136 lines
19 KiB
XML
136 lines
19 KiB
XML
<document id="1EC59A5D166F6D69020280DC0AF2D3A0" ID-DOI="10.5281/zenodo.179369" ID-GBIF-Dataset="f43b5459-3507-44de-ba79-433bad90c141" ID-ISSN="1175-5326" ID-Zenodo-Dep="179369" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459967351429" checkinUser="plazi" docAuthor="Huang, Jiandong, Ren, Dong & Shih, Chungkun" docDate="2007" docId="386E87B3664C9C09FF5F578B010FF807" docLanguage="en" docName="zt01629p050.pdf" docOrigin="Zootaxa 1629" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Epicharmeropsis hexavenulosus Huang, Ren & Shih, sp. nov." docType="treatment" docVersion="9" lastPageNumber="41" masterDocId="C457FFCB664E9C0BFFC8571C052DFFC4" masterDocTitle="New genus and species of Hexagenitidae (Insecta: Ephemeroptera) from Yixian Formation, China" masterLastPageNumber="50" masterPageNumber="39" pageNumber="41" updateTime="1698231180412" updateUser="plazi">
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<mods:mods id="383FA47E1EC9047EB99F1EE0A0DF2C53" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A5F8235F944219124FB7104ECDA17E99">
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<mods:title id="5FCC589AD8D0B03462AD9EEA2CDA92C0">New genus and species of Hexagenitidae (Insecta: Ephemeroptera) from Yixian Formation, China</mods:title>
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</mods:titleInfo>
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<mods:name id="9223AC6984DE1FE15A6074FBED618627" type="personal">
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<mods:roleTerm id="E60D7B26CC87CACC956CF303B252926E">Author</mods:roleTerm>
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<mods:namePart id="F301DE9437198457B73C6D9F08AE279D">Huang, Jiandong</mods:namePart>
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<mods:roleTerm id="010CEBAB286F9AC044F21A6168777759">Author</mods:roleTerm>
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<mods:namePart id="31A50F933E9B0AB3A8DE76F087B0E5C5">Ren, Dong</mods:namePart>
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<mods:name id="1FAC837A163E5D51F790437057B5BD52" type="personal">
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<mods:roleTerm id="4DFA47729FBECF6E1FAAA9248B4626D3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="48077983274FE13C3CACACFF0ADA3DFA">Shih, Chungkun</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0548892B38C3B8C0BC71C4A60C3B676C">text</mods:typeOfResource>
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<mods:titleInfo id="27A6FC001CE4DC8494CBAC2BEC8FA7E6">
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<mods:title id="45ED49A0AB6C9378C1A998A70FF74479">Zootaxa</mods:title>
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<mods:part id="3872727B3FD2808E3448F9ED63AA170F">
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<mods:date id="FB68C286646A568B4F36A86D0D539680">2007</mods:date>
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<mods:detail id="1F42F971422411C4439FA92F42878A2C" type="volume">
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<mods:number id="4A73D44787D0839CA0424C720EB1CA34">1629</mods:number>
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<mods:start id="0FD9EFC0E75715B76A4D68F0B11495ED">39</mods:start>
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<mods:classification id="A4DD6DC779670CB56EC5B889A29A3F19">journal article</mods:classification>
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<mods:identifier id="B4DB9BF672337749B87E0D51CB3A142D" type="DOI">10.5281/zenodo.179369</mods:identifier>
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<mods:identifier id="BFAA60B4CF22C86ECC34A37B68FD5C8F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6A7B1434379A5DC4CADCDFA48236DD55" type="Zenodo-Dep">179369</mods:identifier>
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<treatment id="386E87B3664C9C09FF5F578B010FF807" ID-DOI="http://doi.org/10.5281/zenodo.6249046" ID-GBIF-Taxon="119377461" ID-Zenodo-Dep="6249046" LSID="urn:lsid:plazi:treatment:386E87B3664C9C09FF5F578B010FF807" httpUri="http://treatment.plazi.org/id/386E87B3664C9C09FF5F578B010FF807" lastPageNumber="41" pageId="2" pageNumber="41">
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<subSubSection id="F8DD652E664C9C09FF5F578B0439FF10" pageId="2" pageNumber="41" type="nomenclature">
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<paragraph id="B07836A5664C9C09FF5F578B06AAFF75" blockId="2.[151,903,151,212]" box="[151,903,151,177]" pageId="2" pageNumber="41">
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<heading id="EB3081C9664C9C09FF5F578B06AAFF75" bold="true" box="[151,903,151,177]" fontSize="11" level="1" pageId="2" pageNumber="41" reason="1">
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<emphasis id="82B3EAB7664C9C09FF5F578B06AAFF75" bold="true" box="[151,903,151,177]" pageId="2" pageNumber="41">
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<taxonomicName id="77C74D26664C9C09FF5F578B0633FF75" authority="Huang, Ren & Shih" authorityName="Huang, Ren & Shih" box="[151,798,151,177]" class="Insecta" family="Hexagenitidae" genus="Epicharmeropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="hexavenulosus" status="sp. nov.">
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<emphasis id="82B3EAB7664C9C09FF5F578B0733FF75" bold="true" box="[151,542,151,177]" italics="true" pageId="2" pageNumber="41">Epicharmeropsis hexavenulosus</emphasis>
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Huang, Ren & Shih
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</taxonomicName>
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,
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<taxonomicNameLabel id="998057CC664C9C09FCE2578B06AAFF75" box="[810,903,151,177]" pageId="2" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="B07836A5664C9C09FF5F57A60439FF10" blockId="2.[151,903,151,212]" box="[151,276,186,212]" pageId="2" pageNumber="41">
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(
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<figureCitation id="28FC2A20664C9C09FF5757A60428FF10" box="[159,261,186,212]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,255,1665,1689]" captionStartId-1="4.[151,255,1944,1968]" captionStartId-2="5.[151,255,1909,1933]" captionStartId-3="6.[151,255,1952,1976]" captionTargetBox-0="[378,1210,197,1638]" captionTargetBox-1="[378,1210,880,1919]" captionTargetBox-2="[257,1323,196,1857]" captionTargetBox-3="[270,1330,203,1914]" captionTargetId-0="figure@3.[378,1210,197,1638]" captionTargetId-1="figure@4.[378,1210,880,1921]" captionTargetId-2="figure@5.[238,1354,191,1895]" captionTargetId-3="figure@6.[235,1354,194,1928]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Epicharmeropsis hexavenulosu s gen. et sp. nov., male imago, holotype: A, photograph of part CNU-E-YX- 200701 - 1; B, photograph of counterpart CNU-E-YX- 200701 - 2." captionText-1="FIGURE 2. Epicharmeropsis hexavenulosu s gen. et sp. nov., male imago, holotype CNU-E-YX- 200701 - 1: A, photograph of head and thorax in dorsal view; B, photograph of cubital area on forewing. Without scale." captionText-2="FIGURE 3. Epicharmeropsis hexavenulosu s sp. nov., male imago, holotype: A, body with wings, CNU-E-YX- 200701 - 1; B, forewing, CNU-E-YX- 200701 - 2.; C, hind wing, CNU-E-YX- 200701 - 2. MNs – mesonotal suture, MPs - medioparapsidal suture, LPs – lateroparapsidal suture." captionText-3="FIGURE 4. Epicharmeropsis hexavenulosu s sp. nov., female imago, paratype CNU-E-YX- 2007004: A, photograph of paratype; B, body with wings on lateral view; C, forewing." httpUri-0="https://zenodo.org/record/179370/files/figure.png" httpUri-1="https://zenodo.org/record/179371/files/figure.png" httpUri-2="https://zenodo.org/record/179372/files/figure.png" httpUri-3="https://zenodo.org/record/179373/files/figure.png" pageId="2" pageNumber="41">Fig. 1–4</figureCitation>
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.)
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</paragraph>
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</subSubSection>
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<subSubSection id="F8DD652E664C9C09FF5F56160677FEE0" box="[151,858,266,292]" pageId="2" pageNumber="41" type="etymology">
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<paragraph id="B07836A5664C9C09FF5F56160677FEE0" blockId="2.[151,1437,266,1987]" box="[151,858,266,292]" pageId="2" pageNumber="41">
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<emphasis id="82B3EAB7664C9C09FF5F56160409FEE0" bold="true" box="[151,292,266,292]" pageId="2" pageNumber="41">Etymology.</emphasis>
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Latin prefix
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<emphasis id="82B3EAB7664C9C09FE75561604D3FEE7" box="[445,510,266,291]" italics="true" pageId="2" pageNumber="41">hexa-</emphasis>
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(six) and
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<emphasis id="82B3EAB7664C9C09FDBC561607CAFEE7" box="[628,743,266,291]" italics="true" pageId="2" pageNumber="41">venulosus</emphasis>
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(veinlet).
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</paragraph>
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</subSubSection>
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<subSubSection id="F8DD652E664C9C09FF0F562E0441FDA0" pageId="2" pageNumber="41" type="materials_examined">
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<paragraph id="B07836A5664C9C09FF0F562E05C8FE00" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">
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<emphasis id="82B3EAB7664C9C09FF0F562E046DFE88" bold="true" box="[199,320,306,332]" pageId="2" pageNumber="41">
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<typeStatus id="6F7C8807664C9C09FF0F562E0416FE88" box="[199,315,306,332]" pageId="2" pageNumber="41" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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CNU-E-YX-2007001-1, CNU-E-YX-2007001-2 (
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<figureCitation id="28FC2A20664C9C09FC50562E06D3FE88" box="[920,1022,306,332]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,255,1665,1689]" captionStartId-1="4.[151,255,1944,1968]" captionStartId-2="5.[151,255,1909,1933]" captionTargetBox-0="[378,1210,197,1638]" captionTargetBox-1="[378,1210,880,1919]" captionTargetBox-2="[257,1323,196,1857]" captionTargetId-0="figure@3.[378,1210,197,1638]" captionTargetId-1="figure@4.[378,1210,880,1921]" captionTargetId-2="figure@5.[238,1354,191,1895]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Epicharmeropsis hexavenulosu s gen. et sp. nov., male imago, holotype: A, photograph of part CNU-E-YX- 200701 - 1; B, photograph of counterpart CNU-E-YX- 200701 - 2." captionText-1="FIGURE 2. Epicharmeropsis hexavenulosu s gen. et sp. nov., male imago, holotype CNU-E-YX- 200701 - 1: A, photograph of head and thorax in dorsal view; B, photograph of cubital area on forewing. Without scale." captionText-2="FIGURE 3. Epicharmeropsis hexavenulosu s sp. nov., male imago, holotype: A, body with wings, CNU-E-YX- 200701 - 1; B, forewing, CNU-E-YX- 200701 - 2.; C, hind wing, CNU-E-YX- 200701 - 2. MNs – mesonotal suture, MPs - medioparapsidal suture, LPs – lateroparapsidal suture." httpUri-0="https://zenodo.org/record/179370/files/figure.png" httpUri-1="https://zenodo.org/record/179371/files/figure.png" httpUri-2="https://zenodo.org/record/179372/files/figure.png" pageId="2" pageNumber="41">Fig. 1–3</figureCitation>
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). Well preserved part and counterpart of imago; antennae and tarsi of middle and hind legs are not preserved; the Late Jurassic to the Early Cretaceous, Yixian Formation, Shimen Village, Yangshuling Township, Pingquan County, Hebei Province,
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<collectingCountry id="C8D07635664C9C09FF5F56B605CCFE00" box="[151,225,426,452]" name="China" pageId="2" pageNumber="41">China</collectingCountry>
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.
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</paragraph>
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<paragraph id="B07836A5664C9C09FF0F56CE0441FDA0" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">
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<emphasis id="82B3EAB7664C9C09FF0F56CE0416FE28" bold="true" box="[199,315,466,492]" pageId="2" pageNumber="41">Material.</emphasis>
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<typeStatus id="6F7C8807664C9C09FE8B56CE049DFE28" box="[323,432,466,492]" pageId="2" pageNumber="41" type="holotype">Holotype</typeStatus>
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and from the same locality,
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<typeStatus id="6F7C8807664C9C09FD3756CE0659FE28" box="[767,884,466,492]" pageId="2" pageNumber="41" type="paratype">paratypes</typeStatus>
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: male imago, No. CNU-E-YX-2007021-1, No. CNU-E-YX-2007021-2; male imago, No. CNU-E-YX-2007023; male imago, No. CNU-E-YX-2007024; male imago, No. CNU-E-YX-2007025-1, No. CNU-E-YX-2007025-2; female imago, No. CNU-E-YX- 2007004 (
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<figureCitation id="28FC2A20664C9C09FEC55556047AFDA0" box="[269,343,586,612]" captionStart="FIGURE 4" captionStartId="6.[151,255,1952,1976]" captionTargetBox="[270,1330,203,1914]" captionTargetId="figure@6.[235,1354,194,1928]" captionTargetPageId="6" captionText="FIGURE 4. Epicharmeropsis hexavenulosu s sp. nov., female imago, paratype CNU-E-YX- 2007004: A, photograph of paratype; B, body with wings on lateral view; C, forewing." httpUri="https://zenodo.org/record/179373/files/figure.png" pageId="2" pageNumber="41">Fig. 4</figureCitation>
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.).
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</paragraph>
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</subSubSection>
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<subSubSection id="F8DD652E664C9C09FF0F556E06CBFD73" pageId="2" pageNumber="41" type="diagnosis">
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<paragraph id="B07836A5664C9C09FF0F556E06CBFD73" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">
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<emphasis id="82B3EAB7664C9C09FF0F556E046AFD48" bold="true" box="[199,327,626,652]" pageId="2" pageNumber="41">Diagnosis.</emphasis>
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In addition to the diagnosis for the new genus, the species can be diagnosed as follows: CuA1 of forewing with 6 triads (loop-shaped veinlets) leading to wing margin.
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</paragraph>
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</subSubSection>
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<subSubSection id="F8DD652E664C9C09FF0F55D9010FF807" pageId="2" pageNumber="41" type="description">
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<paragraph id="B07836A5664C9C09FF0F55D9060CFD1B" blockId="2.[151,1437,266,1987]" box="[199,801,709,735]" pageId="2" pageNumber="41">
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<emphasis id="82B3EAB7664C9C09FF0F55D90472FD1B" bold="true" box="[199,351,709,735]" pageId="2" pageNumber="41">Description.</emphasis>
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Male imago (dorsal view) (
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<figureCitation id="28FC2A20664C9C09FD6D55D90621FD1B" box="[677,780,709,735]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,255,1665,1689]" captionStartId-1="4.[151,255,1944,1968]" captionStartId-2="5.[151,255,1909,1933]" captionTargetBox-0="[378,1210,197,1638]" captionTargetBox-1="[378,1210,880,1919]" captionTargetBox-2="[257,1323,196,1857]" captionTargetId-0="figure@3.[378,1210,197,1638]" captionTargetId-1="figure@4.[378,1210,880,1921]" captionTargetId-2="figure@5.[238,1354,191,1895]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Epicharmeropsis hexavenulosu s gen. et sp. nov., male imago, holotype: A, photograph of part CNU-E-YX- 200701 - 1; B, photograph of counterpart CNU-E-YX- 200701 - 2." captionText-1="FIGURE 2. Epicharmeropsis hexavenulosu s gen. et sp. nov., male imago, holotype CNU-E-YX- 200701 - 1: A, photograph of head and thorax in dorsal view; B, photograph of cubital area on forewing. Without scale." captionText-2="FIGURE 3. Epicharmeropsis hexavenulosu s sp. nov., male imago, holotype: A, body with wings, CNU-E-YX- 200701 - 1; B, forewing, CNU-E-YX- 200701 - 2.; C, hind wing, CNU-E-YX- 200701 - 2. MNs – mesonotal suture, MPs - medioparapsidal suture, LPs – lateroparapsidal suture." httpUri-0="https://zenodo.org/record/179370/files/figure.png" httpUri-1="https://zenodo.org/record/179371/files/figure.png" httpUri-2="https://zenodo.org/record/179372/files/figure.png" pageId="2" pageNumber="41">Fig. 1–3</figureCitation>
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.).
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</paragraph>
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<paragraph id="B07836A5664C9C09FF0F55F10186FC93" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">Eyes relatively large, contiguous. Pterothorax well-developed; mesonotal suture (MNs) in anterior part of mesonotum strongly stretched backwards medially, not transverse; paired medioparapsidal sutures (MPs) jointed at middle area of mesonotum, not parallel; metanotum long, with evident scutum.</paragraph>
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<paragraph id="B07836A5664C9C09FF0F547901C4FB64" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">Forewing: broad and triangular with costal brace; costal margin slightly curved; tornoapical (outer) margin longer than basitornal (anal) margin; costal area relatively wide, narrowing toward tip; RA subparallel to SC; membrane slightly thickened at distal part of the field between C and RA; RSa formed two triads, RSp non-branched; MA branched after middle area of wing; MA1 and MA2 uniformly diverging; iMA approximated MA2; MP1 and MP2 diverged at an angle of approximately 30 degree; distinct intercalary veins existing between MP2 and CuA1; CuA bifurcated to CuA1 and CuA2, from the bifurcation a vein iCu formed 6 triads following one another: anterior branch of each triad forms next triad; each of these triads has anterior branch arched by its convexity anteriorly; all branches of these triads go to basitornal margin of wing.</paragraph>
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<paragraph id="B07836A5664C9C09FF0F53B20649FAAF" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">
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<collectingCountry id="C8D07635664C9C09FF0F53B2042FFB0C" box="[199,258,1198,1224]" name="India" pageId="2" pageNumber="41">Hind</collectingCountry>
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wing: broad with fairly obtuse tip; venation relatively profuse; the base of C strongly arched, far from SC; subcostal area broad, more than 3 times as wide as costal area; RA terminated near wing apex; RS formed triad RSa-iRS-RSp, RSa and RSp non-branched; MA bifurcated near middle; MP bifurcated near the base of wing; iMP approximated with MP1; CuA and CuP un-forked; more than 4 A present; anal area broad; numerous crossveins and intercalary veins near wing margin.
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</paragraph>
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<paragraph id="B07836A5664C9C09FF0F52650609FA7F" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">Legs (preserved): relatively sturdy and long; femora longer than tibia; three-segmented tarsi of fore legs preserved, slender; tarsi of middle and hind legs absent.</paragraph>
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<paragraph id="B07836A5664C9C09FF0F52D5049BF9CF" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">Abdomen: the posterolateral projections of the ninth segment, very long and sharp, nearly as long as the tenth abdominal tergum.</paragraph>
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<paragraph id="B07836A5664C9C09FF0F51050770F9F7" blockId="2.[151,1437,266,1987]" box="[199,605,1561,1587]" pageId="2" pageNumber="41">Paracercus, very short; cerci, long.</paragraph>
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<paragraph id="B07836A5664C9C09FF0F515D05F5F917" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">
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Female imago (
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<figureCitation id="28FC2A20664C9C09FEB5515D04EBF99F" box="[381,454,1601,1627]" captionStart="FIGURE 4" captionStartId="6.[151,255,1952,1976]" captionTargetBox="[270,1330,203,1914]" captionTargetId="figure@6.[235,1354,194,1928]" captionTargetPageId="6" captionText="FIGURE 4. Epicharmeropsis hexavenulosu s sp. nov., female imago, paratype CNU-E-YX- 2007004: A, photograph of paratype; B, body with wings on lateral view; C, forewing." httpUri="https://zenodo.org/record/179373/files/figure.png" pageId="2" pageNumber="41">Fig. 4</figureCitation>
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.). Eyes relatively small, separate; mouth apparatus vestigial, only anterior margin of the frons forming a projected lamella; coxa, trochanter and femur of fore and middle legs preserved, sturdy; the posterolateral projections of the ninth segment not extended posterolaterally. Other characters are as in male.
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</paragraph>
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<paragraph id="B07836A5664C9C09FF0F51FD067BF93F" blockId="2.[151,1437,266,1987]" box="[199,854,1761,1787]" pageId="2" pageNumber="41">Male and female subimagoes, and nymph are unknown.</paragraph>
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<paragraph id="B07836A5664C9C09FF0F5014010FF807" blockId="2.[151,1437,266,1987]" pageId="2" pageNumber="41">
|
||
<emphasis id="82B3EAB7664C9C09FF0F50140453F8E6" bold="true" box="[199,382,1800,1826]" pageId="2" pageNumber="41">Measurements</emphasis>
|
||
(mm).
|
||
<typeStatus id="6F7C8807664C9C09FE1D5015076AF8E7" box="[469,583,1801,1827]" pageId="2" pageNumber="41" type="holotype">Holotype</typeStatus>
|
||
: body length (excluding caudalii) 31.0 (head length 3.0, pronotum length 0.5, pterothorax length 9.5, abdomen length 18.0); forewing length 34.0, its width 14.0; hind wing length 18.0, its width 11.0; length of fore leg fragment 27.0 (femur 9.0, tibia 7.0, first tarsus 4.0, second tarsus 5.0, third preserved tarsus 2.0); femora of middle and hind legs 6.0; tarsi of middle and hind legs 5.0.
|
||
<typeStatus id="6F7C8807664C9C09FAE5509D00B4F85F" box="[1325,1433,1921,1947]" pageId="2" pageNumber="41" type="paratype">Paratype</typeStatus>
|
||
: female imago No.CNU-E-YX-2007004, forewing length 34.0, its width 14.5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |