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<document id="226B471622D6D4FA179A092356BCC7FC" ID-DOI="10.11646/zootaxa.5447.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="11119209" ID-ZooBank="3A9842EC-F3D2-4839-9731-AA79A27B25CB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1714984939816" checkinUser="plazi" docAuthor="Kawase, Naoki" docDate="2024" docId="385587AAF7261174FF4CC63EFD0CA06C" docLanguage="en" docName="zootaxa.5447.1.3.pdf" docOrigin="Zootaxa 5447 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5447.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophila furcicauda Kawase 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="72" masterDocId="C46CFFD2F7361165FFDBC75DFF92A619" masterDocTitle="Redescription of Rhyacophila clemens Tsuda 1940 (Trichoptera, Rhyacophilidae), with descriptions of five new, allied species from Japan" masterLastPageNumber="76" masterPageNumber="55" pageNumber="71" updateTime="1715030625241" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="398BC26179F994E1C83F21C79812DDEB">Redescription of Rhyacophila clemens Tsuda 1940 (Trichoptera, Rhyacophilidae), with descriptions of five new, allied species from Japan</mods:title>
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<treatment id="385587AAF7261174FF4CC63EFD0CA06C" ID-DOI="http://doi.org/10.5281/zenodo.11119203" ID-Zenodo-Dep="11119203" LSID="urn:lsid:plazi:treatment:385587AAF7261174FF4CC63EFD0CA06C" httpUri="http://treatment.plazi.org/id/385587AAF7261174FF4CC63EFD0CA06C" lastPageId="17" lastPageNumber="72" pageId="16" pageNumber="71">
<subSubSection id="F8E66537F7261175FF4CC63EFDB3A767" box="[151,545,355,382]" pageId="16" pageNumber="71" type="nomenclature">
<paragraph id="B04336BCF7261175FF4CC63EFDB3A767" blockId="16.[151,545,355,418]" box="[151,545,355,382]" pageId="16" pageNumber="71">
<heading id="EB0B81D0F7261175FF4CC63EFDB3A767" bold="true" box="[151,545,355,382]" fontSize="11" level="1" pageId="16" pageNumber="71" reason="1">
<emphasis id="8288EAAEF7261175FF4CC63EFDB3A767" bold="true" box="[151,545,355,382]" pageId="16" pageNumber="71">
<taxonomicName id="77FC4D3FF7261175FF4CC63EFE2EA767" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[151,444,355,382]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="furcicauda" status="sp. nov.">
<emphasis id="8288EAAEF7261175FF4CC63EFE2EA767" bold="true" box="[151,444,355,382]" italics="true" pageId="16" pageNumber="71">Rhyacophila furcicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F7261175FE18C639FDB3A767" box="[451,545,356,382]" pageId="16" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="F8E66537F7261175FF4CC6DAFC99A7FD" pageId="16" pageNumber="71" type="description">
<paragraph id="B04336BCF7261175FF4CC6DAFEC8A7BB" blockId="16.[151,545,355,418]" box="[151,346,391,418]" pageId="16" pageNumber="71">
(
<figureCitation id="28C72A39F7261175FF44C6DAFEB1A7BB" box="[159,291,391,418]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="16" pageNumber="71">Figs 7A7J</figureCitation>
,
<figureCitation id="28C72A39F7261175FEF4C6D5FEC1A7BB" box="[303,339,392,418]" captionStart="FIGURE 9" captionStartId="19.[151,250,1739,1763]" captionTargetBox="[292,1296,194,1702]" captionTargetId="figure-16@19.[279,1309,181,1715]" captionTargetPageId="19" captionText="FIGURE 9. Distributions of species in the Rhyacophila clemens Species Group in Japan." figureDoi="http://doi.org/10.5281/zenodo.11119229" httpUri="https://zenodo.org/record/11119229/files/figure.png" pageId="16" pageNumber="71">9D</figureCitation>
)
</paragraph>
<paragraph id="B04336BCF7261175FF4CC691FC99A7FD" blockId="16.[151,779,460,484]" box="[151,779,460,484]" pageId="16" pageNumber="71">
<emphasis id="8288EAAEF7261175FF4CC691FE7BA7FD" bold="true" box="[151,489,460,484]" pageId="16" pageNumber="71">
<taxonomicName id="77FC4D3FF7261175FF4CC691FEC1A7FD" box="[151,339,460,484]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="undefined-2">
<emphasis id="8288EAAEF7261175FF4CC691FE89A7FD" bold="true" box="[151,283,460,484]" italics="true" pageId="16" pageNumber="71">Rhyacophila</emphasis>
sp. 2
</taxonomicName>
(
<taxonomicName id="77FC4D3FF7261175FEB9C691FE24A7FD" authorityName="Tsuda" authorityYear="1940" box="[354,438,460,484]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="clemens">
<emphasis id="8288EAAEF7261175FEB9C691FE24A7FD" bold="true" box="[354,438,460,484]" italics="true" pageId="16" pageNumber="71">clemens</emphasis>
</taxonomicName>
gr.):
</emphasis>
<bibRefCitation id="D46D4B4DF7261175FE2BC691FD73A7FD" author="Kawase, N. &amp; Morita, H" box="[496,737,460,484]" pageId="16" pageNumber="71" pagination="31 - 40" refId="ref17648" refString="Kawase, N. &amp; Morita, H (2010) The caddisfly (Insecta, Trichoptera) fauna of the Suzuka Mountains. Biology of Inland Waters, 25, 31 - 40. [in Japanese with English abstract]" type="journal article" year="2010">Kawase &amp; Morita 2010</bibRefCitation>
, 36.
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F7261174FF4CC54FFD9AA240" lastPageId="17" lastPageNumber="72" pageId="16" pageNumber="71" type="diagnosis">
<paragraph id="B04336BCF7261175FF4CC54FFDF3A4A5" blockId="16.[151,1437,530,700]" pageId="16" pageNumber="71">
<emphasis id="8288EAAEF7261175FF4CC54FFE80A435" bold="true" box="[151,274,530,556]" pageId="16" pageNumber="71">Diagnosis.</emphasis>
The male genitalia of
<taxonomicName id="77FC4D3FF7261175FDD6C54EFD36A435" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[525,676,530,556]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="furcicauda" status="sp. nov.">
<emphasis id="8288EAAEF7261175FDD6C54EFD36A435" box="[525,676,530,556]" italics="true" pageId="16" pageNumber="71">R. furcicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F7261175FD70C54EFC93A434" box="[683,769,531,557]" pageId="16" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
are unique among known species in the
<taxonomicName id="77FC4D3FF7261175FB1CC54EFAD2A435" authorityName="Tsuda" authorityYear="1940" box="[1223,1344,530,556]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="clemens">
<emphasis id="8288EAAEF7261175FB1CC54EFAD2A435" box="[1223,1344,530,556]" italics="true" pageId="16" pageNumber="71">R. clemens</emphasis>
</taxonomicName>
Species Group in that the apical segment of each inferior appendage is bifurcate (
<figureCitation id="28C72A39F7261175FC19C56BFB8BA449" box="[962,1049,566,592]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="16" pageNumber="71">Fig. 7A</figureCitation>
) (those of the other species are not bifurcate). The female genitalia of
<taxonomicName id="77FC4D3FF7261175FDFEC506FD2FA46D" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[549,701,602,628]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="furcicauda" status="sp. nov.">
<emphasis id="8288EAAEF7261175FDFEC506FD2FA46D" box="[549,701,602,628]" italics="true" pageId="16" pageNumber="71">R. furcicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F7261175FD1DC506FC8CA46C" box="[710,798,603,629]" pageId="16" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
are similar to those of
<taxonomicName id="77FC4D3FF7261175FBF2C506FB58A46D" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[1065,1226,602,628]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="parvicauda" status="sp. nov.">
<emphasis id="8288EAAEF7261175FBF2C506FB58A46D" box="[1065,1226,602,628]" italics="true" pageId="16" pageNumber="71">R. parvicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F7261175FB09C506FAB8A46C" box="[1234,1322,603,629]" pageId="16" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
in having thick setae densely on the lateral surface of segment VIII but can be distinguished from the latter by the characters given in the diagnosis for
<emphasis id="8288EAAEF7261175FE60C5FEFDF3A4A5" box="[443,609,674,700]" italics="true" pageId="16" pageNumber="71">
<taxonomicName id="77FC4D3FF7261175FE60C5FEFDCFA4A5" authorityName="Kawase" authorityYear="2024" box="[443,605,674,700]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="parvicauda">R. parvicauda</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="E4836634F7261175FF4CC02EFB05A1EE" ID-DOI="http://doi.org/10.5281/zenodo.11119225" ID-Zenodo-Dep="11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="16" pageNumber="71" startId="16.[151,250,1907,1931]" targetBox="[309,1278,737,1871]" targetPageId="16" targetType="figure">
<paragraph id="B04336BCF7261175FF4CC02EFB05A1EE" blockId="16.[151,1437,1907,2040]" pageId="16" pageNumber="71">
<emphasis id="8288EAAEF7261175FF4CC02EFDF9A195" bold="true" box="[151,619,1907,1932]" pageId="16" pageNumber="71">
FIGURE 7.
<taxonomicName id="77FC4D3FF7261175FEC3C02EFD87A192" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[280,533,1907,1931]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="71" phylum="Arthropoda" rank="species" species="furcicauda" status="sp. nov.">
<emphasis id="8288EAAEF7261175FEC3C02EFD87A192" bold="true" box="[280,533,1907,1931]" italics="true" pageId="16" pageNumber="71">Rhyacophila furcicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F7261175FDC0C029FDF9A195" box="[539,619,1908,1932]" pageId="16" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text.
</paragraph>
</caption>
<paragraph id="B04336BCF7271174FF1CC7CBFCE3A6E0" blockId="17.[151,1437,150,1653]" pageId="17" pageNumber="72">
<emphasis id="8288EAAEF7271174FF1CC7CBFE99A6A9" bold="true" box="[199,267,150,176]" pageId="17" pageNumber="72">Adult</emphasis>
. Length of each forewing: male
<quantity id="77049B59F7271174FD5DC7CAFC9DA6A8" box="[646,783,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="7.2" metricValueMin="6.1" pageId="17" pageNumber="72" unit="mm" value="6.65" valueMax="7.2" valueMin="6.1">6.17.2 mm</quantity>
(mean = 6.7, SD = 0.39, n = 8), female
<quantity id="77049B59F7271174FB3FC7CAFAAEA6A8" box="[1252,1340,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="17" pageNumber="72" unit="mm" value="6.7">6.7 mm</quantity>
(n = 1). General morphology and coloration similar to those of
<taxonomicName id="77FC4D3FF7271174FD25C7E6FCE4A6CC" authorityName="Tsuda" authorityYear="1940" box="[766,886,187,213]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="72" phylum="Arthropoda" rank="species" species="clemens">
<emphasis id="8288EAAEF7271174FD25C7E6FCE4A6CC" box="[766,886,187,213]" italics="true" pageId="17" pageNumber="72">R. clemens</emphasis>
</taxonomicName>
, but wing venation similar to
<taxonomicName id="77FC4D3FF7271174FB1EC7E6FABAA6CD" authorityName="Kawase" authorityYear="2024" box="[1221,1320,187,212]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="72" phylum="Arthropoda" rank="species" species="inazui">
<emphasis id="8288EAAEF7271174FB1EC7E6FABAA6CD" box="[1221,1320,187,212]" italics="true" pageId="17" pageNumber="72">R. inazui</emphasis>
</taxonomicName>
(
<figureCitation id="28C72A39F7271174FAECC7E6FA1DA6CC" box="[1335,1423,187,213]" captionStart="FIGURE 4" captionStartId="10.[151,250,1810,1834]" captionTargetBox="[288,1300,430,1774]" captionTargetId="figure-170@10.[275,1313,417,1787]" captionTargetPageId="10" captionText="FIGURE 4. Rhyacophila inazui sp. nov. 4A, male habitus, left dorsolateral (photographed by K. Inazu); 4B, female right forewing (in alcohol), dorsal; 4C4H, male genitalia: 4C, left lateral; 4D, segment X and phallic apparatus, left lateral; 4E, dorsal (left inferior appendage omitted); 4F, ventral (left inferior appendage and apical segment of right inferior appendage omitted); 4G1, apical hinged process of segment X, enlarged, ventral; 4G2, ventral plate of aedeagus, enlarged, ventral; 4H, posterodorsal. 4I4N, female genitalia: 4I, left lateral; 4J, ventral; 4K, segment VIII, left lateral; 4L, same, ventral; 4M, vaginal apparatus, left lateral; 4N, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119217" httpUri="https://zenodo.org/record/11119217/files/figure.png" pageId="17" pageNumber="72">Fig. 4B</figureCitation>
), root of fork I on each forewing clearly longer than that of fork II.
</paragraph>
<paragraph id="B04336BCF7271174FF1CC65EFD83A520" blockId="17.[151,1437,150,1653]" pageId="17" pageNumber="72">
<emphasis id="8288EAAEF7271174FF1CC65EFEE0A705" bold="true" box="[199,370,258,285]" pageId="17" pageNumber="72">Male genitalia</emphasis>
(
<figureCitation id="28C72A39F7271174FE5BC65EFD99A704" box="[384,523,259,285]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7A7F</figureCitation>
). Segment IX dorsal half longer than ventral half in lateral view (
<figureCitation id="28C72A39F7271174FB38C65EFAAEA704" box="[1251,1340,259,285]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7A</figureCitation>
), widely trapezoidal in dorsal view (
<figureCitation id="28C72A39F7271174FE0CC67AFDA7A758" box="[471,565,295,321]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7D</figureCitation>
); apicodorsal lobe extruded posterad, trapezoidal in dorsal view (
<figureCitation id="28C72A39F7271174FAE9C67AFA1DA758" box="[1330,1423,295,321]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7D</figureCitation>
), angular in lateral view (
<figureCitation id="28C72A39F7271174FE47C616FE67A77C" box="[412,501,331,357]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7A</figureCitation>
). Preanal appendages absent. Segment X reverse L-shaped in lateral view, composed of vertical part and horizontal part: vertical part tall in lateral view (
<figureCitation id="28C72A39F7271174FCA7C632FC41A790" box="[892,979,367,393]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7B</figureCitation>
); horizontal part oval in dorsal view (
<figureCitation id="28C72A39F7271174FAB4C632FF2FA7B4" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7D</figureCitation>
), apically bent upwards in lateral view (
<figureCitation id="28C72A39F7271174FDA5C6CEFC99A7B4" box="[638,779,403,429]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7A, 7B</figureCitation>
); apical hinged process with pair of long finger-like lobes, parallel, directed posterodorsad (
<figureCitation id="28C72A39F7271174FDD7C6EAFD58A7C8" box="[524,714,439,465]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7A7D, 7F</figureCitation>
), with pair of round anteroventral corners and small protrusion between them (
<figureCitation id="28C72A39F7271174FE98C686FE0AA7EC" box="[323,408,475,501]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7F</figureCitation>
). Anal sclerites composed of two pairs of lobes, each long and rectangular, but apex of each ventral pair obliquely truncate in lateral view; these two pairs of lobes with common internal root (
<figureCitation id="28C72A39F7271174FB3CC6A2FAD2A400" box="[1255,1344,511,537]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7B</figureCitation>
). Apical band well developed, semi-oval in lateral view, broadly surrounding anus and root of anal sclerites (
<figureCitation id="28C72A39F7271174FADEC57EFA1DA424" box="[1285,1423,547,573]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7B, 7F</figureCitation>
). Tergal band connecting base of apical band to dorsum of phallobase (
<figureCitation id="28C72A39F7271174FC1EC51AFBB3A478" box="[965,1057,583,609]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7B</figureCitation>
). Basal segment of each inferior appendage setose, short, thick, nearly rectangular, posterior half gradually thicker in lateral view (
<figureCitation id="28C72A39F7271174FB3FC536FAADA49C" box="[1252,1343,619,645]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7A</figureCitation>
). Apical segment of each inferior appendage bifurcate in lateral view (
<figureCitation id="28C72A39F7271174FC83C5D2FC75A4B0" box="[856,999,655,681]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7A, 7E</figureCitation>
), covered with hair-like setae sparsely laterally, with dense short spine-like setae on posteromesal margin of ventral fork (
<figureCitation id="28C72A39F7271174FB8EC5EEFA8BA4D4" box="[1109,1305,691,717]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7A, 7D, 7E</figureCitation>
). In phallic apparatus, phallobase short; aedeagus forming narrow tube with sclerotized ventral plate (
<figureCitation id="28C72A39F7271174FBADC58AFB6DA4E8" box="[1142,1279,727,753]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7B, 7C</figureCitation>
); ventral plate horizontal and long and tongue-shaped, posterior margin with small triangular protrusion apicomesally in ventral view (
<figureCitation id="28C72A39F7271174FF06C442FEA0A520" box="[221,306,799,825]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7F</figureCitation>
). Parameres absent.
</paragraph>
<paragraph id="B04336BCF7271174FF1CC41EFF74A22C" blockId="17.[151,1437,150,1653]" pageId="17" pageNumber="72">
<emphasis id="8288EAAEF7271174FF1CC41EFE1EA545" bold="true" box="[199,396,834,861]" pageId="17" pageNumber="72">Female genitalia</emphasis>
(
<figureCitation id="28C72A39F7271174FE40C41EFDB0A544" box="[411,546,835,861]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7G7J</figureCitation>
). Segment VIII annular, dorsal margin almost straight in lateral view (
<figureCitation id="28C72A39F7271174FAEFC41EFA1DA544" box="[1332,1423,835,861]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7G</figureCitation>
); pair of ventral ridges widely separated anteriorly, running subparallel posteriorly (
<figureCitation id="28C72A39F7271174FB8CC43AFB24A598" box="[1111,1206,871,897]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7H</figureCitation>
); with many darkly pigmented and thick setae laterally, with fine pale short setae between ventral ridges (
<figureCitation id="28C72A39F7271174FB99C4D6FB5CA5BC" box="[1090,1230,907,933]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7G, 7H</figureCitation>
); pair of apodemal rods reaching to posterior end of segment VI. Segment IX membranous, with pair of apodemal rods dorsolaterally, extending into abdominal segment VI. Vaginal apparatus simple; posterior process sclerotized, dorsal margin slightly convex in lateral view (
<figureCitation id="28C72A39F7271174FE40C4AAFE7AA208" box="[411,488,1015,1041]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7I</figureCitation>
), apparatus oval in ventral view (
<figureCitation id="28C72A39F7271174FC8CC4AAFC35A208" box="[855,935,1015,1041]" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Fig. 7J</figureCitation>
); processus spermathecae membranous (
<figureCitation id="28C72A39F7271174FAB1C4AAFF4BA22C" captionStart="FIGURE 7" captionStartId="16.[151,250,1907,1931]" captionTargetBox="[309,1278,737,1871]" captionTargetId="figure-194@16.[296,1291,724,1884]" captionTargetPageId="16" captionText="FIGURE 7. Rhyacophila furcicauda sp. nov. 7A7F, male genitalia: 7A, left lateral; 7B, segment X and phallic apparatus, left lateral; 7C, same, posterodorsal; 7D, dorsal (left inferior appendage omitted); 7E, apical segment of right inferior appendage, mesal; 7F, segment X and phallic apparatus, ventral. 7G7J, female genitalia: 7G, segment VIII, left lateral; 7H, same, ventral; 7I, vaginal apparatus, left lateral; 7J, same, ventral. Abbreviation: vr = ventral ridge. Arrows see text." figureDoi="http://doi.org/10.5281/zenodo.11119225" httpUri="https://zenodo.org/record/11119225/files/figure.png" pageId="17" pageNumber="72">Figs 7I, 7J</figureCitation>
).
</paragraph>
<paragraph id="B04336BCF7271174FF1CC362FD9AA240" blockId="17.[151,1437,150,1653]" box="[199,520,1087,1113]" pageId="17" pageNumber="72">
<emphasis id="8288EAAEF7271174FF1CC362FE19A240" bold="true" box="[199,395,1087,1113]" pageId="17" pageNumber="72">Immature stage.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F7271174FF1CC33EFEBAA384" pageId="17" pageNumber="72" type="materials_examined">
<paragraph id="B04336BCF7271174FF1CC33EFC70A2B8" blockId="17.[151,1437,150,1653]" pageId="17" pageNumber="72">
<materialsCitation id="00943CE1F7271174FF1CC33EFC70A2B8" collectingDate="2009-05-31" collectingDateMax="2009-06-26" collectingDateMin="2009-05-31" collectionCode="LBM" collectorName="N. Kawase" country="Japan" county="Honshu" elevation="370" latitude="35.077778" location="Male" longLatPrecision="19" longitude="136.39555" municipality="Yuzurio-cho" pageId="17" pageNumber="72" specimenCode="LBM1410012653" specimenCount="1" stateProvince="Shiga" typeStatus="holotype">
<emphasis id="8288EAAEF7271174FF1CC33EFEA9A264" bold="true" box="[199,315,1123,1149]" pageId="17" pageNumber="72">
<typeStatus id="6F47881EF7271174FF1CC33EFEA4A264" box="[199,310,1123,1149]" pageId="17" pageNumber="72" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="B5236067F7271174FE97C33EFE14A264" LSID="urn:lsid:plazi:treatment:385587AAF7261174FF4CC63EFD0CA06C:B5236067F7271174FE97C33EFE14A264" box="[332,390,1123,1149]" country="Japan" county="Honshu" latitude="35.077778" longLatPrecision="19" longitude="136.39555" municipality="Yuzurio-cho" name="Male" pageId="17" pageNumber="72" stateProvince="Shiga">Male</location>
(in alcohol),
<collectingMunicipality id="5027ACC6F7271174FDE1C33EFD55A264" box="[570,711,1123,1149]" pageId="17" pageNumber="72">Yuzurio-cho</collectingMunicipality>
, Higashi-ômi-shi,
<collectingRegion id="7238F85EF7271174FC70C33EFBA6A264" box="[939,1076,1123,1149]" country="Japan" name="Shiga" pageId="17" pageNumber="72">Shiga Pref.</collectingRegion>
,
<collectingCounty id="59224E30F7271174FB92C33EFB31A264" box="[1097,1187,1123,1149]" pageId="17" pageNumber="72">Honshu</collectingCounty>
,
<collectingCountry id="C8EB762CF7271174FB63C33EFB69A264" box="[1208,1275,1123,1149]" name="Japan" pageId="17" pageNumber="72">Japan</collectingCountry>
,
<geoCoordinate id="D5C8507BF7271174FACBC33FFA0EA264" box="[1296,1436,1122,1149]" degrees="35" direction="north" minutes="4" orientation="latitude" pageId="17" pageNumber="72" precision="15" seconds="40" value="35.077778">35°440” N</geoCoordinate>
<geoCoordinate id="D5C8507BF7271174FF4CC3DAFEBFA2B8" box="[151,301,1159,1185]" degrees="136" direction="east" minutes="23" orientation="longitude" pageId="17" pageNumber="72" precision="15" seconds="44" value="136.39555">136°2344”E</geoCoordinate>
, alt.
<quantity id="77049B59F7271174FEB8C3DAFE3EA2B8" box="[355,428,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="3.7" pageId="17" pageNumber="72" unit="m" value="370.0">
<elevation id="3BD1D18FF7271174FEB8C3DAFE3EA2B8" box="[355,428,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="3.7" pageId="17" pageNumber="72" unit="m" value="370.0">370 m</elevation>
</quantity>
,
<date id="C442107CF7271174FE63C3DAFDE3A2B8" box="[440,625,1159,1185]" pageId="17" pageNumber="72" value="2009-05-31" valueMax="2009-06-26" valueMin="2009-05-31">
<collectingDate id="D406E994F7271174FE63C3DAFDE3A2B8" box="[440,625,1159,1185]" pageId="17" pageNumber="72" value="2009-05-31" valueMax="2009-06-26" valueMin="2009-05-31">31.v26.vi.2009</collectingDate>
</date>
,
<collectorName id="1D09536AF7271174FDA6C3DAFD6AA2B8" box="[637,760,1159,1185]" pageId="17" pageNumber="72">N. Kawase</collectorName>
(
<specimenCode id="E05A9EC7F7271174FCDDC3DBFC44A2B8" box="[774,982,1158,1185]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="17" pageNumber="72">LBM1410012653</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="B04336BCF7271174FF1CC3F6FC4AA314" blockId="17.[151,1437,150,1653]" pageId="17" pageNumber="72">
<materialsCitation id="00943CE1F7271174FF1CC3F6FEA2A2F0" collectingDate="2012-06-09" collectingDateMax="2012-06-22" collectingDateMin="2012-06-09" collectionCode="LBM" collectorName="H. Morita" location="Honshu" pageId="17" pageNumber="72" specimenCode="LBM1410012654- 1410012655" specimenCount="2" specimenCount-male="2" stateProvince="Mie" typeStatus="paratype">
<emphasis id="8288EAAEF7271174FF1CC3F6FE68A2DD" bold="true" box="[199,506,1194,1221]" pageId="17" pageNumber="72">
<typeStatus id="6F47881EF7271174FF1CC3F6FED3A2DC" box="[199,321,1195,1221]" pageId="17" pageNumber="72" type="paratype">Paratypes</typeStatus>
.
<location id="B5236067F7271174FE8EC3F6FE25A2DC" LSID="urn:lsid:plazi:treatment:385587AAF7261174FF4CC63EFD0CA06C:B5236067F7271174FE8EC3F6FE25A2DC" box="[341,439,1195,1221]" name="Honshu" pageId="17" pageNumber="72" stateProvince="Mie">Honshu</location>
:
<collectingRegion id="7238F85EF7271174FE10C3F7FE68A2DD" box="[459,506,1194,1220]" country="Japan" name="Mie" pageId="17" pageNumber="72">Mie</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F7271174FDCAC3F6FDE1A2DC" box="[529,627,1195,1221]" count="2" pageId="17" pageNumber="72" type="male">2 males</specimenCount>
,
<collectorName id="1D09536AF7271174FD5DC3F6FD65A2DC" box="[646,759,1195,1221]" pageId="17" pageNumber="72">Okumano</collectorName>
,
<collectorName id="1D09536AF7271174FCD0C3F6FCCEA2DC" box="[779,860,1195,1221]" pageId="17" pageNumber="72">Iga-shi</collectorName>
,
<date id="C442107CF7271174FCB4C3F6FBB0A2DC" box="[879,1058,1195,1221]" pageId="17" pageNumber="72" value="2012-06-09" valueMax="2012-06-22" valueMin="2012-06-09">
<collectingDate id="D406E994F7271174FCB4C3F6FBB0A2DC" box="[879,1058,1195,1221]" pageId="17" pageNumber="72" value="2012-06-09" valueMax="2012-06-22" valueMin="2012-06-09">9.vi22.vi.2012</collectingDate>
</date>
,
<collectorName id="1D09536AF7271174FBEDC3F6FB3FA2DC" box="[1078,1197,1195,1221]" pageId="17" pageNumber="72">H. Morita</collectorName>
(
<specimenCode id="E05A9EC7F7271174FB1FC3F7FEBAA2F0" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="17" pageNumber="72">LBM1410012654 1410012655</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="00943CE1F7271174FEE0C392FC46A314" collectingDate="2004-05-28" collectingDateMax="2004-05-29" collectingDateMin="2004-05-28" collectionCode="T" collectorName="T. Hattori" location="Ryujin-mura" municipality="Koya-ryujin Skyline" pageId="17" pageNumber="72" specimenCode="LBM1410012656-1410012658" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Wakayama" typeStatus="paratype">
<emphasis id="8288EAAEF7271174FEE0C392FE50A2F0" bold="true" box="[315,450,1231,1257]" pageId="17" pageNumber="72">
<collectingRegion id="7238F85EF7271174FEE0C392FE50A2F0" box="[315,450,1231,1257]" country="Japan" name="Wakayama" pageId="17" pageNumber="72">Wakayama</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F7271174FE09C392FDBEA2F0" box="[466,556,1231,1257]" count="2" pageId="17" pageNumber="72" type="male">2 males</specimenCount>
,
<collectingMunicipality id="5027ACC6F7271174FDE3C392FCB2A2F0" box="[568,800,1231,1257]" pageId="17" pageNumber="72">Kôya-ryujin Skyline</collectingMunicipality>
,
<location id="B5236067F7271174FCF7C392FC2FA2F0" LSID="urn:lsid:plazi:treatment:385587AAF7261174FF4CC63EFD0CA06C:B5236067F7271174FCF7C392FC2FA2F0" box="[812,957,1231,1257]" municipality="Koya-ryujin Skyline" name="Ryujin-mura" pageId="17" pageNumber="72" stateProvince="Wakayama">Ryujin-mura</location>
,
<date id="C442107CF7271174FC12C392FBF6A2F0" box="[969,1124,1231,1257]" pageId="17" pageNumber="72" value="2004-05-28" valueMax="2004-05-29" valueMin="2004-05-28">
<collectingDate id="D406E994F7271174FC12C392FBF6A2F0" box="[969,1124,1231,1257]" pageId="17" pageNumber="72" value="2004-05-28" valueMax="2004-05-29" valueMin="2004-05-28">2829.v.2004</collectingDate>
</date>
, (SPMN-IS-5980059801);
<specimenCount id="A6FAFD35F7271174FF4CC3AEFF62A314" box="[151,240,1267,1293]" count="2" pageId="17" pageNumber="72" type="male">2 males</specimenCount>
,
<specimenCount id="A6FAFD35F7271174FF27C3AEFEF2A314" box="[252,352,1267,1293]" count="1" pageId="17" pageNumber="72" type="female">1 female</specimenCount>
, same data,
<collectionCode id="D6EDAE79F7271174FE3CC3AEFE6BA314" box="[487,505,1267,1293]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="72">T</collectionCode>
.
<collectorName id="1D09536AF7271174FDDFC3AEFDC1A314" box="[516,595,1267,1293]" pageId="17" pageNumber="72">Hattori</collectorName>
(
<specimenCode id="E05A9EC7F7271174FDB9C3AFFC5EA314" box="[610,972,1266,1293]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="17" pageNumber="72">LBM14100126561410012658</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="B04336BCF7271174FF1CC24BFEBAA384" blockId="17.[151,1437,150,1653]" pageId="17" pageNumber="72">
<emphasis id="8288EAAEF7271174FF1CC24BFD11A328" bold="true" box="[199,643,1302,1329]" pageId="17" pageNumber="72">Other specimens examined. Honshu:</emphasis>
<materialsCitation id="00943CE1F7271174FD57C24BFA13A328" box="[652,1409,1302,1329]" collectingDate="2012-05-26" collectingDateMax="2012-06-09" collectingDateMin="2012-05-26" collectorName="H. Morita" location="Iga-shi" municipality="Okumano" pageId="17" pageNumber="72" specimenCount="1" specimenCount-male="1" stateProvince="Mie">
<emphasis id="8288EAAEF7271174FD57C24BFD29A329" bold="true" box="[652,699,1302,1328]" pageId="17" pageNumber="72">
<collectingRegion id="7238F85EF7271174FD57C24BFD29A329" box="[652,699,1302,1328]" country="Japan" name="Mie" pageId="17" pageNumber="72">Mie</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F7271174FD10C24AFC89A328" box="[715,795,1303,1329]" count="1" pageId="17" pageNumber="72" type="male">1 male</specimenCount>
,
<collectingMunicipality id="5027ACC6F7271174FCF3C24AFC0BA328" box="[808,921,1303,1329]" pageId="17" pageNumber="72">Okumano</collectingMunicipality>
,
<location id="B5236067F7271174FC7CC24AFC6AA328" LSID="urn:lsid:plazi:treatment:385587AAF7261174FF4CC63EFD0CA06C:B5236067F7271174FC7CC24AFC6AA328" box="[935,1016,1303,1329]" municipality="Okumano" name="Iga-shi" pageId="17" pageNumber="72" stateProvince="Mie">Iga-shi</location>
,
<date id="C442107CF7271174FBDFC24AFB22A328" box="[1028,1200,1303,1329]" pageId="17" pageNumber="72" value="2012-05-26" valueMax="2012-06-09" valueMin="2012-05-26">
<collectingDate id="D406E994F7271174FBDFC24AFB22A328" box="[1028,1200,1303,1329]" pageId="17" pageNumber="72" value="2012-05-26" valueMax="2012-06-09" valueMin="2012-05-26">26.v9.vi.2012</collectingDate>
</date>
,
<collectorName id="1D09536AF7271174FB66C24AFABFA328" box="[1213,1325,1303,1329]" pageId="17" pageNumber="72">H. Morita</collectorName>
(NKa)
</materialsCitation>
;
<materialsCitation id="00943CE1F7271174FA55C24AFBBDA34C" collectingDate="2006-05-31" collectorName="N. Kawase" latitude="33.95" location="Kumano-shi" longLatPrecision="1202" longitude="136.2" municipality="Hobo-cho" pageId="17" pageNumber="72" specimenCount="1" specimenCount-male="1" stateProvince="Mie">
<specimenCount id="A6FAFD35F7271174FA55C24AFF42A34C" count="1" pageId="17" pageNumber="72" type="male">1 male</specimenCount>
,
<collectingMunicipality id="5027ACC6F7271174FF05C266FEC3A34C" box="[222,337,1339,1365]" pageId="17" pageNumber="72">Hobo-cho</collectingMunicipality>
,
<location id="B5236067F7271174FE85C266FE79A34C" LSID="urn:lsid:plazi:treatment:385587AAF7261174FF4CC63EFD0CA06C:B5236067F7271174FE85C266FE79A34C" box="[350,491,1339,1365]" latitude="33.95" longLatPrecision="1202" longitude="136.2" municipality="Hobo-cho" name="Kumano-shi" pageId="17" pageNumber="72" stateProvince="Mie">Kumano-shi</location>
,
<geoCoordinate id="D5C8507BF7271174FE22C267FDC8A34C" box="[505,602,1338,1365]" degrees="33" direction="north" minutes="57" orientation="latitude" pageId="17" pageNumber="72" precision="925" value="33.95">33°57N</geoCoordinate>
<geoCoordinate id="D5C8507BF7271174FDB8C266FD43A34C" box="[611,721,1339,1365]" degrees="136" direction="east" minutes="12" orientation="longitude" pageId="17" pageNumber="72" precision="925" value="136.2">136°12E</geoCoordinate>
,
<date id="C442107CF7271174FD04C266FCC2A34C" box="[735,848,1339,1365]" pageId="17" pageNumber="72" value="2006-05-31">
<collectingDate id="D406E994F7271174FD04C266FCC2A34C" box="[735,848,1339,1365]" pageId="17" pageNumber="72" value="2006-05-31">31.v.2006</collectingDate>
</date>
,
<collectorName id="1D09536AF7271174FC85C266FC49A34C" box="[862,987,1339,1365]" pageId="17" pageNumber="72">N. Kawase</collectorName>
(NKa)
</materialsCitation>
.
<materialsCitation id="00943CE1F7271174FBE7C266FCD7A360" collectingDate="2005-06-11" collectionCode="SPMN" collectorName="S. Kobayashi" location="Totsukawa-mura" municipality="Kannohgawa" pageId="17" pageNumber="72" specimenCount="1" specimenCount-pupa="1" stateProvince="Nara">
<emphasis id="8288EAAEF7271174FBE7C266FBEBA34C" bold="true" box="[1084,1145,1339,1365]" pageId="17" pageNumber="72">
<collectingRegion id="7238F85EF7271174FBE7C266FBEBA34C" box="[1084,1145,1339,1365]" country="Japan" name="Nara" pageId="17" pageNumber="72">Nara</collectingRegion>
</emphasis>
:
<specimenCount id="A6FAFD35F7271174FB51C266FB45A34C" box="[1162,1239,1339,1365]" count="1" pageId="17" pageNumber="72" type="pupa">1 pupa</specimenCount>
(male),
<collectingMunicipality id="5027ACC6F7271174FAE1C266FF47A360" pageId="17" pageNumber="72">Kannohgawa</collectingMunicipality>
,
<location id="B5236067F7271174FF3EC202FE36A360" LSID="urn:lsid:plazi:treatment:385587AAF7261174FF4CC63EFD0CA06C:B5236067F7271174FF3EC202FE36A360" box="[229,420,1375,1401]" municipality="Kannohgawa" name="Totsukawa-mura" pageId="17" pageNumber="72" stateProvince="Nara">Totsukawa-mura</location>
,
<date id="C442107CF7271174FE68C203FDBFA360" box="[435,557,1374,1401]" pageId="17" pageNumber="72" value="2005-06-11">
<collectingDate id="D406E994F7271174FE68C203FDBFA360" box="[435,557,1374,1401]" pageId="17" pageNumber="72" value="2005-06-11">11.vi.2005</collectingDate>
</date>
,
<collectorName id="1D09536AF7271174FDE6C202FD45A360" box="[573,727,1375,1401]" pageId="17" pageNumber="72">S. Kobayashi</collectorName>
(
<collectionCode id="D6EDAE79F7271174FD30C202FCACA360" box="[747,830,1375,1401]" pageId="17" pageNumber="72">SPMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="00943CE1F7271174FC8EC202FEB6A384" collectingDate="2021-07-22" collectionCode="HI" collectorName="H. Iketake" location="Tenkawa-mura" municipality="Kitozumi" pageId="17" pageNumber="72" specimenCount="1" specimenCount-male="1" stateProvince="Nara">
<specimenCount id="A6FAFD35F7271174FC8EC202FC3AA360" box="[853,936,1375,1401]" count="1" pageId="17" pageNumber="72" type="male">1 male</specimenCount>
,
<collectingMunicipality id="5027ACC6F7271174FC63C202FBB7A360" box="[952,1061,1375,1401]" pageId="17" pageNumber="72">Kitozumi</collectingMunicipality>
,
<location id="B5236067F7271174FBEFC202FB4DA360" LSID="urn:lsid:plazi:treatment:385587AAF7261174FF4CC63EFD0CA06C:B5236067F7271174FBEFC202FB4DA360" box="[1076,1247,1375,1401]" municipality="Kitozumi" name="Tenkawa-mura" pageId="17" pageNumber="72" stateProvince="Nara">Tenkawa-mura</location>
,
<date id="C442107CF7271174FB34C202FAE2A360" box="[1263,1392,1375,1401]" pageId="17" pageNumber="72" value="2021-07-22">
<collectingDate id="D406E994F7271174FB34C202FAE2A360" box="[1263,1392,1375,1401]" pageId="17" pageNumber="72" value="2021-07-22">22.vii.2021</collectingDate>
</date>
,
<collectorName id="1D09536AF7271174FA5AC202FF7BA384" pageId="17" pageNumber="72">H. Iketake</collectorName>
(
<collectionCode id="D6EDAE79F7271174FF22C2DEFE89A384" box="[249,283,1411,1437]" pageId="17" pageNumber="72">HI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F7271174FF1CC2FAFD62A3FC" pageId="17" pageNumber="72" type="etymology">
<paragraph id="B04336BCF7271174FF1CC2FAFD62A3FC" blockId="17.[151,1437,150,1653]" pageId="17" pageNumber="72">
<emphasis id="8288EAAEF7271174FF1CC2FAFEDAA3D8" bold="true" box="[199,328,1447,1473]" pageId="17" pageNumber="72">Etymology</emphasis>
. The species name
<taxonomicName id="77FC4D3FF7271174FDFAC2FAFD3BA3D8" authorityName="Kawase" authorityYear="2024" box="[545,681,1447,1473]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="72" phylum="Arthropoda" rank="species" species="furcicauda">
<emphasis id="8288EAAEF7271174FDFAC2FAFD3BA3D8" box="[545,681,1447,1473]" italics="true" pageId="17" pageNumber="72">“furcicauda</emphasis>
</taxonomicName>
” (= fork tail) is a noun in apposition referring to the bifurcate apical segment of each inferior appendage in male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="F8E66537F7271174FF1CC2B3FD0CA06C" pageId="17" pageNumber="72" type="distribution">
<paragraph id="B04336BCF7271174FF1CC2B3FD75A048" blockId="17.[151,1437,150,1653]" pageId="17" pageNumber="72">
<emphasis id="8288EAAEF7271174FF1CC2B3FE64A011" bold="true" box="[199,502,1518,1545]" pageId="17" pageNumber="72">Distribution and biology.</emphasis>
<taxonomicName id="77FC4D3FF7271174FE24C2B2FC9CA010" authority="Kawase, 2024" authorityName="Kawase" authorityYear="2024" box="[511,782,1519,1545]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="72" phylum="Arthropoda" rank="species" species="furcicauda" status="sp. nov.">
<emphasis id="8288EAAEF7271174FE24C2B2FC9CA010" box="[511,782,1519,1545]" italics="true" pageId="17" pageNumber="72">Rhyacophila furcicauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99BB57D5F7271174FCCCC2B2FCE2A010" box="[791,880,1519,1545]" pageId="17" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
was collected from only Suzuka Mountains and Kii Mountains in Kinki district, central
<collectingCountry id="C8EB762CF7271174FD86C14EFD0FA034" box="[605,669,1555,1581]" name="Japan" pageId="17" pageNumber="72">Japan</collectingCountry>
(
<figureCitation id="28C72A39F7271174FD76C14EFC9BA034" box="[685,777,1555,1581]" captionStart="FIGURE 9" captionStartId="19.[151,250,1739,1763]" captionTargetBox="[292,1296,194,1702]" captionTargetId="figure-16@19.[279,1309,181,1715]" captionTargetPageId="19" captionText="FIGURE 9. Distributions of species in the Rhyacophila clemens Species Group in Japan." figureDoi="http://doi.org/10.5281/zenodo.11119229" httpUri="https://zenodo.org/record/11119229/files/figure.png" pageId="17" pageNumber="72">Fig. 9D</figureCitation>
). Adults of this species were collected near streamside by sweeping or using a Malaise trap in late May to July.
</paragraph>
<paragraph id="B04336BCF7271174FF1CC106FD0CA06C" blockId="17.[151,1437,150,1653]" box="[199,670,1627,1653]" pageId="17" pageNumber="72">
<emphasis id="8288EAAEF7271174FF1CC106FE11A06C" bold="true" box="[199,387,1627,1653]" pageId="17" pageNumber="72">Japanese name.</emphasis>
Nimata-nagare-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>