249 lines
39 KiB
XML
249 lines
39 KiB
XML
<document id="805021F4532D653B87AE680E6D13BE87" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFE4E514531A84FB60ADFE15" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) nitens Cameron 1944" docType="treatment" docVersion="11" lastPageNumber="155" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="154" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="DB20E875BE2E5FB0B56768899B266166">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="1CB528E9BCD59606831F2BACE688297F">Assing, Volker</mods:namePart>
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<mods:title id="728382BD5354EC81494D7E28158E461B">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="A5BB072A5A70EEDBE5429EEB7E5D1681">2017</mods:date>
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<mods:number id="62C8CBBB01D07EF0A92DA318B4163A69">2017-06-30</mods:number>
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<mods:number id="6ECF47865502FE7C24EF42A591274391">67</mods:number>
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<subSubSection id="C3146527FFE4E517531A84FB67BAFBBA" box="[831,1201,1011,1038]" pageId="37" pageNumber="154" type="nomenclature">
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<paragraph id="8BB136ACFFE4E517531A84FB67BAFBBA" blockId="37.[831,1201,1011,1071]" box="[831,1201,1011,1038]" pageId="37" pageNumber="154">
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<taxonomicName id="4C0E4D2FFFE4E517531A84FB67BAFBBA" ID-CoL="7GHZ4" authority="CAMERON, 1944" authorityName="Cameron" authorityYear="1944" box="[831,1201,1011,1038]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="nitens" subGenus="Zyras">
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<emphasis id="B97AEABEFFE4E517531A84FB607EFBBA" box="[831,885,1011,1038]" italics="true" pageId="37" pageNumber="154">Zyras</emphasis>
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(
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<emphasis id="B97AEABEFFE4E51753A684FB60B2FBBA" box="[899,953,1011,1038]" italics="true" pageId="37" pageNumber="154">Zyras</emphasis>
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)
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<emphasis id="B97AEABEFFE4E51753E284FB670DFBBA" box="[967,1030,1011,1038]" italics="true" pageId="37" pageNumber="154">nitens</emphasis>
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<bibRefCitation id="EF9F4B5DFFE4E517542884FB67BAFBBA" author="CAMERON, M." box="[1037,1201,1011,1038]" pageId="37" pageNumber="154" pagination="104 - 108" refId="ref57674" refString="CAMERON, M. 1944: Descriptions of new Staphylinidae (Coleoptera). - Proceedings of the Royal Entomological Society of London (B) 13: 11 - 15, 49 - 52, 104 - 108." type="journal article" year="1944">CAMERON, 1944</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3146527FFE4E517531A8313674CFB9B" box="[831,1095,1051,1071]" pageId="37" pageNumber="154" type="description">
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<paragraph id="8BB136ACFFE4E517531A8313674CFB9B" blockId="37.[831,1201,1011,1071]" box="[831,1095,1051,1071]" pageId="37" pageNumber="154">
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(
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<figureCitation id="13352A29FFE4E517536683136076FB9B" box="[835,893,1051,1071]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="37" pageNumber="154">Figs 10</figureCitation>
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,
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<figureCitation id="13352A29FFE4E51753A683136093FB9B" box="[899,920,1051,1071]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="37" pageNumber="154">71</figureCitation>
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,
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<figureCitation id="13352A29FFE4E51753BA831360B6FB9B" box="[927,957,1051,1071]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="37" pageNumber="154">109</figureCitation>
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,
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<figureCitation id="13352A29FFE4E51753E083136701FB9B" box="[965,1034,1051,1071]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="37" pageNumber="154">216–221</figureCitation>
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,
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<figureCitation id="13352A29FFE4E517543583136749FB9B" box="[1040,1090,1051,1071]" captionStart="Map 8" captionStartId="68.[189,234,1235,1256]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-71@68.[189,1436,232,1226]" captionTargetPageId="68" captionText="Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white" figureDoi="http://doi.org/10.5281/zenodo.5742417" httpUri="https://zenodo.org/record/5742417/files/figure.png" pageId="37" pageNumber="154">Map 8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3146527FFE4E517533F834467A1FBD4" box="[794,1194,1099,1121]" pageId="37" pageNumber="154" type="reference_group">
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<paragraph id="8BB136ACFFE4E517533F834467A1FBD4" blockId="37.[794,1194,1099,1121]" box="[794,1194,1099,1121]" pageId="37" pageNumber="154">
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<treatmentCitationGroup id="AB1E1182FFE4E517533F834467A1FBD4" box="[794,1194,1099,1121]" pageId="37" pageNumber="154">
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<treatmentCitation id="0AAF10BDFFE4E517533F834467ADFBD4" author="CAMERON, M." box="[794,1190,1099,1121]" page="108" pageId="37" pageNumber="154" year="1944">
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||
<taxonomicName id="4C0E4D2FFFE4E517533F834467ADFBD4" ID-CoL="7GHZ4" authority="CAMERON, 1944: 108" authorityName="Cameron" authorityPageNumber="108" authorityYear="1944" box="[794,1190,1099,1121]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="nitens" subGenus="Zyras">
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||
<emphasis id="B97AEABEFFE4E517533F83446045FBD5" box="[794,846,1100,1121]" italics="true" pageId="37" pageNumber="154">Zyras</emphasis>
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||
(
|
||
<emphasis id="B97AEABEFFE4E51753798344609BFBD5" box="[860,912,1100,1121]" italics="true" pageId="37" pageNumber="154">Zyras</emphasis>
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||
)
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||
<emphasis id="B97AEABEFFE4E51753B8834460DEFBD5" box="[925,981,1100,1121]" italics="true" pageId="37" pageNumber="154">nitens</emphasis>
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||
<bibRefCitation id="EF9F4B5DFFE4E51753FE834367ADFBD4" author="CAMERON, M." box="[987,1190,1099,1120]" pageId="37" pageNumber="154" pagination="104 - 108" refId="ref57674" refString="CAMERON, M. 1944: Descriptions of new Staphylinidae (Coleoptera). - Proceedings of the Royal Entomological Society of London (B) 13: 11 - 15, 49 - 52, 104 - 108." type="journal article" year="1944">CAMERON, 1944: 108</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3146527FFE4E517533F838167C0FAB5" pageId="37" pageNumber="154" type="materials_examined">
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||
<paragraph id="8BB136ACFFE4E517533F838167C0FAB5" blockId="37.[794,1399,1161,1281]" pageId="37" pageNumber="154">
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||
<emphasis id="B97AEABEFFE4E517533F8381672FFB15" bold="true" box="[794,1060,1161,1185]" pageId="37" pageNumber="154">Type material examined</emphasis>
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||
:
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<materialsCitation id="3B663CF1FFE4E5175417838167C0FAB5" ID-GBIF-Occurrence="3450162412" collectionCode="BMNH" collectorName="Dr. Cameron & V. Assing" location="Bank of Stream" pageId="37" pageNumber="154" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
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Holotype
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<specimenCount id="9D08FD25FFE4E51754B9838267BAFB15" box="[1180,1201,1162,1185]" pageId="37" pageNumber="154">♂</specimenCount>
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||
: “The Gap,
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<collectingRegion id="49CAF84EFFE4E517551183816035FB76" country="Malaysia" name="Selangor" pageId="37" pageNumber="154">Selangor</collectingRegion>
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||
, F.M.S.,
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||
<collectorName id="26FB537AFFE4E51753BA83A2673BFB75" box="[927,1072,1193,1217]" pageId="37" pageNumber="154">Dr. Cameron</collectorName>
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/
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<location id="8ED16077FFE4E517546A83A167F6FB75" LSID="urn:lsid:plazi:treatment:03A787BAFFE4E514531A84FB60ADFE15:8ED16077FFE4E517546A83A167F6FB75" box="[1103,1277,1193,1217]" name="Bank of Stream" pageId="37" pageNumber="154" stateProvince="Selangor">Bank of Stream</location>
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||
/ Debris /
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||
<taxonomicName id="4C0E4D2FFFE4E517533F83C260BEFB55" authority="Cam." authorityName="Cameron" authorityYear="1944" box="[794,949,1225,1249]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="nitens">
|
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<emphasis id="B97AEABEFFE4E517533F83C2607FFB55" box="[794,884,1226,1249]" italics="true" pageId="37" pageNumber="154">Z. nitens</emphasis>
|
||
Cam.
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||
</taxonomicName>
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||
Type / Holotype / Holotypus
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||
<taxonomicName id="4C0E4D2FFFE4E51754D283C2608BFAB5" authority="Cameron" authorityName="Cameron" authorityYear="1944" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="nitens">
|
||
<emphasis id="B97AEABEFFE4E51754D283C2667DFB55" box="[1271,1398,1226,1249]" italics="true" pageId="37" pageNumber="154">Zyras nitens</emphasis>
|
||
Cameron
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||
</taxonomicName>
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||
, rev.
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||
<collectorName id="26FB537AFFE4E517539083E2671CFAB5" box="[949,1047,1257,1281]" pageId="37" pageNumber="154">V. Assing</collectorName>
|
||
2016” (
|
||
<collectionCode id="ED1FAE69FFE4E517544C83E167B4FAB5" box="[1129,1215,1257,1281]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="37" pageNumber="154">BMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFE4E517533F8221661AF9B5" pageId="37" pageNumber="154" type="discussion">
|
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<paragraph id="8BB136ACFFE4E517533F82216056FA35" blockId="37.[794,1399,1321,1409]" pageId="37" pageNumber="154">
|
||
<emphasis id="B97AEABEFFE4E517533F82216082FAF5" bold="true" box="[794,905,1321,1345]" pageId="37" pageNumber="154">Comment</emphasis>
|
||
: The original description is based on a unique specimen from “
|
||
<collectingRegion id="49CAF84EFFE4E51753FC82416769FAD5" box="[985,1122,1353,1377]" country="Malaysia" name="Selangor" pageId="37" pageNumber="154">SELANGOR</collectingRegion>
|
||
: The Gap” (
|
||
<bibRefCitation id="EF9F4B5DFFE4E517552C82416059FA35" author="CAMERON, M." pageId="37" pageNumber="154" pagination="104 - 108" refId="ref57674" refString="CAMERON, M. 1944: Descriptions of new Staphylinidae (Coleoptera). - Proceedings of the Royal Entomological Society of London (B) 13: 11 - 15, 49 - 52, 104 - 108." type="journal article" year="1944">CAMERON 1944</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE4E517533F82A1661AF9B5" blockId="37.[794,1399,1449,1538]" pageId="37" pageNumber="154">
|
||
<emphasis id="B97AEABEFFE4E517533F82A1676DFA75" bold="true" box="[794,1126,1449,1473]" pageId="37" pageNumber="154">Additional material examined</emphasis>
|
||
:
|
||
<materialsCitation id="3B663CF1FFE4E517545182A16606F9B5" ID-GBIF-Occurrence="3450162382" collectingDate="2004-03" collectingMethod="flight interception trap" collectorName="Maruyama" country="Malaysia" elevation="250" location="Ulu Gombak Field Studies Centre" pageId="37" pageNumber="154" specimenCount="1" stateProvince="Selangor">
|
||
<emphasis id="B97AEABEFFE4E517545182A167DEFA75" bold="true" box="[1140,1237,1449,1473]" pageId="37" pageNumber="154">
|
||
<collectingCountry id="F319763CFFE4E517545182A167DEFA75" box="[1140,1237,1449,1473]" name="Malaysia" pageId="37" pageNumber="154">Malaysia</collectingCountry>
|
||
</emphasis>
|
||
: 1 ♀,
|
||
<collectingRegion id="49CAF84EFFE4E517553382A16678FA75" box="[1302,1395,1449,1473]" country="Malaysia" name="Selangor" pageId="37" pageNumber="154">Selangor</collectingRegion>
|
||
,
|
||
<location id="8ED16077FFE4E517533F82C1678AFA55" LSID="urn:lsid:plazi:treatment:03A787BAFFE4E514531A84FB60ADFE15:8ED16077FFE4E517533F82C1678AFA55" box="[794,1153,1481,1505]" country="Malaysia" name="Ulu Gombak Field Studies Centre" pageId="37" pageNumber="154" stateProvince="Selangor">Ulu Gombak Field Studies Centre</location>
|
||
,
|
||
<quantity id="4CF69B49FFE4E51754AF82C167C4FA55" box="[1162,1231,1481,1505]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="37" pageNumber="154" unit="m" value="250.0">
|
||
<elevation id="0023D19FFFE4E51754AF82C167C4FA55" box="[1162,1231,1481,1505]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="37" pageNumber="154" unit="m" value="250.0">250 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod id="524F4EBBFFE4E51754FD82C16073F9B5" pageId="37" pageNumber="154">flight interception trap</collectingMethod>
|
||
,
|
||
<date id="FFB0106CFFE4E51753A782E160D3F9B5" box="[898,984,1513,1537]" pageId="37" pageNumber="154" value="2004-03">
|
||
<collectingDate id="EFF4E984FFE4E51753A782E160D3F9B5" box="[898,984,1513,1537]" pageId="37" pageNumber="154" value="2004-03">III.2004</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26FB537AFFE4E517542882E2678AF9B6" box="[1037,1153,1514,1538]" pageId="37" pageNumber="154">Maruyama</collectorName>
|
||
et al. (cMar)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFE4E514533F8121618DF795" lastPageId="38" lastPageNumber="155" pageId="37" pageNumber="154" type="description">
|
||
<paragraph id="8BB136ACFFE4E517533F81216720F815" blockId="37.[794,1399,1577,2082]" pageId="37" pageNumber="154">
|
||
<emphasis id="B97AEABEFFE4E517533F812160B8F9F5" bold="true" box="[794,947,1577,1601]" pageId="37" pageNumber="154">Redescription</emphasis>
|
||
: Body length 5.0–
|
||
<quantity id="4CF69B49FFE4E517545E812167C5F9F5" box="[1147,1230,1577,1601]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="37" pageNumber="154" unit="mm" value="5.8">5.8 mm</quantity>
|
||
; length of forebody
|
||
<quantity id="4CF69B49FFE4E5175379814160D6F9D5" box="[860,989,1609,1633]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="37" pageNumber="154" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.7–2.8 mm</quantity>
|
||
. Coloration (
|
||
<figureCitation id="13352A29FFE4E5175450814167CFF9D5" box="[1141,1220,1609,1633]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="37" pageNumber="154">Figs 10</figureCitation>
|
||
,
|
||
<figureCitation id="13352A29FFE4E51754F0814167FBF9D5" box="[1237,1264,1609,1633]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="37" pageNumber="154">71</figureCitation>
|
||
,
|
||
<figureCitation id="13352A29FFE4E517552481416620F9D5" box="[1281,1323,1609,1633]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="37" pageNumber="154">109</figureCitation>
|
||
): head and pronotum dark-brown; elytra brown or dark-reddish with the postero-lateral portions slightly and diffusely darker; abdomen dark-brown, with tergite II, the anterior and antero-lateral portions of tergites III–VI, and most of the paratergites yellowish to yellowish-brown, and with segments IX–X dark-yellowish; legs yellowish with the apices of the meso- and metafemora slightly darker; antennae brown with the basal 2–3 antennomeres pale-reddish and XI pale yellowish to dark-yellowish; maxillary palpi yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE4E514533F80A162F2FE35" blockId="37.[794,1399,1577,2082]" lastBlockId="38.[189,794,233,1665]" lastPageId="38" lastPageNumber="155" pageId="37" pageNumber="154">
|
||
Head (
|
||
<figureCitation id="13352A29FFE4E517534280A160BAF875" box="[871,945,1961,1985]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="37" pageNumber="154">Fig. 71</figureCitation>
|
||
) moderately transverse, median portion extensively impunctate; punctures in lateral portions sparse and fine to moderately coarse. Eyes at least twice as long as distance from posterior margin of eye to posterior constriction of head in dorsal view. Antenna (
|
||
<figureCitation id="13352A29FFE7E51452E287E16019FEB5" box="[711,786,233,257]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="38" pageNumber="155">Fig. 10</figureCitation>
|
||
)
|
||
<quantity id="4CF69B49FFE7E514509886016204FE95" box="[189,271,265,289]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="38" pageNumber="155" unit="mm" value="2.4">2.4 mm</quantity>
|
||
long and slender; antennomeres IV–VIII oblong, IX approximately as long as broad to weakly oblong, X very weakly transverse, and XI slightly shorter than the combined length of IX and X.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE7E514509886826255FD55" blockId="38.[189,794,233,1665]" pageId="38" pageNumber="155">
|
||
Pronotum (
|
||
<figureCitation id="13352A29FFE7E514511D86816288FE15" box="[312,387,393,417]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="38" pageNumber="155">Fig. 71</figureCitation>
|
||
) 1.09–1.12 times as broad as long and 1.1–1.2 times as broad as head, broadest in anterior half, weakly tapering posteriad; lateral margins weakly sinuate in posterior half in dorsal view; punctation coarse and irregularly distributed, on either side of midline with extensive impunctate areas; lateral margins each with four long and erect black setae, one of them inserting near anterior and one near posterior angle, and two in between; anterior margin with an additional long seta on either side; pubescence of disc moderately long, suberect, and pale.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE7E514509885E161A8FC15" blockId="38.[189,794,233,1665]" pageId="38" pageNumber="155">
|
||
Elytra (
|
||
<figureCitation id="13352A29FFE7E514513785E16257FCB5" box="[274,348,745,769]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="38" pageNumber="155">Fig. 71</figureCitation>
|
||
) 0.77–0.79 times as long as pronotum; punctation coarse, moderately dense anteriorly and rather sparse near posterior margins; pubescence suberect, rather long, and pale on disc, dark and stouter at lateral margins. Hind wings present. Metatarsomere I longer than the combined length of II and III.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE7E514509884A1627EFA15" blockId="38.[189,794,233,1665]" pageId="38" pageNumber="155">
|
||
Abdomen (
|
||
<figureCitation id="13352A29FFE7E514511384A1629BFC75" box="[310,400,937,961]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="38" pageNumber="155">Fig. 109</figureCitation>
|
||
) as broad as, or slightly broader than elytra, with rather deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of rather weakly defined grooves; tergites III–VI with a lateral setiferous puncture on either side and with a transverse row of six setiferous punctures at or near posterior margin, tergite VI additionally with scattered non-setiferous punctures anteriorly; tergite VII anteriorly with a narrow transverse band of non-setiferous punctures and posteriorly with two transverse rows of sparse setiferous punctures, posterior margin with palisade fringe; tergite VIII (
|
||
<figureCitation id="13352A29FFE7E514529D82016019FA95" box="[696,786,1289,1313]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="38" pageNumber="155">Fig. 219</figureCitation>
|
||
) with approximately twelve long dark setae in posterior fourth, arranged in two transverse rows, posterior margin convex; all sternites with numerous long dark setae posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE7E514509882A26214F935" blockId="38.[189,794,233,1665]" pageId="38" pageNumber="155">
|
||
♂: sternite VIII (
|
||
<figureCitation id="13352A29FFE7E514515C82A162DBFA75" box="[377,464,1449,1473]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="38" pageNumber="155">Fig. 220</figureCitation>
|
||
) apically conspicuously acute, apex somewhat bifid (
|
||
<figureCitation id="13352A29FFE7E514519682C16107FA55" box="[435,524,1481,1505]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="38" pageNumber="155">Fig. 221</figureCitation>
|
||
); median lobe of aedeagus (
|
||
<figureCitation id="13352A29FFE7E51450D382E1628DF9B5" box="[246,390,1513,1537]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="38" pageNumber="155">Figs 216–217</figureCitation>
|
||
)
|
||
<quantity id="4CF69B49FFE7E51451BE82E162F1F9B5" box="[411,506,1513,1537]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="38" pageNumber="155" unit="mm" value="0.84">0.84 mm</quantity>
|
||
long and with a ventral process of distinctive shape, particularly in lateral view; paramere (
|
||
<figureCitation id="13352A29FFE7E514511181216287F9F5" box="[308,396,1577,1601]" captionStart="Figs 208–226" captionStartId="60.[189,229,1991,2013]" captionTargetBox="[196,1429,238,1961]" captionTargetId="figure-130@60.[189,1436,232,1971]" captionText="Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226):medianlobeofaedeagusinlateralandinventralview(208–213,216–217,222–223);paramere (214,218,224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5888376" httpUri="https://zenodo.org/record/5888376/files/figure.png" pageId="38" pageNumber="155">Fig. 218</figureCitation>
|
||
) approximately
|
||
<quantity id="4CF69B49FFE7E514526581216199F9F5" box="[576,658,1577,1601]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="155" unit="mm" value="0.9">0.9 mm</quantity>
|
||
long, apical lobe distinctly modified, long, depressed, and apically rounded.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE7E514509881A1618DF795" blockId="38.[189,794,1705,2081]" pageId="38" pageNumber="155">
|
||
<emphasis id="B97AEABEFFE7E514509881A1629BF975" bold="true" box="[189,400,1705,1729]" pageId="38" pageNumber="155">Comparative notes</emphasis>
|
||
: As can be inferred from the slender habitus, the long and slender appendages, the glossy appearance, and particularly by the posteriorly strongly produced male sternite VIII, the morphology of the aedeagus (shape of apex of ventral process; long and flattened apical lobe of paramere),
|
||
<taxonomicName id="4C0E4D2FFFE7E51452318042617AF8D5" authorityName="Cameron" authorityYear="1944" box="[532,625,1866,1889]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="nitens">
|
||
<emphasis id="B97AEABEFFE7E51452318042617AF8D5" box="[532,625,1866,1889]" italics="true" pageId="38" pageNumber="155">Z. nitens</emphasis>
|
||
</taxonomicName>
|
||
is closely allied to
|
||
<taxonomicName id="4C0E4D2FFFE7E51450FD80626250F835" authorityName="Assing" authorityYear="2017" box="[216,347,1898,1921]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="truncatus">
|
||
<emphasis id="B97AEABEFFE7E51450FD80626250F835" box="[216,347,1898,1921]" italics="true" pageId="38" pageNumber="155">Z. truncatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0E4D2FFFE7E5145143806262FEF835" authorityName="SCHEERPELTZ" authorityYear="1965" box="[358,501,1897,1921]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="glabricollis">
|
||
<emphasis id="B97AEABEFFE7E5145143806262FEF835" box="[358,501,1897,1921]" italics="true" pageId="38" pageNumber="155">Z. glabricollis</emphasis>
|
||
</taxonomicName>
|
||
, and allied species. Among the species of this group, it is characterized by large eyes, the morphology of the antennae, coarse punctation of the elytra, the punctation pattern of the pronotum, and particularly by the distinctive shapes of the male sternite VIII and the median lobe of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFE7E514536587E160ADFE15" pageId="38" pageNumber="155" type="distribution">
|
||
<paragraph id="8BB136ACFFE7E514536587E160ADFE15" blockId="38.[832,1436,233,417]" pageId="38" pageNumber="155">
|
||
<emphasis id="B97AEABEFFE7E514536587E167C5FEB5" bold="true" box="[832,1230,233,257]" pageId="38" pageNumber="155">Distribution and natural history</emphasis>
|
||
: This species is currently known from two close localities in
|
||
<collectingRegion id="49CAF84EFFE7E514557D8601606EFEF6" country="Malaysia" name="Selangor" pageId="38" pageNumber="155">Selangor</collectingRegion>
|
||
, Peninsular
|
||
<collectingCountry id="F319763CFFE7E51453D28621675CFEF5" box="[1015,1111,297,321]" name="Malaysia" pageId="38" pageNumber="155">Malaysia</collectingCountry>
|
||
(
|
||
<figureCitation id="13352A29FFE7E5145449862167BFFEF5" box="[1132,1204,297,321]" captionStart="Map 8" captionStartId="68.[189,234,1235,1256]" captionTargetBox="[189,1436,232,1226]" captionTargetId="figure-71@68.[189,1436,232,1226]" captionTargetPageId="68" captionText="Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white" figureDoi="http://doi.org/10.5281/zenodo.5742417" httpUri="https://zenodo.org/record/5742417/files/figure.png" pageId="38" pageNumber="155">Map 8</figureCitation>
|
||
). The
|
||
<typeStatus id="54B5880EFFE7E51455228621666DFEF5" box="[1287,1382,297,321]" pageId="38" pageNumber="155" type="holotype">holotype</typeStatus>
|
||
was collected from debris on a stream bank, the additional specimen with a flight interception trap at an altitude of
|
||
<specimenCount id="9D08FD25FFE7E5145379868160ADFE15" box="[860,934,393,417]" pageId="38" pageNumber="155" type="generic">
|
||
<quantity id="4CF69B49FFE7E5145379868160A9FE15" box="[860,930,393,417]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="155" unit="m" value="250.0">250 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |