treatments-xml/data/38/55/87/385587AAF73F1169FF4CC15AFD1BA7B4.xml
2024-06-21 12:33:38 +02:00

964 lines
138 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="E0A74C42481FC534EF0E6EE8359596FD" ID-DOI="10.11646/zootaxa.5447.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="11119209" ID-ZooBank="3A9842EC-F3D2-4839-9731-AA79A27B25CB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1714984939816" checkinUser="plazi" docAuthor="Kawase, Naoki" docDate="2024" docId="385587AAF73F1169FF4CC15AFD1BA7B4" docLanguage="en" docName="zootaxa.5447.1.3.pdf" docOrigin="Zootaxa 5447 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5447.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophila inazui Kawase 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="67" masterDocId="C46CFFD2F7361165FFDBC75DFF92A619" masterDocTitle="Redescription of Rhyacophila clemens Tsuda 1940 (Trichoptera, Rhyacophilidae), with descriptions of five new, allied species from Japan" masterLastPageNumber="76" masterPageNumber="55" pageNumber="64" updateTime="1715030625241" updateUser="diego" zenodo-license-document="CLOSED">
<mods:mods id="2FD15404E9FC2B055ED2F3CD2595FB47" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7C5A13AF2CF2BBAB7A7EAB4D8696103C">
<mods:title id="919A0AB36A09762029911DF7A1C4672E">Redescription of Rhyacophila clemens Tsuda 1940 (Trichoptera, Rhyacophilidae), with descriptions of five new, allied species from Japan</mods:title>
</mods:titleInfo>
<mods:name id="E26F23A0135236DE2B06C468E8000210" type="personal">
<mods:role id="17E5F4C047A9242D3E5F67180DBC3949">
<mods:roleTerm id="3709A17ED9EBA908EEED4E07515BCEBB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF32EBD87D77FB491E2F32C0929212AE">Kawase, Naoki</mods:namePart>
</mods:name>
<mods:typeOfResource id="336DDD835D3E5BFE4F96036A689315AF">text</mods:typeOfResource>
<mods:relatedItem id="7AD710811DB97AFD051EE9CB9415809E" type="host">
<mods:titleInfo id="650207412D72A92A317AB5810184D26D">
<mods:title id="E8C934A35313CF08E1D0BCC90ED37156">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="651D4C659B186F70C63483ADEFCE3387">
<mods:date id="C4B434D4229E589E2D6DFD5CDAE6C185">2024</mods:date>
<mods:detail id="E6A87ACCBFE407F15DA708081FCE7ACD" type="pubDate">
<mods:number id="D76B796334802EF9F12AD7A16868536A">2024-05-03</mods:number>
</mods:detail>
<mods:detail id="B69E7ECA685D3ABC032F602B2BD27794" type="volume">
<mods:number id="5FAD182F27F53858866A749DCAA0608A">5447</mods:number>
</mods:detail>
<mods:detail id="0C5056C70379F45813EE3B7DE27F6C2B" type="issue">
<mods:number id="3136503937C3BC64DE72CEA926E2F71B">1</mods:number>
</mods:detail>
<mods:extent id="C68E5512FD3BBD393B29580473696F6A" unit="page">
<mods:start id="07315802343090F05EAEC0ECDC25D404">55</mods:start>
<mods:end id="7298C9012932341881B916BE36E89B6E">76</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="86F0B7A8176642249E931ACBF84A0A8A">
<mods:url id="39E6684F21FF0C2D5D92107646119A72">http://dx.doi.org/10.11646/zootaxa.5447.1.3</mods:url>
</mods:location>
<mods:classification id="B005A7C8B943D70C9406C06AD0182A9B">journal article</mods:classification>
<mods:identifier id="13C611C2C2C1504EEE5CD42257F462D2" type="DOI">10.11646/zootaxa.5447.1.3</mods:identifier>
<mods:identifier id="B6A32E2767AC1385FB6A2129CEEDD6C5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D7CEC2111F7CEBF2D42237C99905332D" type="Zenodo-Dep">11119209</mods:identifier>
<mods:identifier id="855FF11968475896082D8D27D324FAC8" type="ZooBank">3A9842EC-F3D2-4839-9731-AA79A27B25CB</mods:identifier>
</mods:mods>
<treatment id="385587AAF73F1169FF4CC15AFD1BA7B4" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4" httpUri="http://treatment.plazi.org/id/385587AAF73F1169FF4CC15AFD1BA7B4" lastPageId="12" lastPageNumber="67" pageId="9" pageNumber="64">
<subSubSection id="F8E66537F73F116CFF4CC15AFE7AA03B" box="[151,488,1543,1570]" pageId="9" pageNumber="64" type="nomenclature">
<paragraph id="B04336BCF73F116CFF4CC15AFE7AA03B" blockId="9.[151,488,1543,1606]" box="[151,488,1543,1570]" pageId="9" pageNumber="64">
<heading id="EB0B81D0F73F116CFF4CC15AFE7AA03B" bold="true" box="[151,488,1543,1570]" fontSize="11" level="1" pageId="9" pageNumber="64" reason="1">
<emphasis id="8288EAAEF73F116CFF4CC15AFE7AA03B" bold="true" box="[151,488,1543,1570]" pageId="9" pageNumber="64">
<taxonomicName id="77FC4D3FF73F116CFF4CC15AFE11A03B" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[151,387,1543,1570]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="inazui" status="sp. nov.">
<emphasis id="8288EAAEF73F116CFF4CC15AFE11A03B" bold="true" box="[151,387,1543,1570]" italics="true" pageId="9" pageNumber="64">Rhyacophila inazui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73F116CFE51C155FE7AA03B" box="[394,488,1544,1570]" pageId="9" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F73F116CFF4CC171FEF0A05F" box="[151,354,1580,1606]" pageId="9" pageNumber="64" type="description">
<paragraph id="B04336BCF73F116CFF4CC171FEF0A05F" blockId="9.[151,488,1543,1606]" box="[151,354,1580,1606]" pageId="9" pageNumber="64">
(
<figureCitation id="28C72A39F73F116CFF44C171FEBFA05F" box="[159,301,1580,1606]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="9" pageNumber="64">Figs 4A4N</figureCitation>
,
<figureCitation id="28C72A39F73F116CFEE3C171FEC8A05F" box="[312,346,1580,1606]" captionStart="FIGURE 9" captionStartId="19.[151,250,1739,1763]" captionTargetBox="[292,1296,194,1702]" captionTargetId="figure-16@19.[279,1309,181,1715]" captionTargetPageId="19" captionText="FIGURE 9. Distributions of species in the Rhyacophila clemens Species Group in Japan." figureDoi="http://doi.org/10.5281/zenodo.11119229" httpUri="https://zenodo.org/record/11119229/files/figure.png" pageId="9" pageNumber="64">9C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F73F116CFF4CC136FC83A0D9" pageId="9" pageNumber="64" type="reference_group">
<paragraph id="B04336BCF73F116CFF4CC136FCB9A0BB" blockId="9.[151,1370,1643,1790]" pageId="9" pageNumber="64">
<taxonomicName id="77FC4D3FF73F116CFF4CC136FEDCA09A" box="[151,334,1643,1667]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="undefined-2">
<emphasis id="8288EAAEF73F116CFF4CC136FE85A09A" box="[151,279,1643,1667]" italics="true" pageId="9" pageNumber="64">Rhyacophila</emphasis>
sp. 2
</taxonomicName>
(
<taxonomicName id="77FC4D3FF73F116CFE87C136FE3CA09A" ID-CoL="4SNQ4" authorityName="Tsuda" authorityYear="1940" box="[348,430,1643,1667]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="clemens">
<emphasis id="8288EAAEF73F116CFE87C136FE3CA09A" box="[348,430,1643,1667]" italics="true" pageId="9" pageNumber="64">clemens</emphasis>
</taxonomicName>
gr.):
<bibRefCitation id="D46D4B4DF73F116CFE3FC131FDCAA09A" author="Inazu, K." box="[484,600,1643,1667]" pageId="9" pageNumber="64" pagination="59 - 104" refId="ref17485" refString="Inazu, K. (2008) Collection record of caddisflies in the Itoi River Hyogo over 5 years (2001 - 2005) - Notes on emergence periods, classification, distribution, and ecology of adults and larvae. Hyogo Freshwater Biology, 60, 59 - 104. [in Japanese]" type="journal article" year="2008">Inazu 2008</bibRefCitation>
, 65, 88, 99, wing markings, flight period;
<bibRefCitation id="D46D4B4DF73F116CFBD8C131FB77A09A" author="Inazu, K. &amp; Nishida, A." box="[1027,1253,1643,1667]" pageId="9" pageNumber="64" pagination="169 - 212" refId="ref17540" refString="Inazu, K. &amp; Nishida, A. (2011) Trichoptera. In: Hyogo Freshwater Biology Society (Ed.), Hyogo no kawa no ikimono zukan [Illustrated Guide to River Organisms of Hyogo Prefecture]. Hyogo Freshwater Biology Society, Himeji, pp. 169 - 212. [in Japanese]" type="book chapter" year="2011">Inazu &amp; Nishida 2011</bibRefCitation>
, 176, male.
<taxonomicName id="77FC4D3FF73F116CFF4CC1D7FEDCA0BB" box="[151,334,1674,1698]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="8288EAAEF73F116CFF4CC1D7FE85A0BB" box="[151,279,1674,1698]" italics="true" pageId="9" pageNumber="64">Rhyacophila</emphasis>
sp. 1
</taxonomicName>
(
<taxonomicName id="77FC4D3FF73F116CFE87C1D7FE3CA0BB" ID-CoL="4SNQ4" authorityName="Tsuda" authorityYear="1940" box="[348,430,1674,1698]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="clemens">
<emphasis id="8288EAAEF73F116CFE87C1D7FE3CA0BB" box="[348,430,1674,1698]" italics="true" pageId="9" pageNumber="64">clemens</emphasis>
</taxonomicName>
gr.):
<bibRefCitation id="D46D4B4DF73F116CFE3FC1D7FD47A0BB" author="Kawase, N. &amp; Morita, H" box="[484,725,1674,1698]" pageId="9" pageNumber="64" pagination="31 - 40" refId="ref17648" refString="Kawase, N. &amp; Morita, H (2010) The caddisfly (Insecta, Trichoptera) fauna of the Suzuka Mountains. Biology of Inland Waters, 25, 31 - 40. [in Japanese with English abstract]" type="journal article" year="2010">Kawase &amp; Morita 2010</bibRefCitation>
, 36, list.
</paragraph>
<paragraph id="B04336BCF73F116CFF4CC1F5FC83A0D9" blockId="9.[151,1370,1643,1790]" box="[151,785,1704,1729]" pageId="9" pageNumber="64">
<taxonomicName id="77FC4D3FF73F116CFF4CC1F5FEA9A0D9" box="[151,315,1704,1728]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8288EAAEF73F116CFF4CC1F5FE85A0D9" box="[151,279,1704,1728]" italics="true" pageId="9" pageNumber="64">Rhyacophila</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="77FC4D3FF73F116CFE92C1F5FE32A0D9" ID-CoL="4SNQ4" authorityName="Tsuda" authorityYear="1940" box="[329,416,1704,1728]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="clemens">
<emphasis id="8288EAAEF73F116CFE92C1F5FE32A0D9" box="[329,416,1704,1728]" italics="true" pageId="9" pageNumber="64">Clemens</emphasis>
</taxonomicName>
gr.):
<bibRefCitation id="D46D4B4DF73F116CFE0CC1F4FD55A0D8" author="Kawase, N. &amp; Morita, H." box="[471,711,1704,1729]" pageId="9" pageNumber="64" pagination="1 - 10" refId="ref17690" refString="Kawase, N. &amp; Morita, H. (2014) The caddisfly (Insecta: Trichoptera) fauna of the upstream area of the Takatoki River, Shiga Prefecture, central Japan. Biology of Inland Waters, 28, 1 - 10. [in Japanese with English abstract]" type="journal article" year="2014">Kawase &amp; Morita 2014</bibRefCitation>
, 4, list.
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F73F116CFF4CC19AFD0DA0E7" pageId="9" pageNumber="64" type="description">
<paragraph id="B04336BCF73F116CFF4CC19AFDD4A0C6" blockId="9.[151,1370,1643,1790]" box="[151,582,1735,1759]" pageId="9" pageNumber="64">
<taxonomicName id="77FC4D3FF73F116CFF4CC19AFEC6A0C6" box="[151,340,1735,1759]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="undefined-E">
<emphasis id="8288EAAEF73F116CFF4CC19AFE85A0C6" box="[151,279,1735,1759]" italics="true" pageId="9" pageNumber="64">Rhyacophila</emphasis>
sp. E
</taxonomicName>
:
<bibRefCitation id="D46D4B4DF73F116CFE85C19AFE77A0C6" author="Nojima, K." box="[350,485,1735,1759]" pageId="9" pageNumber="64" pagination="107 - 131" refId="ref18109" refString="Nojima, K. (2017) Caddisflies (Insecta: Trichoptera) in Okayama Prefecture, western Japan. Biology of Inland Waters, 32, 107 - 131. [in Japanese with English abstract]" type="journal article" year="2017">Nojima 2017</bibRefCitation>
, 111, list.
</paragraph>
<paragraph id="B04336BCF73F116CFF4CC1BBFD0DA0E7" blockId="9.[151,1370,1643,1790]" box="[151,671,1766,1790]" pageId="9" pageNumber="64">
<taxonomicName id="77FC4D3FF73F116CFF4CC1BBFEA9A0E4" box="[151,315,1766,1790]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8288EAAEF73F116CFF4CC1BBFE85A0E7" box="[151,279,1766,1790]" italics="true" pageId="9" pageNumber="64">Rhyacophila</emphasis>
sp.
</taxonomicName>
13:
<bibRefCitation id="D46D4B4DF73F116CFEBCC1BBFDDBA0E7" author="Yamamoto, E. &amp; Nozaki, T. &amp; Kuhara, N. &amp; Ito, T." box="[359,585,1766,1790]" pageId="9" pageNumber="64" pagination="67 - 76" refId="ref18786" refString="Yamamoto, E., Nozaki, T., Kuhara, N. &amp; Ito, T. (2021) Caddisflies (Trichoptera) collected from a mountain stream, Namakusadani, in Shikoku, western Japan. In: Yamamoto Institute of Forest Biology (Ed.), Aquatic Insects of Odamiyama and Nature of Matsubara-izumi. Ecosystem Conservation Society of Ehime, Ehime, pp. 67 - 76. [in Japanese]" type="book chapter" year="2021">
Yamamoto
<emphasis id="8288EAAEF73F116CFE00C1BAFD95A0E7" box="[475,519,1766,1790]" italics="true" pageId="9" pageNumber="64">et al</emphasis>
. 2021
</bibRefCitation>
, 70, list.
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F73F116EFF4CC07BFD9AA264" lastPageId="11" lastPageNumber="66" pageId="9" pageNumber="64" type="diagnosis">
<paragraph id="B04336BCF73F116FFF4CC07BFB77A790" blockId="9.[151,1437,1830,2037]" lastBlockId="10.[151,1437,151,393]" lastPageId="10" lastPageNumber="65" pageId="9" pageNumber="64">
<emphasis id="8288EAAEF73F116CFF4CC07BFE80A159" bold="true" box="[151,274,1830,1856]" pageId="9" pageNumber="64">Diagnosis.</emphasis>
Adults of this species are unique in the
<taxonomicName id="77FC4D3FF73F116CFD37C07AFCFAA158" authorityName="Tsuda" authorityYear="1940" box="[748,872,1831,1857]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="clemens">
<emphasis id="8288EAAEF73F116CFD37C07AFCFAA158" box="[748,872,1831,1857]" italics="true" pageId="9" pageNumber="64">R. clemens</emphasis>
</taxonomicName>
Species Group in having distinct pale yellowish spots along the posterior margin of each forewing (
<figureCitation id="28C72A39F73F116CFD01C016FCFAA17C" box="[730,872,1867,1893]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="9" pageNumber="64">Figs 4A, 4B</figureCitation>
). The male genitalia of
<taxonomicName id="77FC4D3FF73F116CFBADC016FB4BA17D" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[1142,1241,1867,1892]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="inazui" status="sp. nov.">
<emphasis id="8288EAAEF73F116CFBADC016FB4BA17D" box="[1142,1241,1867,1892]" italics="true" pageId="9" pageNumber="64">R. inazui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73F116CFB39C016FAABA17C" box="[1250,1337,1867,1893]" pageId="9" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
are very similar to those of
<taxonomicName id="77FC4D3FF73F116CFEB0C032FD85A190" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[363,535,1903,1929]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="angulicauda" status="sp. nov.">
<emphasis id="8288EAAEF73F116CFEB0C032FD85A190" box="[363,535,1903,1929]" italics="true" pageId="9" pageNumber="64">R. angulicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73F116CFDC4C032FDE4A190" box="[543,630,1903,1929]" pageId="9" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="77FC4D3FF73F116CFD76C032FCDFA190" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[685,845,1903,1929]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="parvicauda" status="sp. nov.">
<emphasis id="8288EAAEF73F116CFD76C032FCDFA190" box="[685,845,1903,1929]" italics="true" pageId="9" pageNumber="64">R. parvicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73F116CFC8FC032FC39A190" box="[852,939,1903,1929]" pageId="9" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
in having a U-shaped apical hinged process of segment X in ventral view, but can be distinguished from the latter two by the size of the apicodorsal lobe of segment IX: The apicodorsal lobe of segment IX is indistinct in the present species (
<figureCitation id="28C72A39F73F116CFB94C0EAFB48A1C8" box="[1103,1242,1975,2001]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="9" pageNumber="64">Figs 4C, 4E</figureCitation>
), but is large and trapezoidal in
<taxonomicName id="77FC4D3FF73F116CFEE7C086FE7BA1EC" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[316,489,2011,2037]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="angulicauda" status="sp. nov.">
<emphasis id="8288EAAEF73F116CFEE7C086FE7BA1EC" box="[316,489,2011,2037]" italics="true" pageId="9" pageNumber="64">R. angulicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73F116CFE29C086FDD9A1EC" box="[498,587,2011,2037]" pageId="9" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
(arrows indicated in
<figureCitation id="28C72A39F73F116CFC9AC086FC5CA1EC" box="[833,974,2010,2037]" captionStart="FIGURE 5" captionStartId="13.[151,250,1525,1549]" captionTargetBox="[284,1304,286,1489]" captionTargetId="figure-44@13.[271,1316,273,1501]" captionTargetPageId="13" captionText="FIGURE 5. Rhyacophila angulicauda sp. nov. 5A5D, male genitalia: 5A, left lateral; 5B, segment X and phallic apparatus, left lateral; 5C, dorsal (left inferior appendage omitted); 5D, ventral (left inferior appendage and apex of apical segment of right inferior appendage omitted). 5E5I, female genitalia: 5E, left lateral; 5F, segment VIII, left lateral; 5G, same, ventral; 5H, vaginal apparatus, left lateral; 5I, same, ventral. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119221" httpUri="https://zenodo.org/record/11119221/files/figure.png" pageId="9" pageNumber="64">Figs 5A, 5C</figureCitation>
) and small trapezoidal in
<taxonomicName id="77FC4D3FF73F116CFB20C086FA0FA1EC" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[1275,1437,2011,2037]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="parvicauda" status="sp. nov.">
<emphasis id="8288EAAEF73F116CFB20C086FA0FA1EC" box="[1275,1437,2011,2037]" italics="true" pageId="9" pageNumber="64">R. parvicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73C116FFF4CC7CAFF7DA6A8" box="[151,239,151,177]" pageId="10" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="28C72A39F73C116FFF26C7CAFE1EA6A8" box="[253,396,151,177]" captionStart="FIGURE 6" captionStartId="15.[151,250,1368,1392]" captionTargetBox="[298,1289,250,1332]" captionTargetId="figure-43@15.[285,1301,237,1345]" captionTargetPageId="15" captionText="FIGURE 6. Rhyacophila parvicauda sp. nov. 6A6D, male genitalia: 6A, left lateral; 6B, segment X and phallic apparatus, left lateral; 6C, dorsal (left inferior appendage omitted); 6D, ventral (left inferior appendage omitted). 6E6H, female genitalia: 6E, segment VIII, left lateral; 6F, same, ventral; 6G, vaginal apparatus, left lateral; 6H, same, ventral. Abbreviation: vr = ventral ridge.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119223" httpUri="https://zenodo.org/record/11119223/files/figure.png" pageId="10" pageNumber="65">Figs 6A, 6C</figureCitation>
). The female genitalia of
<taxonomicName id="77FC4D3FF73C116FFD75C7CAFC80A6A9" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[686,786,151,176]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="inazui" status="sp. nov.">
<emphasis id="8288EAAEF73C116FFD75C7CAFC80A6A9" box="[686,786,151,176]" italics="true" pageId="10" pageNumber="65">R. inazui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73C116FFCC1C7CAFCE0A6A8" box="[794,882,151,177]" pageId="10" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
are somewhat similar to those of
<taxonomicName id="77FC4D3FF73C116FFB2BC7CAFA0EA6A8" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[1264,1436,151,177]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="angulicauda" status="sp. nov.">
<emphasis id="8288EAAEF73C116FFB2BC7CAFA0EA6A8" box="[1264,1436,151,177]" italics="true" pageId="10" pageNumber="65">R. angulicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73C116FFF4CC7E6FF60A6CC" box="[151,242,187,213]" pageId="10" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="77FC4D3FF73C116FFEDBC7E6FE33A6CC" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[256,417,187,213]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="parvicauda" status="sp. nov.">
<emphasis id="8288EAAEF73C116FFEDBC7E6FE33A6CC" box="[256,417,187,213]" italics="true" pageId="10" pageNumber="65">R. parvicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73C116FFE71C7E6FD97A6CC" box="[426,517,187,213]" pageId="10" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="77FC4D3FF73C116FFD9FC7E6FD4FA6CC" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[580,733,187,213]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="furcicauda" status="sp. nov.">
<emphasis id="8288EAAEF73C116FFD9FC7E6FD4FA6CC" box="[580,733,187,213]" italics="true" pageId="10" pageNumber="65">R. furcicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73C116FFD3CC7E6FCD2A6CC" box="[743,832,187,213]" pageId="10" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
in having almost straight dorsal margins of segment VIII in lateral view, but can be distinguished from the latter three species by the combination of the following two characteristics: (1) the setae on the lateral surface of segment VIII are hair like (
<figureCitation id="28C72A39F73C116FFBF0C65EFB18A704" box="[1067,1162,259,285]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="10" pageNumber="65">Figs 4K</figureCitation>
,
<figureCitation id="28C72A39F73C116FFB4CC65FFB24A704" box="[1175,1206,258,285]" captionStart="FIGURE 5" captionStartId="13.[151,250,1525,1549]" captionTargetBox="[284,1304,286,1489]" captionTargetId="figure-44@13.[271,1316,273,1501]" captionTargetPageId="13" captionText="FIGURE 5. Rhyacophila angulicauda sp. nov. 5A5D, male genitalia: 5A, left lateral; 5B, segment X and phallic apparatus, left lateral; 5C, dorsal (left inferior appendage omitted); 5D, ventral (left inferior appendage and apex of apical segment of right inferior appendage omitted). 5E5I, female genitalia: 5E, left lateral; 5F, segment VIII, left lateral; 5G, same, ventral; 5H, vaginal apparatus, left lateral; 5I, same, ventral. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119221" httpUri="https://zenodo.org/record/11119221/files/figure.png" pageId="10" pageNumber="65">5F</figureCitation>
) in
<taxonomicName id="77FC4D3FF73C116FFB3EC65EFAD8A705" authorityName="Kawase" authorityYear="2024" box="[1253,1354,259,284]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="inazui">
<emphasis id="8288EAAEF73C116FFB3EC65EFAD8A705" box="[1253,1354,259,284]" italics="true" pageId="10" pageNumber="65">R. inazui</emphasis>
</taxonomicName>
and
<taxonomicName id="77FC4D3FF73C116FFA5FC65EFEB1A758" authorityName="Kawase" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="angulicauda">
<emphasis id="8288EAAEF73C116FFA5FC65EFEB1A758" italics="true" pageId="10" pageNumber="65">R. angulicauda</emphasis>
</taxonomicName>
, but thick, spine-like in
<taxonomicName id="77FC4D3FF73C116FFDE9C67AFD43A758" authorityName="Kawase" authorityYear="2024" box="[562,721,295,321]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="parvicauda">
<emphasis id="8288EAAEF73C116FFDE9C67AFD43A758" box="[562,721,295,321]" italics="true" pageId="10" pageNumber="65">R. parvicauda</emphasis>
</taxonomicName>
and
<taxonomicName id="77FC4D3FF73C116FFCDDC67AFC0FA758" authorityName="Kawase" authorityYear="2024" box="[774,925,295,321]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="furcicauda">
<emphasis id="8288EAAEF73C116FFCDDC67AFC0FA758" box="[774,925,295,321]" italics="true" pageId="10" pageNumber="65">R. furcicauda</emphasis>
</taxonomicName>
(
<figureCitation id="28C72A39F73C116FFC70C67AFB97A758" box="[939,1029,295,321]" captionStart="FIGURE 6" captionStartId="15.[151,250,1368,1392]" captionTargetBox="[298,1289,250,1332]" captionTargetId="figure-43@15.[285,1301,237,1345]" captionTargetPageId="15" captionText="FIGURE 6. Rhyacophila parvicauda sp. nov. 6A6D, male genitalia: 6A, left lateral; 6B, segment X and phallic apparatus, left lateral; 6C, dorsal (left inferior appendage omitted); 6D, ventral (left inferior appendage omitted). 6E6H, female genitalia: 6E, segment VIII, left lateral; 6F, same, ventral; 6G, vaginal apparatus, left lateral; 6H, same, ventral. Abbreviation: vr = ventral ridge.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119223" httpUri="https://zenodo.org/record/11119223/files/figure.png" pageId="10" pageNumber="65">Figs 6E</figureCitation>
,
<figureCitation id="28C72A39F73C116FFBCBC67AFBA4A758" box="[1040,1078,295,321]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="10" pageNumber="65">7G</figureCitation>
); and (2) a pair of ventral ridges extend posterolaterad and can be seen clearly in lateral view in
<taxonomicName id="77FC4D3FF73C116FFC8AC616FC21A77D" authorityName="Kawase" authorityYear="2024" box="[849,947,331,356]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="inazui">
<emphasis id="8288EAAEF73C116FFC8AC616FC21A77D" box="[849,947,331,356]" italics="true" pageId="10" pageNumber="65">R. inazui</emphasis>
</taxonomicName>
and
<taxonomicName id="77FC4D3FF73C116FFC33C616FB14A77C" authorityName="Kawase" authorityYear="2024" box="[1000,1158,331,357]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="parvicauda">
<emphasis id="8288EAAEF73C116FFC33C616FB14A77C" box="[1000,1158,331,357]" italics="true" pageId="10" pageNumber="65">R. parvicauda</emphasis>
</taxonomicName>
(
<figureCitation id="28C72A39F73C116FFB48C616FB7DA77C" box="[1171,1263,331,357]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="10" pageNumber="65">Figs 4K</figureCitation>
,
<figureCitation id="28C72A39F73C116FFB21C616FA88A77C" box="[1274,1306,331,357]" captionStart="FIGURE 6" captionStartId="15.[151,250,1368,1392]" captionTargetBox="[298,1289,250,1332]" captionTargetId="figure-43@15.[285,1301,237,1345]" captionTargetPageId="15" captionText="FIGURE 6. Rhyacophila parvicauda sp. nov. 6A6D, male genitalia: 6A, left lateral; 6B, segment X and phallic apparatus, left lateral; 6C, dorsal (left inferior appendage omitted); 6D, ventral (left inferior appendage omitted). 6E6H, female genitalia: 6E, segment VIII, left lateral; 6F, same, ventral; 6G, vaginal apparatus, left lateral; 6H, same, ventral. Abbreviation: vr = ventral ridge.Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119223" httpUri="https://zenodo.org/record/11119223/files/figure.png" pageId="10" pageNumber="65">6E</figureCitation>
) but extend posteroventrad and are invisible in lateral view in
<taxonomicName id="77FC4D3FF73C116FFD12C632FCE6A790" authorityName="Kawase" authorityYear="2024" box="[713,884,367,393]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="angulicauda">
<emphasis id="8288EAAEF73C116FFD12C632FCE6A790" box="[713,884,367,393]" italics="true" pageId="10" pageNumber="65">R. angulicauda</emphasis>
</taxonomicName>
and
<taxonomicName id="77FC4D3FF73C116FFC71C632FBD3A790" authorityName="Kawase" authorityYear="2024" box="[938,1089,367,393]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="furcicauda">
<emphasis id="8288EAAEF73C116FFC71C632FBD3A790" box="[938,1089,367,393]" italics="true" pageId="10" pageNumber="65">R. furcicauda</emphasis>
</taxonomicName>
(
<figureCitation id="28C72A39F73C116FFB8BC632FB35A790" box="[1104,1191,366,393]" captionStart="FIGURE 5" captionStartId="13.[151,250,1525,1549]" captionTargetBox="[284,1304,286,1489]" captionTargetId="figure-44@13.[271,1316,273,1501]" captionTargetPageId="13" captionText="FIGURE 5. Rhyacophila angulicauda sp. nov. 5A5D, male genitalia: 5A, left lateral; 5B, segment X and phallic apparatus, left lateral; 5C, dorsal (left inferior appendage omitted); 5D, ventral (left inferior appendage and apex of apical segment of right inferior appendage omitted). 5E5I, female genitalia: 5E, left lateral; 5F, segment VIII, left lateral; 5G, same, ventral; 5H, vaginal apparatus, left lateral; 5I, same, ventral. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119221" httpUri="https://zenodo.org/record/11119221/files/figure.png" pageId="10" pageNumber="65">Figs 5F</figureCitation>
,
<figureCitation id="28C72A39F73C116FFB69C632FB4AA790" box="[1202,1240,367,393]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="10" pageNumber="65">7G</figureCitation>
).
</paragraph>
<caption id="E4836634F73C116FFF4CC04FFBBFA1C7" ID-DOI="http://doi.org/10.5281/zenodo.11119217" ID-Zenodo-Dep="11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="10" pageNumber="65" startId="10.[151,250,1810,1834]" targetBox="[288,1300,430,1774]" targetPageId="10" targetType="figure">
<paragraph id="B04336BCF73C116FFF4CC04FFBBFA1C7" blockId="10.[151,1437,1810,2015]" pageId="10" pageNumber="65">
<emphasis id="8288EAAEF73C116FFF4CC04FFDDBA132" bold="true" box="[151,585,1810,1835]" pageId="10" pageNumber="65">
FIGURE 4.
<taxonomicName id="77FC4D3FF73C116FFEC4C04FFE7FA132" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[287,493,1810,1835]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="inazui" status="sp. nov.">
<emphasis id="8288EAAEF73C116FFEC4C04FFE7FA132" bold="true" box="[287,493,1810,1835]" italics="true" pageId="10" pageNumber="65">Rhyacophila inazui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73C116FFE2DC04EFDDBA132" box="[502,585,1811,1835]" pageId="10" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text.
</paragraph>
</caption>
<paragraph id="B04336BCF73D116EFF1CC7CBFF5AA758" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="66">
<emphasis id="8288EAAEF73D116EFF1CC7CBFE99A6A9" bold="true" box="[199,267,150,176]" pageId="11" pageNumber="66">Adult</emphasis>
(
<figureCitation id="28C72A39F73D116EFEC6C7CAFE3DA6A8" box="[285,431,151,177]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4A, 4B</figureCitation>
). Length of each forewing: male
<quantity id="77049B59F73D116EFCE9C7CAFC2EA6A8" box="[818,956,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="7.8" metricValueMin="6.8" pageId="11" pageNumber="66" unit="mm" value="7.3" valueMax="7.8" valueMin="6.8">6.87.8 mm</quantity>
(mean = 7.3, SD = 0.30, n = 17), female
<quantity id="77049B59F73D116EFF4CC7E6FE8CA6CC" box="[151,286,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="8.55" metricValueMax="9.3" metricValueMin="7.8" pageId="11" pageNumber="66" unit="mm" value="8.55" valueMax="9.3" valueMin="7.8">7.89.3 mm</quantity>
(mean = 8.2, SD = 0.47, n = 8). General morphology and coloration similar to those of
<taxonomicName id="77FC4D3FF73D116EFB2AC7E6FAF8A6CC" authorityName="Tsuda" authorityYear="1940" box="[1265,1386,187,213]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="66" phylum="Arthropoda" rank="species" species="clemens">
<emphasis id="8288EAAEF73D116EFB2AC7E6FAF8A6CC" box="[1265,1386,187,213]" italics="true" pageId="11" pageNumber="66">R. clemens</emphasis>
</taxonomicName>
, but forewings bearing pale yellowish speckles more densely from base to apical margins (
<figureCitation id="28C72A39F73D116EFBB8C782FB2CA6E0" box="[1123,1214,223,249]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4A</figureCitation>
), distinct pale spots visible even in alcohol (
<figureCitation id="28C72A39F73D116EFE7BC65EFE6AA704" box="[416,504,259,285]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4B</figureCitation>
). In venation, root of fork I on each forewing clearly longer than that of fork II (
<figureCitation id="28C72A39F73D116EFAB4C65EFF29A758" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="B04336BCF73D116EFF1CC616FF74A544" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="66">
<emphasis id="8288EAAEF73D116EFF1CC616FEE6A77D" bold="true" box="[199,372,330,357]" pageId="11" pageNumber="66">Male genitalia</emphasis>
(
<figureCitation id="28C72A39F73D116EFE5FC616FD87A77C" box="[388,533,331,357]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4C4H</figureCitation>
). Segment IX rectangular in dorsal view with apicodorsal lobe very short (
<figureCitation id="28C72A39F73D116EFAB4C616FF2BA790" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4E</figureCitation>
), dorsal half longer than ventral half in lateral view (
<figureCitation id="28C72A39F73D116EFCCCC632FCE3A790" box="[791,881,367,393]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4C</figureCitation>
). Preanal appendages absent. Segment X reverse L-shaped in lateral view composed of vertical part and horizontal part: vertical part tall in lateral view (
<figureCitation id="28C72A39F73D116EFAE8C6CEFA1DA7B4" box="[1331,1423,403,429]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4D</figureCitation>
); horizontal part nearly oval in dorsal view with shallow incisions anteriorly and posteriorly in dorsal view (
<figureCitation id="28C72A39F73D116EFAE3C6EAFA1DA7C8" box="[1336,1423,439,465]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4E</figureCitation>
); apical hinged process U-shaped in ventral view, with pair of slender finger-like lobes, directed posterodorsad (
<figureCitation id="28C72A39F73D116EFAB1C686FE8AA400" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4C4F, 4G</figureCitation>
<figureCitation id="28C72A39F73D116EFEC3C6A2FEB0A400" box="[280,290,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1690,1714]" captionTargetBox="[279,1307,246,1655]" captionTargetId="figure-51@2.[271,1316,234,1667]" captionTargetPageId="2" captionText="FIGURE 1. Rhyacophila clemens Tsuda 1940. 1A1J: 1A, male habitus, left dorsolateral (photographed by K. Inazu); 1B, male head and pro-, meso- and metanota, dorsal; 1C, male head, left lateral; 1D, male right wings, dorsal; 1E, lectotype (in glass tube) and labels preserved in INHS; 1F, male right wings, paralectotype (balsam mounted slide) preserved in OMNH; 1G1J, male genitalia: 1G, left lateral; 1H, segment X and phallic apparatus, left lateral; 1I, same, posterodorsal; 1J, phallic apparatus, left dorsolateral. Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; IX, X = abdominal segments IX, X; A1, A2, A3 = anal veins 13; ab = apical band; aed = aedeagus; adl = apicodorsal lobe of segment IX; ahp = apical hinged process of segment X; as = anal sclerite; asia = apical segment of inferior appendage (paired); bsia = basal segment of inferior appendage (paired); C = costa; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; hp = horizontal part of segment X; M1, M2, M3, M4 = median veins 14; m-cu = crossvein between median and cubital veins; phb = phallobase; R1, R2, R3, R4, R5 = radial veins 15; r-m = crossvein between radial and median veins; Sc = subcosta; tb = tergal band; TC = thyridial cell; vp = vertical part of segment X; vpl = ventral plate of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11119211" httpUri="https://zenodo.org/record/11119211/files/figure.png" pageId="11" pageNumber="66">1</figureCitation>
), with pair of round anteroventral corners, bearing small median protrusion anteroventrally (arrows in
<figureCitation id="28C72A39F73D116EFF4CC57EFEB1A424" box="[151,291,547,573]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4F, 4G</figureCitation>
<figureCitation id="28C72A39F73D116EFEF8C57EFEBFA424" box="[291,301,547,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1690,1714]" captionTargetBox="[279,1307,246,1655]" captionTargetId="figure-51@2.[271,1316,234,1667]" captionTargetPageId="2" captionText="FIGURE 1. Rhyacophila clemens Tsuda 1940. 1A1J: 1A, male habitus, left dorsolateral (photographed by K. Inazu); 1B, male head and pro-, meso- and metanota, dorsal; 1C, male head, left lateral; 1D, male right wings, dorsal; 1E, lectotype (in glass tube) and labels preserved in INHS; 1F, male right wings, paralectotype (balsam mounted slide) preserved in OMNH; 1G1J, male genitalia: 1G, left lateral; 1H, segment X and phallic apparatus, left lateral; 1I, same, posterodorsal; 1J, phallic apparatus, left dorsolateral. Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; IX, X = abdominal segments IX, X; A1, A2, A3 = anal veins 13; ab = apical band; aed = aedeagus; adl = apicodorsal lobe of segment IX; ahp = apical hinged process of segment X; as = anal sclerite; asia = apical segment of inferior appendage (paired); bsia = basal segment of inferior appendage (paired); C = costa; Cu1, Cu1a, Cu1b = cubital veins 1, 1a, 1b; hp = horizontal part of segment X; M1, M2, M3, M4 = median veins 14; m-cu = crossvein between median and cubital veins; phb = phallobase; R1, R2, R3, R4, R5 = radial veins 15; r-m = crossvein between radial and median veins; Sc = subcosta; tb = tergal band; TC = thyridial cell; vp = vertical part of segment X; vpl = ventral plate of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11119211" httpUri="https://zenodo.org/record/11119211/files/figure.png" pageId="11" pageNumber="66">1</figureCitation>
). Anal sclerites composed of two pairs of lobes, each long rectangular in lateral view (
<figureCitation id="28C72A39F73D116EFAD3C57EFAF1A424" box="[1288,1379,547,573]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4D</figureCitation>
), but apex of dorsal lobe weakly incised mesally in lateral view, bearing common internal root (
<figureCitation id="28C72A39F73D116EFBADC51AFB42A478" box="[1142,1232,583,609]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4D</figureCitation>
). Apical band well developed, semi-oval in lateral view, broadly surrounding anus and base of anal sclerites (
<figureCitation id="28C72A39F73D116EFB52C536FAD1A49C" box="[1161,1347,619,645]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4D, 4F, 4H</figureCitation>
). Tergal band connecting base of apical band to dorsum of phallobase (
<figureCitation id="28C72A39F73D116EFC8DC5D2FC20A4B0" box="[854,946,655,681]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4D</figureCitation>
). Basal segment of each inferior appendage setose, elongate, posterior half gradually thicker in lateral view (
<figureCitation id="28C72A39F73D116EFC80C5EEFC20A4D4" box="[859,946,691,717]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4C</figureCitation>
). Apical segment of each inferior appendage rounded triangular in lateral view (
<figureCitation id="28C72A39F73D116EFDF3C58AFD13A4E8" box="[552,641,727,753]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4C</figureCitation>
), covered with hair-like setae mesally (
<figureCitation id="28C72A39F73D116EFB9EC58AFB0DA4E8" box="[1093,1183,727,753]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4E</figureCitation>
). In phallic apparatus, phallobase short (
<figureCitation id="28C72A39F73D116EFE85C5A6FE2BA50C" box="[350,441,763,789]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4D</figureCitation>
); aedeagus forming narrow tube with sclerotized ventral plate (
<figureCitation id="28C72A39F73D116EFB5BC5A6FAA8A50C" box="[1152,1338,763,789]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4D, 4F, 4H</figureCitation>
); ventral plate horizontal and tongue-shaped with posterior margin shallowly concave in ventral view (
<figureCitation id="28C72A39F73D116EFB77C442FA97A520" box="[1196,1285,799,825]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4G</figureCitation>
<figureCitation id="28C72A39F73D116EFADDC442FA87A520" box="[1286,1301,799,825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1552,1576]" captionTargetBox="[284,1304,193,1516]" captionTargetId="figure-16@3.[271,1316,181,1528]" captionTargetPageId="3" captionText="FIGURE 2. Rhyacophila clemens Tsuda 1940. 2A2E, male genitalia: 2A, dorsal (left inferior appendage omitted); 2B, ventral (left inferior appendage and part of right inferior appendage omitted); 2C2D, variations of apical hinged process of segment X, ventral (2C1, 2C2); variations of ventral plate of aedeagus, ventral (2D1, 2D2) (2C1, 2D1 collected from Hokkaido; 2C2, 2D2 from Ibaraki); 2E, apical band and tergal band, ventral; 2F2K, female genitalia: 2F, left lateral; 2G, ventral; 2H12H3, variations of segment VIII, left lateral (2H1 collected from Hokkaido; 2H2, same Kanagawa; 2H3, same Nagano); 2I, segment VIII, ventral; 2J, vaginal apparatus, left lateral; 2K, same, ventral.Abbreviations: VI, VII, VIII, IX, X, XI = abdominal segments VI, VII, VIII, IX, X, XI; ab = apical band; adl = apicodorsal lobe of segment IX; as = anal sclerite; ahp = apical hinged process of segment X; asia = apical segment of inferior appendage (paired); bsia = basal segment of inferior appendage (paired); hp = horizontal part of segment X; phb = phallobase; pp = posterior process; ps = processus spermathecae; tb = tergal band; vp = vertical part of segment X; vpl = ventral plate of aedeagus; vr = ventral ridge (paired). Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119213" httpUri="https://zenodo.org/record/11119213/files/figure.png" pageId="11" pageNumber="66">2</figureCitation>
). Parameres absent.
</paragraph>
<paragraph id="B04336BCF73D116EFF1CC43AFDE5A240" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="66">
<emphasis id="8288EAAEF73D116EFF1CC43AFE02A599" bold="true" box="[199,400,870,897]" pageId="11" pageNumber="66">Female genitalia</emphasis>
(
<figureCitation id="28C72A39F73D116EFE78C43AFDBEA598" box="[419,556,871,897]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4I4N</figureCitation>
). Segment VIII annular, dorsal margin almost straight in lateral view (
<figureCitation id="28C72A39F73D116EFAB1C43AFF74A5BC" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4I, 4K</figureCitation>
), slightly flattened ventrally; with pair of distinct ventral ridges extending posterolaterad in lateral view, gradually separating from anterior to posterior in lateral and ventral views (
<figureCitation id="28C72A39F73D116EFBD8C4F2FB1AA5D0" box="[1027,1160,943,969]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4I4L</figureCitation>
); pair of apodemal rods reaching posterior end of segment VI (
<figureCitation id="28C72A39F73D116EFD9FC48EFD00A5F4" box="[580,658,979,1005]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4I</figureCitation>
). Segment IX membranous, with pair of apodemal rods dorsolaterally extending into abdominal segment VII (
<figureCitation id="28C72A39F73D116EFD85C4AAFD3DA208" box="[606,687,1015,1041]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4I</figureCitation>
). Vaginal apparatus simple; posterior process sclerotized, smooth, rectangular, about three times as long as wide in lateral view (
<figureCitation id="28C72A39F73D116EFC8CC346FC2AA22C" box="[855,952,1051,1077]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4M</figureCitation>
), oval in ventral view (
<figureCitation id="28C72A39F73D116EFB64C346FA88A22C" box="[1215,1306,1051,1077]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Fig. 4N</figureCitation>
); processus spermathecae membranous (
<figureCitation id="28C72A39F73D116EFE0DC362FDF8A240" box="[470,618,1087,1113]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="11" pageNumber="66">Figs 4M, 4N</figureCitation>
).
</paragraph>
<paragraph id="B04336BCF73D116EFF1CC33EFD9AA264" blockId="11.[151,1437,150,2013]" box="[199,520,1123,1149]" pageId="11" pageNumber="66">
<emphasis id="8288EAAEF73D116EFF1CC33EFE16A264" bold="true" box="[199,388,1123,1149]" pageId="11" pageNumber="66">Immature stage</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F73D1169FF1CC3DAFBFCA6CC" lastPageId="12" lastPageNumber="67" pageId="11" pageNumber="66" type="materials_examined">
<paragraph id="B04336BCF73D116EFF1CC3DAFB85A2DC" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="66">
<materialsCitation id="00943CE1F73D116EFF1CC3DAFB85A2DC" collectingDate="2004-04-16" collectionCode="LBM" collectorName="K. Inazu" country="Japan" county="Honshu" elevation="250" latitude="35.405834" location="Male" longLatPrecision="19" longitude="134.92029" municipality="Asago-shi" pageId="11" pageNumber="66" specimenCode="LBM1410012601" specimenCount="1" stateProvince="Hyogo" typeStatus="holotype">
<emphasis id="8288EAAEF73D116EFF1CC3DAFEA9A2B8" bold="true" box="[199,315,1159,1185]" pageId="11" pageNumber="66">
<typeStatus id="6F47881EF73D116EFF1CC3DAFEA4A2B8" box="[199,310,1159,1185]" pageId="11" pageNumber="66" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="B5236067F73D116EFE9AC3DAFEE9A2B8" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFE9AC3DAFEE9A2B8" box="[321,379,1159,1185]" country="Japan" county="Honshu" latitude="35.405834" longLatPrecision="19" longitude="134.92029" municipality="Asago-shi" name="Male" pageId="11" pageNumber="66" stateProvince="Hyogo">Male</location>
(in alcohol),
<location id="B5236067F73D116EFDD6C3DAFD1CA2B8" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFDD6C3DAFD1CA2B8" box="[525,654,1159,1185]" country="Japan" county="Honshu" latitude="35.405834" longLatPrecision="19" longitude="134.92029" municipality="Asago-shi" name="Amago-ike" pageId="11" pageNumber="66" stateProvince="Hyogo">Amago-ike</location>
,
<location id="B5236067F73D116EFD4CC3DAFC8EA2B8" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFD4CC3DAFC8EA2B8" box="[663,796,1159,1185]" country="Japan" county="Honshu" latitude="35.405834" longLatPrecision="19" longitude="134.92029" municipality="Asago-shi" name="Takenouchi" pageId="11" pageNumber="66" stateProvince="Hyogo">Takenouchi</location>
,
<location id="B5236067F73D116EFCFEC3DAFC47A2B8" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFCFEC3DAFC47A2B8" box="[805,981,1159,1185]" country="Japan" county="Honshu" latitude="35.405834" longLatPrecision="19" longitude="134.92029" municipality="Asago-shi" name="Wadayama-cho" pageId="11" pageNumber="66" stateProvince="Hyogo">Wadayama-cho</location>
,
<collectingMunicipality id="5027ACC6F73D116EFC05C3DAFBC0A2B8" box="[990,1106,1159,1185]" pageId="11" pageNumber="66">Asago-shi</collectingMunicipality>
,
<collectingRegion id="7238F85EF73D116EFB86C3DAFB7AA2B8" box="[1117,1256,1159,1185]" country="Japan" name="Hyogo" pageId="11" pageNumber="66">Hyogo Pref.</collectingRegion>
,
<collectingCounty id="59224E30F73D116EFB2AC3DAFAD9A2B8" box="[1265,1355,1159,1185]" pageId="11" pageNumber="66">Honshu</collectingCounty>
,
<collectingCountry id="C8EB762CF73D116EFA8EC3DAFA0AA2B8" box="[1365,1432,1159,1185]" name="Japan" pageId="11" pageNumber="66">Japan</collectingCountry>
,
<geoCoordinate id="D5C8507BF73D116EFF4CC3F7FEB2A2DC" box="[151,288,1194,1221]" degrees="35" direction="north" minutes="24" orientation="latitude" pageId="11" pageNumber="66" precision="15" seconds="21" value="35.405834">35°2421”N</geoCoordinate>
<geoCoordinate id="D5C8507BF73D116EFEFCC3F7FE2CA2DC" box="[295,446,1194,1221]" degrees="134" direction="east" minutes="55" orientation="longitude" pageId="11" pageNumber="66" precision="15" seconds="13" value="134.92029">134°5513”E</geoCoordinate>
, alt.
<quantity id="77049B59F73D116EFE28C3F7FDAEA2DC" box="[499,572,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="66" unit="m" value="250.0">
<elevation id="3BD1D18FF73D116EFE28C3F7FDAEA2DC" box="[499,572,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="66" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<date id="C442107CF73D116EFD93C3F6FD52A2DC" box="[584,704,1195,1221]" pageId="11" pageNumber="66" value="2004-04-16">
<collectingDate id="D406E994F73D116EFD93C3F6FD52A2DC" box="[584,704,1195,1221]" pageId="11" pageNumber="66" value="2004-04-16">16.iv.2004</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFD17C3F6FCBEA2DC" box="[716,812,1195,1221]" pageId="11" pageNumber="66">K. Inazu</collectorName>
(
<specimenCode id="E05A9EC7F73D116EFCE0C3F6FB98A2DC" box="[827,1034,1195,1221]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="11" pageNumber="66">LBM1410012601</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="B04336BCF73D116EFF1CC392FEA6A010" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="66">
<materialsCitation id="00943CE1F73D116EFF1CC392FB81A314" collectingDate="1986-04-20" collectingDateMax="1994-04-03" collectingDateMin="1986-04-20" collectionCode="OMNH, T" collectorName="I. Togashi" county="Shizuoka-shi" location="Honshu" municipality="Oma" pageId="11" pageNumber="66" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<emphasis id="8288EAAEF73D116EFF1CC392FDA7A2F1" bold="true" box="[199,565,1230,1257]" pageId="11" pageNumber="66">
<typeStatus id="6F47881EF73D116EFF1CC392FED3A2F0" box="[199,321,1231,1257]" pageId="11" pageNumber="66" type="paratype">Paratypes</typeStatus>
.
<location id="B5236067F73D116EFE8FC392FE24A2F0" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFE8FC392FE24A2F0" box="[340,438,1231,1257]" county="Shizuoka-shi" municipality="Oma" name="Honshu" pageId="11" pageNumber="66">Honshu</location>
:
<location id="B5236067F73D116EFE13C393FDA7A2F1" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFE13C393FDA7A2F1" box="[456,565,1230,1256]" county="Shizuoka-shi" municipality="Oma" name="Ishikawa" pageId="11" pageNumber="66">Ishikawa</location>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFD91C392FD0DA2F0" box="[586,671,1231,1257]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
,
<location id="B5236067F73D116EFD6AC392FCA0A2F0" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFD6AC392FCA0A2F0" box="[689,818,1231,1257]" county="Shizuoka-shi" municipality="Oma" name="Sannomiya" pageId="11" pageNumber="66">Sannomiya</location>
,
<location id="B5236067F73D116EFC98C392FC7BA2F0" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFC98C392FC7BA2F0" box="[835,1001,1231,1257]" county="Shizuoka-shi" municipality="Oma" name="Tsurugi-machi" pageId="11" pageNumber="66">Tsurugi-machi</location>
,
<location id="B5236067F73D116EFC20C392FB18A2F0" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFC20C392FB18A2F0" box="[1019,1162,1231,1257]" county="Shizuoka-shi" municipality="Oma" name="Hakusan-shi" pageId="11" pageNumber="66">Hakusan-shi</location>
,
<date id="C442107CF73D116EFB47C392FA87A2F0" box="[1180,1301,1231,1257]" pageId="11" pageNumber="66" value="1986-04-20">
<collectingDate id="D406E994F73D116EFB47C392FA87A2F0" box="[1180,1301,1231,1257]" pageId="11" pageNumber="66" value="1986-04-20">20.iv.1986</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFAFDC392FA0EA2F0" box="[1318,1436,1231,1257]" pageId="11" pageNumber="66">I. Togashi</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFF44C3AEFF69A314" box="[159,251,1267,1293]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="11" pageNumber="66" type="Museum">OMNH</collectionCode>
).
<emphasis id="8288EAAEF73D116EFED0C3AFFEEBA315" bold="true" box="[267,377,1266,1292]" pageId="11" pageNumber="66">
<location id="B5236067F73D116EFED0C3AFFEEBA315" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFED0C3AFFEEBA315" box="[267,377,1266,1292]" county="Shizuoka-shi" municipality="Oma" name="Shizuoka" pageId="11" pageNumber="66">Shizuoka</location>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFE5CC3AEFE4CA314" box="[391,478,1267,1293]" count="2" pageId="11" pageNumber="66" type="male">2 males</specimenCount>
,
<location id="B5236067F73D116EFE3CC3AEFD36A314" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFE3CC3AEFD36A314" box="[487,676,1267,1293]" county="Shizuoka-shi" municipality="Oma" name="Warashina-gawa" pageId="11" pageNumber="66">Warashina-gawa</location>
,
<collectingMunicipality id="5027ACC6F73D116EFD77C3A9FD77A316" box="[684,741,1268,1295]" pageId="11" pageNumber="66">Ôma</collectingMunicipality>
,
<collectingCounty id="59224E30F73D116EFD34C3AEFC16A314" box="[751,900,1267,1293]" pageId="11" pageNumber="66">Shizuoka-shi</collectingCounty>
,
<date id="C442107CF73D116EFC56C3AEFC6AA314" box="[909,1016,1267,1293]" pageId="11" pageNumber="66" value="1994-04-03">
<collectingDate id="D406E994F73D116EFC56C3AEFC6AA314" box="[909,1016,1267,1293]" pageId="11" pageNumber="66" value="1994-04-03">3.iv.1994</collectingDate>
</date>
,
<collectionCode id="D6EDAE79F73D116EFBDAC3AEFB81A314" box="[1025,1043,1267,1293]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="66">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFBC7C3AEFCB9A328" collectingDate="2004-03-24" collectionCode="T" collectorName="Utaugenotaki" county="Fujieda-shi" location="Hattori" municipality="Kurata" pageId="11" pageNumber="66" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="B5236067F73D116EFBC7C3AEFBF9A314" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFBC7C3AEFBF9A314" box="[1052,1131,1267,1293]" county="Fujieda-shi" municipality="Kurata" name="Hattori" pageId="11" pageNumber="66">Hattori</location>
(SPMN-IS-5975259753);
<specimenCount id="A6FAFD35F73D116EFF4CC24AFF7BA328" box="[151,233,1303,1329]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
,
<collectorName id="1D09536AF73D116EFF23C24AFE00A328" box="[248,402,1303,1329]" pageId="11" pageNumber="66">Utaugenotaki</collectorName>
,
<collectingMunicipality id="5027ACC6F73D116EFE7AC24AFE62A328" box="[417,496,1303,1329]" pageId="11" pageNumber="66">Kurata</collectingMunicipality>
,
<collectingCounty id="59224E30F73D116EFE24C24AFD12A328" box="[511,640,1303,1329]" pageId="11" pageNumber="66">Fujieda-shi</collectingCounty>
,
<date id="C442107CF73D116EFD54C24AFC99A328" box="[655,779,1303,1329]" pageId="11" pageNumber="66" value="2004-03-24">
<collectingDate id="D406E994F73D116EFD54C24AFC99A328" box="[655,779,1303,1329]" pageId="11" pageNumber="66" value="2004-03-24">24.iii.2004</collectingDate>
</date>
,
<collectionCode id="D6EDAE79F73D116EFCC2C24AFCB9A328" box="[793,811,1303,1329]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="66">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFCE1C24AFD0AA34C" collectingDate="2012-04-12" collectionCode="T" collectorName="Sugino" county="Nagahama-shi" location="Hattori" municipality="Kinomoto-cho" pageId="11" pageNumber="66" specimenCount="16" specimenCount-male="16" typeStatus="paratype">
<location id="B5236067F73D116EFCE1C24AFC1BA328" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFCE1C24AFC1BA328" box="[826,905,1303,1329]" county="Nagahama-shi" municipality="Kinomoto-cho" name="Hattori" pageId="11" pageNumber="66">Hattori</location>
(SPMN-IS-59754).
<emphasis id="8288EAAEF73D116EFBADC24BFB2BA329" bold="true" box="[1142,1209,1302,1328]" pageId="11" pageNumber="66">
<location id="B5236067F73D116EFBADC24BFB2BA329" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFBADC24BFB2BA329" box="[1142,1209,1302,1328]" county="Nagahama-shi" municipality="Kinomoto-cho" name="Shiga" pageId="11" pageNumber="66">Shiga</location>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFB17C24AFAA5A328" box="[1228,1335,1303,1329]" count="16" pageId="11" pageNumber="66" type="male">16 males</specimenCount>
,
<collectorName id="1D09536AF73D116EFA9DC24AFA0AA328" box="[1350,1432,1303,1329]" pageId="11" pageNumber="66">Sugino</collectorName>
,
<collectingMunicipality id="5027ACC6F73D116EFF4CC266FEAFA34C" box="[151,317,1339,1365]" pageId="11" pageNumber="66">Kinomoto-cho</collectingMunicipality>
,
<collectingCounty id="59224E30F73D116EFE97C266FE63A34C" box="[332,497,1339,1365]" pageId="11" pageNumber="66">Nagahama-shi</collectingCounty>
,
<date id="C442107CF73D116EFE24C266FDE5A34C" box="[511,631,1339,1365]" pageId="11" pageNumber="66" value="2012-04-12">
<collectingDate id="D406E994F73D116EFE24C266FDE5A34C" box="[511,631,1339,1365]" pageId="11" pageNumber="66" value="2012-04-12">12.iv.2012</collectingDate>
</date>
,
<collectionCode id="D6EDAE79F73D116EFD5DC266FD0AA34C" box="[646,664,1339,1365]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="66">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFD7CC266FE91A384" collectingDate="1990-04-10" collectingDateMax="1990-04-11" collectingDateMin="1990-04-10" collectionCode="OMNH" collectorName="K. Tanida" county="Osaka" location="Hattori" municipality="Kyoto-shi" pageId="11" pageNumber="66" specimenCode="LBM1410012602-1410012617" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<location id="B5236067F73D116EFD7CC266FD64A34C" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFD7CC266FD64A34C" box="[679,758,1339,1365]" county="Osaka" municipality="Kyoto-shi" name="Hattori" pageId="11" pageNumber="66">Hattori</location>
(
<specimenCode id="E05A9EC7F73D116EFCD3C266FBE3A34C" box="[776,1137,1339,1365]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="11" pageNumber="66">LBM14100126021410012617</specimenCode>
).
<emphasis id="8288EAAEF73D116EFB5CC266FB42A34C" bold="true" box="[1159,1232,1339,1365]" pageId="11" pageNumber="66">
<location id="B5236067F73D116EFB5CC266FB42A34C" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFB5CC266FB42A34C" box="[1159,1232,1339,1365]" county="Osaka" municipality="Kyoto-shi" name="Kyoto" pageId="11" pageNumber="66">Kyoto</location>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFB39C266FAA6A34C" box="[1250,1332,1339,1365]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
,
<location id="B5236067F73D116EFA98C266FA0AA34C" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFA98C266FA0AA34C" box="[1347,1432,1339,1365]" county="Osaka" municipality="Kyoto-shi" name="Kibune" pageId="11" pageNumber="66">Kibune</location>
,
<collectingMunicipality id="5027ACC6F73D116EFF4CC202FE98A360" box="[151,266,1375,1401]" pageId="11" pageNumber="66">Kyoto-shi</collectingMunicipality>
,
<date id="C442107CF73D116EFECFC202FE19A360" box="[276,395,1375,1401]" pageId="11" pageNumber="66" value="1990-04-11">
<collectingDate id="D406E994F73D116EFECFC202FE19A360" box="[276,395,1375,1401]" pageId="11" pageNumber="66" value="1990-04-11">11.iv.1990</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFE4EC202FD90A360" box="[405,514,1375,1401]" pageId="11" pageNumber="66">K. Tanida</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFDD4C202FDF9A360" box="[527,619,1375,1401]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="11" pageNumber="66" type="Museum">OMNH</collectionCode>
).
<emphasis id="8288EAAEF73D116EFDA7C203FD5AA361" bold="true" box="[636,712,1374,1400]" pageId="11" pageNumber="66">
<collectingCounty id="59224E30F73D116EFDA7C203FD5AA361" box="[636,712,1374,1400]" pageId="11" pageNumber="66">Osaka</collectingCounty>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFD0EC202FC8DA360" box="[725,799,1375,1401]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
&amp;
<specimenCount id="A6FAFD35F73D116EFC9BC202FC32A360" box="[832,928,1375,1401]" count="1" pageId="11" pageNumber="66" type="female">1 female</specimenCount>
(couple), same locality,
<date id="C442107CF73D116EFB75C202FAB4A360" box="[1198,1318,1375,1401]" pageId="11" pageNumber="66" value="1990-04-10">
<collectingDate id="D406E994F73D116EFB75C202FAB4A360" box="[1198,1318,1375,1401]" pageId="11" pageNumber="66" value="1990-04-10">10.iv.1990</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFAEBC202FA0EA360" box="[1328,1436,1375,1401]" pageId="11" pageNumber="66">K. Tanida</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFF44C2DEFF69A384" box="[159,251,1411,1437]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="11" pageNumber="66" type="Museum">OMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D116EFED4C2DEFECCA3D8" collectingDate="1987-04-20" collectionCode="OMNH, Hyogo" collectorName="Ishi-kawa &amp; K. Tanida" location="Takihata-horikoshi" pageId="11" pageNumber="66" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A6FAFD35F73D116EFED4C2DEFEC8A384" box="[271,346,1411,1437]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
&amp;
<specimenCount id="A6FAFD35F73D116EFE5BC2DEFE73A384" box="[384,481,1411,1437]" count="1" pageId="11" pageNumber="66" type="female">1 female</specimenCount>
(couple),
<collectorName id="1D09536AF73D116EFD8EC2DEFD54A384" box="[597,710,1411,1437]" pageId="11" pageNumber="66">Ishi-kawa</collectorName>
,
<location id="B5236067F73D116EFD09C2DEFC3AA384" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFD09C2DEFC3AA384" box="[722,936,1411,1437]" name="Takihata-horikoshi" pageId="11" pageNumber="66">Takihata-horikoshi</location>
, Kawachi-nagano-shi,
<date id="C442107CF73D116EFB71C2DEFAB0A384" box="[1194,1314,1411,1437]" pageId="11" pageNumber="66" value="1987-04-20">
<collectingDate id="D406E994F73D116EFB71C2DEFAB0A384" box="[1194,1314,1411,1437]" pageId="11" pageNumber="66" value="1987-04-20">20.iv.1987</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFAF5C2DEFA0EA384" box="[1326,1436,1411,1437]" pageId="11" pageNumber="66">K. Tanida</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFF44C2FAFF69A3D8" box="[159,251,1447,1473]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="11" pageNumber="66" type="Museum">OMNH</collectionCode>
).
<emphasis id="8288EAAEF73D116EFECBC2FAFECCA3D8" bold="true" box="[272,350,1447,1473]" pageId="11" pageNumber="66">
<collectionCode id="D6EDAE79F73D116EFECBC2FAFECCA3D8" box="[272,350,1447,1473]" country="Japan" httpUri="http://grbio.org/cool/vy23-a9cj" name="Museum of Nature and Human Activities" pageId="11" pageNumber="66">Hyogo</collectionCode>
</emphasis>
</materialsCitation>
:
<materialsCitation id="00943CE1F73D116EFEB4C2FAFEA2A010" collectingDate="2017-04-23" collectionCode="LBM" collectorName="Shiraka-keikoku &amp; K. Nojima" country="Japan" county="Kaganino-cho" elevation="250" latitude="35.405834" location="Okayama" longLatPrecision="19" longitude="134.92029" municipality="Tomi-nishidani" pageId="11" pageNumber="66" specimenCode="LBM1410012618-1410012620, LBM1410012621- 1410012629" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Hyogo" typeStatus="paratype">
<specimenCount id="A6FAFD35F73D116EFEB4C2FAFE52A3D8" box="[367,448,1447,1473]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
,
<specimenCount id="A6FAFD35F73D116EFE15C2FAFDD2A3D8" box="[462,576,1447,1473]" count="2" pageId="11" pageNumber="66" type="female">2 females</specimenCount>
, same data as the holotype (
<specimenCode id="E05A9EC7F73D116EFC51C2FAFB61A3D8" box="[906,1267,1447,1473]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="11" pageNumber="66">LBM14100126181410012620</specimenCode>
).
<emphasis id="8288EAAEF73D116EFAD3C2FBFAEFA3D9" bold="true" box="[1288,1405,1446,1472]" pageId="11" pageNumber="66">
<location id="B5236067F73D116EFAD3C2FBFAEFA3D9" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFAD3C2FBFAEFA3D9" box="[1288,1405,1446,1472]" county="Kaganino-cho" municipality="Tomi-nishidani" name="Okayama" pageId="11" pageNumber="66">Okayama</location>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFA55C2FBFF49A3FC" count="5" pageId="11" pageNumber="66" type="male">5 males</specimenCount>
,
<specimenCount id="A6FAFD35F73D116EFF37C296FEF2A3FC" box="[236,352,1483,1509]" count="4" pageId="11" pageNumber="66" type="female">4 females</specimenCount>
,
<collectorName id="1D09536AF73D116EFEABC296FDBEA3FC" box="[368,556,1483,1509]" pageId="11" pageNumber="66">Shiraka-keikoku</collectorName>
,
<collectingMunicipality id="5027ACC6F73D116EFDE7C296FD78A3FC" box="[572,746,1483,1509]" pageId="11" pageNumber="66">Tomi-nishidani</collectingMunicipality>
,
<collectingCounty id="59224E30F73D116EFD21C296FC0EA3FC" box="[762,924,1483,1509]" pageId="11" pageNumber="66">Kaganino-cho</collectingCounty>
,
<date id="C442107CF73D116EFC70C296FBB6A3FC" box="[939,1060,1483,1509]" pageId="11" pageNumber="66" value="2017-04-23">
<collectingDate id="D406E994F73D116EFC70C296FBB6A3FC" box="[939,1060,1483,1509]" pageId="11" pageNumber="66" value="2017-04-23">23.iv.2017</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFBEEC296FB22A3FC" box="[1077,1200,1483,1509]" pageId="11" pageNumber="66">K. Nojima</collectorName>
(
<specimenCode id="E05A9EC7F73D116EFB1FC296FEBAA010" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="11" pageNumber="66">LBM1410012621 1410012629</specimenCode>
)
</materialsCitation>
;
</paragraph>
<paragraph id="B04336BCF73D1169FF1CC14FFBFCA6CC" blockId="11.[151,1437,150,2013]" lastBlockId="12.[151,1436,151,429]" lastPageId="12" lastPageNumber="67" pageId="11" pageNumber="66">
<emphasis id="8288EAAEF73D116EFF1CC14FFD1BA034" bold="true" box="[199,649,1554,1581]" pageId="11" pageNumber="66">Other specimens examined. Honshu:</emphasis>
<materialsCitation id="00943CE1F73D116EFD4FC14FFF68A048" collectingDate="1995-03-21" collectionCode="T, SPMN" collectorName="T. Hattori" location="Shizuoka" pageId="11" pageNumber="66" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka">
<emphasis id="8288EAAEF73D116EFD4FC14FFC90A035" bold="true" box="[660,770,1554,1580]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFD4FC14FFC90A035" box="[660,770,1554,1580]" country="Japan" name="Shizuoka" pageId="11" pageNumber="66">Shizuoka</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFCCEC14EFCF5A034" box="[789,871,1555,1581]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
,
<collectorName id="1D09536AF73D116EFCACC14EFC7CA034" box="[887,1006,1555,1581]" pageId="11" pageNumber="66">Nakazawa</collectorName>
,
<collectorName id="1D09536AF73D116EFC26C14EFB00A034" box="[1021,1170,1555,1581]" pageId="11" pageNumber="66">Shizuoka-shi</collectorName>
,
<date id="C442107CF73D116EFB7AC14FFA8FA034" box="[1185,1309,1554,1581]" pageId="11" pageNumber="66" value="1995-03-21">
<collectingDate id="D406E994F73D116EFB7AC14FFA8FA034" box="[1185,1309,1554,1581]" pageId="11" pageNumber="66" value="1995-03-21">21.iii.1995</collectingDate>
</date>
,
<collectionCode id="D6EDAE79F73D116EFAF7C14EFAACA034" box="[1324,1342,1555,1581]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="66">T</collectionCode>
.
<collectorName id="1D09536AF73D116EFA96C14EFA0EA034" box="[1357,1436,1555,1581]" pageId="11" pageNumber="66">Hattori</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFF44C16AFF60A048" box="[159,242,1591,1617]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D116EFEDCC16BFB59A048" box="[263,1227,1590,1617]" collectingDate="2023-04-10" collectingDateMax="2023-04-20" collectingDateMin="2023-04-10" collectorName="N. Kawase" location="Ibigawa-cho" municipality="Sakauchi-sakamoto" pageId="11" pageNumber="66" specimenCount="1" specimenCount-female="1" stateProvince="Gifu">
<emphasis id="8288EAAEF73D116EFEDCC16BFEAFA049" bold="true" box="[263,317,1590,1616]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFEDCC16BFEAFA049" box="[263,317,1590,1616]" country="Japan" name="Gifu" pageId="11" pageNumber="66">Gifu</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFE94C16AFE27A048" box="[335,437,1591,1617]" count="1" pageId="11" pageNumber="66" type="female">1 female</specimenCount>
,
<collectingMunicipality id="5027ACC6F73D116EFE19C16AFD32A048" box="[450,672,1591,1617]" pageId="11" pageNumber="66">Sakauchi-sakamoto</collectingMunicipality>
,
<location id="B5236067F73D116EFD75C16AFCAFA048" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFD75C16AFCAFA048" box="[686,829,1591,1617]" municipality="Sakauchi-sakamoto" name="Ibigawa-cho" pageId="11" pageNumber="66" stateProvince="Gifu">Ibigawa-cho</location>
,
<date id="C442107CF73D116EFC91C16AFC7EA048" box="[842,1004,1591,1617]" pageId="11" pageNumber="66" value="2023-04-10" valueMax="2023-04-20" valueMin="2023-04-10">
<collectingDate id="D406E994F73D116EFC91C16AFC7EA048" box="[842,1004,1591,1617]" pageId="11" pageNumber="66" value="2023-04-10" valueMax="2023-04-20" valueMin="2023-04-10">1020.iv.2023</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFC21C16AFBE5A048" box="[1018,1143,1591,1617]" pageId="11" pageNumber="66">N. Kawase</collectorName>
(NKa)
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFB03C16BFB98A06C" collectingDate="2008-04-06" collectingDateMax="2008-04-16" collectingDateMin="2008-04-06" collectorName="H. Morita" county="Kanzan" location="Yokkaichi-shi" municipality="Suizawa-cho" pageId="11" pageNumber="66" specimenCount="15" specimenCount-female="5" specimenCount-male="10" stateProvince="Mie">
<emphasis id="8288EAAEF73D116EFB03C16BFA95A049" bold="true" box="[1240,1287,1590,1616]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFB03C16BFA95A049" box="[1240,1287,1590,1616]" country="Japan" name="Mie" pageId="11" pageNumber="66">Mie</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFAC3C16AFA13A048" box="[1304,1409,1591,1617]" count="10" pageId="11" pageNumber="66" type="male">10 males</specimenCount>
,
<specimenCount id="A6FAFD35F73D116EFA55C16BFF63A06C" count="5" pageId="11" pageNumber="66" type="female">5 females</specimenCount>
,
<collectingCounty id="59224E30F73D116EFF26C106FEC7A06C" box="[253,341,1627,1653]" pageId="11" pageNumber="66">Kanzan</collectingCounty>
,
<collectingMunicipality id="5027ACC6F73D116EFEBBC106FE61A06C" box="[352,499,1627,1653]" pageId="11" pageNumber="66">Suizawa-cho</collectingMunicipality>
,
<location id="B5236067F73D116EFE25C106FD0FA06C" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFE25C106FD0FA06C" box="[510,669,1627,1653]" county="Kanzan" municipality="Suizawa-cho" name="Yokkaichi-shi" pageId="11" pageNumber="66" stateProvince="Mie">Yokkaichi-shi</location>
,
<date id="C442107CF73D116EFD72C106FCAFA06C" box="[681,829,1627,1653]" pageId="11" pageNumber="66" value="2008-04-06" valueMax="2008-04-16" valueMin="2008-04-06">
<collectingDate id="D406E994F73D116EFD72C106FCAFA06C" box="[681,829,1627,1653]" pageId="11" pageNumber="66" value="2008-04-06" valueMax="2008-04-16" valueMin="2008-04-06">616.iv.2008</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFC92C106FC25A06C" box="[841,951,1627,1653]" pageId="11" pageNumber="66">H. Morita</collectorName>
(NKa)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D116EFBCDC106FC81A080" collectingDate="2009-04-06" collectingDateMax="2009-04-16" collectingDateMin="2009-04-06" collectionCode="SPMN" collectorName="H. Morita" pageId="11" pageNumber="66" specimenCount="3" specimenCount-male="3" stateProvince="Mie">
<specimenCount id="A6FAFD35F73D116EFBCDC106FBFDA06C" box="[1046,1135,1627,1653]" count="3" pageId="11" pageNumber="66" type="male">3 males</specimenCount>
,
<collectorName id="1D09536AF73D116EFBA0C106FABBA06C" box="[1147,1321,1627,1653]" pageId="11" pageNumber="66">Miyazuma-kyo</collectorName>
,
<collectorName id="1D09536AF73D116EFAEFC106FF50A080" pageId="11" pageNumber="66">Suizawa-cho</collectorName>
,
<collectorName id="1D09536AF73D116EFF15C122FEFFA080" box="[206,365,1663,1689]" pageId="11" pageNumber="66">Yokkaichi-shi</collectorName>
,
<date id="C442107CF73D116EFEA1C122FDB9A080" box="[378,555,1663,1689]" pageId="11" pageNumber="66" value="2009-04-06" valueMax="2009-04-16" valueMin="2009-04-06">
<collectingDate id="D406E994F73D116EFEA1C122FDB9A080" box="[378,555,1663,1689]" pageId="11" pageNumber="66" value="2009-04-06" valueMax="2009-04-16" valueMin="2009-04-06">6.iv16.iv.2009</collectingDate>
</date>
, H.
<collectorName id="1D09536AF73D116EFD87C122FD3AA080" box="[604,680,1663,1689]" pageId="11" pageNumber="66">Morita</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFD63C122FC99A080" box="[696,779,1663,1689]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFCC4C123FD4CA0A4" collectingDate="2010-04-26" collectingDateMax="2010-05-17" collectingDateMin="2010-04-26" collectorName="N. Kawase" county="Nakanokawachi" location="Nagahama-shi" municipality="Yogo-cho" pageId="11" pageNumber="66" specimenCount="18" specimenCount-female="2" specimenCount-male="16" stateProvince="Shiga">
<emphasis id="8288EAAEF73D116EFCC4C123FCF0A081" bold="true" box="[799,866,1662,1688]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFCC4C123FCF0A081" box="[799,866,1662,1688]" country="Japan" name="Shiga" pageId="11" pageNumber="66">Shiga</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFCA9C122FC49A080" box="[882,987,1663,1689]" count="16" pageId="11" pageNumber="66" type="male">16 males</specimenCount>
,
<specimenCount id="A6FAFD35F73D116EFC33C122FBCAA080" box="[1000,1112,1663,1689]" count="2" pageId="11" pageNumber="66" type="female">2 females</specimenCount>
,
<collectingCounty id="59224E30F73D116EFBBEC122FA8EA080" box="[1125,1308,1663,1689]" pageId="11" pageNumber="66">Nakanokawachi</collectingCounty>
,
<collectingMunicipality id="5027ACC6F73D116EFAF3C122FA0AA080" box="[1320,1432,1663,1689]" pageId="11" pageNumber="66">Yogo-cho</collectingMunicipality>
,
<location id="B5236067F73D116EFF4CC1FEFEAEA0A4" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFF4CC1FEFEAEA0A4" box="[151,316,1699,1725]" county="Nakanokawachi" municipality="Yogo-cho" name="Nagahama-shi" pageId="11" pageNumber="66" stateProvince="Shiga">Nagahama-shi</location>
,
<date id="C442107CF73D116EFE92C1FEFD92A0A4" box="[329,512,1699,1725]" pageId="11" pageNumber="66" value="2010-04-26" valueMax="2010-05-17" valueMin="2010-04-26">
<collectingDate id="D406E994F73D116EFE92C1FEFD92A0A4" box="[329,512,1699,1725]" pageId="11" pageNumber="66" value="2010-04-26" valueMax="2010-05-17" valueMin="2010-04-26">26.iv17.v.2010</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFDD6C1FEFD18A0A4" box="[525,650,1699,1725]" pageId="11" pageNumber="66">N. Kawase</collectorName>
(NKa)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D116EFD30C1FEFED1A0F8" collectingDate="2005-04-21" collectorName="N. Kawase" county="Okawara" location="Koka-shi" municipality="Tsuchiyama-cho" pageId="11" pageNumber="66" specimenCount="1" specimenCount-male="1" stateProvince="Shiga">
<specimenCount id="A6FAFD35F73D116EFD30C1FEFCA9A0A4" box="[747,827,1699,1725]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
,
<collectingCounty id="59224E30F73D116EFC93C1F9FC22A0A6" box="[840,944,1700,1727]" pageId="11" pageNumber="66">Ôkawara</collectingCounty>
,
<collectingMunicipality id="5027ACC6F73D116EFC67C1FEFBEAA0A4" box="[956,1144,1699,1725]" pageId="11" pageNumber="66">Tsuchiyama-cho</collectingMunicipality>
,
<location id="B5236067F73D116EFB5EC1FEFB7DA0A4" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFB5EC1FEFB7DA0A4" box="[1157,1263,1699,1725]" county="Okawara" municipality="Tsuchiyama-cho" name="Koka-shi" pageId="11" pageNumber="66" stateProvince="Shiga">Kôka-shi</location>
,
<date id="C442107CF73D116EFB20C1FFFAE6A0A4" box="[1275,1396,1698,1725]" pageId="11" pageNumber="66" value="2005-04-21">
<collectingDate id="D406E994F73D116EFB20C1FFFAE6A0A4" box="[1275,1396,1698,1725]" pageId="11" pageNumber="66" value="2005-04-21">21.iv.2005</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFA5AC1FEFF62A0F8" pageId="11" pageNumber="66">N. Kawase</collectorName>
(NKa)
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFE94C19AFF68A11C" collectingDate="2002-04-05" collectionCode="SPMN" collectorName="K. Inazu" location="Hyogo" pageId="11" pageNumber="66" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hyogo">
<emphasis id="8288EAAEF73D116EFE94C19AFE0FA0F8" bold="true" box="[335,413,1735,1761]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFE94C19AFE0FA0F8" box="[335,413,1735,1761]" country="Japan" name="Hyogo" pageId="11" pageNumber="66">Hyogo</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFE77C19AFE69A0F8" box="[428,507,1735,1761]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
,
<specimenCount id="A6FAFD35F73D116EFDD3C19AFDFFA0F8" box="[520,621,1735,1761]" count="1" pageId="11" pageNumber="66" type="female">1 female</specimenCount>
,
<collectorName id="1D09536AF73D116EFDA2C19AFD75A0F8" box="[633,743,1735,1761]" pageId="11" pageNumber="66">Itoi-gawa</collectorName>
,
<collectorName id="1D09536AF73D116EFD28C19AFC31A0F8" box="[755,931,1735,1761]" pageId="11" pageNumber="66">Wadayama-cho</collectorName>
,
<collectorName id="1D09536AF73D116EFC75C19AFBB0A0F8" box="[942,1058,1735,1761]" pageId="11" pageNumber="66">Asago-shi</collectorName>
,
<date id="C442107CF73D116EFBF4C19BFB08A0F8" box="[1071,1178,1734,1761]" pageId="11" pageNumber="66" value="2002-04-05">
<collectingDate id="D406E994F73D116EFBF4C19BFB08A0F8" box="[1071,1178,1734,1761]" pageId="11" pageNumber="66" value="2002-04-05">5.iv.2002</collectingDate>
</date>
, K.
<collectorName id="1D09536AF73D116EFB11C19AFA9AA0F8" box="[1226,1288,1735,1761]" pageId="11" pageNumber="66">Inazu</collectorName>
(in glycerin) (
<collectionCode id="D6EDAE79F73D116EFF44C1B6FF60A11C" box="[159,242,1771,1797]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D116EFEDDC1B6FC5CA11C" box="[262,974,1771,1797]" collectingDate="2004-04-06" collectionCode="SPMN" collectorName="Inazu" location="Hyogo" pageId="11" pageNumber="66" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Hyogo">
<specimenCount id="A6FAFD35F73D116EFEDDC1B6FEF2A11C" box="[262,352,1771,1797]" count="2" pageId="11" pageNumber="66" type="male">2 males</specimenCount>
,
<specimenCount id="A6FAFD35F73D116EFEB7C1B6FE4EA11C" box="[364,476,1771,1797]" count="2" pageId="11" pageNumber="66" type="female">2 females</specimenCount>
, same locality,
<date id="C442107CF73D116EFD50C1B6FD67A11C" box="[651,757,1771,1797]" pageId="11" pageNumber="66" value="2004-04-06">
<collectingDate id="D406E994F73D116EFD50C1B6FD67A11C" box="[651,757,1771,1797]" pageId="11" pageNumber="66" value="2004-04-06">6.iv.2004</collectingDate>
</date>
, K.
<collectorName id="1D09536AF73D116EFCFEC1B6FCF1A11C" box="[805,867,1771,1797]" pageId="11" pageNumber="66">Inazu</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFCA8C1B6FC54A11C" box="[883,966,1771,1797]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D116EFC01C1B7FE0BA130" collectingDate="1984-04-25" collectionCode="SPMN" collectorName="H. Nishimoto" pageId="11" pageNumber="66" specimenCount="5" specimenCount-male="5" stateProvince="Hyogo">
<specimenCount id="A6FAFD35F73D116EFC01C1B7FBA6A11C" box="[986,1076,1770,1797]" count="5" pageId="11" pageNumber="66" type="male">5 males</specimenCount>
,
<collectorName id="1D09536AF73D116EFB9BC1B6FB09A11C" box="[1088,1179,1771,1797]" pageId="11" pageNumber="66">Wakasu</collectorName>
,
<collectorName id="1D09536AF73D116EFB7CC1B6FA86A11C" box="[1191,1300,1771,1797]" pageId="11" pageNumber="66">Sayô-cho</collectorName>
,
<date id="C442107CF73D116EFAC4C1B7FA05A11C" box="[1311,1431,1770,1797]" pageId="11" pageNumber="66" value="1984-04-25">
<collectingDate id="D406E994F73D116EFAC4C1B7FA05A11C" box="[1311,1431,1770,1797]" pageId="11" pageNumber="66" value="1984-04-25">25.iv.1984</collectingDate>
</date>
, H.
<collectorName id="1D09536AF73D116EFF61C052FEA2A130" box="[186,304,1807,1833]" pageId="11" pageNumber="66">Nishimoto</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFEE4C052FE00A130" box="[319,402,1807,1833]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFE7FC053FADFA130" box="[420,1357,1806,1833]" collectingDate="1985-04-25" collectionCode="SPMN" collectorName="Takihata &amp; Nishimoto" location="Osaka" pageId="11" pageNumber="66" specimenCount="2" specimenCount-male="2" stateProvince="Osaka">
<emphasis id="8288EAAEF73D116EFE7FC053FE62A131" bold="true" box="[420,496,1806,1832]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFE7FC053FE62A131" box="[420,496,1806,1832]" country="Japan" name="Osaka" pageId="11" pageNumber="66">Osaka</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFE24C052FDCAA130" box="[511,600,1807,1833]" count="2" pageId="11" pageNumber="66" type="male">2 males</specimenCount>
,
<collectorName id="1D09536AF73D116EFDB8C052FD54A130" box="[611,710,1807,1833]" pageId="11" pageNumber="66">Takihata</collectorName>
, Kawachi-nagano-shi,
<date id="C442107CF73D116EFC1CC053FBD2A130" box="[967,1088,1806,1833]" pageId="11" pageNumber="66" value="1985-04-25">
<collectingDate id="D406E994F73D116EFC1CC053FBD2A130" box="[967,1088,1806,1833]" pageId="11" pageNumber="66" value="1985-04-25">25.iv.1985</collectingDate>
</date>
, H.
<collectorName id="1D09536AF73D116EFBB6C052FB71A130" box="[1133,1251,1807,1833]" pageId="11" pageNumber="66">Nishimoto</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFB28C052FAD4A130" box="[1267,1350,1807,1833]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFA83C052FD66A154" collectingDate="1984-04-12" collectionCode="SPMN" collectorName="Gojyo-shi &amp; Nishimoto" location="Nara" pageId="11" pageNumber="66" specimenCount="1" specimenCount-male="1" stateProvince="Nara">
<emphasis id="8288EAAEF73D116EFA83C052FA07A130" bold="true" box="[1368,1429,1807,1833]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFA83C052FA07A130" box="[1368,1429,1807,1833]" country="Japan" name="Nara" pageId="11" pageNumber="66">Nara</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFF4CC06EFF77A154" box="[151,229,1843,1869]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
,
<collectorName id="1D09536AF73D116EFF2BC06EFEF1A154" box="[240,355,1843,1869]" pageId="11" pageNumber="66">Gojyô-shi</collectorName>
,
<date id="C442107CF73D116EFEB5C06EFE75A154" box="[366,487,1843,1869]" pageId="11" pageNumber="66" value="1984-04-12">
<collectingDate id="D406E994F73D116EFEB5C06EFE75A154" box="[366,487,1843,1869]" pageId="11" pageNumber="66" value="1984-04-12">12.iv.1984</collectingDate>
</date>
, H.
<collectorName id="1D09536AF73D116EFDCFC06EFD18A154" box="[532,650,1843,1869]" pageId="11" pageNumber="66">Nishimoto</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFD41C06EFD7FA154" box="[666,749,1843,1869]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFD25C06FFEF7A168" collectingDate="1984-04-24" collectionCode="SPMN" collectorName="Wakasa-cho &amp; Nishimoto" location="Tottori" pageId="11" pageNumber="66" specimenCount="3" specimenCount-male="3" stateProvince="Tottori">
<emphasis id="8288EAAEF73D116EFD25C06FFCC3A155" bold="true" box="[766,849,1842,1868]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFD25C06FFCC3A155" box="[766,849,1842,1868]" country="Japan" name="Tottori" pageId="11" pageNumber="66">Tottori</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFCBBC06EFC2BA154" box="[864,953,1843,1869]" count="3" pageId="11" pageNumber="66" type="male">3 males</specimenCount>
,
<collectorName id="1D09536AF73D116EFC1FC06EFBDDA154" box="[964,1103,1843,1869]" pageId="11" pageNumber="66">Wakasa-cho</collectorName>
,
<date id="C442107CF73D116EFB81C06EFB41A154" box="[1114,1235,1843,1869]" pageId="11" pageNumber="66" value="1984-04-24">
<collectingDate id="D406E994F73D116EFB81C06EFB41A154" box="[1114,1235,1843,1869]" pageId="11" pageNumber="66" value="1984-04-24">24.iv.1984</collectingDate>
</date>
, H.
<collectorName id="1D09536AF73D116EFADBC06EFAE4A154" box="[1280,1398,1843,1869]" pageId="11" pageNumber="66">Nishimoto</collectorName>
(in glycerin) (
<collectionCode id="D6EDAE79F73D116EFED1C00AFECFA168" box="[266,349,1879,1905]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D116EFEB4C00AFCABA168" box="[367,825,1879,1905]" collectingDate="1984-04-24" collectionCode="SPMN" collectorName="Wakasa-cho &amp; Nishimoto" location="Tottori" pageId="11" pageNumber="66" specimenCount="3" specimenCount-male="3" stateProvince="Tottori">
<specimenCount id="A6FAFD35F73D116EFEB4C00AFE55A168" box="[367,455,1879,1905]" count="3" pageId="11" pageNumber="66" type="male">3 males</specimenCount>
, same data (in glycerin) (
<collectionCode id="D6EDAE79F73D116EFD05C00AFCA3A168" box="[734,817,1879,1905]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="00943CE1F73D116EFC99C00BFDB5A18C" collectingDate="2019-04-13" collectorName="K. Nojima" county="Ombara" location="Kagamino-cho" municipality="Kamisaibara" pageId="11" pageNumber="66" specimenCount="1" specimenCount-male="1" stateProvince="Okayama">
<emphasis id="8288EAAEF73D116EFC99C00BFC25A169" bold="true" box="[834,951,1878,1904]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFC99C00BFC25A169" box="[834,951,1878,1904]" country="Japan" name="Okayama" pageId="11" pageNumber="66">Okayama</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFC1EC00AFB80A168" box="[965,1042,1879,1905]" count="1" pageId="11" pageNumber="66" type="male">1 male</specimenCount>
,
<collectingCounty id="59224E30F73D116EFBC7C00AFBEBA168" box="[1052,1145,1879,1905]" pageId="11" pageNumber="66">Ombara</collectingCounty>
,
<collectingMunicipality id="5027ACC6F73D116EFB5FC00AFA86A168" box="[1156,1300,1879,1905]" pageId="11" pageNumber="66">Kamisaibara</collectingMunicipality>
,
<location id="B5236067F73D116EFAC5C00AFF50A18C" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFAC5C00AFF50A18C" county="Ombara" municipality="Kamisaibara" name="Kagamino-cho" pageId="11" pageNumber="66" stateProvince="Okayama">Kagamino-cho</location>
,
<date id="C442107CF73D116EFF0AC026FEDBA18C" box="[209,329,1915,1941]" pageId="11" pageNumber="66" value="2019-04-13">
<collectingDate id="D406E994F73D116EFF0AC026FEDBA18C" box="[209,329,1915,1941]" pageId="11" pageNumber="66" value="2019-04-13">13.iv.2019</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFE83C026FE40A18C" box="[344,466,1915,1941]" pageId="11" pageNumber="66">K. Nojima</collectorName>
(NKa)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D116EFDEDC026FA05A18D" box="[566,1431,1914,1943]" collectingDate="2017-05-07" collectorName="K. Nojima &amp; Shikoku" location="Ogaya" pageId="11" pageNumber="66" specimenCount="4" specimenCount-male="4" stateProvince="Okayama">
<specimenCount id="A6FAFD35F73D116EFDEDC026FD00A18C" box="[566,658,1915,1941]" count="4" pageId="11" pageNumber="66" type="male">4 males</specimenCount>
,
<location id="B5236067F73D116EFD7BC021FD7EA18E" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFD7BC021FD7EA18E" box="[672,748,1916,1943]" name="Ogaya" pageId="11" pageNumber="66" stateProvince="Okayama">Ôgaya</location>
, Nishi-awakura-son,
<date id="C442107CF73D116EFC3AC026FBD6A18C" box="[993,1092,1915,1941]" pageId="11" pageNumber="66" value="2017-05-07">
<collectingDate id="D406E994F73D116EFC3AC026FBD6A18C" box="[993,1092,1915,1941]" pageId="11" pageNumber="66" value="2017-05-07">7.v.2017</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFB88C026FB5EA18C" box="[1107,1228,1915,1941]" pageId="11" pageNumber="66">K. Nojima</collectorName>
(NKa).
<collectorName id="1D09536AF73D116EFAF4C027FA05A18D" box="[1327,1431,1914,1940]" pageId="11" pageNumber="66">
<emphasis id="8288EAAEF73D116EFAF4C027FA05A18D" bold="true" box="[1327,1431,1914,1940]" pageId="11" pageNumber="66">Shikoku</emphasis>
</collectorName>
</materialsCitation>
<emphasis id="8288EAAEF73D116EFA4CC027FA0EA18D" bold="true" box="[1431,1436,1914,1940]" pageId="11" pageNumber="66">:</emphasis>
<materialsCitation id="00943CE1F73D116EFF4CC0C3FAB1A1A0" box="[151,1315,1950,1977]" collectingDate="2018-05-01" collectingDateMax="2018-05-10" collectingDateMin="2018-05-01" collectorName="E. Yamamoto" county="Izugataniyama" location="Kumakogen-cho" municipality="Nishidani" pageId="11" pageNumber="66" specimenCount="1" specimenCount-female="1" stateProvince="Ehime">
<emphasis id="8288EAAEF73D116EFF4CC0C3FF77A1A1" bold="true" box="[151,229,1950,1976]" pageId="11" pageNumber="66">
<collectingRegion id="7238F85EF73D116EFF4CC0C3FF77A1A1" box="[151,229,1950,1976]" country="Japan" name="Ehime" pageId="11" pageNumber="66">Ehime</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73D116EFF27C0C2FEFAA1A0" box="[252,360,1951,1977]" count="1" pageId="11" pageNumber="66" type="female">1 female</specimenCount>
,
<collectingCounty id="59224E30F73D116EFEA0C0C2FDB1A1A0" box="[379,547,1951,1977]" pageId="11" pageNumber="66">Izugataniyama</collectingCounty>
,
<collectingMunicipality id="5027ACC6F73D116EFDEDC0C2FD34A1A0" box="[566,678,1951,1977]" pageId="11" pageNumber="66">Nishidani</collectingMunicipality>
,
<location id="B5236067F73D116EFD62C0C2FCE5A1A0" LSID="urn:lsid:plazi:treatment:385587AAF73F1169FF4CC15AFD1BA7B4:B5236067F73D116EFD62C0C2FCE5A1A0" box="[697,887,1951,1977]" county="Izugataniyama" municipality="Nishidani" name="Kumakogen-cho" pageId="11" pageNumber="66" stateProvince="Ehime">Kumakôgen-cho</location>
,
<date id="C442107CF73D116EFC52C0C2FB84A1A0" box="[905,1046,1951,1977]" pageId="11" pageNumber="66" value="2018-05-01" valueMax="2018-05-10" valueMin="2018-05-01">
<collectingDate id="D406E994F73D116EFC52C0C2FB84A1A0" box="[905,1046,1951,1977]" pageId="11" pageNumber="66" value="2018-05-01" valueMax="2018-05-10" valueMin="2018-05-01">110.v.2018</collectingDate>
</date>
,
<collectorName id="1D09536AF73D116EFBF1C0C2FB58A1A0" box="[1066,1226,1951,1977]" pageId="11" pageNumber="66">E. Yamamoto</collectorName>
(NKa)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D116EFAECC0C2FA8FA1C4" collectingDate="2000-04-21" collectionCode="T, SPMN" collectorName="Namakusa-dani &amp; Odamiyama &amp; Uchiko-cho &amp; Yamamoto &amp; Ito &amp; A. Ohkawa" pageId="11" pageNumber="66" specimenCount="2" specimenCount-male="2" stateProvince="Ehime">
<specimenCount id="A6FAFD35F73D116EFAECC0C2FA0AA1A0" box="[1335,1432,1951,1977]" count="2" pageId="11" pageNumber="66" type="male">2 males</specimenCount>
,
<collectorName id="1D09536AF73D116EFF4CC09EFED8A1C4" box="[151,330,1987,2013]" pageId="11" pageNumber="66">Namakusa-dani</collectorName>
,
<collectorName id="1D09536AF73D116EFE82C09EFE77A1C4" box="[345,485,1987,2013]" pageId="11" pageNumber="66">Odamiyama</collectorName>
,
<collectorName id="1D09536AF73D116EFE2FC09EFDE9A1C4" box="[500,635,1987,2013]" pageId="11" pageNumber="66">Uchiko-cho</collectorName>
,
<date id="C442107CF73D116EFD50C09EFC91A1C4" box="[651,771,1987,2013]" pageId="11" pageNumber="66" value="2000-04-21">
<collectingDate id="D406E994F73D116EFD50C09EFC91A1C4" box="[651,771,1987,2013]" pageId="11" pageNumber="66" value="2000-04-21">21.iv.2000</collectingDate>
</date>
, E.
<collectorName id="1D09536AF73D116EFCEEC09EFC20A1C4" box="[821,946,1987,2013]" pageId="11" pageNumber="66">Yamamoto</collectorName>
,
<collectionCode id="D6EDAE79F73D116EFC1AC09EFC41A1C4" box="[961,979,1987,2013]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="66">T</collectionCode>
.
<collectorName id="1D09536AF73D116EFC39C09EFB93A1C4" box="[994,1025,1987,2013]" pageId="11" pageNumber="66">Ito</collectorName>
&amp;
<collectorName id="1D09536AF73D116EFBF7C09EFB3DA1C4" box="[1068,1199,1987,2013]" pageId="11" pageNumber="66">A. Ohkawa</collectorName>
(
<collectionCode id="D6EDAE79F73D116EFB18C09EFA84A1C4" box="[1219,1302,1987,2013]" pageId="11" pageNumber="66">SPMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="00943CE1F73D1169FAF6C09EFCD2A6A8" collectingDate="2000-04-29" collectionCode="SPMN" collectorName="Yamamoto" lastPageId="12" lastPageNumber="67" pageId="11" pageNumber="66" specimenCount="11" specimenCount-female="1" specimenCount-male="10" stateProvince="Ehime">
<specimenCount id="A6FAFD35F73D116EFAF6C09EFA0AA1C4" box="[1325,1432,1987,2013]" count="10" pageId="11" pageNumber="66" type="male">10 males</specimenCount>
,
<specimenCount id="A6FAFD35F73A1169FF4CC7CAFF6CA6A8" box="[151,254,151,177]" count="1" pageId="12" pageNumber="67" type="female">1 female</specimenCount>
, same locality,
<date id="C442107CF73A1169FE69C7CAFDB9A6A8" box="[434,555,151,177]" pageId="12" pageNumber="67" value="2000-04-29">
<collectingDate id="D406E994F73A1169FE69C7CAFDB9A6A8" box="[434,555,151,177]" pageId="12" pageNumber="67" value="2000-04-29">29.iv.2000</collectingDate>
</date>
, E.
<collectorName id="1D09536AF73A1169FD81C7CAFD46A6A8" box="[602,724,151,177]" pageId="12" pageNumber="67">Yamamoto</collectorName>
(
<collectionCode id="D6EDAE79F73A1169FD3EC7CAFCAAA6A8" box="[741,824,151,177]" pageId="12" pageNumber="67">SPMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="00943CE1F73A1169FC95C7CAFEF1A6CC" collectingDate="2020-04-21" collectingDateMax="2020-04-30" collectingDateMin="2020-04-21" collectionCode="SPMN" collectorName="E. Yamamoto" pageId="12" pageNumber="67" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Ehime">
<specimenCount id="A6FAFD35F73A1169FC95C7CAFC0DA6A8" box="[846,927,151,177]" count="1" pageId="12" pageNumber="67" type="male">1 male</specimenCount>
,
<specimenCount id="A6FAFD35F73A1169FC76C7CAFB8DA6A8" box="[941,1055,151,177]" count="3" pageId="12" pageNumber="67" type="female">3 females</specimenCount>
, same locality,
<date id="C442107CF73A1169FB08C7CAFAE4A6A8" box="[1235,1398,151,177]" pageId="12" pageNumber="67" value="2020-04-21" valueMax="2020-04-30" valueMin="2020-04-21">
<collectingDate id="D406E994F73A1169FB08C7CAFAE4A6A8" box="[1235,1398,151,177]" pageId="12" pageNumber="67" value="2020-04-21" valueMax="2020-04-30" valueMin="2020-04-21">2130.iv.2020</collectingDate>
</date>
,
<collectorName id="1D09536AF73A1169FA5FC7CAFE83A6CC" pageId="12" pageNumber="67">E. Yamamoto</collectorName>
(NKa)
</materialsCitation>
.
<materialsCitation id="00943CE1F73A1169FEB6C7E7FBF8A6CC" box="[365,1130,186,213]" collectingDate="2004-04-12" collectionCode="SPMN" collectorName="K. Nio" location="Kochi" pageId="12" pageNumber="67" specimenCount="1" specimenCount-male="1" stateProvince="Kochi">
<emphasis id="8288EAAEF73A1169FEB6C7E7FE27A6CD" bold="true" box="[365,437,186,212]" pageId="12" pageNumber="67">
<collectingRegion id="7238F85EF73A1169FEB6C7E7FE27A6CD" box="[365,437,186,212]" country="Japan" name="Kochi" pageId="12" pageNumber="67">Kochi</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F73A1169FE1FC7E6FD80A6CC" box="[452,530,187,213]" count="1" pageId="12" pageNumber="67" type="male">1 male</specimenCount>
,
<collectorName id="1D09536AF73A1169FDC6C7E6FD18A6CC" box="[541,650,187,213]" pageId="12" pageNumber="67">Befu-kyo</collectorName>
,
<collectorName id="1D09536AF73A1169FD4EC7E6FCB7A6CC" box="[661,805,187,213]" pageId="12" pageNumber="67">Monobe-son</collectorName>
,
<date id="C442107CF73A1169FCEBC7E6FC3BA6CC" box="[816,937,187,213]" pageId="12" pageNumber="67" value="2004-04-12">
<collectingDate id="D406E994F73A1169FCEBC7E6FC3BA6CC" box="[816,937,187,213]" pageId="12" pageNumber="67" value="2004-04-12">12.iv.2004</collectingDate>
</date>
, K.
<collectorName id="1D09536AF73A1169FC0CC7E6FB93A6CC" box="[983,1025,187,213]" pageId="12" pageNumber="67">Nio</collectorName>
(
<collectionCode id="D6EDAE79F73A1169FBD4C7E6FBF0A6CC" box="[1039,1122,187,213]" pageId="12" pageNumber="67">SPMN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F73A1169FF1CC783FB29A704" pageId="12" pageNumber="67" type="distribution">
<paragraph id="B04336BCF73A1169FF1CC783FB29A704" blockId="12.[151,1436,151,429]" pageId="12" pageNumber="67">
<emphasis id="8288EAAEF73A1169FF1CC783FE62A6E1" bold="true" box="[199,496,222,249]" pageId="12" pageNumber="67">Distribution and biology.</emphasis>
<taxonomicName id="77FC4D3FF73A1169FE2CC782FD5DA6E1" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[503,719,223,249]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="67" phylum="Arthropoda" rank="species" species="inazui" status="sp. nov.">
<emphasis id="8288EAAEF73A1169FE2CC782FD5DA6E1" box="[503,719,223,249]" italics="true" pageId="12" pageNumber="67">Rhyacophila inazui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F73A1169FD0DC782FCBEA6E0" box="[726,812,223,249]" pageId="12" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
is distributed in central to western Honshu and Shikoku (
<figureCitation id="28C72A39F73A1169FF44C65EFF6AA704" box="[159,248,259,285]" captionStart="FIGURE 9" captionStartId="19.[151,250,1739,1763]" captionTargetBox="[292,1296,194,1702]" captionTargetId="figure-16@19.[279,1309,181,1715]" captionTargetPageId="19" captionText="FIGURE 9. Distributions of species in the Rhyacophila clemens Species Group in Japan." figureDoi="http://doi.org/10.5281/zenodo.11119229" httpUri="https://zenodo.org/record/11119229/files/figure.png" pageId="12" pageNumber="67">Fig. 9C</figureCitation>
). The adults were collected near mountain streams in spring, late March to early May.
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F73A1169FF1CC67AFD1BA7B4" pageId="12" pageNumber="67" type="etymology">
<paragraph id="B04336BCF73A1169FF1CC67AFE27A790" blockId="12.[151,1436,151,429]" pageId="12" pageNumber="67">
<emphasis id="8288EAAEF73A1169FF1CC67AFEDCA758" bold="true" box="[199,334,295,321]" pageId="12" pageNumber="67">Etymology.</emphasis>
This species is dedicated to Mr. Kazuyuki Inazu, who mentioned this species as a possibly undescribed species based on its early flight season and wing color pattern (
<bibRefCitation id="D46D4B4DF73A1169FC2DC616FB13A77C" author="Inazu, K." box="[1014,1153,331,357]" pageId="12" pageNumber="67" pagination="59 - 104" refId="ref17485" refString="Inazu, K. (2008) Collection record of caddisflies in the Itoi River Hyogo over 5 years (2001 - 2005) - Notes on emergence periods, classification, distribution, and ecology of adults and larvae. Hyogo Freshwater Biology, 60, 59 - 104. [in Japanese]" type="journal article" year="2008">Inazu, 2008</bibRefCitation>
). The species epithet is a noun in the genitive case.
</paragraph>
<paragraph id="B04336BCF73A1169FF1CC6CEFD1BA7B4" blockId="12.[151,1436,151,429]" box="[199,649,403,429]" pageId="12" pageNumber="67">
<emphasis id="8288EAAEF73A1169FF1CC6CEFEEEA7B4" bold="true" box="[199,380,403,429]" pageId="12" pageNumber="67">Japanese name</emphasis>
. Inazu-nagare-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>