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<document id="A7F818030CAE4BED7904077B699FA329" ID-DOI="10.1111/j.1096-3642.2011.00779.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5407022" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630346869281" checkinUser="marcus" docAuthor="Griswold, Charles E., Wood, Hannah Marie &amp; Carmichael, Anthea D." docDate="2012" docId="977847656976FFBB05217231BAEBFD01" docLanguage="en" docName="j.1096-3642.2011.00779.x.pdf" docOrigin="Zoological Journal of the Linnean Society 164 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00779.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Manampoka Griswold &amp; Wood &amp; Carmichael 2012, GEN. NOV." docType="treatment" docVersion="3" lastPageNumber="797" masterDocId="6B413F1D6932FFFE046B7016BA1DFFC9" masterDocTitle="The lace web spiders (Araneae, Phyxelididae) of Madagascar: phylogeny, biogeography and taxonomy" masterLastPageNumber="810" masterPageNumber="728" pageNumber="796" updateTime="1699238303782" updateUser="plazi">
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<mods:title id="B8FDB6B386E16EE3EDEAC89903119B75">The lace web spiders (Araneae, Phyxelididae) of Madagascar: phylogeny, biogeography and taxonomy</mods:title>
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<mods:namePart id="F671953A5E615578835906F3017788B1">Griswold, Charles E.</mods:namePart>
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<mods:namePart id="69DE190D50AA80D8282837216327BBF7">Wood, Hannah Marie</mods:namePart>
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<mods:namePart id="544C2E269684466ED7FA8CB109E909D7">Carmichael, Anthea D.</mods:namePart>
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<mods:title id="B06193831B5A1711BBDD443841EABEF4">Zoological Journal of the Linnean Society</mods:title>
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<subSubSection id="57CBA5F86976FFBA05217231B879FDF5" box="[330,612,551,572]" pageId="68" pageNumber="796" type="nomenclature">
<paragraph id="1F6EF6736976FFBA05217231B879FDF5" blockId="68.[330,612,551,572]" box="[330,612,551,572]" pageId="68" pageNumber="796">
<heading id="4426411F6976FFBA05217231B879FDF5" box="[330,612,551,572]" centered="true" fontSize="9" level="2" pageId="68" pageNumber="796" reason="4">
<taxonomicName id="D8D18DF06976FFBA05217231BBFCFDF2" ID-CoL="62ZMJ" ID-ENA="1161402" authority="Griswold &amp; Wood &amp; Carmichael, 2012" authorityName="Griswold &amp; Wood &amp; Carmichael" authorityYear="2012" box="[330,481,551,572]" class="Arachnida" family="Phyxelididae" genus="Manampoka" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus" status="gen. nov.">MANAMPOKA</taxonomicName>
<emphasis id="2DA52A616976FFBA0580723CB879FDF5" bold="true" box="[491,612,551,572]" pageId="68" pageNumber="796">
<taxonomicNameLabel id="3696971A6976FFBA0580723CB879FDF5" box="[491,612,551,572]" pageId="68" pageNumber="796" rank="genus">GEN. NOV.</taxonomicNameLabel>
</emphasis>
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<paragraph id="1F6EF6736976FFBA04CF7258BB3AFD4B" blockId="68.[164,779,589,642]" pageId="68" pageNumber="796">
<emphasis id="2DA52A616976FFBA04CF7258B858FDAA" box="[164,581,589,611]" italics="true" pageId="68" pageNumber="796">
<typeStatus id="C06A48D16976FFBA04CF7258BAC5FDAA" box="[164,216,590,611]" pageId="68" pageNumber="796">Type</typeStatus>
species:
<taxonomicName id="D8D18DF06976FFBA053D725BB858FDAA" authority="Griswold &amp; Wood &amp; Carmichael, 2012" authorityName="Griswold &amp; Wood &amp; Carmichael" authorityYear="2012" box="[342,581,589,611]" class="Arachnida" family="Phyxelididae" genus="Manampoka" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="species" species="atsimo" status="sp. nov.">Manampoka atsimo</taxonomicName>
</emphasis>
<taxonomicNameLabel id="3696971A6976FFBA06317258B8A2FDAA" box="[602,703,590,611]" pageId="68" pageNumber="796" rank="species">sp. nov.</taxonomicNameLabel>
, here designated.
</paragraph>
<paragraph id="1F6EF6736976FFBA04CF72BDB916FD16" blockId="68.[164,779,683,735]" pageId="68" pageNumber="796">
<emphasis id="2DA52A616976FFBA04CF72BDBB3FFD09" box="[164,290,683,704]" italics="true" pageId="68" pageNumber="796">Etymology:</emphasis>
The generic name is from the Malagasy verb meaning to surprise, and is feminine in gender.
</paragraph>
<paragraph id="1F6EF6736976FFBA04CF731FBBC7FB03" blockId="68.[164,779,776,1226]" pageId="68" pageNumber="796">
<emphasis id="2DA52A616976FFBA04CF731FBAC2FCD7" box="[164,223,777,798]" italics="true" pageId="68" pageNumber="796">Note:</emphasis>
These large phyxelidids resemble
<taxonomicName id="D8D18DF06976FFBA06E5731EB917FCD4" authorityName="GRISWOLD" authorityYear="1990" box="[654,778,776,797]" class="Arachnida" family="Phyxelididae" genus="Ambohima" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA06E5731EB917FCD4" box="[654,778,776,797]" italics="true" pageId="68" pageNumber="796">Ambohima</emphasis>
</taxonomicName>
but possess a mosaic of derived characters. Like other Malagasy phyxelidids the male metatarsal claspers are prolateral (
<figureCitation id="87EAEAF66976FFBA050B7372BBF9FCB3" box="[352,484,868,890]" captionStart="Figure 44" captionStartId="47.[144,223,1215,1234]" captionTargetBox="[144,1424,196,1185]" captionTargetId="figure-183@47.[144,1424,196,1185]" captionTargetPageId="47" captionText="Figure 44. Photographs of male claspers of Manampoka and Rahavavy. A–C, Manampoka atsimo, holotype, CASENT9031164, right metatarsus II. A, dorsal. B, dorsal, close up. C, prolateral. D, E, Rahavavy fanivelona from Vohiparara, CASENT9016975, left metatarsus I. D, prolateral. E, prolateral, close up. F, G. Rahavavy malagasyana from Andasibe, CASENT9016182, right metatarsus I. F, dorsal. E, dorsal, close up. H, I, Rahavavy ida from Talatakely, CASENT9003423, right metatarsus I. H, prolateral, right, close up. I, dorsal, left, close up." figureDoi="http://doi.org/10.5281/zenodo.5407110" httpUri="https://zenodo.org/record/5407110/files/figure.png" pageId="68" pageNumber="796">Fig. 44A–C</figureCitation>
), not retrolateral, as in African and Asian phyxelidids. The presence of a clasper on male metatarsus II is a potential synapomorphy shared with
<taxonomicName id="D8D18DF06976FFBA05FF73D6B80DFC1C" authorityName="GRISWOLD" authorityYear="1990" box="[404,528,960,981]" class="Arachnida" family="Phyxelididae" genus="Ambohima" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA05FF73D6B80DFC1C" box="[404,528,960,981]" italics="true" pageId="68" pageNumber="796">Ambohima</emphasis>
</taxonomicName>
and the female vulva (
<figureCitation id="87EAEAF66976FFBA04C773C8BB12FC3D" box="[172,271,990,1012]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="68" pageNumber="796">Fig. 26H</figureCitation>
) with an anterodorsally projecting capsule containing an internal spiral chamber is a potential synapomorphy with
<taxonomicName id="D8D18DF06976FFBA05F4740DB80CFBF9" authorityName="Griswold &amp; Wood &amp; Carmichael" authorityYear="2012" box="[415,529,1051,1072]" class="Arachnida" family="Phyxelididae" genus="Rahavavy" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA05F4740DB80CFBF9" box="[415,529,1051,1072]" italics="true" pageId="68" pageNumber="796">Rahavavy</emphasis>
</taxonomicName>
. In some molecular phylogenetic analyses this species grouped with
<taxonomicName id="D8D18DF06976FFBA04CF744FBB0BFBA7" authorityName="Griswold &amp; Wood &amp; Carmichael" authorityYear="2012" box="[164,278,1113,1134]" class="Arachnida" family="Phyxelididae" genus="Rahavavy" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA04CF744FBB0BFBA7" box="[164,278,1113,1134]" italics="true" pageId="68" pageNumber="796">Rahavavy</emphasis>
</taxonomicName>
, but never with high support values, and it never grouped with
<taxonomicName id="D8D18DF06976FFBA05C37461B839FB45" authorityName="GRISWOLD" authorityYear="1990" box="[424,548,1143,1164]" class="Arachnida" family="Phyxelididae" genus="Ambohima" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA05C37461B839FB45" box="[424,548,1143,1164]" italics="true" pageId="68" pageNumber="796">Ambohima</emphasis>
</taxonomicName>
. As morphological data are equivocal, we propose a separate, new genus for this surprising species.
</paragraph>
<paragraph id="1F6EF6736976FFBA04CF74E2BB80F93A" blockId="68.[164,779,1267,1779]" pageId="68" pageNumber="796">
<emphasis id="2DA52A616976FFBA04CF74E2BB06FAC0" box="[164,283,1268,1289]" italics="true" pageId="68" pageNumber="796">Diagnosis:</emphasis>
Males have a unique combination of characters: the palpal bulb (
<figureCitation id="87EAEAF66976FFBA05A67504B849FAEE" box="[461,596,1298,1320]" captionStart="Figure 17" captionStartId="19.[144,223,1433,1452]" captionTargetBox="[369,1198,197,1398]" captionTargetId="figure-137@19.[367,1200,195,1403]" captionTargetPageId="19" captionText="Figure 17. Left male palpus of Manampoka atsimo, holotype from Manangotry, CASENT9031164, tibia and tarsus. A, prolateral. B, ventral. C, retrolateral. Illustrations by GM." figureDoi="http://doi.org/10.5281/zenodo.5407056" httpUri="https://zenodo.org/record/5407056/files/figure.png" pageId="68" pageNumber="796">Figs 17A–C</figureCitation>
,
<figureCitation id="87EAEAF66976FFBA060D7504B8AFFAE1" box="[614,690,1298,1320]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="68" pageNumber="796">26A–F</figureCitation>
) has a small median apophysis (like
<taxonomicName id="D8D18DF06976FFBA067B7526B89FFA8C" authorityName="Griswold &amp; Wood &amp; Carmichael" authorityYear="2012" box="[528,642,1328,1349]" class="Arachnida" family="Phyxelididae" genus="Rahavavy" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA067B7526B89FFA8C" box="[528,642,1328,1349]" italics="true" pageId="68" pageNumber="796">Rahavavy</emphasis>
</taxonomicName>
but unlike
<taxonomicName id="D8D18DF06976FFBA04CF7559BB3DFAAD" authorityName="GRISWOLD" authorityYear="1990" box="[164,288,1359,1380]" class="Arachnida" family="Phyxelididae" genus="Ambohima" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA04CF7559BB3DFAAD" box="[164,288,1359,1380]" italics="true" pageId="68" pageNumber="796">Ambohima</emphasis>
</taxonomicName>
) and male metatarsi II (
<figureCitation id="87EAEAF66976FFBA06267559B8D3FAAC" box="[589,718,1359,1381]" captionStart="Figure 44" captionStartId="47.[144,223,1215,1234]" captionTargetBox="[144,1424,196,1185]" captionTargetId="figure-183@47.[144,1424,196,1185]" captionTargetPageId="47" captionText="Figure 44. Photographs of male claspers of Manampoka and Rahavavy. A–C, Manampoka atsimo, holotype, CASENT9031164, right metatarsus II. A, dorsal. B, dorsal, close up. C, prolateral. D, E, Rahavavy fanivelona from Vohiparara, CASENT9016975, left metatarsus I. D, prolateral. E, prolateral, close up. F, G. Rahavavy malagasyana from Andasibe, CASENT9016182, right metatarsus I. F, dorsal. E, dorsal, close up. H, I, Rahavavy ida from Talatakely, CASENT9003423, right metatarsus I. H, prolateral, right, close up. I, dorsal, left, close up." figureDoi="http://doi.org/10.5281/zenodo.5407110" httpUri="https://zenodo.org/record/5407110/files/figure.png" pageId="68" pageNumber="796">Fig. 44A–C</figureCitation>
) and probably I (legs I are missing from the unique male) have a prolateral, spinule-lined concavity at midsegment, with a short, prolateral process surmounted by a stout clasping spine at the base of this concavity (like
<taxonomicName id="D8D18DF06976FFBA048F75FEBB7DFA34" authorityName="GRISWOLD" authorityYear="1990" box="[228,352,1512,1533]" class="Arachnida" family="Phyxelididae" genus="Ambohima" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA048F75FEBB7DFA34" box="[228,352,1512,1533]" italics="true" pageId="68" pageNumber="796">Ambohima</emphasis>
</taxonomicName>
but unlike
<taxonomicName id="D8D18DF06976FFBA059C75FEB874FA34" authorityName="Griswold &amp; Wood &amp; Carmichael" authorityYear="2012" box="[503,617,1512,1533]" class="Arachnida" family="Phyxelididae" genus="Rahavavy" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA059C75FEB874FA34" box="[503,617,1512,1533]" italics="true" pageId="68" pageNumber="796">Rahavavy</emphasis>
</taxonomicName>
). The female vulva (
<figureCitation id="87EAEAF66976FFBA04927611BB41F9D4" box="[249,348,1543,1565]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="68" pageNumber="796">Fig. 26H</figureCitation>
) is like that of most
<taxonomicName id="D8D18DF06976FFBA061F7611B8FBF9D5" authorityName="Griswold &amp; Wood &amp; Carmichael" authorityYear="2012" box="[628,742,1543,1564]" class="Arachnida" family="Phyxelididae" genus="Rahavavy" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA061F7611B8FBF9D5" box="[628,742,1543,1564]" italics="true" pageId="68" pageNumber="796">Rahavavy</emphasis>
</taxonomicName>
in having an anterodorsally projecting capsule containing an internal spiralled chamber. The epigynum (
<figureCitation id="87EAEAF66976FFBA04C77674BB12F9B1" box="[172,271,1634,1656]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="68" pageNumber="796">Fig. 26G</figureCitation>
) differs from
<taxonomicName id="D8D18DF06976FFBA05C27674B838F9BE" authorityName="GRISWOLD" authorityYear="1990" box="[425,549,1634,1655]" class="Arachnida" family="Phyxelididae" genus="Ambohima" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA05C27674B838F9BE" box="[425,549,1634,1655]" italics="true" pageId="68" pageNumber="796">Ambohima</emphasis>
</taxonomicName>
in that the anterior margins of the median lobe are transverse rather than oblique, and from
<taxonomicName id="D8D18DF06976FFBA05DA76B6B83EF97C" authorityName="Griswold &amp; Wood &amp; Carmichael" authorityYear="2012" box="[433,547,1696,1717]" class="Arachnida" family="Phyxelididae" genus="Rahavavy" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA05DA76B6B83EF97C" box="[433,547,1696,1717]" italics="true" pageId="68" pageNumber="796">Rahavavy</emphasis>
</taxonomicName>
in that the external copulatory openings are large and the median lobe is prolonged posteriorly.
</paragraph>
<paragraph id="1F6EF6736976FFBA04CF770AB917F8A6" blockId="68.[164,779,1820,1903]" pageId="68" pageNumber="796">
<emphasis id="2DA52A616976FFBA04CF770ABB7AF8F8" box="[164,359,1820,1841]" italics="true" pageId="68" pageNumber="796">Synapomorphies:</emphasis>
The cuspule-like median apophysis on the male palpal bulb (
<figureCitation id="87EAEAF66976FFBA05A1772CB84EF886" box="[458,595,1850,1872]" captionStart="Figure 17" captionStartId="19.[144,223,1433,1452]" captionTargetBox="[369,1198,197,1398]" captionTargetId="figure-137@19.[367,1200,195,1403]" captionTargetPageId="19" captionText="Figure 17. Left male palpus of Manampoka atsimo, holotype from Manangotry, CASENT9031164, tibia and tarsus. A, prolateral. B, ventral. C, retrolateral. Illustrations by GM." figureDoi="http://doi.org/10.5281/zenodo.5407056" httpUri="https://zenodo.org/record/5407056/files/figure.png" pageId="68" pageNumber="796">Figs 17A, C</figureCitation>
,
<figureCitation id="87EAEAF66976FFBA060A772CB893F899" box="[609,654,1850,1872]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="68" pageNumber="796">26B</figureCitation>
) and male palpal tibia with a retrolateral lobe of the apex (
<collectionCode id="79C06EB66976FFBA06B4774FB91EF8A6" box="[735,771,1881,1903]" pageId="68" pageNumber="796">RL</collectionCode>
)
</paragraph>
<paragraph id="1F6EF6736976FFBA075170D0BF7AFF33" blockId="68.[826,1441,198,250]" pageId="68" pageNumber="796">
that is longer than the dorsal tibial apophysis (
<collectionCode id="79C06EB66976FFBA010870D0BF87FF15" box="[1379,1434,198,220]" pageId="68" pageNumber="796">DTA</collectionCode>
) (
<figureCitation id="87EAEAF66976FFBA072970F2B9B8FF33" box="[834,933,228,250]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="68" pageNumber="796">Fig. 26D</figureCitation>
) are synapomorphies for
<taxonomicName id="D8D18DF06976FFBA00BA70F2BF7DFF30" authorityName="Griswold &amp; Wood &amp; Carmichael" authorityYear="2012" box="[1233,1376,228,249]" class="Arachnida" family="Phyxelididae" genus="Manampoka" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="796" phylum="Arthropoda" rank="genus">
<emphasis id="2DA52A616976FFBA00BA70F2BF7DFF30" box="[1233,1376,228,249]" italics="true" pageId="68" pageNumber="796">Manampoka</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1F6EF6736976FFBB07517134BAE6FDC7" blockId="68.[826,1441,289,1903]" lastBlockId="69.[144,759,198,526]" lastPageId="69" lastPageNumber="797" pageId="68" pageNumber="796">
<emphasis id="2DA52A616976FFBA07517134B9DFFEFE" box="[826,962,290,311]" italics="true" pageId="68" pageNumber="796">Description:</emphasis>
Total length 5.40–7.00. Carapace length 1.30–1.38 times width, height 0.30–0.49 width, thoracic fovea length twice width, length 0.06–0.20 carapace length; PER straight to slightly recurved, AER straight, ocular area width 2.50–3.10 times length; clypeal height 1.40–2.20 times
<collectionCode id="79C06EB66976FFBA00D971ADBEC6FE19" box="[1202,1243,443,464]" country="Australia" name="Australian Museum" pageId="68" pageNumber="796" type="Museum">AM</collectionCode>
diameter; chelicerae smooth, length 4.60–5.50 times clypeal height, with small basal boss, pro- and retromargins of fang furrow with 5–7 heterogeneous teeth; sternum length 1.02–1.36 times width, posterior apex a blunt point; labium rectangular. Legs elongate, female femur I length 1.70–2.00 times carapace length, male femur II 2.40 times carapace length; leg formula 1423; male metatarsi II (
<figureCitation id="87EAEAF66976FFBA078F72B9BE78FD0F" box="[996,1125,687,710]" captionStart="Figure 44" captionStartId="47.[144,223,1215,1234]" captionTargetBox="[144,1424,196,1185]" captionTargetId="figure-183@47.[144,1424,196,1185]" captionTargetPageId="47" captionText="Figure 44. Photographs of male claspers of Manampoka and Rahavavy. A–C, Manampoka atsimo, holotype, CASENT9031164, right metatarsus II. A, dorsal. B, dorsal, close up. C, prolateral. D, E, Rahavavy fanivelona from Vohiparara, CASENT9016975, left metatarsus I. D, prolateral. E, prolateral, close up. F, G. Rahavavy malagasyana from Andasibe, CASENT9016182, right metatarsus I. F, dorsal. E, dorsal, close up. H, I, Rahavavy ida from Talatakely, CASENT9003423, right metatarsus I. H, prolateral, right, close up. I, dorsal, left, close up." figureDoi="http://doi.org/10.5281/zenodo.5407110" httpUri="https://zenodo.org/record/5407110/files/figure.png" pageId="68" pageNumber="796">Fig. 44A–C</figureCitation>
) (and probably I) having prolateral, spinule-lined concavity at midsegment, at base of concavity a short, prolateral process surmounted by stout clasping spine, legs otherwise unmodified; metatarsi lacking apical combs; calamistrum subapical, origin at 0.45 from metatarsus base, length 0.55 that of segment. Male spination: palp: femur d0-0-0-1; leg I: (unknown); leg II: femur d1-1- 0-0, p0-1-1-1, r0-1-1–1, tibia p0-1-1-0, r0-1-1-1, metatarsus d1-0-0-0, p0-1(clasper)-0–1, v2-0-0-1, r0-0-0-1; leg
<collectionCode id="79C06EB66976FFBA070173F4B996FC3E" box="[874,907,994,1015]" pageId="68" pageNumber="796">III</collectionCode>
: femur d1-1-0-0, p0-0-1–1, r0-1-1–1, patella d0-1, tibia d1-0-0-0, p0-1-1-0, r0-0-0-1, metatarsus d1-0-0-0, p0-1-0-1, v1-2-0-2; leg
<collectionCode id="79C06EB66976FFBA00D67409BEC6FBFD" box="[1213,1243,1055,1076]" pageId="68" pageNumber="796">IV</collectionCode>
: femur d1-1-0-0, r0-0-0-1, tibia d1-1-1-0, p0-0-1-0, v0-0-0-1, metatarsus d1-1-0-0, p0-0-0-1, r0-0-0-1, v1-0-1–1. Female spination: palp: femur d0-0-1–1, patella d0-0-1, tibia d0-0- 0-1, p0-1-1-0, tarsus p1-0-2–2-3, v0-0-0-1-1-1-1-1-1, r1-0-0-1-0; leg I: femur d1-0-0-0, p0-1-1-1, r0-0-1–1, tibia p0-1-1-0, v0-0-2-0, r0-1-1-0, metatarsus p1-0-0-1, v1-0-2-2, r1-0-0-0; leg II: femur d1-1-1-0, p0-1-1–1, r0-1-1–1-1, tibia d1-0-0-0, p0-1-1-0, v0-1-0-2, r0-1-1-0, metatarsus p1-0-0-1, v0-0-2–2, r0-1-0-0; leg
<collectionCode id="79C06EB66976FFBA015B7524BF4CFA8E" box="[1328,1361,1330,1351]" pageId="68" pageNumber="796">III</collectionCode>
: femur d1-1-0-0, p0-0-0-1, r0-0-0-1, tibia d1-0-0-0, p0-1-1-0, v0-1-0-2, r0-1-1-0, metatarsus p0-1-1–1, v2-0-2–2, r0-1-1-0; leg
<collectionCode id="79C06EB66976FFBA07B97598B9EDFA6A" box="[978,1008,1422,1443]" pageId="68" pageNumber="796">IV</collectionCode>
: femur d1-1-1-0, r0-0-0-1, tibia p0-1- 1-0, v0-0-0-2, r0-1-1-0, metatarsus p0-1-0-0, v1-2-1–2. Tracheae not examined. Spinnerets not examined with
<collectionCode id="79C06EB66976FFBA071E75FCB9B3FA36" box="[885,942,1514,1535]" pageId="68" pageNumber="796">SEM</collectionCode>
but light microscopic examination suggests that these are typical of
<taxonomicName id="D8D18DF06976FFBA0016761EBF1FF9D7" box="[1149,1282,1544,1566]" class="Arachnida" family="Phyxelididae" kingdom="Animalia" order="Araneae" pageId="68" pageNumber="742" phylum="Arthropoda" rank="tribe" tribe="Phyxelidini">Phyxelidini</taxonomicName>
, e.g. divided cribellum, an encircling row of paracribellar spigots on the
<collectionCode id="79C06EB66976FFBA07E57653B9D5F993" box="[910,968,1605,1626]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="68" pageNumber="796">PMS</collectionCode>
, and a black, stout seta retroapically on the PLS. Palpal femur with anterobasal group of slender setae set in enlarged bases (
<figureCitation id="87EAEAF66976FFBA009B7694BF4FF951" box="[1264,1362,1666,1688]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="68" pageNumber="796">Fig. 26F</figureCitation>
); male palpal tibia (
<figureCitation id="87EAEAF66976FFBA078876B7BE6DF97E" box="[995,1136,1697,1719]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="68" pageNumber="796">Fig. 26D, E</figureCitation>
) with dorsoapical blade (
<collectionCode id="79C06EB66976FFBA072A76D6B965F91F" box="[833,888,1728,1750]" pageId="68" pageNumber="796">DTA</collectionCode>
) and retroapical lobe (
<collectionCode id="79C06EB66976FFBA00A976D6BEF5F91F" box="[1218,1256,1728,1750]" pageId="68" pageNumber="796">RL</collectionCode>
); palpal bulb (
<figureCitation id="87EAEAF66976FFBA072976C8B9D4F93A" box="[834,969,1758,1780]" captionStart="Figure 17" captionStartId="19.[144,223,1433,1452]" captionTargetBox="[369,1198,197,1398]" captionTargetId="figure-137@19.[367,1200,195,1403]" captionTargetPageId="19" captionText="Figure 17. Left male palpus of Manampoka atsimo, holotype from Manangotry, CASENT9031164, tibia and tarsus. A, prolateral. B, ventral. C, retrolateral. Illustrations by GM." figureDoi="http://doi.org/10.5281/zenodo.5407056" httpUri="https://zenodo.org/record/5407056/files/figure.png" pageId="68" pageNumber="796">Figs 17A–C</figureCitation>
,
<figureCitation id="87EAEAF66976FFBA07B576C8BE36F93D" box="[990,1067,1758,1780]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="68" pageNumber="796">26A–C</figureCitation>
) length twice width, embolus base retromedian, reservoir course in embolus base straight except initial loop, embolus a slender threadlike spine, lamellar pars pendula slender, conductor greatly hypertrophied, with embolic groove completely encircling bulb, apex pointing retrolaterally; median apophysis a small, pointed cuspule, arising retromedially near embolus base. Epigynum (
<figureCitation id="87EAEAF66977FFBB04F37134BAE7FEF1" box="[152,250,290,312]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="69" pageNumber="797">Fig. 26G</figureCitation>
) with broad, semicircular median lobe (
<collectionCode id="79C06EB66977FFBB06D47134B8F6FEF1" box="[703,747,290,312]" country="France" name="Musee de Lectoure" pageId="69" pageNumber="797">ML</collectionCode>
), anterior margins transverse, copulatory openings large; vulva (
<figureCitation id="87EAEAF66977FFBB052B7149BBD5FEBD" box="[320,456,351,373]" captionStart="Figure 26" captionStartId="28.[164,243,1356,1375]" captionTargetBox="[323,1284,196,1326]" captionTargetId="figure-151@28.[323,1284,196,1326]" captionTargetPageId="28" captionText="Figure 26. Manampoka atsimo, from Manangotry, CASENT9031164. A–F, right male palpus of holotype. G–I, vulva of female. A, retrolateral. B, ventral. C, prolateral. D, tibia, dorsal. E, apex of tibia, retrolateral. F, femur, prolateral. G, epigynum, ventral. H, spermathecae, dorsal. I, poreplate on left spermatheca, dorsal. DTA: dorsal tibial apophysis, RL: retrolateral lobe of tibial apex, SP: spermathecal poreplate." figureDoi="http://doi.org/10.5281/zenodo.5407074" httpUri="https://zenodo.org/record/5407074/files/figure.png" pageId="69" pageNumber="797">Fig. 26H, I</figureCitation>
) with copulatory ducts arising mesally, extending laterally to enter vulval capsule posteriorly, large poreplate posteriorly on vulval capsule, anterodorsally projecting capsule containing internal spiral chamber, fertilization duct posterior.
</paragraph>
<paragraph id="1F6EF6736977FFBB04FB7221BBA3FD85" blockId="69.[144,446,566,588]" box="[144,446,566,588]" pageId="69" pageNumber="797">
<emphasis id="2DA52A616977FFBB04FB7221BB39FD85" box="[144,292,567,588]" italics="true" pageId="69" pageNumber="797">Composition:</emphasis>
One species.
</paragraph>
<paragraph id="1F6EF6736977FFBB04FB7263BAEBFD01" blockId="69.[144,759,629,712]" pageId="69" pageNumber="797">
<emphasis id="2DA52A616977FFBB04FB7263BB3CFD43" box="[144,289,629,650]" italics="true" pageId="69" pageNumber="797">Distribution:</emphasis>
Known only from the
<typeStatus id="C06A48D16977FFBB062B7263B86CFD43" box="[576,625,629,650]" pageId="69" pageNumber="797">type</typeStatus>
locality in Parc National Andohahela, southern
<collectingCountry id="67C6B6E36977FFBB06077285B8EAFD60" box="[620,759,659,681]" name="Madagascar" pageId="69" pageNumber="797">Madagascar</collectingCountry>
(
<figureCitation id="87EAEAF66977FFBB04F372A4BAF4FD01" box="[152,233,690,712]" captionStart="Figure 59" captionStartId="61.[144,223,1092,1111]" captionTargetBox="[147,757,200,1058]" captionTargetId="figure-668@61.[144,760,197,1062]" captionTargetPageId="61" captionText="Figure 59. Distribution of Madagascar Phyxelididae." figureDoi="http://doi.org/10.5281/zenodo.5407138" httpUri="https://zenodo.org/record/5407138/files/figure.png" pageId="69" pageNumber="797">Fig. 59</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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