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<document id="9B476E60CBBE506B34CD1C197DE4D789" ID-DOI="10.5281/zenodo.170887" ID-GBIF-Dataset="9260f7ec-a335-4fea-b6b5-95921cc686f1" ID-ISSN="1175­5326" ID-Zenodo-Dep="170887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459716181808" checkinUser="plazi" docAuthor="Poorani, J. &amp; Booth, R. G." docDate="2005" docId="F76187B74417560DFE892BB3FD86F9C4" docLanguage="en" docName="zt00875.pdf" docOrigin="Zootaxa 875" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Oenopia excellens Crotch, comb. nov." docType="treatment" docVersion="10" lastPageNumber="4" masterDocId="0B58FFCF4416560EFF812967FF93FFCA" masterDocTitle="On the identity of Dysis excellens Crotch (Coleoptera: Coccinellidae: Coccinellini), a little known Oriental lady beetle" masterLastPageNumber="5" masterPageNumber="1" pageNumber="2" updateTime="1698208014518" updateUser="plazi">
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<mods:title id="A8971852B4E304A6FE3E1EBB8CA0F180">On the identity of Dysis excellens Crotch (Coleoptera: Coccinellidae: Coccinellini), a little known Oriental lady beetle</mods:title>
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<mods:namePart id="F5B1767441EB93BC7AD687C5D16CD2E8">Poorani, J.</mods:namePart>
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<mods:namePart id="D22AC67CE0D081E1BF3FF1D7B3EC3DDE">Booth, R. G.</mods:namePart>
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<mods:date id="7CF40009201162665D0FE5FAD2E3A198">2005</mods:date>
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<mods:identifier id="7FFACE6607BEDCF05AD4C45E3E797D9E" type="DOI">10.5281/zenodo.170887</mods:identifier>
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<mods:identifier id="F234188E6EA52883DD26B8B0B4593D59" type="ISSN">1175­5326</mods:identifier>
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<treatment id="F76187B74417560DFE892BB3FD86F9C4" ID-DOI="http://doi.org/10.5281/zenodo.6266816" ID-GBIF-Taxon="119334663" ID-Zenodo-Dep="6266816" LSID="urn:lsid:plazi:treatment:F76187B74417560DFE892BB3FD86F9C4" httpUri="http://treatment.plazi.org/id/F76187B74417560DFE892BB3FD86F9C4" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<subSubSection id="37D2652A4417560FFE892BB3FCE6FD24" box="[264,885,724,750]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="7F7736A14417560FFE892BB3FCE6FD24" blockId="1.[264,885,724,750]" box="[264,885,724,750]" pageId="1" pageNumber="2">
<heading id="243F81CD4417560FFE892BB3FCE6FD24" bold="true" box="[264,885,724,750]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="4DBCEAB34417560FFE892BB3FCE6FD24" bold="true" box="[264,885,724,750]" pageId="1" pageNumber="2">
<taxonomicName id="B8C84D224417560FFE892BB3FDCBFD24" authority="Crotch" authorityName="Crotch" box="[264,600,724,750]" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="excellens" status="comb. nov.">
<emphasis id="4DBCEAB34417560FFE892BB3FE76FD24" bold="true" box="[264,485,724,750]" italics="true" pageId="1" pageNumber="2">Oenopia excellens</emphasis>
(Crotch)
</taxonomicName>
,
<taxonomicNameLabel id="568F57C84417560FFDE42BB3FD78FD24" box="[613,747,724,750]" pageId="1" pageNumber="2">comb. nov.</taxonomicNameLabel>
(
<figureCitation id="E7F32A244417560FFD7A2BB3FCFFFD24" box="[763,876,724,750]" captionStart="FIGURES 1 4" captionStartId="2.[264,383,1671,1695]" captionTargetBox="[296,1306,390,1652]" captionTargetId="figure@2.[274,1320,840,1669]" captionTargetPageId="2" captionText="FIGURES 1 4. Oenopia excellens (Crotch): 1 A. Female, dorsal habitus; 1 B. Male, dorsal habitus; 2. Female spermatheca; 3 4. Male genitalia: 3. Tegmen, ventral view; 4. Sipho." httpUri="https://zenodo.org/record/170888/files/figure.png" pageId="1" pageNumber="2">Figs. 14</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="37D2652A4417560FFE892A78FC0EFC9F" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="7F7736A14417560FFE892A78FC0EFC9F" blockId="1.[264,1203,798,854]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="5FD8118F4417560FFE892A78FC0EFC9F" pageId="1" pageNumber="2">
<taxonomicName id="B8C84D224417560FFE892A78FDE0FCFF" authority="Crotch, 1874: 160" authorityName="Crotch" authorityPageNumber="160" authorityYear="1874" box="[264,627,798,822]" class="Insecta" family="Coccinellidae" genus="Dysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="4DBCEAB34417560FFE892A78FE3FFCFC" box="[264,428,799,822]" italics="true" pageId="1" pageNumber="2">Dysis excellens</emphasis>
<treatmentCitation id="FE6910B04417560FFE332A79FDE0FCFF" author="Crotch" box="[434,627,798,821]" page="160" pageId="1" pageNumber="2" year="1874">
<bibRefCitation id="1B594B504417560FFE332A79FDD2FCFF" author="Crotch" box="[434,577,798,821]" pageId="1" pageNumber="5" refString="Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae, E. W. Janson, London. 311 pp." type="book" year="1874">Crotch, 1874</bibRefCitation>
: 160
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="A07388034417560FFD002A79FD49FCFF" box="[641,730,798,821]" pageId="1" pageNumber="2" type="syntype">Syntype</typeStatus>
Ψ, BMNH;
<typeStatus id="A07388034417560FFCDF2A79FC00FCFF" box="[862,915,798,821]" pageId="1" pageNumber="2">Type</typeStatus>
locality: “
<collectingCountry id="07DF76314417560FFB832A79FB0DFCFF" box="[1026,1182,798,821]" name="Vietnam" pageId="1" pageNumber="2">Cochin China</collectingCountry>
”).
<taxonomicName id="B8C84D224417560FFE892A58FD24FC9F" authority="Korschefsky, 1932: 298" authorityName="Korschefsky" authorityPageNumber="298" authorityYear="1932" box="[264,695,830,854]" class="Insecta" family="Coccinellidae" genus="Dysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="4DBCEAB34417560FFE892A58FE3FFC9C" box="[264,428,831,854]" italics="true" pageId="1" pageNumber="2">Dysis excellens</emphasis>
:
<treatmentCitation id="FE6910B04417560FFE3B2A59FD24FC9F" author="Korschefsky" box="[442,695,830,853]" page="298" pageId="1" pageNumber="2" year="1932">
<bibRefCitation id="1B594B504417560FFE3B2A59FD16FC9F" author="Korschefsky" box="[442,645,830,853]" pageId="1" pageNumber="5" refString="Korschefsky, R. (1932) Coleopterorum Catalogus. Pars 120. Coccinellidae II. Berlin, 435 pp." type="book" year="1932">Korschefsky, 1932</bibRefCitation>
: 298
</treatmentCitation>
</taxonomicName>
.—Gordon, 1985: 20.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="37D2652A4417560DFE892AE1FBDAFE64" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph id="7F7736A14417560FFE892AE1FE38FAC3" blockId="1.[264,1324,902,1889]" pageId="1" pageNumber="2">
<emphasis id="4DBCEAB34417560FFE892AE1FE0BFC6A" bold="true" box="[264,408,902,928]" pageId="1" pageNumber="2">Description</emphasis>
. Length:
<quantity id="B8309B444417560FFD932AE0FD50FC6B" box="[530,707,903,929]" metricMagnitude="-3" metricUnit="m" metricValue="2.9699999999999998" metricValueMax="3.06" metricValueMin="2.88" pageId="1" pageNumber="2" unit="mm" value="2.9699999999999998" valueMax="3.06" valueMin="2.88">2.883.06 mm</quantity>
, breadth:
<quantity id="B8309B444417560FFCBC2AE0FC3DFC6B" box="[829,942,903,929]" metricMagnitude="-3" metricUnit="m" metricValue="2.52" pageId="1" pageNumber="2" unit="mm" value="2.52">2.52 mm</quantity>
.
<emphasis id="4DBCEAB34417560FFC3D2AE1FB84FC6A" bold="true" box="[956,1047,902,928]" pageId="1" pageNumber="2">Female</emphasis>
: Form (
<figureCitation id="E7F32A244417560FFBFC2AE0FB51FC6B" box="[1149,1218,903,929]" captionStart="FIGURES 1 4" captionStartId="2.[264,383,1671,1695]" captionTargetBox="[296,1306,390,1652]" captionTargetId="figure@2.[274,1320,840,1669]" captionTargetPageId="2" captionText="FIGURES 1 4. Oenopia excellens (Crotch): 1 A. Female, dorsal habitus; 1 B. Male, dorsal habitus; 2. Female spermatheca; 3 4. Male genitalia: 3. Tegmen, ventral view; 4. Sipho." httpUri="https://zenodo.org/record/170888/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
A) oval, about 1.2 x as long as broad, convex, head with few long hairs just below clypeal margin and on either side of eyes. Head black, antennae and mouthparts yellowish to pitchy brown. Pronotum black, sides and anterolateral corners white when alive, turning luteous to yellow when dead. Scutellum black. Elytra black with four reddish­orange or yellow spots arranged in two rows, one in either half of elytra, anterior pair large and transverse, posterior pair more or less oval and much smaller. Ventral side with pronotal hypomera and mesosternal epimera whitish to luteous yellow, legs dark brown except femoral apices, tibiae, and tarsi much lighter brown, rest of ventral side more or less completely dark pitchy brown.
</paragraph>
<paragraph id="7F7736A14417560FFEB92C70FD0DFA63" blockId="1.[264,1324,902,1889]" pageId="1" pageNumber="2">Head sparsely punctate, interspaces between punctures with reticulate microsculpture. Pronotum evenly punctate, interspaces between punctures smooth and shiny. Elytra with punctation denser and deeper than that on pronotum, interspaces between punctures with conspicuous, deep microsculpture.</paragraph>
<paragraph id="7F7736A14417560FFEB92CD0FCE9F953" blockId="1.[264,1324,902,1889]" pageId="1" pageNumber="2">
Prosternal intercoxal process with paired carinae, anteriorly slightly divergent. Anterior margin of mesosternum medially shallowly emarginate. Postcoxal line on abdominal sternite I incomplete, without a clearly demarcated associate line. Apical border of fifth abdominal sternite truncate, of sixth evenly arcuate. Spermatheca (
<figureCitation id="E7F32A244417560FFB9D2F48FBF0F983" box="[1052,1123,1583,1609]" captionStart="FIGURES 1 4" captionStartId="2.[264,383,1671,1695]" captionTargetBox="[296,1306,390,1652]" captionTargetId="figure@2.[274,1320,840,1669]" captionTargetPageId="2" captionText="FIGURES 1 4. Oenopia excellens (Crotch): 1 A. Female, dorsal habitus; 1 B. Male, dorsal habitus; 2. Female spermatheca; 3 4. Male genitalia: 3. Tegmen, ventral view; 4. Sipho." httpUri="https://zenodo.org/record/170888/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
) with ramus and cornu together very long and S­shaped, ramus very elongate and gooseneck­shaped, distally swollen and bulbous; infundibulum bell­shaped.
</paragraph>
<paragraph id="7F7736A14417560FFEB92FC1FAB8F8AB" blockId="1.[264,1324,902,1889]" pageId="1" pageNumber="2">
<emphasis id="4DBCEAB34417560FFEB92FC1FEE4F90A" bold="true" box="[312,375,1702,1728]" pageId="1" pageNumber="2">Male</emphasis>
: Body outline (
<figureCitation id="E7F32A244417560FFDAC2FC0FDFCF90B" box="[557,623,1703,1729]" captionStart="FIGURES 1 4" captionStartId="2.[264,383,1671,1695]" captionTargetBox="[296,1306,390,1652]" captionTargetId="figure@2.[274,1320,840,1669]" captionTargetPageId="2" captionText="FIGURES 1 4. Oenopia excellens (Crotch): 1 A. Female, dorsal habitus; 1 B. Male, dorsal habitus; 2. Female spermatheca; 3 4. Male genitalia: 3. Tegmen, ventral view; 4. Sipho." httpUri="https://zenodo.org/record/170888/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
B) subcircular, more rounded than that of female. Head yellow, pronotum and elytra dark pitchy brown to black, with similar pattern as in female. Ventral side with pronotal hypomera, mesosternal epimera, and anterior inner half of elytral epipleura yellow, legs lighter, uniform yellowish brown. Pronotum with interspaces between punctures clearly microreticulate. Elytral punctation similar to that in female.
</paragraph>
<paragraph id="7F7736A14414560CFE89287BFBA3FE94" blockId="2.[264,1323,284,350]" pageId="2" pageNumber="3">
Apical border of fifth abdominal sternite deeply and semicircularly emarginate, of sixth briefly and shallowly emarginate. Genitalia (
<figureCitation id="E7F32A244414560CFC952823FC10FE94" box="[788,899,324,350]" captionStart="FIGURES 1 4" captionStartId="2.[264,383,1671,1695]" captionTargetBox="[296,1306,390,1652]" captionTargetId="figure@2.[274,1320,840,1669]" captionTargetPageId="2" captionText="FIGURES 1 4. Oenopia excellens (Crotch): 1 A. Female, dorsal habitus; 1 B. Male, dorsal habitus; 2. Female spermatheca; 3 4. Male genitalia: 3. Tegmen, ventral view; 4. Sipho." httpUri="https://zenodo.org/record/170888/files/figure.png" pageId="2" pageNumber="3">Figs. 34</figureCitation>
) as illustrated.
</paragraph>
<caption id="2BB766294414560CFE892FE0FBCCF90B" httpUri="https://zenodo.org/record/170888/files/figure.png" pageId="2" pageNumber="3" targetBox="[296,1306,390,1652]" targetPageId="2">
<paragraph id="7F7736A14414560CFE892FE0FBCCF90B" blockId="2.[264,1323,1671,1729]" pageId="2" pageNumber="3">
<emphasis id="4DBCEAB34414560CFE892FE0FE20F955" bold="true" box="[264,435,1671,1695]" pageId="2" pageNumber="3">FIGURES 14.</emphasis>
<taxonomicName id="B8C84D224414560CFE3B2FEFFD70F954" authority="Crotch" authorityName="Crotch" box="[442,739,1671,1695]" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="4DBCEAB34414560CFE3B2FEFFDEDF955" box="[442,638,1672,1695]" italics="true" pageId="2" pageNumber="3">Oenopia excellens</emphasis>
(Crotch)
</taxonomicName>
: 1A. Female, dorsal habitus; 1B. Male, dorsal habitus; 2. Female spermatheca; 34. Male genitalia: 3. Tegmen, ventral view; 4. Sipho.
</paragraph>
</caption>
<paragraph id="7F7736A14414560DFEB92F93FBDAFE64" blockId="2.[264,1323,1780,1887]" lastBlockId="3.[264,1324,283,1550]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="4DBCEAB34414560CFEB92F93FDADF8C4" bold="true" box="[312,574,1780,1806]" pageId="2" pageNumber="3">Specimens examined</emphasis>
: Five.
<typeStatus id="A07388034414560CFD152F92FC9AF8C5" box="[660,777,1781,1807]" pageId="2" pageNumber="3" type="lectotype">Lectotype</typeStatus>
Ψ (here designated)
<typeStatus id="A07388034414560CFB852F92FBADF8C5" box="[1028,1086,1781,1807]" pageId="2" pageNumber="3">Type</typeStatus>
[circular label with red border]/ 25489 [Fry catalogue number]/ Mouhot [?collectors name]/
<collectingCountry id="07DF76314414560CFBE82E7AFB40F8FD" box="[1129,1235,1821,1847]" name="Cambodia" pageId="2" pageNumber="3">Camboja</collectingCountry>
/
<taxonomicName id="B8C84D224414560CFB622E7AFEA4F895" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="excellens" status="sp. nov.">excellens</taxonomicName>
<taxonomicNameLabel id="568F57C84414560CFEC32E22FE17F895" box="[322,388,1861,1887]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
[Crotchs handwriting]/
<typeStatus id="A07388034414560CFD302E22FD78F895" box="[689,747,1861,1887]" pageId="2" pageNumber="3">Type</typeStatus>
/ Fry coll. 1905­100 (BMNH)/
<typeStatus id="A07388034414560CFBE82E22FB4DF895" box="[1129,1246,1861,1887]" pageId="2" pageNumber="3" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="B8C84D224414560DFB692E22FE5FFEFC" authority="Crotch" authorityName="Crotch" class="Insecta" family="Coccinellidae" genus="Dysis" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="3" lastPageNumber="4" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="excellens">Dysis excellens Crotch</taxonomicName>
des. J. Poorani &amp; R.G. Booth 2005.
<emphasis id="4DBCEAB34415560DFCF7287CFB1BFEFF" bold="true" box="[886,1160,283,309]" pageId="3" pageNumber="4">
<typeStatus id="A07388034415560DFCF7287CFC62FEFF" box="[886,1009,283,309]" pageId="3" pageNumber="4" type="lectotype">Lectotype</typeStatus>
designation
</emphasis>
.
<collectingCountry id="07DF76314415560DFB16287BFB4BFEFC" box="[1175,1240,284,310]" name="India" pageId="3" pageNumber="4">India</collectingCountry>
: Nagaland: Piphema,
<date id="0B7610614415560DFE462823FDB0FE94" box="[455,547,324,350]" pageId="3" pageNumber="4" value="1982-07-01">1.vii.82</date>
, S. Sharma, ICAR, rice, 1Ψ (BMNH); Nagaland: Medziphema,
<date id="0B7610614415560DFE89280BFE1AFE4C" box="[264,393,364,390]" pageId="3" pageNumber="4" value="2004-11-25">25.xi.2004</date>
, N.S.Rao, resting on
<taxonomicName id="B8C84D224415560DFDFE280BFCD7FE4F" box="[639,836,364,389]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="4" phylum="Magnoliopsida" rank="species" species="guajava">
<emphasis id="4DBCEAB34415560DFDFE280BFCD7FE4F" box="[639,836,364,389]" italics="true" pageId="3" pageNumber="4">Psidium guajava</emphasis>
</taxonomicName>
, 1Ψ;
<date id="0B7610614415560DFC00280BFC6CFE4C" box="[897,1023,364,390]" pageId="3" pageNumber="4" value="2004-04-10">10.iv.2004</date>
, H.K. Singh, collected on citrus, 1ɗ;
<date id="0B7610614415560DFE0828F3FD92FE64" box="[393,513,404,430]" pageId="3" pageNumber="4" value="2004-03-21">21.3.2004</date>
, H.K. Singh, in netsweeps from rice, 1Ψ (PDBC).
</paragraph>
</subSubSection>
<subSubSection id="37D2652A4415560DFEB928DCFB43FDEC" pageId="3" pageNumber="4" type="distribution">
<paragraph id="7F7736A14415560DFEB928DCFE19FE34" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="4">
<emphasis id="4DBCEAB34415560DFEB928DCFE42FE1F" bold="true" box="[312,465,443,469]" pageId="3" pageNumber="4">Distribution</emphasis>
:
<collectingCountry id="07DF76314415560DFE6528DBFDD4FE1C" box="[484,583,444,470]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
/?
<collectingCountry id="07DF76314415560DFDE528DBFD4CFE1C" box="[612,735,444,470]" name="Cambodia" pageId="3" pageNumber="4">Cambodia</collectingCountry>
(“
<collectingCountry id="07DF76314415560DFD7D28DBFC3DFE1C" box="[764,942,444,470]" name="Vietnam" pageId="3" pageNumber="4">Cochin China</collectingCountry>
”);
<collectingCountry id="07DF76314415560DFC5128DBFB9FFE1C" box="[976,1036,444,470]" name="India" pageId="3" pageNumber="4">India</collectingCountry>
(presently known from Nagaland).
</paragraph>
<paragraph id="7F7736A14415560DFEB92B6CFB43FDEC" blockId="3.[264,1324,283,1550]" box="[312,1232,523,550]" pageId="3" pageNumber="4">
<emphasis id="4DBCEAB34415560DFEB92B6CFE22FDEF" bold="true" box="[312,433,523,549]" pageId="3" pageNumber="4">Host data</emphasis>
: The specimens examined were collected on citrus, guava, and rice.
</paragraph>
</subSubSection>
<subSubSection id="37D2652A4415560DFEB92B54FD86F9C4" pageId="3" pageNumber="4" type="discussion">
<paragraph id="7F7736A14415560DFEB92B54FC56FD0C" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="4">
<emphasis id="4DBCEAB34415560DFEB92B54FE39FD87" bold="true" box="[312,426,563,589]" pageId="3" pageNumber="4">Remarks</emphasis>
: This species is transferred to
<taxonomicName id="B8C84D224415560DFC972B53FCE8FD87" box="[790,891,564,589]" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="4DBCEAB34415560DFC972B53FCE8FD87" box="[790,891,564,589]" italics="true" pageId="3" pageNumber="4">Oenopia</emphasis>
</taxonomicName>
on account of the following characters: (i) short antenna, only slightly longer than frons, (ii) parallel innerocular margins, (iii) shallowly emarginate anterior margin of mesosternum, and (iv) the male and female genitalia, which are more or less characteristic across the genus.
</paragraph>
<paragraph id="7F7736A14415560DFEB92BB3FCD7FCF4" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="4">
Although
<bibRefCitation id="1B594B504415560DFE2E2BB3FDC7FD24" author="Crotch" box="[431,596,724,750]" pageId="3" pageNumber="5" refString="Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae, E. W. Janson, London. 311 pp." type="book" year="1874">Crotch (1874)</bibRefCitation>
gave the locality data of this species as
<collectingCountry id="07DF76314415560DFBB12BB3FB46FD24" box="[1072,1237,724,750]" name="Vietnam" pageId="3" pageNumber="4">Cochin China</collectingCountry>
, nowadays
<collectingCountry id="07DF76314415560DFECB2B9BFE22FCDC" box="[330,433,764,790]" name="Vietnam" pageId="3" pageNumber="4">Vietnam</collectingCountry>
, the specimen locality label indicated
<collectingCountry id="07DF76314415560DFC0C2B9BFC64FCDC" box="[909,1015,764,790]" name="Cambodia" pageId="3" pageNumber="4">Camboja</collectingCountry>
(presumably
<collectingCountry id="07DF76314415560DFB202B9BFA8CFCDC" box="[1185,1311,764,790]" name="Cambodia" pageId="3" pageNumber="4">Cambodia</collectingCountry>
). This is the first record of this species from
<collectingCountry id="07DF76314415560DFC812A43FCD3FCF4" box="[768,832,804,830]" name="India" pageId="3" pageNumber="4">India</collectingCountry>
.
</paragraph>
<paragraph id="7F7736A14415560DFEB92A2BFEAEFBE4" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="4">
This species is remarkably similar in external appearance and coloration to
<taxonomicName id="B8C84D224415560DFB442A2BFDAAFC44" authority="Blackburn, 1892" authorityName="Blackburn" authorityYear="1892" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="4DBCEAB34415560DFB442A2BFEC8FC47" italics="true" pageId="3" pageNumber="4">Oenopia guttata</emphasis>
(
<bibRefCitation id="1B594B504415560DFEEA2A13FDA2FC44" author="Blackburn" box="[363,561,884,910]" pageId="3" pageNumber="5" refString="Blackburn, T. (1892) Further notes on Australian Coleoptera, with descriptions of new genera and species. Transactions of the Royal Society of South Australia, 15, 207 - 261." type="journal article" year="1892">Blackburn, 1892</bibRefCitation>
)
</taxonomicName>
found in
<collectingCountry id="07DF76314415560DFD2A2A13FC89FC44" box="[683,794,884,910]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
, New
<collectingCountry id="07DF76314415560DFCE52A13FC2FFC44" box="[868,956,884,910]" name="Guinea-Bissau" pageId="3" pageNumber="4">Guinea</collectingCountry>
, and the
<collectingCountry id="07DF76314415560DFBA42A13FB3EFC44" box="[1061,1197,884,910]" name="Philippines" pageId="3" pageNumber="4">Philippines</collectingCountry>
. The specimens examined were slightly smaller in size than the range specified for
<taxonomicName id="B8C84D224415560DFB082AFBFA90FC7F" box="[1161,1283,924,949]" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="4DBCEAB34415560DFB082AFBFA90FC7F" box="[1161,1283,924,949]" italics="true" pageId="3" pageNumber="4">O. guttata</emphasis>
</taxonomicName>
by
<bibRefCitation id="1B594B504415560DFE892AA3FE33FC14" author="Pope" box="[264,416,964,990]" pageId="3" pageNumber="5" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 2 (5) [1988], 633 - 735." type="journal article" year="1989">Pope (1989)</bibRefCitation>
. The front pair of elytral spots in the
<typeStatus id="A07388034415560DFC052AA3FC7CFC14" box="[900,1007,964,990]" pageId="3" pageNumber="4" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="B8C84D224415560DFBA22AA3FB24FC17" box="[1059,1207,964,989]" class="Insecta" family="Coccinellidae" genus="Dysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="4DBCEAB34415560DFBA22AA3FB24FC17" box="[1059,1207,964,989]" italics="true" pageId="3" pageNumber="4">O. excellens</emphasis>
</taxonomicName>
is closer together than in
<taxonomicName id="B8C84D224415560DFE4A2A8BFDD0FBCF" box="[459,579,1004,1029]" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="4DBCEAB34415560DFE4A2A8BFDD0FBCF" box="[459,579,1004,1029]" italics="true" pageId="3" pageNumber="4">O. guttata</emphasis>
</taxonomicName>
and there are very slight differences in the strength of punctation.
</paragraph>
<paragraph id="7F7736A14415560DFEB92D5BFE15FA5F" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="4">
However, the male genitalia and female spermatheca are diagnostic and very distinctly different from the illustrations provided by
<bibRefCitation id="1B594B504415560DFC8D2D03FC4BFBB4" author="Bielawski" box="[780,984,1124,1150]" pageId="3" pageNumber="5" refString="Bielawski, R. (1964) Die Arten der Gattung Gyrocaria Timberlake (Coleoptera: Coccinellidae). Bulletin de l'Academie Polonaise des Sciences, Cl. II. Serie des sciences biologiques, 12 (2), 83 - 90." type="journal article" year="1964">Bielawski (1964)</bibRefCitation>
and
<bibRefCitation id="1B594B504415560DFB952D03FB3BFBB4" author="Pope" box="[1044,1192,1124,1150]" pageId="3" pageNumber="5" refString="Pope, R. D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 2 (5) [1988], 633 - 735." type="journal article" year="1989">Pope (1989)</bibRefCitation>
, in the following ways. The apical emargination of the median lobe of the tegmen is deep and parabolic in
<taxonomicName id="B8C84D224415560DFEEF2DD3FE6CFB07" box="[366,511,1204,1229]" class="Insecta" family="Coccinellidae" genus="Dysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="4DBCEAB34415560DFEEF2DD3FE6CFB07" box="[366,511,1204,1229]" italics="true" pageId="3" pageNumber="4">O. excellens</emphasis>
</taxonomicName>
, but shallower and broadly U­shaped in
<taxonomicName id="B8C84D224415560DFC6E2DD3FBFAFB07" box="[1007,1129,1204,1229]" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="4DBCEAB34415560DFC6E2DD3FBFAFB07" box="[1007,1129,1204,1229]" italics="true" pageId="3" pageNumber="4">O. guttata</emphasis>
</taxonomicName>
. The parameres appear to be shorter than the median lobe in
<taxonomicName id="B8C84D224415560DFC992DBBFC3BFB3F" box="[792,936,1244,1269]" class="Insecta" family="Coccinellidae" genus="Dysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="4DBCEAB34415560DFC992DBBFC3BFB3F" box="[792,936,1244,1269]" italics="true" pageId="3" pageNumber="4">O. excellens</emphasis>
</taxonomicName>
, but are comparatively longer in
<taxonomicName id="B8C84D224415560DFE892C63FEECFAD7" box="[264,383,1284,1309]" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="4DBCEAB34415560DFE892C63FEECFAD7" box="[264,383,1284,1309]" italics="true" pageId="3" pageNumber="4">O. guttata</emphasis>
</taxonomicName>
. The ramus of the female spermatheca is very long and gooseneck­shaped in
<taxonomicName id="B8C84D224415560DFA8F2C63FEE0FA8F" class="Insecta" family="Coccinellidae" genus="Dysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="4DBCEAB34415560DFA8F2C63FEE0FA8F" italics="true" pageId="3" pageNumber="4">O. excellens</emphasis>
</taxonomicName>
, but much shorter and straight in
<emphasis id="4DBCEAB34415560DFC902C4BFC01FA8F" box="[785,914,1324,1349]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="B8C84D224415560DFC902C4BFC1EFA8F" box="[785,909,1324,1349]" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guttata">O. guttata</taxonomicName>
.
</emphasis>
Further, the cornu is semicircular and strongly curved in
<taxonomicName id="B8C84D224415560DFD972C33FD36FAA7" box="[534,677,1364,1389]" class="Insecta" family="Coccinellidae" genus="Dysis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="excellens">
<emphasis id="4DBCEAB34415560DFD972C33FD36FAA7" box="[534,677,1364,1389]" italics="true" pageId="3" pageNumber="4">O. excellens</emphasis>
</taxonomicName>
, but much shorter, wider and very shallowly arcuate in
<emphasis id="4DBCEAB34415560DFE892C1BFE15FA5F" box="[264,390,1404,1429]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="B8C84D224415560DFE892C1BFE10FA5F" box="[264,387,1404,1429]" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="guttata">O. guttata</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="7F7736A14415560DFEB92CC3FD86F9C4" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="4">
This species belongs to the
<taxonomicName id="B8C84D224415560DFD052CC3FD6EFA77" box="[644,765,1444,1469]" class="Insecta" family="Coccinellidae" genus="Gyrocaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="4DBCEAB34415560DFD052CC3FD6EFA77" box="[644,765,1444,1469]" italics="true" pageId="3" pageNumber="4">Gyrocaria</emphasis>
</taxonomicName>
group of the genus that are characterized by a more or less convex, circular to oval body outline and other genitalic differences indicated by Vandenberg (1999).
</paragraph>
</subSubSection>
</treatment>
</document>