279 lines
33 KiB
XML
279 lines
33 KiB
XML
<document id="E4915BA42E93E012135D36AF51AFC635" ID-DOI="10.5281/zenodo.189048" ID-GBIF-Dataset="cc3fea04-eca2-4526-a4f8-3cb2502bf9c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="189048" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461099071944" checkinUser="plazi" docAuthor="Wit, Pierre De, Rota, Emilia & Erséus, Christer" docDate="2009" docId="FB45AF01C967876F80E2FADCF82CA4E7" docLanguage="en" docName="zt02165p038.pdf" docOrigin="Zootaxa 2165" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Grania colorata Wit, Rota & Erséus, 2009, sp. n." docType="treatment" docVersion="9" lastPageNumber="27" masterDocId="077CD779C96E87648075FFC3FD76A308" masterDocTitle="Grania (Annelida: Clitellata: Enchytraeidae) of the Great Barrier Reef, Australia, including four new species and a re-description of Grania trichaeta Jamieson, 1977" masterLastPageNumber="38" masterPageNumber="16" pageNumber="25" updateTime="1698590481319" updateUser="plazi">
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<mods:mods id="6DDE5CB83E6B6F03182A1D6362095839" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BBDBD6102F041CCF527D00D8E23E989D">
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<mods:title id="CA8F71B607C6DCDFCE4FFD40C3DDFCF7">Grania (Annelida: Clitellata: Enchytraeidae) of the Great Barrier Reef, Australia, including four new species and a re-description of Grania trichaeta Jamieson, 1977</mods:title>
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</mods:titleInfo>
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<mods:name id="37DB0CEB31C5D5F342B1970A6E427005" type="personal">
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<mods:role id="632F3E7E84D91471DFE39D9AEE41CC20">
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<mods:roleTerm id="121EABA068FE975B4230E269144F36E6">Author</mods:roleTerm>
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<mods:namePart id="CB700C870EBCFD25EBD51A00B57E8479">Wit, Pierre De</mods:namePart>
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</mods:name>
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<mods:name id="E5B0CC929E87C9D9E796AA9CB4F080CE" type="personal">
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<mods:role id="394843D084513C361CA82274D4917E01">
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<mods:roleTerm id="81A9A1FAA9D8EBC96C188DF08D3ACE97">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BC1DC55B874FF623E6E7EC3F578EE32A">Rota, Emilia</mods:namePart>
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</mods:name>
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<mods:name id="3E3062336F7D55A94B6B8F24E0379AE9" type="personal">
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<mods:role id="D0E11184616A18AC7FC548ED9B50F6E0">
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<mods:roleTerm id="D8F41867F72C93B7EE63E9D80C20C800">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7825FB44BE8C564F80F4B470DF7E204D">Erséus, Christer</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="9E153AA065669D8C753E1E9BA87147B4">text</mods:typeOfResource>
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<mods:title id="8C9FE61F758C0C1214F7DA696A43BD36">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="14E0E3C8D254F39CC88E0733C8A9E46A">
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<mods:date id="1302FFDF8972417DF10AF7CF450922F3">2009</mods:date>
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<mods:detail id="E9A4D33CA6DCD4B0A750207B27FDFF2C" type="volume">
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<mods:number id="E80EB9382C70EE311C015824BCF312D6">2165</mods:number>
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<mods:start id="4BCA8C03CEA153C26BB40F1BBD9579B2">16</mods:start>
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<mods:end id="DBF628F77BC34EA369A75A95D5E1EC73">38</mods:end>
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<mods:classification id="C74BBEE772596547BEC0AD7F5A5CE054">journal article</mods:classification>
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<mods:identifier id="F3031BE276849594BBB4F0BFDAEF49B8" type="DOI">10.5281/zenodo.189048</mods:identifier>
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<mods:identifier id="DC670ECA25E31973A8CFED510CE71BA7" type="GBIF-Dataset">cc3fea04-eca2-4526-a4f8-3cb2502bf9c3</mods:identifier>
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<mods:identifier id="88349D300CB349522D5BF7CF2951D81F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5F73ED39BA63B631F40CDD036DA48A9A" type="Zenodo-Dep">189048</mods:identifier>
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<treatment id="FB45AF01C967876F80E2FADCF82CA4E7" ID-DOI="http://doi.org/10.5281/zenodo.6217976" ID-GBIF-Taxon="119621275" ID-Zenodo-Dep="6217976" LSID="urn:lsid:plazi:treatment:FB45AF01C967876F80E2FADCF82CA4E7" httpUri="http://treatment.plazi.org/id/FB45AF01C967876F80E2FADCF82CA4E7" lastPageId="11" lastPageNumber="27" pageId="9" pageNumber="25">
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<subSubSection id="3BF64D9CC967876D80E2FADCFC2FA654" pageId="9" pageNumber="25" type="nomenclature">
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<paragraph id="73531E17C967876D80E2FADCFCD7A631" blockId="9.[151,417,1311,1372]" box="[151,417,1311,1337]" pageId="9" pageNumber="25">
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<heading id="281BA97BC967876D80E2FADCFCD7A631" bold="true" box="[151,417,1311,1337]" fontSize="11" level="1" pageId="9" pageNumber="25" reason="1">
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<emphasis id="4198C205C967876D80E2FADCFCD7A631" bold="true" box="[151,417,1311,1337]" pageId="9" pageNumber="25">
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<taxonomicName id="B4EC6594C967876D80E2FADCFC2EA631" ID-CoL="3H7YR" box="[151,344,1311,1337]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="9" pageNumber="25" phylum="Annelida" rank="species" species="colorata" status="sp. nov.">
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<emphasis id="4198C205C967876D80E2FADCFC2EA631" bold="true" box="[151,344,1311,1337]" italics="true" pageId="9" pageNumber="25">Grania colorata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5AAB7F7EC967876D812AFADCFCD7A631" box="[351,417,1311,1337]" pageId="9" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="73531E17C967876D80E2FA81FC2FA654" blockId="9.[151,417,1311,1372]" box="[151,345,1346,1372]" pageId="9" pageNumber="25">
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(
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<figureCitation id="EBD70292C967876D80EAFA81FD85A654" box="[159,243,1346,1372]" captionStart="FIGURE 6" captionStartId="10.[151,255,1607,1631]" captionTargetBox="[272,1315,473,1575]" captionTargetId="figure@10.[258,1328,465,1584]" captionTargetPageId="10" captionText="FIGURE 6. Grania colorata sp. n. (Holotype, stn. L 19). A: Chaetae. B: Clitellar gland cell pattern, dorsal view, anterior end to the top. C: Penial apparatus, side view, anterior end to the top." httpUri="https://zenodo.org/record/189054/files/figure.png" pageId="9" pageNumber="25">Figs. 6</figureCitation>
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,
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<figureCitation id="EBD70292C967876D808AFA81FC79A654" box="[255,271,1346,1372]" captionStart="FIGURE 7" captionStartId="11.[151,256,1050,1074]" captionTargetBox="[231,1354,347,1024]" captionTargetId="figure@11.[231,1355,341,1025]" captionTargetPageId="11" captionText="FIGURE 7. Grania colorata sp. n. (Holotype, stn. L 19). Segment V with spermatheca, side view, anterior end to the right." httpUri="https://zenodo.org/record/189055/files/figure.png" pageId="9" pageNumber="25">7</figureCitation>
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,
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<figureCitation id="EBD70292C967876D8169FA81FC4FA654" box="[284,313,1346,1372]" captionStart="FIGURE 10" captionStartId="16.[151,255,1976,2000]" captionTargetBox="[455,1130,429,1948]" captionTargetId="figure@16.[455,1131,429,1950]" captionTargetPageId="16" captionText="FIGURE 10. Chaetae. A: G. breviductus sp. n.. B: G. regina sp. n.. C: G. ho m o c ha e ta sp. n.. D: G. c o lor at a sp. n.. E: G. trichaeta Jamieson 1977." httpUri="https://zenodo.org/record/189058/files/figure.png" pageId="9" pageNumber="25">10</figureCitation>
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D)
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</paragraph>
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</subSubSection>
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<subSubSection id="3BF64D9CC967876D80E2FA4CFC47A424" pageId="9" pageNumber="25" type="materials_examined">
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<paragraph id="73531E17C967876D80E2FA4CFC62A6FF" blockId="9.[151,1437,1423,2030]" pageId="9" pageNumber="25">
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<emphasis id="4198C205C967876D80E2FA4CFC64A6A1" bold="true" box="[151,274,1423,1449]" pageId="9" pageNumber="25">
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<typeStatus id="AC57A0B5C967876D80E2FA4CFC78A6A1" box="[151,270,1423,1449]" pageId="9" pageNumber="25" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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AMS
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<typeStatus id="AC57A0B5C967876D8114FA53FCE5A6A2" box="[353,403,1424,1450]" pageId="9" pageNumber="25">type</typeStatus>
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coll. W.35545, a whole-mounted front end from Lizard Island, stn. L19, also serving as voucher specimen for
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<collectionCode id="15FD86D2C967876D81C4FA75FC90A6D8" LSID="urn:lsid:biocol.org:col:14498" box="[433,486,1462,1488]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="9" pageNumber="25">COI</collectionCode>
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barcode (GenBank accession no.
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<accessionNumber id="6CBF83F4C967876D83E0FA75F950A6D8" box="[917,1062,1462,1488]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GQ247639" pageId="9" pageNumber="25">GQ247639</accessionNumber>
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); posterior end used for
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<collectionCode id="15FD86D2C967876D852EFA75F8EBA6D8" LSID="urn:lsid:biocol.org:col:15536" box="[1371,1437,1462,1488]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="9" pageNumber="25">DNA</collectionCode>
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extraction.
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</paragraph>
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<paragraph id="73531E17C967876D80B0F9C0FE85A59A" blockId="9.[151,1437,1423,2030]" pageId="9" pageNumber="25">
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<emphasis id="4198C205C967876D80B0F9C0FC39A515" bold="true" box="[197,335,1539,1565]" pageId="9" pageNumber="25">
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<typeStatus id="AC57A0B5C967876D80B0F9C0FC3DA515" box="[197,331,1539,1565]" pageId="9" pageNumber="25" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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AMS
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<typeStatus id="AC57A0B5C967876D81D4F9C7FCA3A516" box="[417,469,1540,1566]" pageId="9" pageNumber="25">type</typeStatus>
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coll. W.35546, whole-mounted front end from Lizard Island, stn. L21, voucher specimen for
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<collectionCode id="15FD86D2C967876D8143F9E9FC1EA54C" LSID="urn:lsid:biocol.org:col:14498" box="[310,360,1578,1604]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="9" pageNumber="25">COI</collectionCode>
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barcode (GenBank accession no.
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<accessionNumber id="6CBF83F4C967876D8280F9E9FE0AA54C" box="[757,892,1578,1604]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GQ247641" pageId="9" pageNumber="25">GQ247641</accessionNumber>
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). AMS
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<typeStatus id="AC57A0B5C967876D83A3F9E9F97EA54C" box="[982,1032,1578,1604]" pageId="9" pageNumber="25">type</typeStatus>
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coll. W.35547-W.35553, 7 wholemounted specimens: 1 each from stns. L1, L2, L10, L11, L12, L25 and L28.
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<collectionCode id="15FD86D2C967876D845FF992F909A563" box="[1066,1151,1617,1643]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="9" pageNumber="25">SMNH</collectionCode>
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<typeStatus id="AC57A0B5C967876D84F2F992F9CCA563" box="[1159,1210,1617,1643]" pageId="9" pageNumber="25">type</typeStatus>
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coll. 7768-7772, 5 whole-mounted specimens, of which 3 are from stn. L10 and 2 from L11.
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</paragraph>
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<paragraph id="73531E17C967876D80B0F95DFC47A424" blockId="9.[151,1437,1423,2030]" pageId="9" pageNumber="25">
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<emphasis id="4198C205C967876D80B0F95DFF68A5B0" bold="true" box="[197,542,1694,1720]" pageId="9" pageNumber="25">Other material examined:</emphasis>
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<collectionCode id="15FD86D2C967876D825DF95DFFF6A5B0" box="[552,640,1694,1720]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="9" pageNumber="25">SMNH</collectionCode>
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main coll. 105500-105539, 40 whole-mounted specimens from Lizard Island (stns. L8 (12), L11 (15), L20 (4), L21 (9)). First author’s collection: 38 whole-mounted specimens from Lizard Island (stns. L1 (1), L2 (17), L5 (2), L9 (3), L10 (7), L16 (1), L19 (1), L24 (1), L27 (2), L29 (3)).
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</paragraph>
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</subSubSection>
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<subSubSection id="3BF64D9CC967876F80B0F8FBF9D1A22E" lastPageId="11" lastPageNumber="27" pageId="9" pageNumber="25" type="description">
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<paragraph id="73531E17C967876E80B0F8FBFCBBA292" blockId="9.[151,1437,1423,2030]" lastBlockId="10.[151,1436,152,410]" lastPageId="10" lastPageNumber="26" pageId="9" pageNumber="25">
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<emphasis id="4198C205C967876D80B0F8FBFC17A45A" bold="true" box="[197,353,1848,1874]" pageId="9" pageNumber="25">Description:</emphasis>
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Living specimens greenish-yellow. Body
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<quantity id="B414B3F2C967876D832FF8FAFE9FA45B" box="[858,1001,1849,1875]" metricMagnitude="-3" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.1" metricValueMin="4.3" pageId="9" pageNumber="25" unit="mm" value="4.699999999999999" valueMax="5.1" valueMin="4.3">4.3–5.1 mm</quantity>
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long (n=30),
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<quantity id="B414B3F2C967876D84FAF8FAF84AA45B" box="[1167,1340,1849,1875]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="9" pageNumber="25" unit="mm" value="0.1" valueMax="0.12" valueMin="0.08">0.08–0.12 mm</quantity>
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wide at III,
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<quantity id="B414B3F2C967876D80B7F8A3FC1AA472" box="[194,364,1888,1914]" metricMagnitude="-4" metricUnit="m" metricValue="1.1500000000000001" metricValueMax="1.4000000000000001" metricValueMin="0.8999999999999999" pageId="9" pageNumber="25" unit="mm" value="0.115" valueMax="0.14" valueMin="0.09">0.09–0.14 mm</quantity>
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at clitellum (n=30). Segment number 35–38 (n=30). Prostomium rounded, 40–60 µm wide, 30–45 µm long (n=30); epidermis 6–14 µm thick dorsally and anteriorly (n=30), 5–11 µm ventrally (n=30). Peristomium 70–90 µm wide at 1/2 (n=30). Ventral chaetae commencing in XIV (XV in one specimen); lateral chaetae commencing in XVI–XVIII. Chaetae of uniform size throughout body, 55–70 µm long (n=30); chaetae L-shaped: shaft thickest at base and sharply pointed distally, foot 10–15 µm long, with broad instep, flat sole and conspicuous heel; chaetal index=
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<geoCoordinate id="16D878D0C964876E82B1FF7DFE66A3D0" box="[708,784,190,216]" direction="north" orientation="latitude" pageId="10" pageNumber="26" precision="555" value="4.63">4.63 n</geoCoordinate>
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=30, sd=0.791 (
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<figureCitation id="EBD70292C964876E83BBFF7DF954A3D0" box="[974,1058,190,216]" captionStart="FIGURE 6" captionStartId="10.[151,255,1607,1631]" captionTargetBox="[272,1315,473,1575]" captionTargetId="figure@10.[258,1328,465,1584]" captionTargetPageId="10" captionText="FIGURE 6. Grania colorata sp. n. (Holotype, stn. L 19). A: Chaetae. B: Clitellar gland cell pattern, dorsal view, anterior end to the top. C: Penial apparatus, side view, anterior end to the top." httpUri="https://zenodo.org/record/189054/files/figure.png" pageId="10" pageNumber="26">Figs. 6</figureCitation>
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A, 10D). Epidermal gland cells inconspicuous. Clitellum 8–14 µm thick, extending from anterior of XII to mid XIII, with an irregular pattern of granular gland cells interspersed with hyaline cells at a frequency of about 4:3 (
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<figureCitation id="EBD70292C964876E8417FECFF9DFA22E" box="[1122,1193,268,294]" captionStart="FIGURE 6" captionStartId="10.[151,255,1607,1631]" captionTargetBox="[272,1315,473,1575]" captionTargetId="figure@10.[258,1328,465,1584]" captionTargetPageId="10" captionText="FIGURE 6. Grania colorata sp. n. (Holotype, stn. L 19). A: Chaetae. B: Clitellar gland cell pattern, dorsal view, anterior end to the top. C: Penial apparatus, side view, anterior end to the top." httpUri="https://zenodo.org/record/189054/files/figure.png" pageId="10" pageNumber="26">Fig. 6</figureCitation>
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B), except near male pores where hyaline cells are absent, and midventrally, where no gland cells are present. Copulatory gland in XIV in some specimens. Spermathecal pores lateral, located immediately behind 4/5. Male pores located ventrolaterally in mid XII.
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</paragraph>
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<caption id="27934E9FC964876E80E2F984FE00A589" httpUri="https://zenodo.org/record/189054/files/figure.png" pageId="10" pageNumber="26" targetBox="[272,1315,473,1575]" targetPageId="10">
|
||
<paragraph id="73531E17C964876E80E2F984FE00A589" blockId="10.[151,1436,1607,1665]" pageId="10" pageNumber="26">
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||
<emphasis id="4198C205C964876E80E2F984FC6FA557" bold="true" box="[151,281,1607,1631]" pageId="10" pageNumber="26">FIGURE 6.</emphasis>
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<taxonomicName id="B4EC6594C964876E816AF98BFCBBA557" box="[287,461,1608,1631]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="26" phylum="Annelida" rank="species" species="colorata" status="sp. nov.">
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<emphasis id="4198C205C964876E816AF98BFCBBA557" box="[287,461,1608,1631]" italics="true" pageId="10" pageNumber="26">Grania colorata</emphasis>
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</taxonomicName>
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<emphasis id="4198C205C964876E81A6F984FF79A557" bold="true" box="[467,527,1607,1631]" pageId="10" pageNumber="26">
|
||
<taxonomicNameLabel id="5AAB7F7EC964876E81A6F984FF79A557" box="[467,527,1607,1631]" pageId="10" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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(Holotype, stn. L19). A: Chaetae. B: Clitellar gland cell pattern, dorsal view, anterior end to the top. C: Penial apparatus, side view, anterior end to the top.
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||
</paragraph>
|
||
</caption>
|
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<paragraph id="73531E17C964876F80B0F973F9D1A22E" blockId="10.[151,1437,1712,2008]" lastBlockId="11.[151,1436,152,294]" lastPageId="11" lastPageNumber="27" pageId="10" pageNumber="26">
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||
Brain posteriorly indented. Head organ absent. Pharyngeal glands in IV–VI; dorsal lobes present in IV–VI, ventral lobes present in V (2 pairs) and VI (2 pairs); not connected dorsally. First pair of nephridia at 7/8. Dorsal blood vessel commencing in XVI–XIX. Chloragogen cells small (5–7 µm tall). Coelomocytes oval, small with stained nucleus, in high densities anterior to clitellum. Sperm sac extending posteriorly from clitellum as far back as XV; loose sperm packages also present in X–XII. Sperm funnels of uniform width, 45–50 µm wide, 1.5 times as long as wide. Heads of spermatozoa 13–17 µm long. Vasa deferentia unmodified, loosely coiled in XII; 5 µm wide, internally ciliated. Penial apparatuses (
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<figureCitation id="EBD70292C964876E84E6F85BF9ACA4BA" box="[1171,1242,1944,1970]" captionStart="FIGURE 6" captionStartId="10.[151,255,1607,1631]" captionTargetBox="[272,1315,473,1575]" captionTargetId="figure@10.[258,1328,465,1584]" captionTargetPageId="10" captionText="FIGURE 6. Grania colorata sp. n. (Holotype, stn. L 19). A: Chaetae. B: Clitellar gland cell pattern, dorsal view, anterior end to the top. C: Penial apparatus, side view, anterior end to the top." httpUri="https://zenodo.org/record/189054/files/figure.png" pageId="10" pageNumber="26">Fig. 6</figureCitation>
|
||
C) with uniform oval glandular structures, 40–55 µm long, 20–40 µm wide; vasa deferentia opening into epidermal invaginations; stylets absent (penial bulb
|
||
<typeStatus id="AC57A0B5C965876F8208FF5BFFC6A3BA" box="[637,688,152,178]" pageId="11" pageNumber="27">type</typeStatus>
|
||
3). Egg sac reaching as far back as XIX. Spermathecae (
|
||
<figureCitation id="EBD70292C965876F8539FF5BF8E4A3BA" box="[1356,1426,152,178]" captionStart="FIGURE 7" captionStartId="11.[151,256,1050,1074]" captionTargetBox="[231,1354,347,1024]" captionTargetId="figure@11.[231,1355,341,1025]" captionTargetPageId="11" captionText="FIGURE 7. Grania colorata sp. n. (Holotype, stn. L 19). Segment V with spermatheca, side view, anterior end to the right." httpUri="https://zenodo.org/record/189055/files/figure.png" pageId="11" pageNumber="27">Fig. 7</figureCitation>
|
||
) attached to oesophagus near 5/6; ampullae roughly spherical, relatively small (30–45 µm in diameter), ducts of uniform width, 50–70 µm long and 7–10 µm wide; very few sperm rings in ampullar walls, most of sperm freely dispersed in bundles throughout ampullar lumen; no glands at spermathecal pores.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="27934E9FC965876F80E2FBD9FDA7A75C" httpUri="https://zenodo.org/record/189055/files/figure.png" pageId="11" pageNumber="27" targetBox="[231,1354,347,1024]" targetPageId="11">
|
||
<paragraph id="73531E17C965876F80E2FBD9FDA7A75C" blockId="11.[151,1436,1050,1108]" pageId="11" pageNumber="27">
|
||
<emphasis id="4198C205C965876F80E2FBD9FC6DA73A" bold="true" box="[151,283,1050,1074]" pageId="11" pageNumber="27">FIGURE 7.</emphasis>
|
||
<taxonomicName id="B4EC6594C965876F8156FBD8FCA3A73A" box="[291,469,1051,1074]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="colorata" status="sp. nov.">
|
||
<emphasis id="4198C205C965876F8156FBD8FCA3A73A" box="[291,469,1051,1074]" italics="true" pageId="11" pageNumber="27">Grania colorata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4198C205C965876F81A9FBD9FF6CA73A" bold="true" box="[476,538,1050,1074]" pageId="11" pageNumber="27">
|
||
<taxonomicNameLabel id="5AAB7F7EC965876F81A9FBD9FF6CA73A" box="[476,538,1050,1074]" pageId="11" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype, stn. L19). Segment V with spermatheca, side view, anterior end to the right.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="3BF64D9CC965876F80B0FB41FF2EA7CB" pageId="11" pageNumber="27" type="etymology">
|
||
<paragraph id="73531E17C965876F80B0FB41FF2EA7CB" blockId="11.[151,1437,1154,2031]" pageId="11" pageNumber="27">
|
||
<emphasis id="4198C205C965876F80B0FB41FC20A794" bold="true" box="[197,342,1154,1180]" pageId="11" pageNumber="27">Etymology:</emphasis>
|
||
The Latin
|
||
<taxonomicName id="B4EC6594C965876F81ACFB40FF4BA794" box="[473,573,1155,1180]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="colorata">
|
||
<emphasis id="4198C205C965876F81ACFB40FF4BA794" box="[473,573,1155,1180]" italics="true" pageId="11" pageNumber="27">colorata</emphasis>
|
||
</taxonomicName>
|
||
refers to the conspicuous greenish-yellow coloration of the living worm, which is an unusual feature in
|
||
<taxonomicName id="B4EC6594C965876F8188FB6AFF27A7CA" box="[509,593,1193,1218]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="genus">
|
||
<emphasis id="4198C205C965876F8188FB6AFF27A7CA" box="[509,593,1193,1218]" italics="true" pageId="11" pageNumber="27">Grania</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3BF64D9CC965876F80B0FB13F8EAA4C1" pageId="11" pageNumber="27" type="discussion">
|
||
<paragraph id="73531E17C965876F80B0FB13FF26A5E9" blockId="11.[151,1437,1154,2031]" pageId="11" pageNumber="27">
|
||
<emphasis id="4198C205C965876F80B0FB13FC37A7E2" bold="true" box="[197,321,1232,1258]" pageId="11" pageNumber="27">Remarks:</emphasis>
|
||
The peculiar coloration of this species resembles that of
|
||
<taxonomicName id="B4EC6594C965876F839DFB13F8F4A7E2" authority="De Wit & Erseus, 2007" authorityName="De Wit & Erseus" authorityYear="2007" box="[1000,1410,1232,1258]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="galbina">
|
||
<emphasis id="4198C205C965876F839DFB13F915A7E1" box="[1000,1123,1232,1257]" italics="true" pageId="11" pageNumber="27">G. galbina</emphasis>
|
||
<bibRefCitation id="177D63E6C965876F841EFB13F8F4A7E2" author="De" box="[1131,1410,1232,1258]" pageId="11" pageNumber="35" refString="De Wit, P. & Erseus, C. (2007) Seven new species of Grania (Annelida: Clitellata: Enchytraeidae) from New Caledonia, South Pacific Ocean. Zootaxa, 1426, 27 - 50." type="journal article" year="2007">De Wit & Erséus, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, a recently described species from
|
||
<collectingCountry id="0BFB5E87C965876F8251FB34FF92A619" box="[548,740,1271,1297]" name="New Caledonia" pageId="11" pageNumber="27">New Caledonia</collectingCountry>
|
||
, and preliminary molecular analyses also support that
|
||
<taxonomicName id="B4EC6594C965876F85F6FB34FD87A63E" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="galbina">
|
||
<emphasis id="4198C205C965876F85F6FB34FD87A63E" italics="true" pageId="11" pageNumber="27">G. galbina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="4198C205C965876F815FFADEFCDCA63E" box="[298,426,1309,1334]" italics="true" pageId="11" pageNumber="27">G. c o l o r a t a</emphasis>
|
||
are closely related (De Wit,
|
||
<emphasis id="4198C205C965876F8375FADEFEE3A63E" box="[768,917,1309,1334]" italics="true" pageId="11" pageNumber="27">unpubl. data</emphasis>
|
||
). The two species are easily distinguishable from each other, however, by the body size, where
|
||
<emphasis id="4198C205C965876F8285FA87FE06A655" box="[752,880,1348,1373]" italics="true" pageId="11" pageNumber="27">G. c o l o r a t a</emphasis>
|
||
is much smaller than
|
||
<emphasis id="4198C205C965876F8402FA87F99BA655" box="[1143,1261,1348,1373]" italics="true" pageId="11" pageNumber="27">G. g a l b i n a</emphasis>
|
||
both in length and segment number (maximum length of
|
||
<emphasis id="4198C205C965876F82EEFAA8FE69A68C" box="[667,799,1387,1412]" italics="true" pageId="11" pageNumber="27">G. c o l o r a t a</emphasis>
|
||
is
|
||
<quantity id="B414B3F2C965876F833DFAA9FED2A68C" box="[840,932,1386,1412]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="11" pageNumber="27" unit="mm" value="5.1">5.1 mm</quantity>
|
||
with 38 segments; minimum length of
|
||
<taxonomicName id="B4EC6594C965876F85F6FAA8FD83A6A2" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="galbina">
|
||
<emphasis id="4198C205C965876F85F6FAA8FD83A6A2" italics="true" pageId="11" pageNumber="27">G. galbina</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<quantity id="B414B3F2C965876F816EFA52FC0FA6A3" box="[283,377,1425,1451]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="11" pageNumber="27" unit="mm" value="6.4">6.4 mm</quantity>
|
||
with 51 segments), the chaetal distribution (
|
||
<taxonomicName id="B4EC6594C965876F83D0FA52F950A6A2" box="[933,1062,1425,1450]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="galbina">
|
||
<emphasis id="4198C205C965876F83D0FA52F950A6A2" box="[933,1062,1425,1450]" italics="true" pageId="11" pageNumber="27">G. galbina</emphasis>
|
||
</taxonomicName>
|
||
possesses pre-clitellar ventral chaetae,
|
||
<taxonomicName id="B4EC6594C965876F808BFA7BFCF5A6D9" box="[254,387,1464,1489]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="colorata">
|
||
<emphasis id="4198C205C965876F808BFA7BFCF5A6D9" box="[254,387,1464,1489]" italics="true" pageId="11" pageNumber="27">G. colorata</emphasis>
|
||
</taxonomicName>
|
||
does not) and the sperm distribution in the spermathecal ampullae. In
|
||
<taxonomicName id="B4EC6594C965876F84BEFA7BF856A6D9" box="[1227,1312,1464,1489]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="genus">
|
||
<emphasis id="4198C205C965876F84BEFA7BF856A6D9" box="[1227,1312,1464,1489]" italics="true" pageId="11" pageNumber="27">Grania</emphasis>
|
||
</taxonomicName>
|
||
the sperm are usually segregated in hollow spherical compartments in the walls of the ampullae, which gives the impression of the sperm being organized in rings.
|
||
<taxonomicName id="B4EC6594C965876F8297F9C6FEEEA516" box="[738,920,1541,1566]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="galbina">
|
||
<emphasis id="4198C205C965876F8297F9C6FEEEA516" box="[738,920,1541,1566]" italics="true" pageId="11" pageNumber="27">Grania galbina</emphasis>
|
||
</taxonomicName>
|
||
has many such rings in its spermathecae. In
|
||
<taxonomicName id="B4EC6594C965876F80E2F9EFFC6DA54D" box="[151,283,1580,1605]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="colorata">
|
||
<emphasis id="4198C205C965876F80E2F9EFFC6DA54D" box="[151,283,1580,1605]" italics="true" pageId="11" pageNumber="27">G. colorata</emphasis>
|
||
</taxonomicName>
|
||
, however, bundles of sperm are distributed mostly in the lumen of the ampullae.
|
||
<taxonomicName id="B4EC6594C965876F84A9F9EFF8EAA54D" box="[1244,1436,1580,1605]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="colorata">
|
||
<emphasis id="4198C205C965876F84A9F9EFF8EAA54D" box="[1244,1436,1580,1605]" italics="true" pageId="11" pageNumber="27">Grania colorata</emphasis>
|
||
</taxonomicName>
|
||
is also similar to, but clearly distinguishable from,
|
||
<taxonomicName id="B4EC6594C965876F8286F990FED2A564" box="[755,932,1619,1644]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="homochaeta" status="sp. nov.">
|
||
<emphasis id="4198C205C965876F8286F990FED2A564" box="[755,932,1619,1644]" italics="true" pageId="11" pageNumber="27">G. homochaeta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4198C205C965876F83D9F991FE98A564" bold="true" box="[940,1006,1618,1644]" pageId="11" pageNumber="27">
|
||
<taxonomicNameLabel id="5AAB7F7EC965876F83D9F991FE98A564" box="[940,1006,1618,1644]" pageId="11" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
described herein from Heron Island (see above for details). Interestingly,
|
||
<taxonomicName id="B4EC6594C965876F8227F9BAFFA1A59A" box="[594,727,1657,1682]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="colorata">
|
||
<emphasis id="4198C205C965876F8227F9BAFFA1A59A" box="[594,727,1657,1682]" italics="true" pageId="11" pageNumber="27">G. colorata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B4EC6594C965876F8292F9BAFE14A59A" box="[743,866,1657,1682]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="galbina">
|
||
<emphasis id="4198C205C965876F8292F9BAFE14A59A" box="[743,866,1657,1682]" italics="true" pageId="11" pageNumber="27">G. galbina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B4EC6594C965876F83E8F9BAF938A59A" box="[925,1102,1657,1682]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="homochaeta">
|
||
<emphasis id="4198C205C965876F83E8F9BAF938A59A" box="[925,1102,1657,1682]" italics="true" pageId="11" pageNumber="27">G. homochaeta</emphasis>
|
||
</taxonomicName>
|
||
all lack ventral lobes of the pharyngeal glands in IV. This is an uncommon feature within
|
||
<taxonomicName id="B4EC6594C965876F83C6F963F978A5B1" box="[947,1038,1696,1721]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="genus">
|
||
<emphasis id="4198C205C965876F83C6F963F978A5B1" box="[947,1038,1696,1721]" italics="true" pageId="11" pageNumber="27">Grania</emphasis>
|
||
</taxonomicName>
|
||
, and further supports the close relationship between these three taxa.
|
||
</paragraph>
|
||
<paragraph id="73531E17C965876F80B0F92EF8EAA4C1" blockId="11.[151,1437,1154,2031]" pageId="11" pageNumber="27">
|
||
Another interesting characteristic of
|
||
<taxonomicName id="B4EC6594C965876F820EF92DFE76A40F" box="[635,768,1774,1799]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="species" species="colorata">
|
||
<emphasis id="4198C205C965876F820EF92DFE76A40F" box="[635,768,1774,1799]" italics="true" pageId="11" pageNumber="27">G. colorata</emphasis>
|
||
</taxonomicName>
|
||
, not seen in any other member of
|
||
<taxonomicName id="B4EC6594C965876F84E8F92DF984A40F" box="[1181,1266,1774,1799]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="genus">
|
||
<emphasis id="4198C205C965876F84E8F92DF984A40F" box="[1181,1266,1774,1799]" italics="true" pageId="11" pageNumber="27">Grania</emphasis>
|
||
</taxonomicName>
|
||
to date, is the presence of large masses of developing sperm in segments X–XII; this was observed in all specimens studied. In fixed
|
||
<taxonomicName id="B4EC6594C965876F8089F8F8FC27A45C" box="[252,337,1851,1876]" class="Clitellata" family="Enchytraeidae" genus="Grania" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="27" phylum="Annelida" rank="genus">
|
||
<emphasis id="4198C205C965876F8089F8F8FC27A45C" box="[252,337,1851,1876]" italics="true" pageId="11" pageNumber="27">Grania</emphasis>
|
||
</taxonomicName>
|
||
material it is not uncommon to observe limited amounts of developing sperm in front of the sperm sac, due to rupture or partial emptying of the sperm sac during fixation. That this would have happened consistently to all 91 individuals studied is highly unlikely, however, suggesting that this species uses its coelomic cavity as a storage place for maturing sperm which might not fit into the relatively small sperm sac.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3BF64D9CC965876F80B3F816F82CA4E7" box="[198,1370,2005,2031]" pageId="11" pageNumber="27" type="distribution">
|
||
<paragraph id="73531E17C965876F80B3F816F82CA4E7" blockId="11.[151,1437,1154,2031]" box="[198,1370,2005,2031]" pageId="11" pageNumber="27">
|
||
<emphasis id="4198C205C965876F80B3F816FF76A4E7" bold="true" box="[198,512,2005,2031]" pageId="11" pageNumber="27">Distribution and habitat:</emphasis>
|
||
Lizard Island, Great Barrier Reef, subtidal (to
|
||
<quantity id="B414B3F2C965876F8453F816F920A4E7" box="[1062,1110,2005,2031]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="27" unit="m" value="7.0">7 m</quantity>
|
||
), heterogeneous sand.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |