||
</date>
|
||
at 9h40,
|
||
<location id="BF0D602B6162FFC7FEC54662FD8AF8EA" LSID="urn:lsid:plazi:treatment:327B87E66162FFCBFDB4447DFC5CFCEB:BF0D602B6162FFC7FEC54662FD8AF8EA" box="[261,560,1844,1870]" country="Venezuela" latitude="5.1805553" longLatPrecision="21" longitude="-62.296944" name="summit of Abakapa-tepui" pageId="2" pageNumber="3" stateProvince="Bolivar State">summit of Abakapá-tepui</location>
|
||
,
|
||
<collectingRegion id="7816F8126162FFC7FDF94662FD6FF8EA" box="[569,725,1844,1870]" country="Venezuela" name="Bolivar" pageId="2" pageNumber="3">Bolívar State</collectingRegion>
|
||
,
|
||
<collectingCountry id="C2C576606162FFC7FD1F4662FCE1F8EA" box="[735,859,1844,1870]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
|
||
,
|
||
<geoCoordinate id="DFE650376162FFC7FCA44662FC57F8EB" box="[868,1005,1844,1871]" degrees="05" direction="north" minutes="10" orientation="latitude" pageId="2" pageNumber="3" precision="15" seconds="50" value="5.1805553">05°10'50''N</geoCoordinate>
|
||
<geoCoordinate id="DFE650376162FFC7FC334662FB2BF8EA" box="[1011,1169,1844,1870]" degrees="062" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="49" value="-62.296944">062°17'49''W</geoCoordinate>
|
||
,
|
||
<quantity id="7D2A9B156162FFC7FB5C4662FB49F8EB" box="[1180,1267,1844,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.242" pageId="2" pageNumber="3" unit="m" value="2242.0">
|
||
<elevation id="31FFD1C36162FFC7FB5C4662FB49F8EB" box="[1180,1267,1844,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.242" pageId="2" pageNumber="3" unit="m" value="2242.0">2242 m</elevation>
|
||
</quantity>
|
||
elevation);
|
||
</materialsCitation>
|
||
<materialsCitation id="0ABA3CAD6162FFC7FF7D460AFA84F838" ID-GBIF-Occurrence="2575931622" collectingDate="2011-05-06" collectionCode="IRSNB, PK" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2200" latitude="5.1833334" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.292778" pageId="2" pageNumber="3" specimenCode="IRSNB 2675" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Bolivar State" typeStatus="paratopotype">
|
||
<specimenCount id="ACD4FD796162FFC7FF7D460AFE8BF8D1" box="[189,305,1883,1910]" pageId="2" pageNumber="3" type="juvenile">1 juvenile</specimenCount>
|
||
<specimenCount id="ACD4FD796162FFC7FEF8460AFEF4F8D2" box="[312,334,1884,1910]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
|
||
(
|
||
<specimenCode id="EA749E8B6162FFC7FE9D460AFE40F8D2" box="[349,506,1883,1910]" pageId="2" pageNumber="3">IRSNB 2675</specimenCode>
|
||
, field number PK 3577) collected by
|
||
<collectorName id="172753266162FFC7FC6C460DFBC4F8D1" box="[940,1150,1883,1910]" pageId="2" pageNumber="3">Philippe J.R. Kok</collectorName>
|
||
,
|
||
<date id="CE6C10306162FFC7FB4A460DFAA8F8D2" box="[1162,1298,1883,1910]" pageId="2" pageNumber="3" value="2011-05-06">
|
||
<collectingDate id="DE28E9D86162FFC7FB4A460DFAA8F8D2" box="[1162,1298,1883,1910]" pageId="2" pageNumber="3" value="2011-05-06">06-05-2011</collectingDate>
|
||
</date>
|
||
at 9h10,
|
||
<location id="BF0D602B6162FFC7FF7D46D4FE54F838" LSID="urn:lsid:plazi:treatment:327B87E66162FFCBFDB4447DFC5CFCEB:BF0D602B6162FFC7FF7D46D4FE54F838" box="[189,494,1922,1948]" country="Venezuela" latitude="5.1833334" longLatPrecision="21" longitude="-62.292778" name="summit of Abakapa-tepui" pageId="2" pageNumber="3" stateProvince="Bolivar State">summit of Abakapá-tepui</location>
|
||
,
|
||
<collectingRegion id="7816F8126162FFC7FE3B46D4FD20F839" box="[507,666,1922,1949]" country="Venezuela" name="Bolivar" pageId="2" pageNumber="3">Bolívar State</collectingRegion>
|
||
,
|
||
<collectingCountry id="C2C576606162FFC7FD6646D4FC98F838" box="[678,802,1922,1948]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
|
||
,
|
||
<geoCoordinate id="DFE650376162FFC7FCEF46D4FC0DF839" box="[815,951,1922,1949]" degrees="05" direction="north" minutes="11" orientation="latitude" pageId="2" pageNumber="3" precision="15" seconds="00" value="5.1833334">05°11'00''N</geoCoordinate>
|
||
<geoCoordinate id="DFE650376162FFC7FC0046D4FBE5F838" box="[960,1119,1922,1948]" degrees="062" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="34" value="-62.292778">062°17'34''W</geoCoordinate>
|
||
,
|
||
<quantity id="7D2A9B156162FFC7FBAC46D5FB7CF839" box="[1132,1222,1923,1949]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">
|
||
<elevation id="31FFD1C36162FFC7FBAC46D5FB7CF839" box="[1132,1222,1923,1949]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">2200 m</elevation>
|
||
</quantity>
|
||
elevation
|
||
</materialsCitation>
|
||
; and
|
||
<materialsCitation id="0ABA3CAD6162FFC7FF7D46FCFACDF84F" ID-GBIF-Occurrence="2575931619" collectingDate="2011-05-09" collectionCode="IRSNB, PK" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2204" latitude="5.185278" location="summit of Abakapa-tepui" longLatPrecision="21" longitude="-62.288612" pageId="2" pageNumber="3" specimenCode="IRSNB 2676" specimenCount="1" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Bolivar State">
|
||
<specimenCount id="ACD4FD796162FFC7FF7D46FCFE28F860" box="[189,402,1962,1988]" pageId="2" pageNumber="3">1 other juvenile ♂</specimenCount>
|
||
(
|
||
<specimenCode id="EA749E8B6162FFC7FE6146FCFD84F867" box="[417,574,1961,1988]" pageId="2" pageNumber="3">IRSNB 2676</specimenCode>
|
||
, field number PK 3589) collected by
|
||
<collectorName id="172753266162FFC7FC3346FCFB7CF860" box="[1011,1222,1962,1988]" pageId="2" pageNumber="3">Philippe J.R. Kok</collectorName>
|
||
,
|
||
<date id="CE6C10306162FFC7FB1246FFFAE0F860" box="[1234,1370,1961,1988]" pageId="2" pageNumber="3" value="2011-05-09">
|
||
<collectingDate id="DE28E9D86162FFC7FB1246FFFAE0F860" box="[1234,1370,1961,1988]" pageId="2" pageNumber="3" value="2011-05-09">09-05-2011</collectingDate>
|
||
</date>
|
||
at 9h00,
|
||
<location id="BF0D602B6162FFC7FEC64687FD89F84F" LSID="urn:lsid:plazi:treatment:327B87E66162FFCBFDB4447DFC5CFCEB:BF0D602B6162FFC7FEC64687FD89F84F" box="[262,563,2001,2027]" country="Venezuela" latitude="5.185278" longLatPrecision="21" longitude="-62.288612" name="summit of Abakapa-tepui" pageId="2" pageNumber="3" stateProvince="Bolivar State">summit of Abakapá-tepui</location>
|
||
,
|
||
<collectingRegion id="7816F8126162FFC7FDFE4687FD61F84F" box="[574,731,2001,2027]" country="Venezuela" name="Bolivar" pageId="2" pageNumber="3">Bolívar State</collectingRegion>
|
||
,
|
||
<collectingCountry id="C2C576606162FFC7FD254687FCDBF84F" box="[741,865,2001,2027]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
|
||
,
|
||
<geoCoordinate id="DFE650376162FFC7FCAC4686FC4EF84F" box="[876,1012,2000,2027]" degrees="05" direction="north" minutes="11" orientation="latitude" pageId="2" pageNumber="3" precision="15" seconds="07" value="5.185278">05°11'07''N</geoCoordinate>
|
||
<geoCoordinate id="DFE650376162FFC7FC3B4687FB22F84F" box="[1019,1176,2001,2027]" degrees="062" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="19" value="-62.288612">062°17'19''W</geoCoordinate>
|
||
,
|
||
<quantity id="7D2A9B156162FFC7FB644687FB47F84F" box="[1188,1277,2001,2027]" metricMagnitude="3" metricUnit="m" metricValue="2.2039999999999997" pageId="2" pageNumber="3" unit="m" value="2204.0">
|
||
<elevation id="31FFD1C36162FFC7FB644687FB47F84F" box="[1188,1277,2001,2027]" metricMagnitude="3" metricUnit="m" metricValue="2.2039999999999997" pageId="2" pageNumber="3" unit="m" value="2204.0">2204 m</elevation>
|
||
</quantity>
|
||
elevation.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="EEAD66786163FFC6FF7D4631FE00F84F" ID-DOI="http://doi.org/10.5281/zenodo.3711437" ID-Zenodo-Dep="3711437" httpUri="https://zenodo.org/record/3711437/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1895,1921]" targetBox="[207,1380,265,1855]" targetPageId="3">
|
||
<paragraph id="BA6D36F06163FFC6FF7D4631FE00F84F" blockId="3.[189,1399,1895,2027]" pageId="3" pageNumber="4">
|
||
<emphasis id="88A6EAE26163FFC6FF7D4631FEB1F825" bold="true" box="[189,267,1895,1921]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
|
||
<taxonomicName id="7DD24D736163FFC6FED24631FE4CF825" authority="Kok & Rivas, 2011" authorityName="Kok & Rivas" authorityYear="2011" box="[274,502,1895,1921]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="mcdiarmidi" status="sp. nov.">
|
||
<emphasis id="88A6EAE26163FFC6FED24631FE4CF825" box="[274,502,1895,1921]" italics="true" pageId="3" pageNumber="4">Anadia mcdiarmidi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="939557996163FFC6FE3E463EFDE3F826" box="[510,601,1896,1922]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="88A6EAE26163FFC6FDA04631FDCFF825" bold="true" box="[608,629,1895,1921]" pageId="3" pageNumber="4">A</emphasis>
|
||
. IRSNB 2677, ♂ holotype in life.
|
||
<emphasis id="88A6EAE26163FFC6FBCD4631FB9BF825" bold="true" box="[1037,1057,1895,1921]" pageId="3" pageNumber="4">B</emphasis>
|
||
. IRSNB 2675, juvenile male paratype in life.
|
||
<emphasis id="88A6EAE26163FFC6FE4446DCFE23F800" bold="true" box="[388,409,1930,1956]" pageId="3" pageNumber="4">C</emphasis>
|
||
. Ventral view of the anaesthetized holotype (grid squares = 5 mm).
|
||
<emphasis id="88A6EAE26163FFC6FB1146DDFB5CF801" bold="true" box="[1233,1254,1931,1957]" pageId="3" pageNumber="4">D</emphasis>
|
||
. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="F2C8657B6164FFC1FF7D405DFD44FD6D" pageId="4" pageNumber="5" type="etymology">
|
||
<paragraph id="BA6D36F06164FFC1FF7D405DFEFEFE81" blockId="4.[189,1399,267,399]" box="[189,324,267,293]" pageId="4" pageNumber="5">
|
||
<heading id="E125819C6164FFC1FF7D405DFEFEFE81" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="88A6EAE26164FFC1FF7D405DFEFEFE81" bold="true" box="[189,324,267,293]" pageId="4" pageNumber="5">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F06164FFC1FF7D4078FEE4FE2B" blockId="4.[189,1399,267,399]" pageId="4" pageNumber="5">The specific epithet is a noun in the genitive case, honouring Roy McDiarmid, North American herpetologist (Smithsonian Institution), for his contribution to the knowledge of the Pantepui herpetofauna.</paragraph>
|
||
<paragraph id="BA6D36F06164FFC1FF7D40E1FE18FE75" blockId="4.[189,1399,439,714]" box="[189,418,439,465]" pageId="4" pageNumber="5">
|
||
<heading id="E125819C6164FFC1FF7D40E1FE18FE75" bold="true" box="[189,418,439,465]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="88A6EAE26164FFC1FF7D40E1FE18FE75" bold="true" box="[189,418,439,465]" pageId="4" pageNumber="5">Generic allocation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F06164FFC1FF7D408CFD44FD6D" blockId="4.[189,1399,439,714]" pageId="4" pageNumber="5">
|
||
The new species is placed in the genus
|
||
<taxonomicName id="7DD24D736164FFC1FD66408DFD41FE51" authorityName="Gray" authorityYear="1845" box="[678,763,475,501]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="88A6EAE26164FFC1FD66408DFD41FE51" box="[678,763,475,501]" italics="true" pageId="4" pageNumber="5">Anadia</emphasis>
|
||
</taxonomicName>
|
||
because it fits the 11 diagnostic traits for the genus proposed by
|
||
<bibRefCitation id="DE434B016164FFC1FE9640A8FDB3FDBC" author="Oftedal O. T." box="[342,521,510,536]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="4" pageNumber="5" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
: dorsal scales smooth, essentially homogeneous; no distinct rows of much smaller scales between lateral scales; ventral scales smooth, quadrangular; head scales smooth, without striation; single frontonasal separating nasals; paired prefrontals; several large pigmented palpebrals; gular scales smooth, essentially homogeneous, with no distinctly enlarged longitudinal rows; 7-10 large preanal scales [maximum
|
||
<quantity id="7D2A9B156164FFC1FE3743DAFD9DFD02" box="[503,551,652,678]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="4" pageNumber="5" unit="in" value="9.0">9 in</quantity>
|
||
<bibRefCitation id="DE434B016164FFC1FDF143DAFD5CFD02" author="Oftedal O. T." box="[561,742,651,678]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="4" pageNumber="5" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
]; femoral pores conspicuous in adult males; and limbs well developed, pentadactyl with all toes clawed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F2C8657B6164FFC2FF7D43A7FA82FAA8" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="diagnosis">
|
||
<paragraph id="BA6D36F06164FFC1FF7D43A7FE51FCAF" blockId="4.[189,1399,753,1028]" box="[189,491,753,779]" pageId="4" pageNumber="5">
|
||
<heading id="E125819C6164FFC1FF7D43A7FE51FCAF" bold="true" box="[189,491,753,779]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="88A6EAE26164FFC1FF7D43A7FE51FCAF" bold="true" box="[189,491,753,779]" pageId="4" pageNumber="5">Definition and diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F06164FFC2FF7D4243FBBFFEED" blockId="4.[189,1399,753,1028]" lastBlockId="7.[189,1398,267,330]" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
|
||
A fairly robust species currently not assigned to any species group. The new species is characterized by the following combination of characters: (1) size small, body robust (maximum known SVL
|
||
<quantity id="7D2A9B156164FFC1FF7D420AFE96FCD2" box="[189,300,860,886]" metricMagnitude="-2" metricUnit="m" metricValue="6.22" pageId="4" pageNumber="5" unit="mm" value="62.2">62.2 mm</quantity>
|
||
); (2) tail longer than SVL; (3) dorsal scales small, quadrangular; (4) middorsal scales 53-57; (5) suboculars large, unequal in size, with sometimes one scale slightly protruding downward between 4
|
||
<superScript id="4DA79BB86164FFC1FF0C42F7FF63FC0B" attach="left" box="[204,217,929,943]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
||
and 5
|
||
<superScript id="4DA79BB86164FFC1FEE342F7FE8AFC0B" attach="left" box="[291,304,929,943]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
||
supralabial, suboculars separated from palpebrals by 2-3 rows of mostly inconspicuous tiny scales, a few of them distinctly enlarged with angular downward protrusion between suboculars; (6) nasal entire, without sub-nostril groove; (7) 3-4 large pigmented palpebrals; (8) body uniform beige or greyish to bluish brown in life, devoid of any conspicuous pattern in males; (9) absence of any dark line on side of head in adult males; (10) venter immaculate golden grey in life; (11) femoral pores 9-10 on each side in males, unknown in females; (12) preanal pores absent; (13) hemipenis globose, weakly bilobed, bordered by numerous flounces (>20, including about eight medioproximal asulcate flounces) bearing comblike rows of minute weakly mineralized spinules, sulcus spermaticus divided in the crotch by a small fleshy bumplike structure, sulcus branches barely detectable.
|
||
</paragraph>
|
||
<caption id="EEAD66786164FFC1FF7D46FBFC2DF84F" ID-DOI="http://doi.org/10.5281/zenodo.3711439" ID-Zenodo-Dep="3711439" httpUri="https://zenodo.org/record/3711439/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1965,1991]" targetBox="[189,1372,1072,1925]" targetPageId="4">
|
||
<paragraph id="BA6D36F06164FFC1FF7D46FBFC2DF84F" blockId="4.[189,1398,1965,2028]" pageId="4" pageNumber="5">
|
||
<emphasis id="88A6EAE26164FFC1FF7D46FBFEB3F863" bold="true" box="[189,265,1965,1991]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
|
||
<taxonomicName id="7DD24D736164FFC1FED046FBFE48F863" authority="Kok & Rivas, 2011" authorityName="Kok & Rivas" authorityYear="2011" box="[272,498,1965,1991]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="mcdiarmidi" status="sp. nov.">
|
||
<emphasis id="88A6EAE26164FFC1FED046FBFE48F863" box="[272,498,1965,1991]" italics="true" pageId="4" pageNumber="5">Anadia mcdiarmidi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="939557996164FFC1FE3846F8FDE8F86C" box="[504,594,1966,1992]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Ventral, dorsal, and lateral views of the head of the holotype (IRSNB 2677). Scale lines = 5 mm. Photographs by Philippe J.R. Kok.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EEAD66786165FFC0FF7D46DCFB55F84F" ID-DOI="http://doi.org/10.5281/zenodo.3711441" ID-Zenodo-Dep="3711441" httpUri="https://zenodo.org/record/3711441/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1930,1956]" targetBox="[189,1399,467,1890]" targetPageId="5">
|
||
<paragraph id="BA6D36F06165FFC0FF7D46DCFB55F84F" blockId="5.[189,1399,1930,2028]" pageId="5" pageNumber="6">
|
||
<emphasis id="88A6EAE26165FFC0FF7D46DCFEB5F800" bold="true" box="[189,271,1930,1956]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
|
||
<taxonomicName id="7DD24D736165FFC0FEDC46DCFDBEF800" authority="Kok & Rivas, 2011" authorityName="Kok & Rivas" authorityYear="2011" box="[284,516,1930,1956]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="mcdiarmidi" status="sp. nov.">
|
||
<emphasis id="88A6EAE26165FFC0FEDC46DCFDBEF800" box="[284,516,1930,1956]" italics="true" pageId="5" pageNumber="6">Anadia mcdiarmidi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="939557996165FFC0FDD146DDFDCAF801" box="[529,624,1931,1957]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Subpelvic region of the preserved ♂ holotype (IRSNB 2677), showing preanal scales and femoral pores, and trunk of the preserved holotype in lateral view (head to left), showing scalation at midbody. Scale lines = 5 mm. Photographs by Philippe J.R. Kok.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EEAD66786166FFC3FF7D42BEFD48FB82" ID-DOI="http://doi.org/10.5281/zenodo.14766542" ID-Zenodo-Dep="14766542" httpUri="https://zenodo.org/record/14766542/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1000,1026]" targetBox="[189,1399,1085,1787]" targetPageId="6">
|
||
<paragraph id="BA6D36F06166FFC3FF7D42BEFD48FB82" pageId="6" pageNumber="7">
|
||
<emphasis id="88A6EAE26166FFC3FF7D42BEFEB0FBA6" bold="true" box="[189,266,1000,1026]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
|
||
<emphasis id="88A6EAE26166FFC3FED142BEFE4EFBA6" box="[273,500,1000,1026]" italics="true" pageId="6" pageNumber="7">Anadia mcdiarmidi</emphasis>
|
||
sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EEAD66786166FFC3FF7D4676FACDF84F" ID-DOI="http://doi.org/10.5281/zenodo.3711443" ID-Zenodo-Dep="3711443" httpUri="https://zenodo.org/record/3711443/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1824,1850]" targetBox="[189,1399,1085,1787]" targetPageId="6">
|
||
<paragraph id="BA6D36F06166FFC3FF7D4676FACDF84F" blockId="6.[189,1399,1824,2027]" pageId="6" pageNumber="7">
|
||
<emphasis id="88A6EAE26166FFC3FF7D4676FEB0F89F" bold="true" box="[189,266,1824,1851]" pageId="6" pageNumber="7">Fig. 5.</emphasis>
|
||
<taxonomicName id="7DD24D736166FFC3FED24677FE4FF89F" authority="Kok & Rivas, 2011" authorityName="Kok & Rivas" authorityYear="2011" box="[274,501,1825,1851]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="mcdiarmidi" status="sp. nov.">
|
||
<emphasis id="88A6EAE26166FFC3FED24677FE4FF89F" box="[274,501,1825,1851]" italics="true" pageId="6" pageNumber="7">Anadia mcdiarmidi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="939557996166FFC3FE3D4677FDEDF89F" box="[509,599,1825,1851]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BA6D36F06167FFC2FF7D4027FA82FAA8" blockId="7.[189,1399,369,1292]" pageId="7" pageNumber="8">
|
||
The new taxon is immediately distinguished from
|
||
<taxonomicName id="7DD24D736167FFC2FD364027FC14FE2F" authorityName="Gray" authorityYear="1845" box="[758,942,369,395]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ocellata">
|
||
<emphasis id="88A6EAE26167FFC2FD364027FC14FE2F" box="[758,942,369,395]" italics="true" pageId="7" pageNumber="8">Anadia ocellata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7DD24D736167FFC2FC7A4024FB98FE2F" authorityName="Boulenger" authorityYear="1913" box="[954,1058,370,396]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="88A6EAE26167FFC2FC7A4024FB98FE2F" box="[954,1058,370,396]" italics="true" pageId="7" pageNumber="8">A. vittata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7DD24D736167FFC2FBEE4024FB74FE2F" baseAuthorityName="Gunther" baseAuthorityYear="1859" box="[1070,1230,369,396]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="rhombifera">
|
||
<emphasis id="88A6EAE26167FFC2FBEE4024FB74FE2F" box="[1070,1230,369,396]" italics="true" pageId="7" pageNumber="8">A. rhombifera</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="7DD24D736167FFC2FAC94024FACDFE2F" authorityName="Oftedal" authorityYear="1974" box="[1289,1399,370,396]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="petersi">
|
||
<emphasis id="88A6EAE26167FFC2FAC94024FACDFE2F" box="[1289,1399,370,396]" italics="true" pageId="7" pageNumber="8">A. petersi</emphasis>
|
||
</taxonomicName>
|
||
[all members of the
|
||
<taxonomicName id="7DD24D736167FFC2FE6140C3FDBAFE0B" authorityName="Gray" authorityYear="1845" box="[417,512,405,431]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ocellata">
|
||
<emphasis id="88A6EAE26167FFC2FE6140C3FDBAFE0B" box="[417,512,405,431]" italics="true" pageId="7" pageNumber="8">ocellata</emphasis>
|
||
</taxonomicName>
|
||
group of
|
||
<bibRefCitation id="DE434B016167FFC2FDAC40C3FCA1FE0B" author="Oftedal O. T." box="[620,795,404,431]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="7" pageNumber="8" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
] and
|
||
<taxonomicName id="7DD24D736167FFC2FC9640C3FC48FE0B" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[854,1010,405,431]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bogotensis">
|
||
<emphasis id="88A6EAE26167FFC2FC9640C3FC48FE0B" box="[854,1010,405,431]" italics="true" pageId="7" pageNumber="8">A. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
[single member of the
|
||
<taxonomicName id="7DD24D736167FFC2FB3940C3FACCFE0B" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[1273,1398,405,431]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bogotensis">
|
||
<emphasis id="88A6EAE26167FFC2FB3940C3FACCFE0B" box="[1273,1398,405,431]" italics="true" pageId="7" pageNumber="8">bogotensis</emphasis>
|
||
</taxonomicName>
|
||
group of
|
||
<bibRefCitation id="DE434B016167FFC2FEE940EFFE60FE77" author="Oftedal O. T." box="[297,474,440,467]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="7" pageNumber="8" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
] in having quadrangular dorsal scales (subhexagonal in species of the
|
||
<taxonomicName id="7DD24D736167FFC2FAD840EFFACDFE77" authorityName="Gray" authorityYear="1845" box="[1304,1399,441,467]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ocellata">
|
||
<emphasis id="88A6EAE26167FFC2FAD840EFFACDFE77" box="[1304,1399,441,467]" italics="true" pageId="7" pageNumber="8">ocellata</emphasis>
|
||
</taxonomicName>
|
||
group, imbricate in
|
||
<taxonomicName id="7DD24D736167FFC2FE64408BFDFBFE53" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[420,577,477,503]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bogotensis">
|
||
<emphasis id="88A6EAE26167FFC2FE64408BFDFBFE53" box="[420,577,477,503]" italics="true" pageId="7" pageNumber="8">A. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
), and in lacking a subnostril groove or divided nasal (nasal divided or subnostril groove present in species of the
|
||
<taxonomicName id="7DD24D736167FFC2FD7E4357FCA7FDBF" authorityName="Gray" authorityYear="1845" box="[702,797,513,539]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ocellata">
|
||
<emphasis id="88A6EAE26167FFC2FD7E4357FCA7FDBF" box="[702,797,513,539]" italics="true" pageId="7" pageNumber="8">ocellata</emphasis>
|
||
</taxonomicName>
|
||
group and in
|
||
<taxonomicName id="7DD24D736167FFC2FC064357FBDCFDBF" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[966,1126,513,539]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bogotensis">
|
||
<emphasis id="88A6EAE26167FFC2FC064357FBDCFDBF" box="[966,1126,513,539]" italics="true" pageId="7" pageNumber="8">A. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="7DD24D736167FFC2FB034357FACCFDBF" authorityName="Harris & Ayala" authorityYear="1987" box="[1219,1398,513,539]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="altaserrania">
|
||
<emphasis id="88A6EAE26167FFC2FB034357FACCFDBF" box="[1219,1398,513,539]" italics="true" pageId="7" pageNumber="8">A. altaserrania</emphasis>
|
||
</taxonomicName>
|
||
[a member of the
|
||
<taxonomicName id="7DD24D736167FFC2FE584372FDB5FD9A" authorityName="Boulenger" authorityYear="1903" box="[408,527,548,574]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bitaeniata">
|
||
<emphasis id="88A6EAE26167FFC2FE584372FDB5FD9A" box="[408,527,548,574]" italics="true" pageId="7" pageNumber="8">bitaeniata</emphasis>
|
||
</taxonomicName>
|
||
group of
|
||
<bibRefCitation id="DE434B016167FFC2FD4B4372FC80FD9A" author="Oftedal O. T." box="[651,826,548,574]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="7" pageNumber="8" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
according to
|
||
<bibRefCitation id="DE434B016167FFC2FC244372FB4FFD9B" author="Harris D. M. & Ayala S. C." box="[996,1269,548,575]" firstAuthor="Harris" journalOrPublisher="Herpetologica" pageId="7" pageNumber="8" pagination="182 - 190" part="43" refId="ref7801" refString="Harris D. M. & Ayala S. C. 1987. A new Anadia (Sauria: Teiidae) from Colombia and restoration of Anadia pamplonensis Dunn to species status. Herpetologica 43 (2): 182 - 190." title="A new Anadia (Sauria: Teiidae) from Colombia and restoration of Anadia pamplonensis Dunn to species status" type="journal article" year="1987">Harris & Ayala (1987)</bibRefCitation>
|
||
] in having prefrontals (lacking in
|
||
<taxonomicName id="7DD24D736167FFC2FE0B431FFDC7FDC6" authorityName="Harris & Ayala" authorityYear="1987" box="[459,637,584,611]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="altaserrania">
|
||
<emphasis id="88A6EAE26167FFC2FE0B431FFDC7FDC6" box="[459,637,584,611]" italics="true" pageId="7" pageNumber="8">A. altaserrania</emphasis>
|
||
</taxonomicName>
|
||
), in having more femoral pores in males (
|
||
<quantity id="7D2A9B156167FFC2FBB1431EFB7CFDC6" box="[1137,1222,584,610]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="7" pageNumber="8" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9-10 in</quantity>
|
||
<emphasis id="88A6EAE26167FFC2FB0E431FFF67FD22" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="7DD24D736167FFC2FB0E431FFACCFDC6" authorityName="Kok & Rivas" authorityYear="2011" box="[1230,1398,584,611]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcdiarmidi">A. mcdiarmidi</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
<quantity id="7D2A9B156167FFC2FF25433AFE90FD22" box="[229,298,620,646]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="7" pageNumber="8" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3-4 in</quantity>
|
||
<taxonomicName id="7DD24D736167FFC2FEF2433AFE5EFD22" authorityName="Harris & Ayala" authorityYear="1987" box="[306,484,620,646]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="altaserrania">
|
||
<emphasis id="88A6EAE26167FFC2FEF2433AFE5EFD22" box="[306,484,620,646]" italics="true" pageId="7" pageNumber="8">A. altaserrania</emphasis>
|
||
</taxonomicName>
|
||
), and in having a higher number of middorsal scales (
|
||
<quantity id="7D2A9B156167FFC2FBA2433AFB7CFD22" box="[1122,1222,620,647]" metricMagnitude="0" metricUnit="m" metricValue="1.3969999999999998" metricValueMax="1.4478" metricValueMin="1.3462" pageId="7" pageNumber="8" unit="in" value="55.0" valueMax="57.0" valueMin="53.0">53-57 in</quantity>
|
||
<emphasis id="88A6EAE26167FFC2FB0E433AFF67FD0E" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="7DD24D736167FFC2FB0E433AFACCFD22" authorityName="Kok & Rivas" authorityYear="2011" box="[1230,1398,620,646]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcdiarmidi">A. mcdiarmidi</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
<quantity id="7D2A9B156167FFC2FF2543C6FEF2FD0E" box="[229,328,656,683]" metricMagnitude="0" metricUnit="m" metricValue="1.1684" metricValueMax="1.1938" metricValueMin="1.143" pageId="7" pageNumber="8" unit="in" value="46.0" valueMax="47.0" valueMin="45.0">45-47 in</quantity>
|
||
<taxonomicName id="7DD24D736167FFC2FE9043C6FDB8FD0E" authorityName="Harris & Ayala" authorityYear="1987" box="[336,514,656,682]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="altaserrania">
|
||
<emphasis id="88A6EAE26167FFC2FE9043C6FDB8FD0E" box="[336,514,656,682]" italics="true" pageId="7" pageNumber="8">A. altaserrania</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="7DD24D736167FFC2FD9D43C6FD4FFD0E" authorityName="Boulenger" authorityYear="1903" box="[605,757,656,682]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bitaeniata">
|
||
<emphasis id="88A6EAE26167FFC2FD9D43C6FD4FFD0E" box="[605,757,656,682]" italics="true" pageId="7" pageNumber="8">A. bitaeniata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7DD24D736167FFC2FCC443C6FC79FD0E" baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[772,963,656,682]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="brevifrontalis">
|
||
<emphasis id="88A6EAE26167FFC2FCC443C6FC79FD0E" box="[772,963,656,682]" italics="true" pageId="7" pageNumber="8">A. brevifrontalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7DD24D736167FFC2FC1243C6FBF3FD0E" authorityName="La Marca & Garcia-Perez" authorityYear="1990" box="[978,1097,656,682]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="hobarti">
|
||
<emphasis id="88A6EAE26167FFC2FC1243C6FBF3FD0E" box="[978,1097,656,682]" italics="true" pageId="7" pageNumber="8">A. hobarti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7DD24D736167FFC2FB9943C6FAA1FD0E" authorityName="Dunn" authorityYear="1944" box="[1113,1307,656,682]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pamplonensis">
|
||
<emphasis id="88A6EAE26167FFC2FB9943C6FAA1FD0E" box="[1113,1307,656,682]" italics="true" pageId="7" pageNumber="8">A. pamplonensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="7DD24D736167FFC2FA9D43C6FE90FD6A" authorityName="Ruthven" authorityYear="1926" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pulchella">
|
||
<emphasis id="88A6EAE26167FFC2FA9D43C6FE90FD6A" italics="true" pageId="7" pageNumber="8">A. pulchella</emphasis>
|
||
</taxonomicName>
|
||
[all members of the
|
||
<taxonomicName id="7DD24D736167FFC2FDD943E2FD2AFD6A" authorityName="Boulenger" authorityYear="1903" box="[537,656,692,718]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bitaeniata">
|
||
<emphasis id="88A6EAE26167FFC2FDD943E2FD2AFD6A" box="[537,656,692,718]" italics="true" pageId="7" pageNumber="8">bitaeniata</emphasis>
|
||
</taxonomicName>
|
||
group of
|
||
<bibRefCitation id="DE434B016167FFC2FCC043E2FC16FD6A" author="Oftedal O. T." box="[768,940,692,718]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="7" pageNumber="8" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
according to
|
||
<bibRefCitation id="DE434B016167FFC2FB8A43E2FEB4FD56" author="La Marca E. & Garcia-Perez J. E." firstAuthor="La Marca" journalOrPublisher="Herpetologica" pageId="7" pageNumber="8" pagination="275 - 282" part="46" refId="ref7844" refString="La Marca E. & Garcia-Perez J. E. 1990. A new species of high Andean Anadia (Sauria: Teiidae) from Paramo El Riecito, Estado Trujillo, Venezuela. Herpetologica 46 (3): 275 - 282." title="A new species of high Andean Anadia (Sauria: Teiidae) from Paramo El Riecito, Estado Trujillo, Venezuela" type="journal article" year="1990">La Marca & García-Pérez (1990)</bibRefCitation>
|
||
] in having a higher number of middorsal scales (
|
||
<quantity id="7D2A9B156167FFC2FCA64381FC77FD55" box="[870,973,727,754]" metricMagnitude="0" metricUnit="m" metricValue="1.3969999999999998" metricValueMax="1.4478" metricValueMin="1.3462" pageId="7" pageNumber="8" unit="in" value="55.0" valueMax="57.0" valueMin="53.0">53-57 in</quantity>
|
||
<emphasis id="88A6EAE26167FFC2FC17438EFB16FD55" box="[983,1196,728,754]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="7DD24D736167FFC2FC17438EFB38FD56" authorityName="Kok & Rivas" authorityYear="2011" box="[983,1154,728,754]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcdiarmidi">A. mcdiarmidi</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
maximum
|
||
<quantity id="7D2A9B156167FFC2FAF84381FACDFD55" box="[1336,1399,727,754]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="7" pageNumber="8" unit="in" value="50.0">50 in</quantity>
|
||
species of the
|
||
<taxonomicName id="7DD24D736167FFC2FEA743ADFE64FCB1" authorityName="Boulenger" authorityYear="1903" box="[359,478,763,789]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bitaeniata">
|
||
<emphasis id="88A6EAE26167FFC2FEA743ADFE64FCB1" box="[359,478,763,789]" italics="true" pageId="7" pageNumber="8">bitaeniata</emphasis>
|
||
</taxonomicName>
|
||
group); from
|
||
<taxonomicName id="7DD24D736167FFC2FD4743AAFD4BFCB1" authorityName="Schmidt" authorityYear="1932" box="[647,753,763,790]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="blakei">
|
||
<emphasis id="88A6EAE26167FFC2FD4743AAFD4BFCB1" box="[647,753,763,790]" italics="true" pageId="7" pageNumber="8">A. blakei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7DD24D736167FFC2FCEC43AAFC68FCB1" baseAuthorityName="Gray" baseAuthorityYear="1846" box="[812,978,764,790]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="marmorata">
|
||
<emphasis id="88A6EAE26167FFC2FCEC43AAFC68FCB1" box="[812,978,764,790]" italics="true" pageId="7" pageNumber="8">A. marmorata</emphasis>
|
||
</taxonomicName>
|
||
[members of the
|
||
<taxonomicName id="7DD24D736167FFC2FB6543AAFA93FCB1" baseAuthorityName="Gray" baseAuthorityYear="1846" box="[1189,1321,764,789]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="marmorata">
|
||
<emphasis id="88A6EAE26167FFC2FB6543AAFA93FCB1" box="[1189,1321,764,789]" italics="true" pageId="7" pageNumber="8">marmorata</emphasis>
|
||
</taxonomicName>
|
||
group of
|
||
<bibRefCitation id="DE434B016167FFC2FF1E4249FE2AFC9E" author="Oftedal O. T." box="[222,400,799,826]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="7" pageNumber="8" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
] in having a higher number of middorsal scales (more than
|
||
<quantity id="7D2A9B156167FFC2FB9E4249FB26FC9D" box="[1118,1180,799,826]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="7" pageNumber="8" unit="in" value="50.0">50 in</quantity>
|
||
<emphasis id="88A6EAE26167FFC2FB644276FACCFC9D" box="[1188,1398,799,826]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="7DD24D736167FFC2FB644276FAF7FC9D" authorityName="Kok & Rivas" authorityYear="2011" box="[1188,1357,799,826]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcdiarmidi">A. mcdiarmidi</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
less than
|
||
<quantity id="7D2A9B156167FFC2FEED4215FED0FCF9" box="[301,362,835,862]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="7" pageNumber="8" unit="in" value="35.0">35 in</quantity>
|
||
<taxonomicName id="7DD24D736167FFC2FEB34215FE67FCF9" authorityName="Schmidt" authorityYear="1932" box="[371,477,835,861]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="blakei">
|
||
<emphasis id="88A6EAE26167FFC2FEB34215FE67FCF9" box="[371,477,835,861]" italics="true" pageId="7" pageNumber="8">A. blakei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7DD24D736167FFC2FDD94215FD05FCF9" baseAuthorityName="Gray" baseAuthorityYear="1846" box="[537,703,835,861]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="marmorata">
|
||
<emphasis id="88A6EAE26167FFC2FDD94215FD05FCF9" box="[537,703,835,861]" italics="true" pageId="7" pageNumber="8">A. marmorata</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="7DD24D736167FFC2FCDC4215FC68FCF9" authorityName="Rueda-Almonacid & Caicedo" authorityYear="2004" box="[796,978,835,861]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bumanguesa">
|
||
<emphasis id="88A6EAE26167FFC2FCDC4215FC68FCF9" box="[796,978,835,861]" italics="true" pageId="7" pageNumber="8">A. bumanguesa</emphasis>
|
||
</taxonomicName>
|
||
[a member of the
|
||
<taxonomicName id="7DD24D736167FFC2FB724215FA93FCF9" authorityName="Boulenger" authorityYear="1903" box="[1202,1321,835,861]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bitaeniata">
|
||
<emphasis id="88A6EAE26167FFC2FB724215FA93FCF9" box="[1202,1321,835,861]" italics="true" pageId="7" pageNumber="8">bitaeniata</emphasis>
|
||
</taxonomicName>
|
||
group of
|
||
<bibRefCitation id="DE434B016167FFC2FF214231FE28FC25" author="Oftedal O. T." box="[225,402,871,897]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="7" pageNumber="8" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
according to
|
||
<bibRefCitation id="DE434B016167FFC2FDFE4231FC44FC25" author="Rueda-Almonacid J. V. & Caicedo J. R." box="[574,1022,871,897]" firstAuthor="Rueda-Almonacid" journalOrPublisher="Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales" pageId="7" pageNumber="8" pagination="281 - 284" part="28" refId="ref8593" refString="Rueda-Almonacid J. V. & Caicedo J. R. 2004. Una nueva especie de Anadia (Reptilia: Sauria: Gymnophthalmidae) para el noroccidente de la cordillera oriental de Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 28 (107): 281 - 284." title="Una nueva especie de Anadia (Reptilia: Sauria: Gymnophthalmidae) para el noroccidente de la cordillera oriental de Colombia" type="journal article" year="2004">Rueda-Almonacid & Caicedo (2004)</bibRefCitation>
|
||
, but a possible synonym of
|
||
<taxonomicName id="7DD24D736167FFC2FA9D4231FEB1FC01" authorityName="Nieden" authorityYear="1914" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="steyeri">
|
||
<emphasis id="88A6EAE26167FFC2FA9D4231FEB1FC01" italics="true" pageId="7" pageNumber="8">A. steyeri</emphasis>
|
||
</taxonomicName>
|
||
and thus a member of the
|
||
<taxonomicName id="7DD24D736167FFC2FD9042DAFD24FC01" authorityName="Nieden" authorityYear="1914" box="[592,670,908,933]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="steyeri">
|
||
<emphasis id="88A6EAE26167FFC2FD9042DAFD24FC01" box="[592,670,908,933]" italics="true" pageId="7" pageNumber="8">steyeri</emphasis>
|
||
</taxonomicName>
|
||
group of
|
||
<bibRefCitation id="DE434B016167FFC2FCD642DDFC7FFC01" author="Oftedal O. T." box="[790,965,907,933]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="7" pageNumber="8" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
according to Rivas
|
||
<emphasis id="88A6EAE26167FFC2FB7742DAFB4EFC01" box="[1207,1268,907,933]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||
(in press)] in having a lower number of scales around midbody (
|
||
<quantity id="7D2A9B156167FFC2FCF842F8FC27FC6C" box="[824,925,942,969]" metricMagnitude="-1" metricUnit="m" metricValue="9.017" metricValueMax="9.144" metricValueMin="8.89" pageId="7" pageNumber="8" unit="in" value="35.5" valueMax="36.0" valueMin="35.0">35-36 in</quantity>
|
||
<emphasis id="88A6EAE26167FFC2FC6542F9FBCCFC6C" box="[933,1142,943,969]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="7DD24D736167FFC2FC6542F9FBF4FC6D" authorityName="Kok & Rivas" authorityYear="2011" box="[933,1102,943,969]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcdiarmidi">A. mcdiarmidi</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
<quantity id="7D2A9B156167FFC2FBBE42F9FB01FC6C" box="[1150,1211,942,969]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" pageId="7" pageNumber="8" unit="in" value="41.0">41 in</quantity>
|
||
<taxonomicName id="7DD24D736167FFC2FB0342F9FA89FC6C" authorityName="Nieden" authorityYear="1914" box="[1219,1331,943,969]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="steyeri">
|
||
<emphasis id="88A6EAE26167FFC2FB0342F9FA89FC6C" box="[1219,1331,943,969]" italics="true" pageId="7" pageNumber="8">A. steyeri</emphasis>
|
||
</taxonomicName>
|
||
), and in having pigmented palpebrals (unpigmented in
|
||
<taxonomicName id="7DD24D736167FFC2FCC44285FC00FC48" authorityName="Rueda-Almonacid & Caicedo" authorityYear="2004" box="[772,954,978,1005]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bumanguesa">
|
||
<emphasis id="88A6EAE26167FFC2FCC44285FC00FC48" box="[772,954,978,1005]" italics="true" pageId="7" pageNumber="8">A. bumanguesa</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="7DD24D736167FFC2FBD64285FB08FC48" authorityName="Rivas, La Marca & Oliveros" authorityYear="1999" box="[1046,1202,979,1005]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pariaensis">
|
||
<emphasis id="88A6EAE26167FFC2FBD64285FB08FC48" box="[1046,1202,979,1005]" italics="true" pageId="7" pageNumber="8">A. pariaensis</emphasis>
|
||
</taxonomicName>
|
||
(not assigned to any species group in the original description) in having a much lower number of middorsal scales (
|
||
<quantity id="7D2A9B156167FFC2FA8E42A0FF43FB90" metricMagnitude="0" metricUnit="m" metricValue="1.4097" metricValueMax="1.4478" metricValueMin="1.3716" pageId="7" pageNumber="8" unit="in" value="55.5" valueMax="57.0" valueMin="54.0">54- 57 in</quantity>
|
||
<emphasis id="88A6EAE26167FFC2FEC1454CFE6BFB90" box="[257,465,1050,1076]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="7DD24D736167FFC2FEC1454CFE13FB90" authorityName="Kok & Rivas" authorityYear="2011" box="[257,425,1050,1076]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcdiarmidi">A. mcdiarmidi</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
<quantity id="7D2A9B156167FFC2FE18454CFD86FB90" box="[472,572,1050,1076]" metricMagnitude="0" metricUnit="m" metricValue="1.8034" metricValueMax="1.8288" metricValueMin="1.778" pageId="7" pageNumber="8" unit="in" value="71.0" valueMax="72.0" valueMin="70.0">70-72 in</quantity>
|
||
<taxonomicName id="7DD24D736167FFC2FD83454CFD64FB90" authorityName="Rivas, La Marca & Oliveros" authorityYear="1999" box="[579,734,1050,1076]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="pariaensis">
|
||
<emphasis id="88A6EAE26167FFC2FD83454CFD64FB90" box="[579,734,1050,1076]" italics="true" pageId="7" pageNumber="8">A. pariaensis</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="7DD24D736167FFC2FCF9454CFC12FB90" authorityName="Nieden" authorityYear="1914" box="[825,936,1050,1076]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="steyeri">
|
||
<emphasis id="88A6EAE26167FFC2FCF9454CFC12FB90" box="[825,936,1050,1076]" italics="true" pageId="7" pageNumber="8">A. steyeri</emphasis>
|
||
</taxonomicName>
|
||
[single member of the
|
||
<taxonomicName id="7DD24D736167FFC2FB7C454DFAB0FB90" authorityName="Nieden" authorityYear="1914" box="[1212,1290,1051,1076]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="steyeri">
|
||
<emphasis id="88A6EAE26167FFC2FB7C454DFAB0FB90" box="[1212,1290,1051,1076]" italics="true" pageId="7" pageNumber="8">steyeri</emphasis>
|
||
</taxonomicName>
|
||
group of
|
||
<bibRefCitation id="DE434B016167FFC2FF7D4568FED7FBFC" author="Oftedal O. T." box="[189,365,1085,1112]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="7" pageNumber="8" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
] in having a lower number of scales around midbody (
|
||
<quantity id="7D2A9B156167FFC2FC304568FBEEFBFC" box="[1008,1108,1086,1113]" metricMagnitude="-1" metricUnit="m" metricValue="9.017" metricValueMax="9.144" metricValueMin="8.89" pageId="7" pageNumber="8" unit="in" value="35.5" valueMax="36.0" valueMin="35.0">35-36 in</quantity>
|
||
<emphasis id="88A6EAE26167FFC2FB9B4568FA90FBFC" box="[1115,1322,1086,1112]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="7DD24D736167FFC2FB9B4568FAB9FBFC" authorityName="Kok & Rivas" authorityYear="2011" box="[1115,1283,1086,1112]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcdiarmidi">A. mcdiarmidi</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
<quantity id="7D2A9B156167FFC2FAF24568FF6EFBD8" metricMagnitude="0" metricUnit="m" metricValue="1.0795" metricValueMax="1.143" metricValueMin="1.016" pageId="7" pageNumber="8" unit="in" value="42.5" valueMax="45.0" valueMin="40.0">40-45 in</quantity>
|
||
<taxonomicName id="7DD24D736167FFC2FF1E4534FEEAFBD8" authorityName="Nieden" authorityYear="1914" box="[222,336,1122,1148]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="steyeri">
|
||
<emphasis id="88A6EAE26167FFC2FF1E4534FEEAFBD8" box="[222,336,1122,1148]" italics="true" pageId="7" pageNumber="8">A. steyeri</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName id="7DD24D736167FFC2FE244534FDCDFBD8" authorityName="Myers, Rivas & Jadin" authorityYear="2009" box="[484,631,1122,1148]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="escalerae">
|
||
<emphasis id="88A6EAE26167FFC2FE244534FDCDFBD8" box="[484,631,1122,1148]" italics="true" pageId="7" pageNumber="8">A. escalerae</emphasis>
|
||
</taxonomicName>
|
||
(not assigned to any species group in the original description), the geographically closest relative, in having a robust body (slender in
|
||
<taxonomicName id="7DD24D736167FFC2FBC045D0FB2BFB3B" authorityName="Myers, Rivas & Jadin" authorityYear="2009" box="[1024,1169,1157,1184]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="escalerae">
|
||
<emphasis id="88A6EAE26167FFC2FBC045D0FB2BFB3B" box="[1024,1169,1157,1184]" italics="true" pageId="7" pageNumber="8">A. escalerae</emphasis>
|
||
</taxonomicName>
|
||
), in having smooth dorsal scales (weakly keeled in
|
||
<taxonomicName id="7DD24D736167FFC2FDF845FCFD70FB67" authorityName="Myers, Rivas & Jadin" authorityYear="2009" box="[568,714,1193,1220]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="escalerae">
|
||
<emphasis id="88A6EAE26167FFC2FDF845FCFD70FB67" box="[568,714,1193,1220]" italics="true" pageId="7" pageNumber="8">A. escalerae</emphasis>
|
||
</taxonomicName>
|
||
), in lacking dorsolateral stripes (present in
|
||
<taxonomicName id="7DD24D736167FFC2FB1345FCFADFFB67" authorityName="Myers, Rivas & Jadin" authorityYear="2009" box="[1235,1381,1193,1220]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="escalerae">
|
||
<emphasis id="88A6EAE26167FFC2FB1345FCFADFFB67" box="[1235,1381,1193,1220]" italics="true" pageId="7" pageNumber="8">A. escalerae</emphasis>
|
||
</taxonomicName>
|
||
), in having a lower number of middorsal scales (
|
||
<quantity id="7D2A9B156167FFC2FD30459BFCECFB43" box="[752,854,1229,1256]" metricMagnitude="0" metricUnit="m" metricValue="1.4097" metricValueMax="1.4478" metricValueMin="1.3716" pageId="7" pageNumber="8" unit="in" value="55.5" valueMax="57.0" valueMin="54.0">54-57 in</quantity>
|
||
<emphasis id="88A6EAE26167FFC2FC9F4598FB8BFB43" box="[863,1073,1229,1256]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="7DD24D736167FFC2FC9F4598FBB2FB43" authorityName="Kok & Rivas" authorityYear="2011" box="[863,1032,1229,1256]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcdiarmidi">A. mcdiarmidi</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
<quantity id="7D2A9B156167FFC2FBFA459BFBCDFB43" box="[1082,1143,1229,1255]" metricMagnitude="0" metricUnit="m" metricValue="1.6002" pageId="7" pageNumber="8" unit="in" value="63.0">63 in</quantity>
|
||
<taxonomicName id="7DD24D736167FFC2FB404598FAA8FB43" authorityName="Myers, Rivas & Jadin" authorityYear="2009" box="[1152,1298,1229,1256]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="escalerae">
|
||
<emphasis id="88A6EAE26167FFC2FB404598FAA8FB43" box="[1152,1298,1229,1256]" italics="true" pageId="7" pageNumber="8">A. escalerae</emphasis>
|
||
</taxonomicName>
|
||
), and in having a lower number of scales around midbody (
|
||
<quantity id="7D2A9B156167FFC2FCD245A7FCCCFAAF" box="[786,886,1265,1292]" metricMagnitude="-1" metricUnit="m" metricValue="9.017" metricValueMax="9.144" metricValueMin="8.89" pageId="7" pageNumber="8" unit="in" value="35.5" valueMax="36.0" valueMin="35.0">35-36 in</quantity>
|
||
<emphasis id="88A6EAE26167FFC2FCBD45A7FBF6FAAF" box="[893,1100,1265,1291]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="7DD24D736167FFC2FCBD45A7FB9EFAAF" authorityName="Kok & Rivas" authorityYear="2011" box="[893,1060,1265,1291]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="mcdiarmidi">A. mcdiarmidi</taxonomicName>
|
||
vs.
|
||
</emphasis>
|
||
<quantity id="7D2A9B156167FFC2FB9345A7FB35FAAF" box="[1107,1167,1265,1291]" metricMagnitude="0" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="8" unit="in" value="40.0">40 in</quantity>
|
||
<taxonomicName id="7DD24D736167FFC2FB5645A7FA9CFAAF" authorityName="Myers, Rivas & Jadin" authorityYear="2009" box="[1174,1318,1265,1291]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="escalerae">
|
||
<emphasis id="88A6EAE26167FFC2FB5645A7FA9CFAAF" box="[1174,1318,1265,1291]" italics="true" pageId="7" pageNumber="8">A. escalerae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F2C8657B6167FFCBFF7D4465FCD9FEC8" lastPageId="14" lastPageNumber="15" pageId="7" pageNumber="8" type="description">
|
||
<paragraph id="BA6D36F06167FFC2FF7D4465FDACFAE9" blockId="7.[189,1399,1331,1608]" box="[189,534,1331,1357]" pageId="7" pageNumber="8">
|
||
<heading id="E125819C6167FFC2FF7D4465FDACFAE9" bold="true" box="[189,534,1331,1357]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
|
||
<emphasis id="88A6EAE26167FFC2FF7D4465FDACFAE9" bold="true" box="[189,534,1331,1357]" pageId="7" pageNumber="8">
|
||
Description of the
|
||
<typeStatus id="656988526167FFC2FE644465FDACFAE9" box="[420,534,1331,1357]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F06167FFC2FF7D4401FDB9F9EC" blockId="7.[189,1399,1331,1608]" pageId="7" pageNumber="8">
|
||
An adult male in good condition (
|
||
<figureCitation id="22E92A756167FFC2FD964401FD64FAD5" box="[598,734,1367,1393]" captionStart="Fig" captionStartId="3.[189,232,1895,1921]" captionTargetBox="[207,1380,265,1855]" captionTargetId="figure@3.[207,1380,265,1855]" captionTargetPageId="3" captionText="Fig. 1. Anadia mcdiarmidi sp. nov. A. IRSNB 2677, ♂ holotype in life. B. IRSNB 2675, juvenile male paratype in life. C. Ventral view of the anaesthetized holotype (grid squares = 5 mm). D. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711437" httpUri="https://zenodo.org/record/3711437/files/figure.png" pageId="7" pageNumber="8">Figs 1A, C</figureCitation>
|
||
,
|
||
<figureCitation id="22E92A756167FFC2FD2D4401FD44FAD5" box="[749,766,1367,1393]" captionStart="Fig" captionStartId="4.[189,232,1965,1991]" captionTargetBox="[189,1372,1072,1925]" captionTargetId="figure@4.[189,1398,1072,1925]" captionTargetPageId="4" captionText="Fig. 2. Anadia mcdiarmidi sp. nov. Ventral, dorsal, and lateral views of the head of the holotype (IRSNB 2677). Scale lines = 5 mm. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711439" httpUri="https://zenodo.org/record/3711439/files/figure.png" pageId="7" pageNumber="8">2</figureCitation>
|
||
,
|
||
<figureCitation id="22E92A756167FFC2FCCC4401FCA6FAD5" box="[780,796,1367,1393]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,467,1890]" captionTargetId="figure@5.[189,1399,467,1890]" captionTargetPageId="5" captionText="Fig. 3. Anadia mcdiarmidi sp. nov. Subpelvic region of the preserved ♂ holotype (IRSNB 2677), showing preanal scales and femoral pores, and trunk of the preserved holotype in lateral view (head to left), showing scalation at midbody. Scale lines = 5 mm. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711441" httpUri="https://zenodo.org/record/3711441/files/figure.png" pageId="7" pageNumber="8">3</figureCitation>
|
||
,
|
||
<figureCitation id="22E92A756167FFC2FCEC4401FC87FAD5" box="[812,829,1367,1393]" captionStart="Fig" captionStartId="6.[189,232,1000,1026]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionText="Fig. 4. Anadia mcdiarmidi sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.14766542" httpUri="https://zenodo.org/record/14766542/files/figure.png" pageId="7" pageNumber="8">4</figureCitation>
|
||
, 6), with
|
||
<quantity id="7D2A9B156167FFC2FC724401FBA7FAD5" box="[946,1053,1367,1393]" metricMagnitude="-2" metricUnit="m" metricValue="6.22" pageId="7" pageNumber="8" unit="mm" value="62.2">62.2 mm</quantity>
|
||
SVL and 110.0 mm TL (see also
|
||
<tableCitation id="F750034B6167FFC2FF32442DFEF2FA31" box="[242,328,1403,1429]" captionStart="Table 1" captionStartId="13.[189,258,822,848]" captionTargetId="graphics@13.[191,1397,948,2017]" captionTargetPageId="13" captionText="Table 1. Sex, measurements (in mm) and scale counts of the type series of Anadia mcdiarmidi sp. nov. 1 = tail incomplete or obviously regenerated, 2 = data corresponding to right/left side, other abbreviations are defined in the text." httpUri="http://table.plazi.org/id/EEAD6678616DFFC8FF7D4260FE79FC33" pageId="7" pageNumber="8" tableUuid="EEAD6678616DFFC8FF7D4260FE79FC33">Table 1</tableCitation>
|
||
). A rather robust
|
||
<taxonomicName id="7DD24D736167FFC2FDD4442DFDD3FA31" authorityName="Gray" authorityYear="1845" box="[532,617,1403,1429]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis id="88A6EAE26167FFC2FDD4442DFDD3FA31" box="[532,617,1403,1429]" italics="true" pageId="7" pageNumber="8">Anadia</emphasis>
|
||
</taxonomicName>
|
||
species with moderately long limbs (digits overlap when limbs are adpressed along body). Snout attenuate, flat in profile. HL 25% of SVL, 1.5 times longer than wide, 1.6 times wider than high; head conspicuously wider than neck. Neck long, 68% of HL, 35% of AXG. SAL 93% of AXG, 46% of SVL. Body wider than deep. Tail dorsoventrally flattened, about 1.8 times longer than SVL. Limbs pentadactyl with all digits clawed. Forelimb 27% of SVL, 52% of AXG; hind limb 36% of SVL, 73% of AXG.
|
||
</paragraph>
|
||
<paragraph id="BA6D36F06167FFC2FF7D4726FBB8F8BD" blockId="7.[189,1399,1647,1818]" pageId="7" pageNumber="8">
|
||
Scutellation on the dorsal surface of head is typical for the genus [see fig.
|
||
<quantity id="7D2A9B156167FFC2FBEC4726FBE0F92E" box="[1068,1114,1648,1674]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="8" unit="in" value="1.0">1 in</quantity>
|
||
<bibRefCitation id="DE434B016167FFC2FBA24726FAACF92E" author="Oftedal O. T." box="[1122,1302,1647,1674]" firstAuthor="Oftedal" journalOrPublisher="Arquivos de Zoologia, Sao Paulo" pageId="7" pageNumber="8" pagination="203 - 265" part="25" refId="ref8210" refString="Oftedal O. T. 1974. A revision of the genus Anadia (Sauria, Teiidae). Arquivos de Zoologia, Sao Paulo 25 (4): 203 - 265." title="A revision of the genus Anadia (Sauria, Teiidae)" type="journal article" year="1974">Oftedal (1974)</bibRefCitation>
|
||
], except the condition of the parietals and interparietal (see below and
|
||
<figureCitation id="22E92A756167FFC2FC6847C2FC4FF90A" box="[936,1013,1684,1710]" captionStart="Fig" captionStartId="4.[189,232,1965,1991]" captionTargetBox="[189,1372,1072,1925]" captionTargetId="figure@4.[189,1398,1072,1925]" captionTargetPageId="4" captionText="Fig. 2. Anadia mcdiarmidi sp. nov. Ventral, dorsal, and lateral views of the head of the holotype (IRSNB 2677). Scale lines = 5 mm. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711439" httpUri="https://zenodo.org/record/3711439/files/figure.png" pageId="7" pageNumber="8">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="22E92A756167FFC2FBC347C2FBAEF90A" box="[1027,1044,1684,1710]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[219,1380,874,1814]" captionTargetId="figure@9.[207,1382,855,1890]" captionTargetPageId="9" captionText="Fig. 7. Anadia mcdiarmidi sp. nov. Intrapopulation variation in the condition of the parietals and the interparietal (frontoparietals in grey). Top, from left to right: IRSNB 2677, IRSNB 2676; bottom, from left to right: IRSNB 2675, IRSNB 2674. Drawings not to scale." figureDoi="http://doi.org/10.5281/zenodo.3711449" httpUri="https://zenodo.org/record/3711449/files/figure.png" pageId="7" pageNumber="8">7</figureCitation>
|
||
). The frontal is damaged, the frontoparietals are slightly damaged. Head scales are smooth with a few minute scattered pits, those pits are especially concentrated on the upper and posterior margins of supralabials, on the lower and posterior margins of infralabials, and on the posterior margin of rostral.
|
||
</paragraph>
|
||
<paragraph id="BA6D36F06167FFCDFF7D4617FAE3FEC8" blockId="7.[189,1399,1857,2027]" lastBlockId="8.[189,1399,267,365]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">Rostral much wider than deep, laterally in contact with nasals and first supralabial, dorsally in contact with large frontonasal. Frontonasal with nearly straight anterior margin, posteriorly pointed. Paired prefrontals in contact, medial suture short. Frontal damaged, but obviously wider anteriorly and with blunt anterior point. Paired frontoparietals slightly damaged, with long medial suture, in contact with interparietal, parietals, and two posterior supraoculars. Three large supraoculars subequal in size. Interparietal divided into about two scales unequal in size. Parietals not extending as far as interparietal posteriorly, divided into two scales on the left side, into four scales on the right side. A series of six small to large occipitals (postparietals) circling the common posterior margin of parietals and interparietal.</paragraph>
|
||
<caption id="EEAD66786168FFCDFF7D46DCFD41F84F" ID-DOI="http://doi.org/10.5281/zenodo.3711447" ID-Zenodo-Dep="3711447" httpUri="https://zenodo.org/record/3711447/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1930,1956]" targetBox="[248,1389,501,1830]" targetPageId="8">
|
||
<paragraph id="BA6D36F06168FFCDFF7D46DCFD41F84F" blockId="8.[189,1398,1930,2027]" pageId="8" pageNumber="9">
|
||
<emphasis id="88A6EAE26168FFCDFF7D46DCFEB5F800" bold="true" box="[189,271,1930,1956]" pageId="8" pageNumber="9">Fig. 6.</emphasis>
|
||
<taxonomicName id="7DD24D736168FFCDFEDA46DCFDB8F800" authority="Kok & Rivas, 2011" authorityName="Kok & Rivas" authorityYear="2011" box="[282,514,1930,1956]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="mcdiarmidi" status="sp. nov.">
|
||
<emphasis id="88A6EAE26168FFCDFEDA46DCFDB8F800" box="[282,514,1930,1956]" italics="true" pageId="8" pageNumber="9">Anadia mcdiarmidi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="939557996168FFCDFDCE46DDFDD7F801" box="[526,621,1931,1957]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Intrapopulation variation in dorsal (above) and ventral (below) colour pattern in preserved specimens. From left to right: IRSNB 2677, IRSNB 2674, IRSNB 2676, IRSNB 2675. Photographs by Philippe J.R. Kok.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BA6D36F06169FFCCFF7D405DFD96FD9B" blockId="9.[189,1399,267,575]" pageId="9" pageNumber="10">Nasal scale entire, with a small indentation situated anterodorsally to nostril on the left side. Nasal scale in contact with rostral, posteriorly in contact with prefrontal, and in broad contact with loreal. Loreal large, pentagonal, in broad contact with first superciliary (= presuperciliary) and frenocular, in point contact with preocular. One small preocular, three postoculars. First superciliary large, followed by four shorter superciliaries. No small azygous scale between superciliaries and supraoculars. Frenocular followed posteriorly by three suboculars. Third subocular slightly extending to lip between supralabials 4-5 on the left side only. Suboculars separated from palpebrals by 2-3 poorly defined rows of mostly inconspicuous tiny scales, two of them distinctly enlarged with angular downward protrusion between suboculars. Seven supralabials.</paragraph>
|
||
<paragraph id="BA6D36F06169FFCCFF7D4330FE00FD07" blockId="9.[189,1398,614,675]" pageId="9" pageNumber="10">Eight ciliaries along upper eyelid. Lower eyelid scales translucent, slightly pigmented, with four higherthan-wide palpebrals.</paragraph>
|
||
<paragraph id="BA6D36F06169FFCCFF7D439DFD82FC8F" blockId="9.[189,1399,715,812]" pageId="9" pageNumber="10">Temporal scales subimbricate, smooth, with flat surfaces, slightly larger above and smaller below. Ear opening broad, ovoid, slightly inclined posterodorsally, edged with small, slightly pebblelike scales; tympanum recessed, pigmented.</paragraph>
|
||
<caption id="EEAD66786169FFCCFF7D46DCFC1FF84F" ID-DOI="http://doi.org/10.5281/zenodo.3711449" ID-Zenodo-Dep="3711449" httpUri="https://zenodo.org/record/3711449/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1930,1956]" targetBox="[219,1380,874,1814]" targetPageId="9">
|
||
<paragraph id="BA6D36F06169FFCCFF7D46DCFC1FF84F" blockId="9.[189,1399,1930,2027]" pageId="9" pageNumber="10">
|
||
<emphasis id="88A6EAE26169FFCCFF7D46DCFEB7F800" bold="true" box="[189,269,1930,1956]" pageId="9" pageNumber="10">Fig. 7.</emphasis>
|
||
<taxonomicName id="7DD24D736169FFCCFED646DCFE46F800" authority="Kok & Rivas, 2011" authorityName="Kok & Rivas" authorityYear="2011" box="[278,508,1930,1956]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mcdiarmidi" status="sp. nov.">
|
||
<emphasis id="88A6EAE26169FFCCFED646DCFE46F800" box="[278,508,1930,1956]" italics="true" pageId="9" pageNumber="10">Anadia mcdiarmidi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="939557996169FFCCFDC546DDFDD8F801" box="[517,610,1931,1957]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Intrapopulation variation in the condition of the parietals and the interparietal (frontoparietals in grey). Top, from left to right: IRSNB 2677, IRSNB 2676; bottom, from left to right: IRSNB 2675, IRSNB 2674. Drawings not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BA6D36F0616AFFCFFF7D405DFAD4FE5F" blockId="10.[189,1399,267,508]" pageId="10" pageNumber="11">Underside of head with six infralabials on each side. A large mental followed by a large postmental in contact with first two infralabials. Two pairs of large genials in broad contact medially, in lateral contact with infralabials 3-4; two pair of postgenials, the first pair large, each scale in contact with last genial and infralabials 4-5, the second pair much smaller, each scale in contact with first postgenial and in point contact with fifth infralabial. Gulars arranged in transverse rows, becoming slightly larger posteriad, culminating in a well-defined collar row of eight subequal scales; no distinctly enlarged longitudinal rows. Side of neck between ear and collar pebbled with subequal irregularly shaped juxtaposed scales.</paragraph>
|
||
<paragraph id="BA6D36F0616AFFCFFF7D4375FAA7FD20" blockId="10.[189,1399,547,645]" pageId="10" pageNumber="11">Middorsal scales 54. Dorsal scales on neck subimbricate, irregularly shaped, with rounded surfaces. Dorsal body scales smooth, juxtaposed, mainly quadrangular, a few irregularly shaped, but most are longer than wide, in transverse rows only. Lateral scales smooth, similar to dorsals, but smaller.</paragraph>
|
||
<paragraph id="BA6D36F0616AFFCFFF7D43FAFBE9FCAA" blockId="10.[189,1398,684,782]" pageId="10" pageNumber="11">
|
||
Ventral scales much wider than dorsals, smooth, juxtaposed, quadrangular, slightly longer than wide except for
|
||
<emphasis id="88A6EAE2616AFFCFFE854387FED2FD4E" box="[325,360,721,746]" italics="true" pageId="10" pageNumber="11">ca.</emphasis>
|
||
four median longitudinal rows of square to wider than long scales on belly; in 14 longitudinal rows at midbody and 31 transverse rows between collar and vent.
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616AFFCFFF7D4263FB76FC33" blockId="10.[189,1399,821,919]" pageId="10" pageNumber="11">
|
||
Eight marginal preanal scales anterior to vent, the middle two narrower than the adjacent lateral ones, most outer ones very small. Femoral pores well developed, in swollen pore scales mostly in linear contact; 9 pores on left thigh, 10 on right. Pores do not extend onto preanal area (
|
||
<figureCitation id="22E92A75616AFFCFFBB5422BFB06FC33" box="[1141,1212,893,919]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1399,467,1890]" captionTargetId="figure@5.[189,1399,467,1890]" captionTargetPageId="5" captionText="Fig. 3. Anadia mcdiarmidi sp. nov. Subpelvic region of the preserved ♂ holotype (IRSNB 2677), showing preanal scales and femoral pores, and trunk of the preserved holotype in lateral view (head to left), showing scalation at midbody. Scale lines = 5 mm. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711441" httpUri="https://zenodo.org/record/3711441/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616AFFCFFF7D42E8FBDEFC7C" blockId="10.[189,1124,958,985]" box="[189,1124,958,985]" pageId="10" pageNumber="11">Caudal scales smooth, subimbricate, rectangular, in transverse rows around tail.</paragraph>
|
||
<paragraph id="BA6D36F0616AFFCFFF7D4556FBD0FB21" blockId="10.[189,1398,1024,1157]" pageId="10" pageNumber="11">Scales on dorsal surfaces of arm large, smooth, imbricate; ventral surfaces of arm with similar, but smaller and less imbricate scales. Hind limbs with large, smooth subimbricate to imbricate scales on anterior face of thighs and on lower legs; scales similar but smaller on ventral side of thigh. Dorsal and posterior sides of thigh with much smaller, irregularly shaped juxtaposed scales.</paragraph>
|
||
<paragraph id="BA6D36F0616AFFCFFF7D45FBFBF6FADE" blockId="10.[189,1399,1197,1402]" pageId="10" pageNumber="11">
|
||
Moderate-size, imbricate scales atop hands and feet. Supradigital scales single; upper and lower ungualsheath scales covering base of claws, leaving tips well exposed. Palms and soles with small slightly raised juxtaposed scales (
|
||
<figureCitation id="22E92A75616AFFCFFE3645A2FDFEFAAB" box="[502,580,1268,1295]" captionStart="Fig" captionStartId="6.[189,232,1000,1026]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionText="Fig. 4. Anadia mcdiarmidi sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.14766542" httpUri="https://zenodo.org/record/14766542/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
|
||
). Two enlarged smooth thenar scales at base of pollex. Subdigital lamellae mostly single. Lamellae under first (I) through fifth (V) finger (right/left side): I: 7/6, II: 9/9, III: 11/11, IV: 13/14, V: 10/9. Lamellae under first (I) through fifth (V) toe (right/left side): I: 6/6, II: 10/10, III: 14/12, IV: 17/15, V: 15/6 (large part of Toe V missing on left side).
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616AFFCFFF7D44F4FC6CF9EE" blockId="10.[189,1399,1441,1610]" pageId="10" pageNumber="11">Tongue lanceolate; proximal two-third unpigmented, distal one-third heavily pigmented, including fork. Upper surface behind fork covered with imbricate scalelike papillae that extend onto ventrolateral sides of tongue. Raised medioventral side of tongue with numerous (>10) of thin, oblique, anteriorly converging and pointed (chevronlike) infralingual plicae; anterior pair of plicae swollen, bluntly pointed and larger than those following. Very shallow medioventral groove.</paragraph>
|
||
<paragraph id="BA6D36F0616AFFCFFF7D4724FE37F90B" blockId="10.[189,1399,1650,1711]" pageId="10" pageNumber="11">Anterior maxillary and dentary teeth conical, unicuspid, with no or very feeble recurvature, becoming larger posteriorly.</paragraph>
|
||
<paragraph id="BA6D36F0616AFFC9FF7D4781FB4DFEEF" blockId="10.[189,1399,1751,2027]" lastBlockId="12.[189,1399,267,331]" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
|
||
The left hemipenis (
|
||
<figureCitation id="22E92A75616AFFCFFE6B4781FE4BF956" box="[427,497,1751,1778]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionTargetPageId="6" captionText="Fig. 5. Anadia mcdiarmidi sp. nov. The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711443" httpUri="https://zenodo.org/record/3711443/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
|
||
) extends about four subcaudal rows when adpressed to the tail, and is about 7.0 mm in length and 4.0 mm across its widest point when fully everted (organ not maximally expanded because one of the lobes was slightly damaged during the eversion process). Hemipenial body globose, tapered near the base, weakly and symmetrically bilobed. Sulcus spermaticus broader near crotch, relatively shallow, curving about halfway around the base of the organ, then running medially towards the lobes. Sulcus flanked by a narrow nude area on each side, and divided in the crotch by a small fleshy bumplike structure, from where two narrow, ill-defined and barely detectable sulcus branches seem to extend laterally into the folded thickened 'ear-like' tissues of each lobe towards the apices of the lobes. Lobes are complexly folded, not flattened terminally, and sulcus branches seem to end in small depressions (from which residual seminal fluid was removed before hemipenis preparation).
|
||
</paragraph>
|
||
<caption id="EEAD6678616BFFCEFF7D46DCFEFFF84F" ID-DOI="http://doi.org/10.5281/zenodo.3711451" ID-Zenodo-Dep="3711451" httpUri="https://zenodo.org/record/3711451/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1930,1956]" targetBox="[271,1316,265,1890]" targetPageId="11">
|
||
<paragraph id="BA6D36F0616BFFCEFF7D46DCFEFFF84F" blockId="11.[189,1399,1930,2027]" pageId="11" pageNumber="12">
|
||
<emphasis id="88A6EAE2616BFFCEFF7D46DCFEAAF800" bold="true" box="[189,272,1930,1956]" pageId="11" pageNumber="12">Fig. 8.</emphasis>
|
||
Map of the Guiana Shield region (
|
||
<emphasis id="88A6EAE2616BFFCEFD1046DCFD5FF800" bold="true" box="[720,741,1930,1956]" pageId="11" pageNumber="12">A</emphasis>
|
||
), with enlarged black rectangle corresponding to the Chimantá Massif (
|
||
<emphasis id="88A6EAE2616BFFCEFE5646F8FE10F86C" bold="true" box="[406,426,1966,1992]" pageId="11" pageNumber="12">B</emphasis>
|
||
). Black triangle indicates type locality, black dots indicate other know localities for the species.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BA6D36F0616CFFC9FF7D4022FD5BFCC8" blockId="12.[189,1399,371,876]" pageId="12" pageNumber="13">
|
||
Very base of hemipenis nude, pigmented in life. Lateral and asulcate faces of the organ with a series of roughly equidistant oblique plicae or flounces with weakly calcified spinules. Twenty-three rows of flounces extending along the organ. Basal flounces shorter, almost in straight transverse lines, the three first rows almost continuous across the hemipenial base, except for narrow nude areas on each lateral face and for nude areas edging each side of the sulcus spermaticus; five subsequent rows continuous across hemipenial body (except for nude areas edging each side of the sulcus spermaticus), with the seventh and eight rows distinctly pointing medially towards the apex on the asulcate face (those eight flounces proximal to the median nude space of the asulcate face correspond to the 'medioproximal asulcate flounces'
|
||
<emphasis id="88A6EAE2616CFFC9FE5643CCFE6DFD17" box="[406,471,666,691]" italics="true" pageId="12" pageNumber="13">sensu</emphasis>
|
||
<bibRefCitation id="DE434B01616CFFC9FE2143CCFD07FD10" author="Myers C. W. & Rivas Fuenmayor G. & Jadin R. C." box="[481,701,665,692]" etAl="et al." firstAuthor="Myers" journalOrPublisher="American Museum Novitates" pageId="12" pageNumber="13" pagination="1 - 31" part="3660" refId="ref8112" refString="Myers C. W., Rivas Fuenmayor G. & Jadin R. C. 2009. New species of lizards from Auyantepui and La Escalera in the Venezuelan Guayana, with notes on " microteiid " hemipenes (Squamata: Gymnophthalmidae). American Museum Novitates 3660: 1 - 31. http: // dx. doi. org / 10.1206 / 657.1" title="New species of lizards from Auyantepui and La Escalera in the Venezuelan Guayana, with notes on " microteiid " hemipenes (Squamata: Gymnophthalmidae)" type="journal article" year="2009">
|
||
Myers
|
||
<emphasis id="88A6EAE2616CFFC9FDF643CCFDC9FD17" box="[566,627,665,691]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
); fifteen remaining rows chevron-shaped, also circling the organ, but separated by a broad median nude space on the asulcate face and by the nude areas edging each side of the sulcus spermaticus. The plicae bear comblike rows of minute weakly mineralized spinules that project slightly from the edges of the plicae and usually increase in size towards the lateral faces and the basal part of the organ (staining of the spinules by Alizarin Red is well visible under the microscope although not observable in
|
||
<figureCitation id="22E92A75616CFFC9FD4B4207FD68FCC8" box="[651,722,849,876]" captionStart="Fig" captionStartId="6.[189,232,1824,1850]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionTargetPageId="6" captionText="Fig. 5. Anadia mcdiarmidi sp. nov. The left fully everted—not maximally expanded—hemipenis of the holotype (IRSNB 2677). Left: sulcate side. Centre: lateral view, turned with sulcate side facing left. Right: asulcate side. BLS = fleshy bumplike structure; MPAF = medioproximal asulcate flounces; NSAF = median nude space of the asulcate face; SB = sulcus branches; SF = spinulate flounces; SS = sulcus spermaticus. Scale line = 2 mm. Although staining of the spinules by Alizarin Red is well visible under the microscope, the structures are not clearly observable in the figure. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711443" httpUri="https://zenodo.org/record/3711443/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616CFFC9FF7D42C2FD90FC0A" blockId="12.[189,1399,916,1127]" box="[189,554,916,943]" pageId="12" pageNumber="13">
|
||
<heading id="E125819C616CFFC9FF7D42C2FD90FC0A" bold="true" box="[189,554,916,943]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="88A6EAE2616CFFC9FF7D42C2FD90FC0A" bold="true" box="[189,554,916,943]" pageId="12" pageNumber="13">
|
||
Colour of the
|
||
<typeStatus id="65698852616CFFC9FEAA42C3FE66FC0B" box="[362,476,917,943]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
|
||
in life
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616CFFC9FF7D42ECFA8DFBC3" blockId="12.[189,1399,916,1127]" pageId="12" pageNumber="13">
|
||
Dorsum greyish to bluish brown depending on light intensity, devoid of any conspicuous pattern, but with a few scattered black scales on flanks. Arms and legs, including digits, greyish brown with a few scattered black scales; rear of thighs with a few ill-defined dark markings. Ventral surfaces of head and body golden grey, ventral surfaces of legs and tail bluish grey, palms and soles lemon yellow speckled with brown and yellowish orange scales. Tympanum dark brown, iris pale yellow (
|
||
<figureCitation id="22E92A75616CFFC9FB47451AFAB1FBC3" box="[1159,1291,1100,1127]" captionStart="Fig" captionStartId="3.[189,232,1895,1921]" captionTargetBox="[207,1380,265,1855]" captionTargetId="figure@3.[207,1380,265,1855]" captionTargetPageId="3" captionText="Fig. 1. Anadia mcdiarmidi sp. nov. A. IRSNB 2677, ♂ holotype in life. B. IRSNB 2675, juvenile male paratype in life. C. Ventral view of the anaesthetized holotype (grid squares = 5 mm). D. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711437" httpUri="https://zenodo.org/record/3711437/files/figure.png" pageId="12" pageNumber="13">Figs 1A, C</figureCitation>
|
||
,
|
||
<figureCitation id="22E92A75616CFFC9FAD7451BFA93FBC3" box="[1303,1321,1101,1127]" captionStart="Fig" captionStartId="6.[189,232,1000,1026]" captionTargetBox="[189,1399,1085,1787]" captionTargetId="figure@6.[189,1399,1099,1787]" captionText="Fig. 4. Anadia mcdiarmidi sp. nov. Left hand and right foot of the holotype (IRSNB 2677) in life. Scale line = 5 mm. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.14766542" httpUri="https://zenodo.org/record/14766542/files/figure.png" pageId="12" pageNumber="13">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616CFFC9FF7D45D9FD27FB0D" blockId="12.[189,1399,1167,1378]" box="[189,669,1167,1194]" pageId="12" pageNumber="13">
|
||
<heading id="E125819C616CFFC9FF7D45D9FD27FB0D" bold="true" box="[189,669,1167,1194]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="88A6EAE2616CFFC9FF7D45D9FD27FB0D" bold="true" box="[189,669,1167,1194]" pageId="12" pageNumber="13">
|
||
Colour of the
|
||
<typeStatus id="65698852616CFFC9FEAA45C6FE66FB0E" box="[362,476,1168,1194]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
|
||
in preservative
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616CFFC9FF7D45E2FC31FAC5" blockId="12.[189,1399,1167,1378]" pageId="12" pageNumber="13">
|
||
After approximately two months in preservative, the dorsal surfaces became dark brown (looking slightly peppered when specimen is in liquid), the ventral surfaces are metallic blue, except the anterior surface of head, which is light grey (
|
||
<figureCitation id="22E92A75616CFFC9FDB245A8FD06FABD" box="[626,700,1278,1305]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[248,1389,501,1830]" captionTargetId="figure@8.[189,1398,462,1889]" captionTargetPageId="8" captionText="Fig. 6. Anadia mcdiarmidi sp. nov. Intrapopulation variation in dorsal (above) and ventral (below) colour pattern in preserved specimens. From left to right: IRSNB 2677, IRSNB 2674, IRSNB 2676, IRSNB 2675. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711447" httpUri="https://zenodo.org/record/3711447/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
|
||
). Palms and soles are grey. Tympanum is dark brown. It is worthwhile to note that the ventral skin, and some parts that are light grey (like the anterior underside of head), turn blue or bluer when pressure is applied to the skin.
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616CFFC9FF7D44DCFE6CFA01" blockId="12.[189,1399,1418,1556]" box="[189,470,1418,1445]" pageId="12" pageNumber="13">
|
||
<heading id="E125819C616CFFC9FF7D44DCFE6CFA01" bold="true" box="[189,470,1418,1445]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="88A6EAE2616CFFC9FF7D44DCFE6CFA01" bold="true" box="[189,470,1418,1445]" pageId="12" pageNumber="13">
|
||
Variation in
|
||
<typeStatus id="65698852616CFFC9FE9A44DDFE6CFA01" box="[346,470,1419,1445]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616CFFC9FF7D44F9FE25F9B0" blockId="12.[189,1399,1418,1556]" pageId="12" pageNumber="13">
|
||
The
|
||
<typeStatus id="65698852616CFFC9FF3544E6FEDFFA6E" box="[245,357,1456,1482]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
|
||
consist in
|
||
<specimenCount id="ACD4FD79616CFFC9FE2B44F9FD85FA6D" box="[491,575,1455,1481]" pageId="12" pageNumber="13" type="adult">1 adult</specimenCount>
|
||
<specimenCount id="ACD4FD79616CFFC9FD8944E6FDE5FA6E" box="[585,607,1456,1482]" pageId="12" pageNumber="13" type="male">♂</specimenCount>
|
||
(SVL
|
||
<quantity id="7D2A9B15616CFFC9FD7344F9FC98FA6E" box="[691,802,1455,1482]" metricMagnitude="-2" metricUnit="m" metricValue="6.04" pageId="12" pageNumber="13" unit="mm" value="60.4">60.4 mm</quantity>
|
||
), and
|
||
<specimenCount id="ACD4FD79616CFFC9FCAE44F9FC5FFA6D" box="[878,997,1455,1481]" pageId="12" pageNumber="13" type="juvenile">2 juvenile</specimenCount>
|
||
<specimenCount id="ACD4FD79616CFFC9FC2F44E6FBBFFA6E" box="[1007,1029,1456,1482]" pageId="12" pageNumber="13" type="male">♂</specimenCount>
|
||
(SVL from
|
||
<quantity id="7D2A9B15616CFFC9FB5C44F9FAD1FA6E" box="[1180,1387,1455,1482]" metricMagnitude="-2" metricUnit="m" metricValue="4.04" metricValueMax="4.26" metricValueMin="3.82" pageId="12" pageNumber="13" unit="mm" value="40.4" valueMax="42.6" valueMin="38.2">38.2 to 42.6 mm</quantity>
|
||
).
|
||
<tableCitation id="F750034B616CFFC9FF7D4482FEAFFA4A" box="[189,277,1492,1518]" captionStart="Table 1" captionStartId="13.[189,258,822,848]" captionTargetId="graphics@13.[191,1397,948,2017]" captionTargetPageId="13" captionText="Table 1. Sex, measurements (in mm) and scale counts of the type series of Anadia mcdiarmidi sp. nov. 1 = tail incomplete or obviously regenerated, 2 = data corresponding to right/left side, other abbreviations are defined in the text." httpUri="http://table.plazi.org/id/EEAD6678616DFFC8FF7D4260FE79FC33" pageId="12" pageNumber="13" tableUuid="EEAD6678616DFFC8FF7D4260FE79FC33">Table 1</tableCitation>
|
||
provides measurements of the type series, and
|
||
<figureCitation id="22E92A75616CFFC9FC8B4482FC1EFA4A" box="[843,932,1492,1518]" captionStart="Fig" captionStartId="3.[189,232,1895,1921]" captionTargetBox="[207,1380,265,1855]" captionTargetId="figure@3.[207,1380,265,1855]" captionTargetPageId="3" captionText="Fig. 1. Anadia mcdiarmidi sp. nov. A. IRSNB 2677, ♂ holotype in life. B. IRSNB 2675, juvenile male paratype in life. C. Ventral view of the anaesthetized holotype (grid squares = 5 mm). D. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711437" httpUri="https://zenodo.org/record/3711437/files/figure.png" pageId="12" pageNumber="13">Fig. 1B</figureCitation>
|
||
shows one of the juvenile
|
||
<typeStatus id="65698852616CFFC9FB274483FAEDFA4B" box="[1255,1367,1493,1519]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
|
||
in life (IRSNB 2675).
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616CFFC8FF7D476AFEECFEED" blockId="12.[189,1399,1596,2028]" lastBlockId="13.[189,1398,267,329]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
One of the main differences among specimens of the type series is the occurrence of a presupraocular scale in the three
|
||
<typeStatus id="65698852616CFFC9FE4E4737FE44F9DF" box="[398,510,1633,1659]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
|
||
(absent in the
|
||
<typeStatus id="65698852616CFFC9FD6F4737FCA1F9D8" box="[687,795,1633,1660]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
); that presupraocular scale is divided on both sides in IRSNB 2675. IRSNB 2676 has four suboculars on both sides (
|
||
<emphasis id="88A6EAE2616CFFC9FC1847D0FC4DF93B" box="[984,1015,1670,1695]" italics="true" pageId="12" pageNumber="13">vs.</emphasis>
|
||
three in the
|
||
<typeStatus id="65698852616CFFC9FB5547D3FB46F93B" box="[1173,1276,1669,1695]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
and other
|
||
<typeStatus id="65698852616CFFC9FF7D47FDFE89F961" box="[189,307,1707,1733]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
|
||
). Supralabial counts are consistent, but infralabial counts vary from 5 (n=1) to 7 (n=1, one side only). Pits on head scales are inconspicuous in the two juveniles (
|
||
<emphasis id="88A6EAE2616CFFC9FC274786FBBDF94D" box="[999,1031,1744,1769]" italics="true" pageId="12" pageNumber="13">vs.</emphasis>
|
||
well visible in the adult males). Suboculars are always separated from palpebrals by 2-3 poorly defined rows of inconspicuous tiny scales. In IRSNB 2674 three of these scales (
|
||
<emphasis id="88A6EAE2616CFFC9FD0B464CFD51F897" box="[715,747,1818,1843]" italics="true" pageId="12" pageNumber="13">vs.</emphasis>
|
||
two in the
|
||
<typeStatus id="65698852616CFFC9FCAD464FFC6CF890" box="[877,982,1817,1844]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
) are distinctly enlarged (larger than in
|
||
<typeStatus id="65698852616CFFC9FF1D466BFEFCF8FC" box="[221,326,1853,1880]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
) with angular downward protrusion between suboculars. In both juvenile
|
||
<typeStatus id="65698852616CFFC9FB014668FA8EF8FC" box="[1217,1332,1854,1880]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
|
||
, only one of these scales is only slightly enlarged and slightly protrudes between suboculars. The condition of the parietals and interparietal is highly variable among specimens and none of the specimens examined has complete interparietal and parietals, which are always divided in several smaller scales (see
|
||
<figureCitation id="22E92A75616CFFC9FAF146FAFACDF862" box="[1329,1399,1964,1990]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[219,1380,874,1814]" captionTargetId="figure@9.[207,1382,855,1890]" captionTargetPageId="9" captionText="Fig. 7. Anadia mcdiarmidi sp. nov. Intrapopulation variation in the condition of the parietals and the interparietal (frontoparietals in grey). Top, from left to right: IRSNB 2677, IRSNB 2676; bottom, from left to right: IRSNB 2675, IRSNB 2674. Drawings not to scale." figureDoi="http://doi.org/10.5281/zenodo.3711449" httpUri="https://zenodo.org/record/3711449/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
|
||
for comparison). Number of middorsal scales varies from 53 to 57. Ventrals are in 11 to 14 longitudinal rows and in 29 to 30 transverse rows. The number of femoral pores varies from 8 to 10. Preanals vary from 7 to 10.
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616DFFC8FF7D4026FACDFD9E" blockId="13.[189,1399,368,570]" pageId="13" pageNumber="14">
|
||
The fully everted, maximally expanded, right hemipenis of the adult
|
||
<specimenCount id="ACD4FD79616DFFC8FC084026FC64FE2E" box="[968,990,368,394]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
|
||
<typeStatus id="65698852616DFFC8FC224026FBFDFE2E" box="[994,1095,368,394]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
|
||
(IRSNB 2674) is identical to the holotype's organ, except that maximal expansion makes the sulcus spermaticus bifurcation and the sulcus branches even more difficult to appreciate. Maximal expansion does not highlight hemipenial features that would have been overlooked in the not maximally expanded hemipenis of the
|
||
<typeStatus id="65698852616DFFC8FAC8408FFAC9FE57" box="[1288,1395,473,499]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
. The hemipenis is
|
||
<quantity id="7D2A9B15616DFFC8FE4E40ABFE53FDB3" box="[398,489,509,535]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="13" pageNumber="14" unit="mm" value="6.6">6.6 mm</quantity>
|
||
in length and 4.0 mm across its widest point when everted; there are twentyfour rows of flounces extending along the organ,
|
||
<emphasis id="88A6EAE2616DFFC8FD324377FCAFFD9E" box="[754,789,545,570]" italics="true" pageId="13" pageNumber="14">ca.</emphasis>
|
||
eight of which are medioproximal asulcate flounces.
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616DFFCBFF7D4337FCD9FEC8" blockId="13.[189,1399,609,742]" lastBlockId="14.[189,1399,267,364]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
Colour in life ranges from beige to dark greyish, sometimes bluish, brown dorsally, depending on light intensity. Rear of thighs usually has a few ill-defined dark markings. Some blue markings may be visible around ear, or at the base of the tail, or on ventral parts. Ventral parts range from golden grey to pale yellow. One of the juvenile
|
||
<typeStatus id="65698852616DFFC8FDA3439AFD69FD42" box="[611,723,716,742]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
|
||
(IRSNB 2675, see
|
||
<figureCitation id="22E92A75616DFFC8FC7D439DFBADFD41" box="[957,1047,715,741]" captionStart="Fig" captionStartId="3.[189,232,1895,1921]" captionTargetBox="[207,1380,265,1855]" captionTargetId="figure@3.[207,1380,265,1855]" captionTargetPageId="3" captionText="Fig. 1. Anadia mcdiarmidi sp. nov. A. IRSNB 2677, ♂ holotype in life. B. IRSNB 2675, juvenile male paratype in life. C. Ventral view of the anaesthetized holotype (grid squares = 5 mm). D. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711437" httpUri="https://zenodo.org/record/3711437/files/figure.png" pageId="13" pageNumber="14">Fig. 1B</figureCitation>
|
||
) has two poorly defined dark brown stripes between eye and ear. In preservative all specimens became dark brown dorsally, looking slightly peppered when in liquid. Ventral parts are always metallic blue (
|
||
<figureCitation id="22E92A75616EFFCBFBC14078FBF2FEED" box="[1025,1096,302,329]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[248,1389,501,1830]" captionTargetId="figure@8.[189,1398,462,1889]" captionTargetPageId="8" captionText="Fig. 6. Anadia mcdiarmidi sp. nov. Intrapopulation variation in dorsal (above) and ventral (below) colour pattern in preserved specimens. From left to right: IRSNB 2677, IRSNB 2674, IRSNB 2676, IRSNB 2675. Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711447" httpUri="https://zenodo.org/record/3711447/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
|
||
). Tympanum in preserved specimens varies from dark brown to blue or golden blue.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="EEAD6678616DFFC8FF7D4260FE79FC33" ID-Table-UUID="EEAD6678616DFFC8FF7D4260FE79FC33" httpUri="http://table.plazi.org/id/EEAD6678616DFFC8FF7D4260FE79FC33" pageId="13" pageNumber="14" startId="13.[189,258,822,848]" targetBox="[253,1350,961,2004]" targetIsTable="true" targetPageId="13">
|
||
<paragraph id="BA6D36F0616DFFC8FF7D4260FE79FC33" blockId="13.[189,1399,822,919]" pageId="13" pageNumber="14">
|
||
<emphasis id="88A6EAE2616DFFC8FF7D4260FE9AFCF4" bold="true" box="[189,288,822,848]" pageId="13" pageNumber="14">Table 1.</emphasis>
|
||
Sex, measurements (in mm) and scale counts of the type series of
|
||
<taxonomicName id="7DD24D73616DFFC8FBF14260FAAEFCF4" authority="Kok & Rivas, 2011" authorityName="Kok & Rivas" authorityYear="2011" box="[1073,1300,822,848]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="mcdiarmidi" status="sp. nov.">
|
||
<emphasis id="88A6EAE2616DFFC8FBF14260FAAEFCF4" box="[1073,1300,822,848]" italics="true" pageId="13" pageNumber="14">Anadia mcdiarmidi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="93955799616DFFC8FADC4260FACDFCF4" box="[1308,1399,822,848]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<superScript id="4DA79BB8616DFFC8FF7D4201FF7CFCC1" attach="left" box="[189,198,855,869]" fontSize="6" pageId="13" pageNumber="14">1</superScript>
|
||
= tail incomplete or obviously regenerated,
|
||
<superScript id="4DA79BB8616DFFC8FD054201FD74FCC1" attach="none" box="[709,718,855,869]" fontSize="6" pageId="13" pageNumber="14">2</superScript>
|
||
= data corresponding to right/left side, other abbreviations are defined in the text.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BA6D36F0616DFFC8FF3D4294FA9AF862" pageId="13" pageNumber="14">
|
||
<table id="C8D2C450616D003AFF3D4297FAFCF870" box="[253,1350,961,2004]" gridcols="5" gridrows="21" pageId="13" pageNumber="14">
|
||
<tr id="04E234B2616D003AFF3D4297FAFCFC78" box="[253,1350,961,988]" gridrow="0" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D4297FECBFC78" box="[253,369,961,988]" gridcol="0" gridrow="0" pageId="13" pageNumber="14">Specimen</th>
|
||
<th id="47335DCE616D003AFE1B4297FDC9FC78" box="[475,627,961,988]" gridcol="1" gridrow="0" pageId="13" pageNumber="14">IRSNB 2677</th>
|
||
<th id="47335DCE616D003AFD0B4297FCD9FC78" box="[715,867,961,988]" gridcol="2" gridrow="0" pageId="13" pageNumber="14">IRSNB 2674</th>
|
||
<th id="47335DCE616D003AFC7B4297FBE9FC78" box="[955,1107,961,988]" gridcol="3" gridrow="0" pageId="13" pageNumber="14">IRSNB 2675</th>
|
||
<th id="47335DCE616D003AFB6E4297FAFCFC78" box="[1198,1350,961,988]" gridcol="4" gridrow="0" pageId="13" pageNumber="14">IRSNB 2676</th>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D42A3FAFCFBB4" box="[253,1350,1013,1040]" gridrow="1" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D42A3FECBFBB4" box="[253,369,1013,1040]" gridcol="0" gridrow="1" pageId="13" pageNumber="14">Sex</th>
|
||
<td id="47335DCE616D003AFE1B42A3FDC9FBB4" box="[475,627,1013,1040]" gridcol="1" gridrow="1" pageId="13" pageNumber="14">♂</td>
|
||
<td id="47335DCE616D003AFD0B42A3FCD9FBB4" box="[715,867,1013,1040]" gridcol="2" gridrow="1" pageId="13" pageNumber="14">♂</td>
|
||
<td id="47335DCE616D003AFC7B42A3FBE9FBB4" box="[955,1107,1013,1040]" gridcol="3" gridrow="1" pageId="13" pageNumber="14">Juvenile ♂</td>
|
||
<td id="47335DCE616D003AFB6E42A3FAFCFBB4" box="[1198,1350,1013,1040]" gridcol="4" gridrow="1" pageId="13" pageNumber="14">Juvenile ♂</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D457EFAFCFBE6" box="[253,1350,1064,1090]" gridrow="2" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D457EFECBFBE6" box="[253,369,1064,1090]" gridcol="0" gridrow="2" pageId="13" pageNumber="14">SVL</th>
|
||
<td id="47335DCE616D003AFE1B457EFDC9FBE6" box="[475,627,1064,1090]" gridcol="1" gridrow="2" pageId="13" pageNumber="14">62.2</td>
|
||
<td id="47335DCE616D003AFD0B457EFCD9FBE6" box="[715,867,1064,1090]" gridcol="2" gridrow="2" pageId="13" pageNumber="14">60.4</td>
|
||
<td id="47335DCE616D003AFC7B457EFBE9FBE6" box="[955,1107,1064,1090]" gridcol="3" gridrow="2" pageId="13" pageNumber="14">38.2</td>
|
||
<td id="47335DCE616D003AFB6E457EFAFCFBE6" box="[1198,1350,1064,1090]" gridcol="4" gridrow="2" pageId="13" pageNumber="14">42.6</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D450FFAFCFBD2" box="[253,1350,1113,1142]" gridrow="3" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D450FFECBFBD2" box="[253,369,1113,1142]" gridcol="0" gridrow="3" pageId="13" pageNumber="14">TL</th>
|
||
<td id="47335DCE616D003AFE1B450FFDC9FBD2" box="[475,627,1113,1142]" gridcol="1" gridrow="3" pageId="13" pageNumber="14">110.0</td>
|
||
<td id="47335DCE616D003AFD0B450FFCD9FBD2" box="[715,867,1113,1142]" gridcol="2" gridrow="3" pageId="13" pageNumber="14">57.01</td>
|
||
<td id="47335DCE616D003AFC7B450FFBE9FBD2" box="[955,1107,1113,1142]" gridcol="3" gridrow="3" pageId="13" pageNumber="14">29.71</td>
|
||
<td id="47335DCE616D003AFB6E450FFAFCFBD2" box="[1198,1350,1113,1142]" gridcol="4" gridrow="3" pageId="13" pageNumber="14">74.0</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D45DBFAFCFB0C" box="[253,1350,1165,1192]" gridrow="4" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D45DBFECBFB0C" box="[253,369,1165,1192]" gridcol="0" gridrow="4" pageId="13" pageNumber="14">HL</th>
|
||
<td id="47335DCE616D003AFE1B45DBFDC9FB0C" box="[475,627,1165,1192]" gridcol="1" gridrow="4" pageId="13" pageNumber="14">15.8</td>
|
||
<td id="47335DCE616D003AFD0B45DBFCD9FB0C" box="[715,867,1165,1192]" gridcol="2" gridrow="4" pageId="13" pageNumber="14">13.6</td>
|
||
<td id="47335DCE616D003AFC7B45DBFBE9FB0C" box="[955,1107,1165,1192]" gridcol="3" gridrow="4" pageId="13" pageNumber="14">9.0</td>
|
||
<td id="47335DCE616D003AFB6E45DBFAFCFB0C" box="[1198,1350,1165,1192]" gridcol="4" gridrow="4" pageId="13" pageNumber="14">10.2</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D4596FAFCFB7F" box="[253,1350,1216,1243]" gridrow="5" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D4596FECBFB7F" box="[253,369,1216,1243]" gridcol="0" gridrow="5" pageId="13" pageNumber="14">HW</th>
|
||
<td id="47335DCE616D003AFE1B4596FDC9FB7F" box="[475,627,1216,1243]" gridcol="1" gridrow="5" pageId="13" pageNumber="14">10.8</td>
|
||
<td id="47335DCE616D003AFD0B4596FCD9FB7F" box="[715,867,1216,1243]" gridcol="2" gridrow="5" pageId="13" pageNumber="14">8.7</td>
|
||
<td id="47335DCE616D003AFC7B4596FBE9FB7F" box="[955,1107,1216,1243]" gridcol="3" gridrow="5" pageId="13" pageNumber="14">5.8</td>
|
||
<td id="47335DCE616D003AFB6E4596FAFCFB7F" box="[1198,1350,1216,1243]" gridcol="4" gridrow="5" pageId="13" pageNumber="14">6.6</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D45A5FAFCFAAA" box="[253,1350,1267,1294]" gridrow="6" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D45A5FECBFAAA" box="[253,369,1267,1294]" gridcol="0" gridrow="6" pageId="13" pageNumber="14">HD</th>
|
||
<td id="47335DCE616D003AFE1B45A5FDC9FAAA" box="[475,627,1267,1294]" gridcol="1" gridrow="6" pageId="13" pageNumber="14">6.7</td>
|
||
<td id="47335DCE616D003AFD0B45A5FCD9FAAA" box="[715,867,1267,1294]" gridcol="2" gridrow="6" pageId="13" pageNumber="14">5.7</td>
|
||
<td id="47335DCE616D003AFC7B45A5FBE9FAAA" box="[955,1107,1267,1294]" gridcol="3" gridrow="6" pageId="13" pageNumber="14">3.9</td>
|
||
<td id="47335DCE616D003AFB6E45A5FAFCFAAA" box="[1198,1350,1267,1294]" gridcol="4" gridrow="6" pageId="13" pageNumber="14">4.4</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D4470FAFCFAE4" box="[253,1350,1318,1344]" gridrow="7" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D4470FECBFAE4" box="[253,369,1318,1344]" gridcol="0" gridrow="7" pageId="13" pageNumber="14">SAL</th>
|
||
<td id="47335DCE616D003AFE1B4470FDC9FAE4" box="[475,627,1318,1344]" gridcol="1" gridrow="7" pageId="13" pageNumber="14">28.4</td>
|
||
<td id="47335DCE616D003AFD0B4470FCD9FAE4" box="[715,867,1318,1344]" gridcol="2" gridrow="7" pageId="13" pageNumber="14">23.8</td>
|
||
<td id="47335DCE616D003AFC7B4470FBE9FAE4" box="[955,1107,1318,1344]" gridcol="3" gridrow="7" pageId="13" pageNumber="14">14.6</td>
|
||
<td id="47335DCE616D003AFB6E4470FAFCFAE4" box="[1198,1350,1318,1344]" gridcol="4" gridrow="7" pageId="13" pageNumber="14">18.9</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D440FFAFCFAD0" box="[253,1350,1369,1396]" gridrow="8" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D440FFECBFAD0" box="[253,369,1369,1396]" gridcol="0" gridrow="8" pageId="13" pageNumber="14">NL</th>
|
||
<td id="47335DCE616D003AFE1B440FFDC9FAD0" box="[475,627,1369,1396]" gridcol="1" gridrow="8" pageId="13" pageNumber="14">10.8</td>
|
||
<td id="47335DCE616D003AFD0B440FFCD9FAD0" box="[715,867,1369,1396]" gridcol="2" gridrow="8" pageId="13" pageNumber="14">6.7</td>
|
||
<td id="47335DCE616D003AFC7B440FFBE9FAD0" box="[955,1107,1369,1396]" gridcol="3" gridrow="8" pageId="13" pageNumber="14">5.2</td>
|
||
<td id="47335DCE616D003AFB6E440FFAFCFAD0" box="[1198,1350,1369,1396]" gridcol="4" gridrow="8" pageId="13" pageNumber="14">6.0</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D44DAFAFCFA02" box="[253,1350,1420,1446]" gridrow="9" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D44DAFECBFA02" box="[253,369,1420,1446]" gridcol="0" gridrow="9" pageId="13" pageNumber="14">AXG</th>
|
||
<td id="47335DCE616D003AFE1B44DAFDC9FA02" box="[475,627,1420,1446]" gridcol="1" gridrow="9" pageId="13" pageNumber="14">30.7</td>
|
||
<td id="47335DCE616D003AFD0B44DAFCD9FA02" box="[715,867,1420,1446]" gridcol="2" gridrow="9" pageId="13" pageNumber="14">29.9</td>
|
||
<td id="47335DCE616D003AFC7B44DAFBE9FA02" box="[955,1107,1420,1446]" gridcol="3" gridrow="9" pageId="13" pageNumber="14">18.4</td>
|
||
<td id="47335DCE616D003AFB6E44DAFAFCFA02" box="[1198,1350,1420,1446]" gridcol="4" gridrow="9" pageId="13" pageNumber="14">19.8</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D44E8FAFCFA7C" box="[253,1350,1470,1496]" gridrow="10" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D44E8FECBFA7C" box="[253,369,1470,1496]" gridcol="0" gridrow="10" pageId="13" pageNumber="14">FlL</th>
|
||
<td id="47335DCE616D003AFE1B44E8FDC9FA7C" box="[475,627,1470,1496]" gridcol="1" gridrow="10" pageId="13" pageNumber="14">16.6</td>
|
||
<td id="47335DCE616D003AFD0B44E8FCD9FA7C" box="[715,867,1470,1496]" gridcol="2" gridrow="10" pageId="13" pageNumber="14">16.6</td>
|
||
<td id="47335DCE616D003AFC7B44E8FBE9FA7C" box="[955,1107,1470,1496]" gridcol="3" gridrow="10" pageId="13" pageNumber="14">10.2</td>
|
||
<td id="47335DCE616D003AFB6E44E8FAFCFA7C" box="[1198,1350,1470,1496]" gridcol="4" gridrow="10" pageId="13" pageNumber="14">11.6</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D44A7FAFCF9A8" box="[253,1350,1521,1548]" gridrow="11" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D44A7FECBF9A8" box="[253,369,1521,1548]" gridcol="0" gridrow="11" pageId="13" pageNumber="14">HlL</th>
|
||
<td id="47335DCE616D003AFE1B44A7FDC9F9A8" box="[475,627,1521,1548]" gridcol="1" gridrow="11" pageId="13" pageNumber="14">22.4</td>
|
||
<td id="47335DCE616D003AFD0B44A7FCD9F9A8" box="[715,867,1521,1548]" gridcol="2" gridrow="11" pageId="13" pageNumber="14">18.5</td>
|
||
<td id="47335DCE616D003AFC7B44A7FBE9F9A8" box="[955,1107,1521,1548]" gridcol="3" gridrow="11" pageId="13" pageNumber="14">14.3</td>
|
||
<td id="47335DCE616D003AFB6E44A7FAFCF9A8" box="[1198,1350,1521,1548]" gridcol="4" gridrow="11" pageId="13" pageNumber="14">15.6</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D4774FAFCF99A" box="[253,1350,1570,1598]" gridrow="12" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D4774FECBF99A" box="[253,369,1570,1598]" gridcol="0" gridrow="12" pageId="13" pageNumber="14">SLF</th>
|
||
<td id="47335DCE616D003AFE1B4774FDC9F99A" box="[475,627,1570,1598]" gridcol="1" gridrow="12" pageId="13" pageNumber="14">13/142</td>
|
||
<td id="47335DCE616D003AFD0B4774FCD9F99A" box="[715,867,1570,1598]" gridcol="2" gridrow="12" pageId="13" pageNumber="14">14/132</td>
|
||
<td id="47335DCE616D003AFC7B4774FBE9F99A" box="[955,1107,1570,1598]" gridcol="3" gridrow="12" pageId="13" pageNumber="14">12/132</td>
|
||
<td id="47335DCE616D003AFB6E4774FAFCF99A" box="[1198,1350,1570,1598]" gridcol="4" gridrow="12" pageId="13" pageNumber="14">13/142</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D4703FAFCF9D5" box="[253,1350,1621,1649]" gridrow="13" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D4703FECBF9D5" box="[253,369,1621,1649]" gridcol="0" gridrow="13" pageId="13" pageNumber="14">SLT</th>
|
||
<td id="47335DCE616D003AFE1B4703FDC9F9D5" box="[475,627,1621,1649]" gridcol="1" gridrow="13" pageId="13" pageNumber="14">17/172</td>
|
||
<td id="47335DCE616D003AFD0B4703FCD9F9D5" box="[715,867,1621,1649]" gridcol="2" gridrow="13" pageId="13" pageNumber="14">17/172</td>
|
||
<td id="47335DCE616D003AFC7B4703FBE9F9D5" box="[955,1107,1621,1649]" gridcol="3" gridrow="13" pageId="13" pageNumber="14">17/152</td>
|
||
<td id="47335DCE616D003AFB6E4703FAFCF9D5" box="[1198,1350,1621,1649]" gridcol="4" gridrow="13" pageId="13" pageNumber="14">17/172</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D47DFFAFCF900" box="[253,1350,1673,1700]" gridrow="14" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D47DFFECBF900" box="[253,369,1673,1700]" gridcol="0" gridrow="14" pageId="13" pageNumber="14">MD</th>
|
||
<td id="47335DCE616D003AFE1B47DFFDC9F900" box="[475,627,1673,1700]" gridcol="1" gridrow="14" pageId="13" pageNumber="14">54</td>
|
||
<td id="47335DCE616D003AFD0B47DFFCD9F900" box="[715,867,1673,1700]" gridcol="2" gridrow="14" pageId="13" pageNumber="14">53</td>
|
||
<td id="47335DCE616D003AFC7B47DFFBE9F900" box="[955,1107,1673,1700]" gridcol="3" gridrow="14" pageId="13" pageNumber="14">53</td>
|
||
<td id="47335DCE616D003AFB6E47DFFAFCF900" box="[1198,1350,1673,1700]" gridcol="4" gridrow="14" pageId="13" pageNumber="14">57</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D47EAFAFCF973" box="[253,1350,1724,1751]" gridrow="15" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D47EAFECBF973" box="[253,369,1724,1751]" gridcol="0" gridrow="15" pageId="13" pageNumber="14">VL</th>
|
||
<td id="47335DCE616D003AFE1B47EAFDC9F973" box="[475,627,1724,1751]" gridcol="1" gridrow="15" pageId="13" pageNumber="14">14</td>
|
||
<td id="47335DCE616D003AFD0B47EAFCD9F973" box="[715,867,1724,1751]" gridcol="2" gridrow="15" pageId="13" pageNumber="14">12</td>
|
||
<td id="47335DCE616D003AFC7B47EAFBE9F973" box="[955,1107,1724,1751]" gridcol="3" gridrow="15" pageId="13" pageNumber="14">14</td>
|
||
<td id="47335DCE616D003AFB6E47EAFAFCF973" box="[1198,1350,1724,1751]" gridcol="4" gridrow="15" pageId="13" pageNumber="14">11</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D47B9FAFCF8AD" box="[253,1350,1775,1801]" gridrow="16" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D47B9FECBF8AD" box="[253,369,1775,1801]" gridcol="0" gridrow="16" pageId="13" pageNumber="14">VT</th>
|
||
<td id="47335DCE616D003AFE1B47B9FDC9F8AD" box="[475,627,1775,1801]" gridcol="1" gridrow="16" pageId="13" pageNumber="14">31</td>
|
||
<td id="47335DCE616D003AFD0B47B9FCD9F8AD" box="[715,867,1775,1801]" gridcol="2" gridrow="16" pageId="13" pageNumber="14">29</td>
|
||
<td id="47335DCE616D003AFC7B47B9FBE9F8AD" box="[955,1107,1775,1801]" gridcol="3" gridrow="16" pageId="13" pageNumber="14">31</td>
|
||
<td id="47335DCE616D003AFB6E47B9FAFCF8AD" box="[1198,1350,1775,1801]" gridcol="4" gridrow="16" pageId="13" pageNumber="14">30</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D4677FAFCF898" box="[253,1350,1825,1852]" gridrow="17" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D4677FECBF898" box="[253,369,1825,1852]" gridcol="0" gridrow="17" pageId="13" pageNumber="14">SAM</th>
|
||
<td id="47335DCE616D003AFE1B4677FDC9F898" box="[475,627,1825,1852]" gridcol="1" gridrow="17" pageId="13" pageNumber="14">36</td>
|
||
<td id="47335DCE616D003AFD0B4677FCD9F898" box="[715,867,1825,1852]" gridcol="2" gridrow="17" pageId="13" pageNumber="14">35</td>
|
||
<td id="47335DCE616D003AFC7B4677FBE9F898" box="[955,1107,1825,1852]" gridcol="3" gridrow="17" pageId="13" pageNumber="14">35</td>
|
||
<td id="47335DCE616D003AFB6E4677FAFCF898" box="[1198,1350,1825,1852]" gridcol="4" gridrow="17" pageId="13" pageNumber="14">35</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D4605FAFCF8CB" box="[253,1350,1875,1903]" gridrow="18" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D4605FECBF8CB" box="[253,369,1875,1903]" gridcol="0" gridrow="18" pageId="13" pageNumber="14">SL</th>
|
||
<td id="47335DCE616D003AFE1B4605FDC9F8CB" box="[475,627,1875,1903]" gridcol="1" gridrow="18" pageId="13" pageNumber="14">7/72</td>
|
||
<td id="47335DCE616D003AFD0B4605FCD9F8CB" box="[715,867,1875,1903]" gridcol="2" gridrow="18" pageId="13" pageNumber="14">7/72</td>
|
||
<td id="47335DCE616D003AFC7B4605FBE9F8CB" box="[955,1107,1875,1903]" gridcol="3" gridrow="18" pageId="13" pageNumber="14">7/72</td>
|
||
<td id="47335DCE616D003AFB6E4605FAFCF8CB" box="[1198,1350,1875,1903]" gridcol="4" gridrow="18" pageId="13" pageNumber="14">7/72</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D46D0FAFCF806" box="[253,1350,1926,1954]" gridrow="19" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D46D0FECBF806" box="[253,369,1926,1954]" gridcol="0" gridrow="19" pageId="13" pageNumber="14">IL</th>
|
||
<td id="47335DCE616D003AFE1B46D0FDC9F806" box="[475,627,1926,1954]" gridcol="1" gridrow="19" pageId="13" pageNumber="14">6/62</td>
|
||
<td id="47335DCE616D003AFD0B46D0FCD9F806" box="[715,867,1926,1954]" gridcol="2" gridrow="19" pageId="13" pageNumber="14">6/62</td>
|
||
<td id="47335DCE616D003AFC7B46D0FBE9F806" box="[955,1107,1926,1954]" gridcol="3" gridrow="19" pageId="13" pageNumber="14">7/62</td>
|
||
<td id="47335DCE616D003AFB6E46D0FAFCF806" box="[1198,1350,1926,1954]" gridcol="4" gridrow="19" pageId="13" pageNumber="14">5/52</td>
|
||
</tr>
|
||
<tr id="04E234B2616D003AFF3D46EEFAFCF870" box="[253,1350,1976,2004]" gridrow="20" pageId="13" pageNumber="14">
|
||
<th id="47335DCE616D003AFF3D46EEFECBF870" box="[253,369,1976,2004]" gridcol="0" gridrow="20" pageId="13" pageNumber="14">FP</th>
|
||
<td id="47335DCE616D003AFE1B46EEFDC9F870" box="[475,627,1976,2004]" gridcol="1" gridrow="20" pageId="13" pageNumber="14">10/92</td>
|
||
<td id="47335DCE616D003AFD0B46EEFCD9F870" box="[715,867,1976,2004]" gridcol="2" gridrow="20" pageId="13" pageNumber="14">9/92</td>
|
||
<td id="47335DCE616D003AFC7B46EEFBE9F870" box="[955,1107,1976,2004]" gridcol="3" gridrow="20" pageId="13" pageNumber="14">9/82</td>
|
||
<td id="47335DCE616D003AFB6E46EEFAFCF870" box="[1198,1350,1976,2004]" gridcol="4" gridrow="20" pageId="13" pageNumber="14">10/102</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="F2C8657B616EFFCBFF7D40C5FC5CFCEB" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="BA6D36F0616EFFCBFF7D40C5FE4BFE09" blockId="14.[189,1399,403,570]" box="[189,497,403,429]" pageId="14" pageNumber="15">
|
||
<heading id="E125819C616EFFCBFF7D40C5FE4BFE09" bold="true" box="[189,497,403,429]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="88A6EAE2616EFFCBFF7D40C5FE4BFE09" bold="true" box="[189,497,403,429]" pageId="14" pageNumber="15">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616EFFCBFF7D40E0FD55FD9E" blockId="14.[189,1399,403,570]" pageId="14" pageNumber="15">
|
||
The new species is known only from the Chimantá Massif in
|
||
<collectingCountry id="C2C57660616EFFCBFC7540E0FB94FE74" box="[949,1070,438,464]" name="Venezuela" pageId="14" pageNumber="15">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="22E92A75616EFFCBFB8140E0FB31FE75" box="[1089,1163,438,465]" captionStart="Fig" captionStartId="11.[189,232,1930,1956]" captionTargetBox="[271,1316,265,1890]" captionTargetId="figure@11.[271,1316,265,1890]" captionTargetPageId="11" captionText="Fig. 8. Map of the Guiana Shield region (A), with enlarged black rectangle corresponding to the Chimantá Massif (B). Black triangle indicates type locality, black dots indicate other know localities for the species." figureDoi="http://doi.org/10.5281/zenodo.3711451" httpUri="https://zenodo.org/record/3711451/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
|
||
) where it has been reported from Abakapá-tepui (
|
||
<typeStatus id="65698852616EFFCBFDE2408CFDECFE50" box="[546,598,474,500]" pageId="14" pageNumber="15">type</typeStatus>
|
||
locality), Amurí-tepui, and Murei-tepui (sometimes named Eruodatepui, see Discussion) between
|
||
<emphasis id="88A6EAE2616EFFCBFDED40A8FDEAFDB3" box="[557,592,510,535]" italics="true" pageId="14" pageNumber="15">ca.</emphasis>
|
||
<quantity id="7D2A9B15616EFFCBFD9840ABFD4FFDBC" box="[600,757,509,536]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="14" pageNumber="15" unit="m" value="2350.0" valueMax="2600.0" valueMin="2100.0">2100-2600 m</quantity>
|
||
elevation (
|
||
<bibRefCitation id="DE434B01616EFFCBFCB640ABFB61FDBC" author="McDiarmid R. W. & Donnelly M. A." box="[886,1243,509,536]" editor="Donnelly M. A. & Crother B. I. & Guyer C. & Wake M. H. & White M. E." firstAuthor="McDiarmid" journalOrPublisher="University of Chicago Press, Chicago, Illinois" pageId="14" pageNumber="15" pagination="461 - 560" refId="ref7922" refString="McDiarmid R. W. & Donnelly M. A. 2005. The herpetofauna of the Guayana Highlands: amphibians and reptiles of the Lost World. In: Donnelly M. A., Crother B. I., Guyer C., Wake M. H. & White M. E. (eds.) Ecology and Evolution in the Tropics: A Herpetological Perspective: 461 - 560. University of Chicago Press, Chicago, Illinois." title="The herpetofauna of the Guayana Highlands: amphibians and reptiles of the Lost World" type="book chapter" volumeTitle="Ecology and Evolution in the Tropics: A Herpetological Perspective" year="2005">McDiarmid & Donnelly 2005</bibRefCitation>
|
||
). The species is probably widespread in the Chimantá Massif.
|
||
</paragraph>
|
||
<paragraph id="BA6D36F0616EFFCBFF7D4334FC5CFCEB" blockId="14.[189,1399,610,848]" pageId="14" pageNumber="15">
|
||
<taxonomicName id="7DD24D73616EFFCBFF7D4334FE19FDD8" authorityName="Kok & Rivas" authorityYear="2011" box="[189,419,610,636]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="mcdiarmidi">
|
||
<emphasis id="88A6EAE2616EFFCBFF7D4334FE19FDD8" box="[189,419,610,636]" italics="true" pageId="14" pageNumber="15">Anadia mcdiarmidi</emphasis>
|
||
</taxonomicName>
|
||
is diurnal and inhabits open, mostly flat sandstone areas on tepui summits (
|
||
<figureCitation id="22E92A75616EFFCBFA874334FF59FD3B" captionStart="Fig" captionStartId="3.[189,232,1895,1921]" captionTargetBox="[207,1380,265,1855]" captionTargetId="figure@3.[207,1380,265,1855]" captionTargetPageId="3" captionText="Fig. 1. Anadia mcdiarmidi sp. nov. A. IRSNB 2677, ♂ holotype in life. B. IRSNB 2675, juvenile male paratype in life. C. Ventral view of the anaesthetized holotype (grid squares = 5 mm). D. Summit of Abakapá-tepui, looking NW, showing the macrohabitat of the new species (03-05-2011). Photographs by Philippe J.R. Kok." figureDoi="http://doi.org/10.5281/zenodo.3711437" httpUri="https://zenodo.org/record/3711437/files/figure.png" pageId="14" pageNumber="15">Fig. 1D</figureCitation>
|
||
). The
|
||
<typeStatus id="65698852616EFFCBFEF543D3FE26FD3B" box="[309,412,645,671]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
and two
|
||
<typeStatus id="65698852616EFFCBFDD243D3FD38FD3B" box="[530,642,645,671]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
|
||
were collected during the day under rocks; IRSNB 2674 was found during the day, crawling in a small crack between rocks. Two white fertile eggs (both measuring
|
||
<quantity id="7D2A9B15616EFFCBFF7D439AFE90FD42" box="[189,298,716,742]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="14" pageNumber="15" unit="mm" value="12.3">12.3 mm</quantity>
|
||
) and several older eggshells attributable to the new species were found under a large rock, suggesting that a same suitable nesting site may be reused [as already postulated by
|
||
<bibRefCitation id="DE434B01616EFFCBFB7043B9FAD0FCAD" author="Gorzula S." box="[1200,1386,750,777]" editor="Huber O" firstAuthor="Gorzula" journalOrPublisher="Oscar Todtmann Editores, Caracas." pageId="14" pageNumber="15" pagination="304 - 310" part="280" refId="ref7629" refString="Gorzula S. 1992. La herpetofauna del macizo del Chimanta. In: Huber O (ed.), El Macizo del Chimanta: 267 - 280, 304 - 310. Oscar Todtmann Editores, Caracas." title="La herpetofauna del macizo del Chimanta" type="journal article" volumeTitle="El Macizo del Chimanta: 267" year="1992">Gorzula (1992)</bibRefCitation>
|
||
], and that females lay two eggs at a time. The only other lizard found syntopic with
|
||
<taxonomicName id="7DD24D73616EFFCBFB524244FACCFC88" authorityName="Kok & Rivas" authorityYear="2011" box="[1170,1398,786,812]" class="Reptilia" family="Gymnophthalmidae" genus="Anadia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="mcdiarmidi">
|
||
<emphasis id="88A6EAE2616EFFCBFB524244FACCFC88" box="[1170,1398,786,812]" italics="true" pageId="14" pageNumber="15">Anadia mcdiarmidi</emphasis>
|
||
</taxonomicName>
|
||
was
|
||
<taxonomicName id="7DD24D73616EFFCBFF324260FE5DFCEB" box="[242,487,821,847]" class="Reptilia" family="Gymnophthalmidae" genus="Neusticurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rudis">
|
||
<emphasis id="88A6EAE2616EFFCBFF324260FEC4FCEB" box="[242,382,822,847]" italics="true" pageId="14" pageNumber="15">Neusticurus</emphasis>
|
||
cf.
|
||
<emphasis id="88A6EAE2616EFFCBFE6B4263FE5DFCEB" box="[427,487,821,847]" italics="true" pageId="14" pageNumber="15">rudis</emphasis>
|
||
</taxonomicName>
|
||
, which was also mainly found under rocks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |