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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1120.87526" ID-GBIF-Dataset="76371b49-f4f1-4219-829f-9fc766d2fa60" ID-Pensoft-Pub="1313-2970-1120-1" ID-Pensoft-UUID="DB329BE83C4757FCAFDA96535E8C0D89" ID-ZooBank="711E0A8282AB43B59CC489BD87955061" ModsDocID="1313-2970-1120-1" checkinTime="1662465950044" checkinUser="pensoft" docAuthor="Lee, Chi-Feng &amp; Hsieh, Chia-Hung" docDate="2022" docId="FB00E6380FD858F195AB27478DCFDD56" docLanguage="en" docName="ZooKeys 1120: 1-46" docOrigin="ZooKeys 1120" docPubDate="2022-09-05" docSource="http://dx.doi.org/10.3897/zookeys.1120.87526" docTitle="Gonioctena (Brachyphytodecta) scutellaris Baly 1862, stat. rev." docType="treatment" docVersion="2" id="DB329BE83C4757FCAFDA96535E8C0D89" lastPageNumber="1" masterDocId="DB329BE83C4757FCAFDA96535E8C0D89" masterDocTitle="Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species" masterLastPageNumber="46" masterPageNumber="1" pageNumber="1" updateTime="1662466648910" updateUser="ExternalLinkService">
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<mods:title>Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species</mods:title>
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<mods:namePart>Lee, Chi-Feng</mods:namePart>
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<mods:affiliation>Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan</mods:affiliation>
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<mods:namePart>Hsieh, Chia-Hung</mods:namePart>
<mods:affiliation>Department of Forestry and Nature Conservation, Chinese Culture University, Taipei 111, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">xjh4@ulive.pccu.edu.tw</mods:nameIdentifier>
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<taxonomicName LSID="FB00E638-0FD8-58F1-95AB-27478DCFDD56" authority="Baly, 1862" authorityName="Baly" authorityYear="1862" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brachyphytodecta) scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" status="stat. rev." subGenus="Brachyphytodecta">Gonioctena (Brachyphytodecta) scutellaris Baly, 1862</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">stat. rev.</taxonomicNameLabel>
</paragraph>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) fulva (Motschulsky) A G. (B.) scutellaris, typical form, dorsal view B ditto, ventral view C same species, color variation, syntype of G. dichroa Fairmaire D G. (B.) fulva, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure8" httpUri="https://binary.pensoft.net/fig/738727" pageId="0" pageNumber="1">Figs 8A-C</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) liui sp. nov. A G. (B.) scutellaris, color variation, from Taiwan, dorsal view B ditto, ventral view C ditto, lateral view D G. (B.) liui sp. nov., dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure9" httpUri="https://binary.pensoft.net/fig/738728" pageId="0" pageNumber="1">, 9A-C</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Gonioctena (Brachyphytodecta) scutellaris Baly A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure10" httpUri="https://binary.pensoft.net/fig/738729" pageId="0" pageNumber="1">, 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Natural history of Gonioctena (Brachyphytodecta) scutellaris Baly on host plant, Callerya reticulata (Fabaceae) A eggs B first- and second-instar larvae C fourth-instar larva D pupa E adult, typical form, from Daken (大坑), Taichung F adult, color variation, from Lienhuachih (蓮華池), Nantou." figureDoi="10.3897/zookeys.1120.87526.figure11" httpUri="https://binary.pensoft.net/fig/738730" pageId="0" pageNumber="1">, 11</figureCitation>
</paragraph>
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<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" status="stat. rev.">Gonioctena scutellaris</taxonomicName>
Baly, 1862: 27 (North China); 
<bibRefCitation author="Gressitt, JL" journalOrPublisher="Genus" pageId="0" pageNumber="1" refId="B26" refString="Gressitt, JL, Kimoto, S, 1963. The Chrysomelidae (Coleopt.) of China and Korea, Part 2. Pacific Insects Monograph 1b: 301-1026." title="The Chrysomelidae (Coleopt.) of China and Korea, Part 2. Pacific Insects Monograph 1 b: 301 - 1026." year="1963">Gressitt and Kimoto 1963</bibRefCitation>
: 364 (as synonym of 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" family="Chrysomelidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="G. fulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="fulva" status="stat. rev.">G. fulva</taxonomicName>
; misidentification).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" status="stat. rev.">Phytodecta (s. str.) scutellaris</taxonomicName>
: 
<bibRefCitation author="Weise, J" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/10430680" refId="B64" refString="Weise, J, 1916. Chrysomelidae 12. Chrysomelinae. In: Junk W, Schenkling S (Eds) Coleopterorum Catalogus, Par 68, W. Junk, Berlin, https://www.biodiversitylibrary.org/page/10430680" title="Chrysomelidae 12. Chrysomelinae. In: Junk W, Schenkling S (Eds) Coleopterorum Catalogus, Par 68, W." url="https://www.biodiversitylibrary.org/page/10430680" year="1916">Weise 1916</bibRefCitation>
: 181 (catalogue).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" status="stat. rev.">Phytodecta scutellaris</taxonomicName>
: 
<bibRefCitation author="Chen, SH" journalOrPublisher="These, Faculte de Sciences Universite de Paris, Paris" pageId="0" pageNumber="1" refId="B6" refString="Chen, SH, 1934. Recherches sur les Chrysomelinae de la Chine et du Tonkin. These, Faculte de Sciences Universite de Paris, Paris" title="Recherches sur les Chrysomelinae de la Chine et du Tonkin." year="1934">Chen 1934</bibRefCitation>
: 73.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Asiphytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Asiphytodecta scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" status="stat. rev.">Asiphytodecta scutellaris</taxonomicName>
: 
<bibRefCitation author="Chen, SH" journalOrPublisher="Sinensia" pageId="0" pageNumber="1" pagination="199 - 210" refId="B9" refString="Chen, SH, Young, B, 1941. The coleopterous genus Asiphytodecta Chen. Sinensia 12: 199 - 210" title="The coleopterous genus Asiphytodecta Chen." volume="12" year="1941">Chen and Young 1941</bibRefCitation>
: 206 (key)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena thoracica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thoracica" status="stat. rev.">Gonioctena thoracica</taxonomicName>
Baly, 1862: 27 (North China); 
<bibRefCitation author="Gressitt, JL" journalOrPublisher="Genus" pageId="0" pageNumber="1" refId="B26" refString="Gressitt, JL, Kimoto, S, 1963. The Chrysomelidae (Coleopt.) of China and Korea, Part 2. Pacific Insects Monograph 1b: 301-1026." title="The Chrysomelidae (Coleopt.) of China and Korea, Part 2. Pacific Insects Monograph 1 b: 301 - 1026." year="1963">Gressitt and Kimoto 1963</bibRefCitation>
: 364 (as synonym of 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" family="Chrysomelidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="G. fulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="fulva" status="stat. rev.">G. fulva</taxonomicName>
; misidentification). Syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta thoracicus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thoracicus" status="stat. rev.">Phytodecta (s. str.) thoracicus</taxonomicName>
: 
<bibRefCitation author="Weise, J" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/10430680" refId="B64" refString="Weise, J, 1916. Chrysomelidae 12. Chrysomelinae. In: Junk W, Schenkling S (Eds) Coleopterorum Catalogus, Par 68, W. Junk, Berlin, https://www.biodiversitylibrary.org/page/10430680" title="Chrysomelidae 12. Chrysomelinae. In: Junk W, Schenkling S (Eds) Coleopterorum Catalogus, Par 68, W." url="https://www.biodiversitylibrary.org/page/10430680" year="1916">Weise 1916</bibRefCitation>
: 181 (catalogue).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta thoracicus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thoracicus" status="stat. rev.">Phytodecta thoracicus</taxonomicName>
: 
<bibRefCitation author="Chen, SH" journalOrPublisher="These, Faculte de Sciences Universite de Paris, Paris" pageId="0" pageNumber="1" refId="B6" refString="Chen, SH, 1934. Recherches sur les Chrysomelinae de la Chine et du Tonkin. These, Faculte de Sciences Universite de Paris, Paris" title="Recherches sur les Chrysomelinae de la Chine et du Tonkin." year="1934">Chen 1934</bibRefCitation>
: 73.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Asiphytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Asiphytodecta thoracicus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thoracicus" status="stat. rev.">Asiphytodecta thoracicus</taxonomicName>
: 
<bibRefCitation author="Chen, SH" journalOrPublisher="Sinensia" pageId="0" pageNumber="1" pagination="199 - 210" refId="B9" refString="Chen, SH, Young, B, 1941. The coleopterous genus Asiphytodecta Chen. Sinensia 12: 199 - 210" title="The coleopterous genus Asiphytodecta Chen." volume="12" year="1941">Chen and Young 1941</bibRefCitation>
: 206 (key)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena dichroa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dichroa" status="stat. rev.">Gonioctena dichroa</taxonomicName>
Fairmaire, 1888: 153 (China: Jiangxi); synonymized with 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" family="Chrysomelidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="P. thoracicus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="thoracicus" status="stat. rev.">P. thoracicus</taxonomicName>
by 
<bibRefCitation author="Chen, SH" journalOrPublisher="These, Faculte de Sciences Universite de Paris, Paris" pageId="0" pageNumber="1" refId="B6" refString="Chen, SH, 1934. Recherches sur les Chrysomelinae de la Chine et du Tonkin. These, Faculte de Sciences Universite de Paris, Paris" title="Recherches sur les Chrysomelinae de la Chine et du Tonkin." year="1934">Chen (1934)</bibRefCitation>
. Synonym confirmed
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta dichrous" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dichrous" status="stat. rev.">Phytodecta (s. str.) dichrous</taxonomicName>
: 
<bibRefCitation author="Weise, J" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/10430680" refId="B64" refString="Weise, J, 1916. Chrysomelidae 12. Chrysomelinae. In: Junk W, Schenkling S (Eds) Coleopterorum Catalogus, Par 68, W. Junk, Berlin, https://www.biodiversitylibrary.org/page/10430680" title="Chrysomelidae 12. Chrysomelinae. In: Junk W, Schenkling S (Eds) Coleopterorum Catalogus, Par 68, W." url="https://www.biodiversitylibrary.org/page/10430680" year="1916">Weise 1916</bibRefCitation>
: 181 (catalogue).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta (Asiphytodecta) issikii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="issikii" status="stat. rev." subGenus="Asiphytodecta">Phytodecta (Asiphytodecta) issikii</taxonomicName>
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
, 1958: 69 (Taiwan). Syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Asiphytodecta) issikii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="issikii" status="stat. rev." subGenus="Asiphytodecta">Gonioctena (Asiphytodecta) issikii</taxonomicName>
: 
<bibRefCitation DOI="https://doi.org/10.5109/2358" author="Kimoto, S" journalOrPublisher="Esakia" pageId="0" pageNumber="1" pagination="1 - 68" refId="B37" refString="Kimoto, S, 1969. Notes on the Chrysomelidae from Taiwan II. Esakia 7: 1 - 68, DOI: https://doi.org/10.5109/2358" title="Notes on the Chrysomelidae from Taiwan II." url="https://doi.org/10.5109/2358" volume="7" year="1969">Kimoto 1969</bibRefCitation>
: 22 (additional records in Taiwan)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brchyphytodecta) issikii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="issikii" status="stat. rev." subGenus="Brchyphytodecta">Gonioctena (Brchyphytodecta) issikii</taxonomicName>
: 
<bibRefCitation author="Gruev, B" journalOrPublisher="Entomological Review of Japan" pageId="0" pageNumber="1" pagination="51 - 54" publicationUrl="http://coleoptera.sakura.ne.jp/ERJ/ERJ44(1)1989.pdf" refId="B27" refString="Gruev, B, 1989. Two new species of Gonioctena (Brachyphytodecta) Chevrolat (Coleoptera, Chrysomelidae: Chrysomelinae) from South Asia and a note on Gonioctena issikii Chujo, 1958. Entomological Review of Japan 94: 51 - 54, http://coleoptera.sakura.ne.jp/ERJ/ERJ44(1)1989.pdf" title="Two new species of Gonioctena (Brachyphytodecta) Chevrolat (Coleoptera, Chrysomelidae: Chrysomelinae) from South Asia and a note on Gonioctena issikii Chujo, 1958." url="http://coleoptera.sakura.ne.jp/ERJ/ERJ44(1)1989.pdf" volume="94" year="1989">Gruev 1989</bibRefCitation>
: 54; 
<bibRefCitation author="Kimoto, S" journalOrPublisher="Bulletin of the Institute of Comparative Studies of International Cultures and Societies" pageId="0" pageNumber="1" pagination="1 - 152" refId="B43" refString="Kimoto, S, Chu, Y-I, 1996. Systematic catalog of Chrysomelidae of Taiwan (Insecta: Coleoptera). Bulletin of the Institute of Comparative Studies of International Cultures and Societies 16: 1 - 152" title="Systematic catalog of Chrysomelidae of Taiwan (Insecta: Coleoptera)." volume="16" year="1996">Kimoto and Chu 1996</bibRefCitation>
: 53 (catalogue); 
<bibRefCitation author="Kimoto, S" journalOrPublisher="Tokai University Press, Tokyo" pageId="0" pageNumber="1" refId="B44" refString="Kimoto, S, Takizawa, H, 1997. Leaf beetles (Chrysomelidae) of Taiwan. Tokai University Press, Tokyo" title="Leaf beetles (Chrysomelidae) of Taiwan." year="1997">Kimoto and Takizawa 1997</bibRefCitation>
: 369 (catalogue).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena issikii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="issikii" status="stat. rev.">Gonioctena issikii</taxonomicName>
: 
<bibRefCitation author="Takizawa, H" journalOrPublisher="Miscellaneous Reports of the Hiwa Museum for Natural History" pageId="0" pageNumber="1" pagination="1 - 16" refId="B60" refString="Takizawa, H, Nakamura, S, Kojima, K, 1995. The Taiwanese chrysomelid beetles preserved in Hiwa Museum for Natural History (Chrysomelidae: Coleoptera). Miscellaneous Reports of the Hiwa Museum for Natural History 33: 1 - 16" title="The Taiwanese chrysomelid beetles preserved in Hiwa Museum for Natural History (Chrysomelidae: Coleoptera)." volume="33" year="1995">Takizawa et al. 1995</bibRefCitation>
: 7 (additional records in Taiwan)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brchyphytodecta) foochowensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="foochowensis" status="stat. rev." subGenus="Brchyphytodecta">Gonioctena (Brchyphytodecta) foochowensis</taxonomicName>
Gruev, 1989: 53 (China: Fujian); 
<bibRefCitation author="Ge, S-Q" journalOrPublisher="Apollo Books, Stenstrup" pageId="0" pageNumber="1" refId="B24" refString="Ge, S-Q, 2010. New acts and comments, Chrysomelidae: Chrysomelinae. In: Loebl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Vol. 6. Apollo Books, Stenstrup" title="New acts and comments, Chrysomelidae: Chrysomelinae. In: Loebl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Vol. 6." year="2010">Ge 2010</bibRefCitation>
: 66 (as synonym of 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" family="Chrysomelidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="G. fulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="fulva" status="stat. rev.">G. fulva</taxonomicName>
; misidentification). Syn. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="types">
<paragraph pageId="0" pageNumber="1">Types.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis italics="true" pageId="0" pageNumber="1">Gonioctena scutellaris</emphasis>
</taxonomicName>
. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
♂ (NMHUK, here designated to clarify its identity among other synonyms): &quot;Type [h, w] // Type / H. T. [p, w, circle card with red border] // Baly Coll. [p, w] // 
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Gonioctena</taxonomicName>
/ 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris">Gonioctena scutellaris</taxonomicName>
/ Baly / N: China [h, b]&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena thoracica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thoracica">
<emphasis italics="true" pageId="0" pageNumber="1">Gonioctena thoracica</emphasis>
</taxonomicName>
. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
♂ (NMHUK, here designated for clarifying its identify with other synonyms): &quot;(aedeagus preserved inside a small container) // Type [h, w] // Type / H. T. [p, w, circle card with red border] // Baly Coll. [p, w] // 
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Gonioctena</taxonomicName>
/ 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena thoracica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thoracica">Gonioctena thoracica</taxonomicName>
/ Baly / N: China [h, b] // 
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Gonioctena</taxonomicName>
/ 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena thoracica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thoracica">Gonioctena thoracica</taxonomicName>
Baly [h] / det. H.W. Cho 2013 [p, w] // SYN- / TYPE [p, w, circle card with blue border]&quot;. Paralectotype: 1♀ (NMHUK): &quot;Type [h, w] // Baly Coll. [p, w] // SYN- / TYPE [p, w, circle card with blue border]&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena dichroa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis italics="true" pageId="0" pageNumber="1">Gonioctena dichroa</emphasis>
</taxonomicName>
. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
: 1♂ (MNHN): &quot;MUSEUM PARIS / KIANG-SI / A. DAVID 1875 [p, w] // 406 / 75 [h, w, circle label] // 
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Gonioctena</taxonomicName>
/ 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena dichroa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dichroa">Gonioctena dichroa</taxonomicName>
/ m. [h, w] // TYPE [p, w, red letters] // SYNTYPE [p, r] // 
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Gonioctena</taxonomicName>
/ 
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena dichora" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dichora">Gonioctena dichora</taxonomicName>
Frm. / rev. M. Daccordi, 2005 [p, w] // 
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Gonioctena</taxonomicName>
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena dichora" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dichora">Gonioctena dichora</taxonomicName>
/ Fiarmiare, 1888 / det. H.W. Cho 2014 [p, w] // MNHN, Pairs / EC14213 [p, w]&quot;; 1♀ (MNHN): &quot;MUSEUM PARIS / KIANG-SI / A. DAVID 1875 [p, w] // 406 / 75 [h, w, circle label] // 288 [h, green label] // SYNTYPE [p, r] // 
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Gonioctena</taxonomicName>
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena dichora" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dichora">Gonioctena dichora</taxonomicName>
/ Fiarmiare, 1888 / det. H.W. Cho 2014 [p, w] // MNHN, Pairs / EC14214 [p, w]&quot;. 
<bibRefCitation author="Fairmaire, K" journalOrPublisher="Revue d'Entomologie" pageId="0" pageNumber="1" pagination="111 - 160" publicationUrl="https://www.biodiversitylibrary.org/page/25446429" refId="B21" refString="Fairmaire, K, 1888. Notes sur les coleopteres des enirons de Pekin (2e partie). Revue d'Entomologie 7: 111 - 160, https://www.biodiversitylibrary.org/page/25446429" title="Notes sur les coleopteres des enirons de Pekin (2 e partie)." url="https://www.biodiversitylibrary.org/page/25446429" volume="7" year="1888">Fairmaire (1888)</bibRefCitation>
indicated that this species was described in Beijing (= Pekin) but both types actually were collected from Jiangxi (= Kiang-Si).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta (Asiphytodecta) issikii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="issikii" subGenus="Asiphytodecta">Phytodecta (Asiphytodecta) issikii</taxonomicName>
. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
♂ (TARI, original designation): &quot;Baibara [h] (= Meiyuan, 
<normalizedToken originalValue="眉原">眉原</normalizedToken>
) / FORMOSA [p] / 24.III.1943 [h] / COL. [p] S. Issiki [h, w] // 
<taxonomicName authorityName="Kirby" authorityYear="1837" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Phytodecta</taxonomicName>
/ 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta issikii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="issikii">Phytodecta issikii</taxonomicName>
/ / 
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
[h] / DET. M. CHUJO [p, w] // Holo / Type [h, w; circle card with red letters and border but fade out] // 695 [p, w]&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brchyphytodecta) foochowensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="foochowensis" subGenus="Brchyphytodecta">Gonioctena (Brchyphytodecta) foochowensis</taxonomicName>
. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
♀ (NMHUK, original designation): &quot;CHINA / Foochow / C. R. Kelloqq [h, w] // Field No. [p] / 980 [h, w] // 
<taxonomicName authorityName="Kirby" authorityYear="1837" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Phytodecta</taxonomicName>
/ sp [h] / Det. G. E. Bryant. [p] // Brit. Mus. / 198[p]1-315 [h, w] // 
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Gonioctena</taxonomicName>
/ 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena foochowensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="foochowensis">Gonioctena foochowensis</taxonomicName>
/ Gruev [p, w] // HOLOTYPE [p, r] // 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena dichroa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dichroa">Gonioctena dichroa</taxonomicName>
/ Fairmaire, 1888 / det. H.W. Cho 2014 [p, w]&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material (n = 150).</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Foochow, M. S. Yang" country="China" location="Fujian" specimenCount="13" specimenCount-female="8" specimenCount-male="5" stateProvince="Fujian">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="China">China</collectingCountry>
</emphasis>
. 
<collectingRegion country="China" name="Fujian">Fujian</collectingRegion>
: 
<specimenCount type="male">5♂</specimenCount>
, 
<specimenCount type="female">8♀</specimenCount>
(NMHUK), 
<collectorName>Foochow</collectorName>
(福州), 1935-1938, leg. 
<collectorName>M. S. Yang</collectorName>
</materialsCitation>
; 
<materialsCitation collectingDate="2016-04-28" collectorName="Lienhuachih, A. Li" country="Taiwan" location="Nantou" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Nantou">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
</emphasis>
. 
<collectingRegion country="Taiwan" name="Nantou">Nantou</collectingRegion>
: 
<specimenCount type="male">2♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
(TARI), 
<collectorName>Lienhuachih</collectorName>
(蓮華池), 
<collectingDate value="2016-04-28">28.IV.2016</collectingDate>
, leg. 
<collectorName>A. Li</collectorName>
</materialsCitation>
; 
<materialsCitation collectingDate="2021-04-20" collectorName="Tsaitso, C. - F. Lee" country="Taiwan" location="Island" specimenCount="21" specimenCount-female="7" specimenCount-male="14" stateProvince="Kinmen">
<collectingRegion country="Taiwan" name="Kinmen">Kinmen</collectingRegion>
: 
<collectingRegion country="Taiwan" name="Kinmen">Kinmen</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:FB00E6380FD858F195AB27478DCFDD56:8ADB4EA9B61DAFC039A07469D7233303" country="Taiwan" name="Island" stateProvince="Kinmen">Island</location>
(金門): 
<specimenCount type="male">14♂</specimenCount>
, 
<specimenCount type="female">7♀</specimenCount>
(TARI), 
<collectorName>Tsaitso</collectorName>
trail (蔡厝古道), 
<collectingDate value="2021-04-20">20.IV.2021</collectingDate>
, leg. 
<collectorName>C.-F. Lee</collectorName>
</materialsCitation>
; 
<specimenCount type="male">4♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
(TARI), Taiwushan (太武山), 
<date value="2011-04-14">14.IV.2011</date>
, leg. Y.-J. Chang; 
<materialsCitation collectingDate="2011-05-06" collectingDateMax="2021-04-08" collectingDateMin="2011-05-06" collectorName="C. - F. Lee" country="Taiwan" location="Island" specimenCount="61" specimenCount-female="30" specimenCount-male="31" stateProvince="Kinmen">
<specimenCount type="female">1♀</specimenCount>
(TARI), same but with 
<normalizedToken originalValue="“6.V.2011”">
&quot; 
<collectingDate value="2011-05-06">6.V.2011</collectingDate>
&quot;
</normalizedToken>
; 
<specimenCount type="male">5♂</specimenCount>
, 
<specimenCount type="female">2♀</specimenCount>
(TARI), same but with 
<normalizedToken originalValue="“19.IV.2015”">
&quot; 
<collectingDate value="2015-04-19">19.IV.2015</collectingDate>
&quot;
</normalizedToken>
; 
<specimenCount type="male">2♂</specimenCount>
, 
<specimenCount type="female">4♀</specimenCount>
(TARI), same but with 
<normalizedToken originalValue="“29-30.IV.2015”">
&quot; 
<collectingDate value="2015-04-29" valueMax="2015-04-30" valueMin="2015-04-29">29-30.IV.2015</collectingDate>
&quot;
</normalizedToken>
; 
<specimenCount type="male">24♂</specimenCount>
, 
<specimenCount type="female">23♀</specimenCount>
(TARI), same locality, 
<collectingDate value="2021-04-08">8.IV.2021</collectingDate>
, leg. 
<collectorName>C.-F. Lee</collectorName>
</materialsCitation>
; 
<materialsCitation collectingDate="2016-05-09" collectingDateMax="2021-04-15" collectingDateMin="2016-05-09" collectorName="Y. - J. Chang" country="Taiwan" location="Dadan Island" specimenCount="51" specimenCount-female="28" specimenCount-male="23">
<specimenCount type="male">23♂</specimenCount>
, 
<specimenCount type="female">27♀</specimenCount>
(TARI), same but with 
<normalizedToken originalValue="“15.IV.2021”">
&quot; 
<collectingDate value="2021-04-15">15.IV.2021</collectingDate>
&quot;
</normalizedToken>
; 
<location LSID="urn:lsid:plazi:treatment:FB00E6380FD858F195AB27478DCFDD56:20EAA3581275417EB4F8D46BC5668537" country="Taiwan" name="Dadan Island">Dadan Island</location>
(大膽島): 
<specimenCount type="female">1♀</specimenCount>
(TARI), 
<collectingDate value="2016-05-09">9.V.2016</collectingDate>
, leg. 
<collectorName>Y.-J. Chang</collectorName>
</materialsCitation>
; 
<materialsCitation collectingDate="2018-05-01" collectorName="H. - T. Fang" country="Taiwan" location="Beigan Island" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Matsu Islands">
<collectingRegion country="Taiwan" name="Lienchiang">Matsu Islands</collectingRegion>
: 
<specimenCount type="male">1♂</specimenCount>
, 
<specimenCount type="female">2♀</specimenCount>
(TARI), 
<location LSID="urn:lsid:plazi:treatment:FB00E6380FD858F195AB27478DCFDD56:C84A06A553012EFA9A3FFE69A7B471A6" country="Taiwan" name="Beigan Island" stateProvince="Matsu Islands">Beigan Island</location>
(北竿), 
<collectingDate value="2018-05-01">1.V.2018</collectingDate>
, leg. 
<collectorName>H.-T. Fang</collectorName>
</materialsCitation>
; 
<materialsCitation collectingDate="2009-05-24" collectorName="U. Ong." country="Taiwan" location="Nangan Island" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
(TARI), 
<location LSID="urn:lsid:plazi:treatment:FB00E6380FD858F195AB27478DCFDD56:3929EF9B8E8991C8F736CF3BA4BDE396" country="Taiwan" name="Nangan Island">Nangan Island</location>
(南竿), 
<collectingDate value="2009-05-24">24.V.2009</collectingDate>
, leg. 
<collectorName>U. Ong.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="redescription">
<paragraph pageId="0" pageNumber="1">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="1">
Length 5.3-6.4 mm, width 3.5-4.0 mm. Body color (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) fulva (Motschulsky) A G. (B.) scutellaris, typical form, dorsal view B ditto, ventral view C same species, color variation, syntype of G. dichroa Fairmaire D G. (B.) fulva, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure8" httpUri="https://binary.pensoft.net/fig/738727" pageId="0" pageNumber="1">8A, B</figureCitation>
) yellowish brown; antennomeres V-XI, legs, and scutellum black. Antennae (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Gonioctena (Brachyphytodecta) scutellaris Baly A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure10" httpUri="https://binary.pensoft.net/fig/738729" pageId="0" pageNumber="1">10A</figureCitation>
) with antennomere III slender, IV-VI slightly swollen, VII moderately swollen, VIII-X strongly swollen, XI elongate oval, length ratios of antennomeres I-XI 1.0: 0.6: 0.6: 0.4: 0.4: 0.4: 0.5: 0.5: 0.5: 0.5: 0.9, length to width ratios of antennomeres I-XI 2.3: 1.9: 2.4: 1.7: 1.6: 1.0: 1.1: 1.1: 0.9: 0.9: 1.7. Pronotum 2.2-2.3 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, lateral margins widest at base, convergent anteriorly, anterior angles strongly produced; anterior and lateral margins bordered, lateral margins barely visible in dorsal view; trichobothria absent on both anterior and posterior angles; disc covered with sparse fine punctures and mixed with finer punctures; both sides covered with much larger, denser punctures. Scutellum distinctly wider than long, narrowed posteriorly. Elytra 1.1-1.2 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; lateral margins slightly wider posteriorly, widest near middle; humeral calli well developed; disc covered with rather regular coarse punctures arranged into single stria; interspaces covered with fine, sparse punctures. Hind wing well developed. Aedeagus (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Gonioctena (Brachyphytodecta) scutellaris Baly A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure10" httpUri="https://binary.pensoft.net/fig/738729" pageId="0" pageNumber="1">10B, C</figureCitation>
) with apical process extremely slender and elongate in dorsal view, 0.5 
<normalizedToken originalValue="×">x</normalizedToken>
as long as aedeagus; lateral margins slightly narrowed in basal 1/3; strongly curved at basal 1/3 in lateral view; endophallic sclerite extremely elongate. Gonocoxae (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Gonioctena (Brachyphytodecta) scutellaris Baly A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure10" httpUri="https://binary.pensoft.net/fig/738729" pageId="0" pageNumber="1">10D</figureCitation>
) wide, but apical margin irregular, with several long setae along apical margins. Ventrite VIII (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Gonioctena (Brachyphytodecta) scutellaris Baly A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure10" httpUri="https://binary.pensoft.net/fig/738729" pageId="0" pageNumber="1">10E</figureCitation>
) transverse, with several long setae along outer margin. Receptacle of spermatheca (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Gonioctena (Brachyphytodecta) scutellaris Baly A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure10" httpUri="https://binary.pensoft.net/fig/738729" pageId="0" pageNumber="1">10F</figureCitation>
) slightly swollen, not separated from pump; pump short and curved; sclerotized proximal spermathecal duct moderately short.
</paragraph>
<caption doi="10.3897/zookeys.1120.87526.figure8" httpUri="https://binary.pensoft.net/fig/738727" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
Habitus of 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brachyphytodecta) scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" subGenus="Brachyphytodecta">Gonioctena (Brachyphytodecta) scutellaris</taxonomicName>
Baly and 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) fulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="fulva" subGenus="B.">G. (B.) fulva</taxonomicName>
(Motschulsky) 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) scutellaris" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="scutellaris" subGenus="B.">G. (B.) scutellaris</taxonomicName>
, typical form, dorsal view 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
ditto, ventral view 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
same species, color variation, syntype of 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. dichroa" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="dichroa">
<emphasis italics="true" pageId="0" pageNumber="1">G. dichroa</emphasis>
</taxonomicName>
Fairmaire 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) fulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="fulva" subGenus="B.">G. (B.) fulva</taxonomicName>
, dorsal view 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
ditto, ventral view 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
ditto, lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variations.</paragraph>
<paragraph pageId="0" pageNumber="1">
Many adults possess black elytra with wide yellowish brown borders (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) liui sp. nov. A G. (B.) scutellaris, color variation, from Taiwan, dorsal view B ditto, ventral view C ditto, lateral view D G. (B.) liui sp. nov., dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure9" httpUri="https://binary.pensoft.net/fig/738728" pageId="0" pageNumber="1">9A-C</figureCitation>
) that were described as 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena foochowensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="foochowensis">
<emphasis italics="true" pageId="0" pageNumber="1">Gonioctena foochowensis</emphasis>
</taxonomicName>
and 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. issikii" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="issikii">
<emphasis italics="true" pageId="0" pageNumber="1">G. issikii</emphasis>
</taxonomicName>
. Some with the elytra entirely black were described as 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. dichroa" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="dichroa">
<emphasis italics="true" pageId="0" pageNumber="1">G. dichroa</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) fulva (Motschulsky) A G. (B.) scutellaris, typical form, dorsal view B ditto, ventral view C same species, color variation, syntype of G. dichroa Fairmaire D G. (B.) fulva, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure8" httpUri="https://binary.pensoft.net/fig/738727" pageId="0" pageNumber="1">8C</figureCitation>
) and 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. thoracica" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="thoracica">
<emphasis italics="true" pageId="0" pageNumber="1">G. thoracica</emphasis>
</taxonomicName>
.
</paragraph>
<caption doi="10.3897/zookeys.1120.87526.figure9" httpUri="https://binary.pensoft.net/fig/738728" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
Habitus of 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brachyphytodecta) scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" subGenus="Brachyphytodecta">Gonioctena (Brachyphytodecta) scutellaris</taxonomicName>
Baly and 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) liui" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="liui" subGenus="B.">G. (B.) liui</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) scutellaris" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="scutellaris" subGenus="B.">G. (B.) scutellaris</taxonomicName>
, color variation, from Taiwan, dorsal view 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
ditto, ventral view 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
ditto, lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) liui" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="liui" subGenus="B.">G. (B.) liui</taxonomicName>
sp. nov., dorsal view 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
ditto, ventral view 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
ditto, lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brachyphytodecta) scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" subGenus="Brachyphytodecta">Gonioctena (Brachyphytodecta) scutellaris</taxonomicName>
can be distinguished from the other consubgeneric species, 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) liui" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="liui" subGenus="B.">G. (B.) liui</taxonomicName>
sp. nov. by the following combination of the characters: yellowish brown head and pronotum (Figs 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) fulva (Motschulsky) A G. (B.) scutellaris, typical form, dorsal view B ditto, ventral view C same species, color variation, syntype of G. dichroa Fairmaire D G. (B.) fulva, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure8" httpUri="https://binary.pensoft.net/fig/738727" pageId="0" pageNumber="1">8A, C</figureCitation>
, 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) liui sp. nov. A G. (B.) scutellaris, color variation, from Taiwan, dorsal view B ditto, ventral view C ditto, lateral view D G. (B.) liui sp. nov., dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure9" httpUri="https://binary.pensoft.net/fig/738728" pageId="0" pageNumber="1">9A</figureCitation>
), thoracic and abdominal ventrites (Figs 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) fulva (Motschulsky) A G. (B.) scutellaris, typical form, dorsal view B ditto, ventral view C same species, color variation, syntype of G. dichroa Fairmaire D G. (B.) fulva, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure8" httpUri="https://binary.pensoft.net/fig/738727" pageId="0" pageNumber="1">8B</figureCitation>
, 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) liui sp. nov. A G. (B.) scutellaris, color variation, from Taiwan, dorsal view B ditto, ventral view C ditto, lateral view D G. (B.) liui sp. nov., dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure9" httpUri="https://binary.pensoft.net/fig/738728" pageId="0" pageNumber="1">9B</figureCitation>
) (black head and most parts of pronotum (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) liui sp. nov. A G. (B.) scutellaris, color variation, from Taiwan, dorsal view B ditto, ventral view C ditto, lateral view D G. (B.) liui sp. nov., dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure9" httpUri="https://binary.pensoft.net/fig/738728" pageId="0" pageNumber="1">9D</figureCitation>
), thoracic and abdominal ventrites (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Habitus of Gonioctena (Brachyphytodecta) scutellaris Baly and G. (B.) liui sp. nov. A G. (B.) scutellaris, color variation, from Taiwan, dorsal view B ditto, ventral view C ditto, lateral view D G. (B.) liui sp. nov., dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1120.87526.figure9" httpUri="https://binary.pensoft.net/fig/738728" pageId="0" pageNumber="1">9E</figureCitation>
) in 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) liui" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="liui" subGenus="B.">G. (B.) liui</taxonomicName>
sp. nov.), extremely elongate apical process of aedeagus (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Gonioctena (Brachyphytodecta) scutellaris Baly A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure10" httpUri="https://binary.pensoft.net/fig/738729" pageId="0" pageNumber="1">10B, C</figureCitation>
) (short, wide apical process of aedeagus in 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) liui" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="liui" subGenus="B.">G. (B.) liui</taxonomicName>
sp. nov. (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Diagnostic characters of Gonioctena (Brachyphytodecta) liui sp. nov. A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure12" httpUri="https://binary.pensoft.net/fig/738731" pageId="0" pageNumber="1">12B, C</figureCitation>
)), and wide gonocoxae covered with more setae (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Gonioctena (Brachyphytodecta) scutellaris Baly A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure10" httpUri="https://binary.pensoft.net/fig/738729" pageId="0" pageNumber="1">10D</figureCitation>
) (narrow gonoxae covered with fewer setae in 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. (B.) liui" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="liui" subGenus="B.">G. (B.) liui</taxonomicName>
sp. nov. (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Diagnostic characters of Gonioctena (Brachyphytodecta) liui sp. nov. A antenna B aedeagus, dorsal view C aedeagus, lateral view D gonocoxae E abdominal ventrite VIII F spermatheca." figureDoi="10.3897/zookeys.1120.87526.figure12" httpUri="https://binary.pensoft.net/fig/738731" pageId="0" pageNumber="1">12D</figureCitation>
)).
</paragraph>
<caption doi="10.3897/zookeys.1120.87526.figure10" httpUri="https://binary.pensoft.net/fig/738729" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
Diagnostic characters of 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brachyphytodecta) scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" subGenus="Brachyphytodecta">Gonioctena (Brachyphytodecta) scutellaris</taxonomicName>
Baly 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
antenna 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
aedeagus, dorsal view 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
aedeagus, lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
gonocoxae 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
abdominal ventrite VIII 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
spermatheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="host">
<paragraph pageId="0" pageNumber="1">Host plants.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Chevrolat in Dejean" authorityYear="1836" class="Insecta" family="Fabaceae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" rank="family">Fabaceae</taxonomicName>
: 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Callerya" kingdom="Plantae" lsidName="Callerya reticulata" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis italics="true" pageId="0" pageNumber="1">Callerya reticulata</emphasis>
</taxonomicName>
(Benth.) Schot (new record, present study)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brachyphytodecta) scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" subGenus="Brachyphytodecta">Gonioctena (Brachyphytodecta) scutellaris</taxonomicName>
are presumed to be multivoltine during spring and females are oviparous. In Taiwan, this species seems rare due to unpredictable sprouting times of host plants. Mr. Alex Li (李志穎) collected some adults (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Natural history of Gonioctena (Brachyphytodecta) scutellaris Baly on host plant, Callerya reticulata (Fabaceae) A eggs B first- and second-instar larvae C fourth-instar larva D pupa E adult, typical form, from Daken (大坑), Taichung F adult, color variation, from Lienhuachih (蓮華池), Nantou." figureDoi="10.3897/zookeys.1120.87526.figure11" httpUri="https://binary.pensoft.net/fig/738730" pageId="0" pageNumber="1">11F</figureCitation>
) and mature larvae on April 28, 2016, at Lienhuachih (蓮華池). However, we could not find this species around the same season in the following years because host plants had not sprouted. For example, we could not find sprouts of the host plants in late May 2021 at that location. The populations in small islands are more stable, where host plants began to sprout, and overwintered female laid eggs during middle of March 2021 in Kinmen Islands. They laid 12 eggs into two rows on very young leaves (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Natural history of Gonioctena (Brachyphytodecta) scutellaris Baly on host plant, Callerya reticulata (Fabaceae) A eggs B first- and second-instar larvae C fourth-instar larva D pupa E adult, typical form, from Daken (大坑), Taichung F adult, color variation, from Lienhuachih (蓮華池), Nantou." figureDoi="10.3897/zookeys.1120.87526.figure11" httpUri="https://binary.pensoft.net/fig/738730" pageId="0" pageNumber="1">11A</figureCitation>
). Larvae hatched in three-four days. Early instar larvae (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Natural history of Gonioctena (Brachyphytodecta) scutellaris Baly on host plant, Callerya reticulata (Fabaceae) A eggs B first- and second-instar larvae C fourth-instar larva D pupa E adult, typical form, from Daken (大坑), Taichung F adult, color variation, from Lienhuachih (蓮華池), Nantou." figureDoi="10.3897/zookeys.1120.87526.figure11" httpUri="https://binary.pensoft.net/fig/738730" pageId="0" pageNumber="1">11B</figureCitation>
) fed gregariously on young leaves but became solitary as they matured. The larval duration was nine days. Mature larvae (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Natural history of Gonioctena (Brachyphytodecta) scutellaris Baly on host plant, Callerya reticulata (Fabaceae) A eggs B first- and second-instar larvae C fourth-instar larva D pupa E adult, typical form, from Daken (大坑), Taichung F adult, color variation, from Lienhuachih (蓮華池), Nantou." figureDoi="10.3897/zookeys.1120.87526.figure11" httpUri="https://binary.pensoft.net/fig/738730" pageId="0" pageNumber="1">11C</figureCitation>
) burrowed into soil and built underground chambers for pupation. Duration of the pupal stage (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Natural history of Gonioctena (Brachyphytodecta) scutellaris Baly on host plant, Callerya reticulata (Fabaceae) A eggs B first- and second-instar larvae C fourth-instar larva D pupa E adult, typical form, from Daken (大坑), Taichung F adult, color variation, from Lienhuachih (蓮華池), Nantou." figureDoi="10.3897/zookeys.1120.87526.figure11" httpUri="https://binary.pensoft.net/fig/738730" pageId="0" pageNumber="1">11D</figureCitation>
) was nine days. The newly emerged adults (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Natural history of Gonioctena (Brachyphytodecta) scutellaris Baly on host plant, Callerya reticulata (Fabaceae) A eggs B first- and second-instar larvae C fourth-instar larva D pupa E adult, typical form, from Daken (大坑), Taichung F adult, color variation, from Lienhuachih (蓮華池), Nantou." figureDoi="10.3897/zookeys.1120.87526.figure11" httpUri="https://binary.pensoft.net/fig/738730" pageId="0" pageNumber="1">11F</figureCitation>
) appeared during early April and females started to lay eggs after a week. In Jiangxi, China, females began to lay eggs first and then laid larvae in early April (
<bibRefCitation author="Zhang, S-M" journalOrPublisher="Kunchong Zhishi" pageId="0" pageNumber="1" pagination="132 - 133" refId="B67" refString="Zhang, S-M, Shen, R-W, 1984. Observation on the viviparous habit of the chrysomelid beetle Gonioctena fulva (Mots.). Kunchong Zhishi 21: 132 - 133" title="Observation on the viviparous habit of the chrysomelid beetle Gonioctena fulva (Mots.)." volume="21" year="1984">Zhang and Shen 1984</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1120.87526.figure11" httpUri="https://binary.pensoft.net/fig/738730" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
Natural history of 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Brachyphytodecta) scutellaris" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutellaris" subGenus="Brachyphytodecta">Gonioctena (Brachyphytodecta) scutellaris</taxonomicName>
Baly on host plant, 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Callerya" kingdom="Plantae" lsidName="Callerya reticulata" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="reticulata">
<emphasis italics="true" pageId="0" pageNumber="1">Callerya reticulata</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="Endlicher" authorityYear="1843" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
) 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
eggs 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
first- and second-instar larvae 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
fourth-instar larva 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
pupa 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
adult, typical form, from Daken (大坑), Taichung 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
adult, color variation, from Lienhuachih (蓮華池), Nantou.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Chen, SH" journalOrPublisher="These, Faculte de Sciences Universite de Paris, Paris" pageId="0" pageNumber="1" refId="B6" refString="Chen, SH, 1934. Recherches sur les Chrysomelinae de la Chine et du Tonkin. These, Faculte de Sciences Universite de Paris, Paris" title="Recherches sur les Chrysomelinae de la Chine et du Tonkin." year="1934">Chen (1934)</bibRefCitation>
regarded 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. dichroa" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="dichroa">
<emphasis italics="true" pageId="0" pageNumber="1">G. dichroa</emphasis>
</taxonomicName>
as synonym of 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. thoracica" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="thoracica">
<emphasis italics="true" pageId="0" pageNumber="1">G. thoracica</emphasis>
</taxonomicName>
. 
<bibRefCitation author="Gressitt, JL" journalOrPublisher="Genus" pageId="0" pageNumber="1" refId="B26" refString="Gressitt, JL, Kimoto, S, 1963. The Chrysomelidae (Coleopt.) of China and Korea, Part 2. Pacific Insects Monograph 1b: 301-1026." title="The Chrysomelidae (Coleopt.) of China and Korea, Part 2. Pacific Insects Monograph 1 b: 301 - 1026." year="1963">Gressitt and Kimoto (1963)</bibRefCitation>
synonymized 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. thoracica" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="thoracica">
<emphasis italics="true" pageId="0" pageNumber="1">G. thoracica</emphasis>
</taxonomicName>
and 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. scutellaris" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="scutellaris">
<emphasis italics="true" pageId="0" pageNumber="1">G. scutellaris</emphasis>
</taxonomicName>
with 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" family="Cannabaceae" kingdom="Animalia" lsidName="G. fulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="fulva">
<emphasis italics="true" pageId="0" pageNumber="1">G. fulva</emphasis>
</taxonomicName>
. 
<bibRefCitation author="Ge, S-Q" journalOrPublisher="Apollo Books, Stenstrup" pageId="0" pageNumber="1" refId="B24" refString="Ge, S-Q, 2010. New acts and comments, Chrysomelidae: Chrysomelinae. In: Loebl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Vol. 6. Apollo Books, Stenstrup" title="New acts and comments, Chrysomelidae: Chrysomelinae. In: Loebl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Vol. 6." year="2010">Ge (2010)</bibRefCitation>
treated 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. foochowensis" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="foochowensis">
<emphasis italics="true" pageId="0" pageNumber="1">G. foochowensis</emphasis>
</taxonomicName>
as a synonym of 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" family="Cannabaceae" kingdom="Animalia" lsidName="G. fulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="fulva">
<emphasis italics="true" pageId="0" pageNumber="1">G. fulva</emphasis>
</taxonomicName>
. After examining aedeagi of these species, we concluded that 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. scutellaris" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="scutellaris">
<emphasis italics="true" pageId="0" pageNumber="1">G. scutellaris</emphasis>
</taxonomicName>
is a distinct species, which is removed from synonymy with 
<taxonomicName authorityName="Lee &amp; Hsieh" authorityYear="2022" family="Cannabaceae" kingdom="Animalia" lsidName="G. fulva" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="fulva">
<emphasis italics="true" pageId="0" pageNumber="1">G. fulva</emphasis>
</taxonomicName>
. Moreover, 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. dichora" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="dichora">
<emphasis italics="true" pageId="0" pageNumber="1">G. dichora</emphasis>
</taxonomicName>
, 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. thoracica" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="thoracica">
<emphasis italics="true" pageId="0" pageNumber="1">G. thoracica</emphasis>
</taxonomicName>
, and 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. foochowensis" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="foochowensis">
<emphasis italics="true" pageId="0" pageNumber="1">G. foochowensis</emphasis>
</taxonomicName>
are conspecific with 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. scutellaris" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="scutellaris">
<emphasis italics="true" pageId="0" pageNumber="1">G. scutellaris</emphasis>
</taxonomicName>
. We found that 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. issikii" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="issikii">
<emphasis italics="true" pageId="0" pageNumber="1">G. issikii</emphasis>
</taxonomicName>
is also a junior synonym of 
<taxonomicName family="Cannabaceae" kingdom="Animalia" lsidName="G. scutellaris" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="scutellaris">
<emphasis italics="true" pageId="0" pageNumber="1">G. scutellaris</emphasis>
</taxonomicName>
. The typical color form (entirely yellowish brown elytra) is extremely rare in Kinmen and Matsu islands, and Taiwan. Only three specimens of such form were collected from Kinmen Island (金門島) and Dadan Island (大膽島). Many larvae were brought from Kinmen Island into the laboratory for rearing during March 2021. Only one specimen with the typical color form was among more than 50 adults reared from larvae. In addition, although no specimens of this form were collected from Taiwan, photographs of this form were taken at Darken (大坑), Taichng in May 1, 2020 (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Natural history of Gonioctena (Brachyphytodecta) scutellaris Baly on host plant, Callerya reticulata (Fabaceae) A eggs B first- and second-instar larvae C fourth-instar larva D pupa E adult, typical form, from Daken (大坑), Taichung F adult, color variation, from Lienhuachih (蓮華池), Nantou." figureDoi="10.3897/zookeys.1120.87526.figure11" httpUri="https://binary.pensoft.net/fig/738730" pageId="0" pageNumber="1">11E</figureCitation>
) by Hsien Chung Liu (劉獻宗).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
China, Taiwan, including Kinmen Islands (Kinmen Island 
<normalizedToken originalValue="金門島">金門島</normalizedToken>
, Dadan Island 
<normalizedToken originalValue="大膽島">大膽島</normalizedToken>
) and Matsu Islands (Kaoteng Island 
<normalizedToken originalValue="高登島">高登島</normalizedToken>
, Beigan Island 
<normalizedToken originalValue="北竿島">北竿島</normalizedToken>
, and Nangan Island 
<normalizedToken originalValue="南竿島">南竿島</normalizedToken>
, new records).
</paragraph>
</subSubSection>
</treatment>
</document>