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<document ID-DOI="http://doi.org/10.5281/zenodo.3788199" ID-GBIF-Dataset="18d263ac-9913-4f11-843c-862ca11a38e6" ID-GBIF-Taxon="163755109" ID-ISSN="13132970" ID-Zenodo-Dep="3788199" ID-ZooBank="975251CE-C173-4D80-84B9-C14B870330F9" LSID="urn:lsid:zoobank.org:act:90CDFBB8-88FF-491B-B315-46A782333BBD" approvalRequired="132" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="130" checkinTime="1588000592231" checkinUser="plazi" docAuthor="Engel, Michael" docDate="2010" docId="F0136C1340080170FF06FAD5FBF7FE52" docLanguage="en" docName="ZK_article_2202.pdf" docOrigin="ZooKeys 47 (47)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5b4359" docStyleName="zookeys.2008.journal_article" docTitle="Chlerogella materdonnae Engel 2010, sp. n." docType="treatment" docUuid="90CDFBB8-88FF-491B-B315-46A782333BBD" docUuidSource="ZooBank" docVersion="5" lastPageId="42" lastPageNumber="43" masterDocId="0C2A146B402E015AFF8AFFE5FFA3FFC6" masterDocTitle="Revision of the Bee Genus Chlerogella (Hymenoptera, Halictidae), Part II: South American Species and Generic Diagnosis" masterLastPageNumber="100" masterPageNumber="1" pageId="38" pageNumber="39" updateTime="1643559916529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Bee Genus Chlerogella (Hymenoptera, Halictidae), Part II: South American Species and Generic Diagnosis</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Engel, Michael</mods:namePart>
<mods:affiliation>University of Kansas, Natural History Museum, Lawrence, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-05-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>47</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>100</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.47.416</mods:identifier>
<mods:identifier type="GBIF-Dataset">18d263ac-9913-4f11-843c-862ca11a38e6</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576667</mods:identifier>
<mods:identifier type="ZooBank">975251CE-C173-4D80-84B9-C14B870330F9</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788199" ID-GBIF-Taxon="163755109" ID-Zenodo-Dep="3788199" LSID="urn:lsid:zoobank.org:act:90CDFBB8-88FF-491B-B315-46A782333BBD" httpUri="http://treatment.plazi.org/id/F0136C1340080170FF06FAD5FBF7FE52" lastPageId="42" lastPageNumber="43" pageId="38" pageNumber="39">
<subSubSection pageId="38" pageNumber="39" type="nomenclature">
<paragraph blockId="38.[140,976,1328,1425]" box="[140,593,1328,1355]" pageId="38" pageNumber="39">
<heading box="[140,593,1328,1355]" fontSize="11" level="2" pageId="38" pageNumber="39" reason="3">
<emphasis bold="true" box="[140,593,1328,1355]" pageId="38" pageNumber="39">
<taxonomicName authority="Engel" authorityName="Engel" authorityYear="2010" box="[140,516,1328,1355]" class="Insecta" family="Halictidae" genus="Chlerogella" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="materdonnae" status="sp. n.">
<emphasis bold="true" box="[140,438,1328,1355]" italics="true" pageId="38" pageNumber="39">
<emphasis box="[140,274,1328,1355]" inLineHeading="true" pageId="38" pageNumber="39" reason="1">Chlerogella</emphasis>
materdonnae
</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel box="[528,593,1328,1354]" pageId="38" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="38.[140,976,1328,1425]" box="[140,976,1363,1390]" pageId="38" pageNumber="39">
<uri box="[140,976,1363,1390]" pageId="38" pageNumber="39">
urn:lsid:zoobank.org:act:
<uuid box="[417,976,1363,1390]" pageId="38" pageNumber="39">90CDFBB8-88FF-491B-B315-46A782333BBD</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[140,444,1398,1425]" pageId="38" pageNumber="39" type="description">
<paragraph blockId="38.[140,976,1328,1425]" box="[140,444,1398,1425]" pageId="38" pageNumber="39">Figs 5561, 7071, Map 3</paragraph>
</subSubSection>
<subSubSection lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="materials_examined">
<paragraph blockId="38.[140,1108,1469,1672]" pageId="38" pageNumber="39">
<materialsCitation ID-GBIF-Occurrence="2608994758" collectingDate="1996-02-18" collectionCode="SEMC" collectorName="J. S. Ascher" country="Ecuador" location="Pailon del Diablo" municipality="Rio" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Tungurahua" typeStatus="holotype">
<emphasis bold="true" box="[140,260,1469,1495]" pageId="38" pageNumber="39">
<emphasis box="[140,256,1469,1495]" inLineHeading="true" pageId="38" pageNumber="39" reason="1">
<typeStatus box="[140,256,1469,1495]" pageId="38" pageNumber="39">Holotype</typeStatus>
</emphasis>
.
</emphasis>
♁,
<collectingCountry box="[303,416,1469,1495]" name="Ecuador" pageId="38" pageNumber="39">ECUADOR</collectingCountry>
:
<collectingRegion box="[425,566,1469,1495]" country="Ecuador" name="Tungurahua" pageId="38" pageNumber="39">Tungurahua</collectingRegion>
,
<collectingMunicipality box="[576,617,1469,1495]" pageId="38" pageNumber="39">Rio</collectingMunicipality>
<collectingRegion box="[624,707,1470,1496]" country="Ecuador" name="Pastaza" pageId="38" pageNumber="39">Pastaza</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:F0136C1340080170FF06FAD5FBF7FE52:7D658BDE4008017CFD44FA58FC36FA11" box="[718,917,1469,1495]" country="Ecuador" municipality="Rio" name="Pailon del Diablo" pageId="38" pageNumber="39" stateProvince="Tungurahua">Pailon del Diablo</location>
,
<quantity box="[929,987,1469,1496]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="39" unit="km" value="5.0">5 km</quantity>
SE,
<date pageId="38" pageNumber="39" value="1996-02-18">
<collectingDate pageId="38" pageNumber="39" value="1996-02-18">II-18- 1996</collectingDate>
</date>
,
<collectorName box="[213,338,1504,1530]" pageId="38" pageNumber="39">J.S. Ascher</collectorName>
, flying along forest trail (
<collectionCode box="[620,705,1504,1530]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="38" pageNumber="39" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="38.[140,1108,1469,1672]" pageId="38" pageNumber="39">
<materialsCitation ID-GBIF-Occurrence="2608994750" collectingDate="1986-01-29" collectionCode="COOP" collectorName="M. Cooper" country="Ecuador" elevation="1100" location="Rio Topo" municipality="Valley" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Tungurahua" typeStatus="paratype">
<emphasis bold="true" box="[185,308,1539,1565]" pageId="38" pageNumber="39">
<emphasis box="[185,303,1539,1565]" inLineHeading="true" pageId="38" pageNumber="39" reason="1">
<typeStatus box="[185,303,1539,1565]" pageId="38" pageNumber="39">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry box="[324,437,1540,1566]" name="Ecuador" pageId="38" pageNumber="39">ECUADOR</collectingCountry>
: 1♁,
<collectingRegion box="[516,657,1539,1565]" country="Ecuador" name="Tungurahua" pageId="38" pageNumber="39">Tungurahua</collectingRegion>
,
<collectingRegion box="[678,758,1540,1566]" country="Ecuador" name="Pastaza" pageId="38" pageNumber="39">Pastaza</collectingRegion>
<collectingMunicipality box="[774,843,1539,1565]" pageId="38" pageNumber="39">Valley</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F0136C1340080170FF06FAD5FBF7FE52:7D658BDE4008017CFCD5F9E6FC76F9D8" box="[863,981,1539,1566]" country="Ecuador" municipality="Valley" name="Rio Topo" pageId="38" pageNumber="39" stateProvince="Tungurahua">Rio Topo</location>
,
<quantity box="[1002,1103,1540,1566]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="38" pageNumber="39" unit="m" value="1100.0">
<elevation box="[1002,1103,1540,1566]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="38" pageNumber="39" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date box="[140,250,1575,1601]" pageId="38" pageNumber="39" value="1986-01-29">
<collectingDate box="[140,250,1575,1601]" pageId="38" pageNumber="39" value="1986-01-29">29.i.1986</collectingDate>
</date>
[
<date box="[269,473,1574,1601]" pageId="38" pageNumber="39" value="1986-01-29">
<collectingDate box="[269,473,1574,1601]" pageId="38" pageNumber="39" value="1986-01-29">29 January 1986</collectingDate>
</date>
],
<collectorName box="[497,627,1575,1601]" pageId="38" pageNumber="39">M. Cooper</collectorName>
(
<collectionCode box="[648,736,1575,1601]" pageId="38" pageNumber="39">COOP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608994695" collectingDate="1982-12-20" collectionCode="COOP" collectorName="M. Cooper" country="Ecuador" elevation="1100" location="Rio Topo" municipality="Valley" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Tungurahua" typeStatus="paratype">
<specimenCount box="[760,790,1575,1601]" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
,
<collectingRegion box="[814,955,1575,1601]" country="Ecuador" name="Tungurahua" pageId="38" pageNumber="39">Tungurahua</collectingRegion>
,
<collectingRegion box="[971,1051,1575,1601]" country="Ecuador" name="Pastaza" pageId="38" pageNumber="39">Pastaza</collectingRegion>
<collectingMunicipality pageId="38" pageNumber="39">Valley</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F0136C1340080170FF06FAD5FBF7FE52:7D658BDE4008017CFF33F9AFFE84F9A2" box="[185,295,1610,1636]" country="Ecuador" municipality="Valley" name="Rio Topo" pageId="38" pageNumber="39" stateProvince="Tungurahua">Rio Topo</location>
,
<quantity box="[308,401,1610,1636]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="38" pageNumber="39" unit="m" value="1100.0">
<elevation box="[308,401,1610,1636]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="38" pageNumber="39" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date box="[414,544,1610,1636]" pageId="38" pageNumber="39" value="1982-12-20">
<collectingDate box="[414,544,1610,1636]" pageId="38" pageNumber="39" value="1982-12-20">20.xii.1982</collectingDate>
</date>
[
<date box="[560,788,1609,1636]" pageId="38" pageNumber="39" value="1982-12-20">
<collectingDate box="[560,788,1609,1636]" pageId="38" pageNumber="39" value="1982-12-20">20 December 1982</collectingDate>
</date>
],
<collectorName box="[809,937,1610,1636]" pageId="38" pageNumber="39">M. Cooper</collectorName>
(
<collectionCode box="[954,1042,1610,1636]" pageId="38" pageNumber="39">COOP</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608994684" collectingDate="1983-02-11" collectionCode="SEMC" collectorName="L. Huggert" country="Ecuador" location="El Chaco" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
1♁,
<collectingRegion box="[140,206,1646,1672]" country="Ecuador" name="Napo" pageId="38" pageNumber="39">Napo</collectingRegion>
,
<quantity box="[218,277,1645,1672]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="39" unit="km" value="5.0">5 km</quantity>
NE
<location LSID="urn:lsid:plazi:treatment:F0136C1340080170FF06FAD5FBF7FE52:7D658BDE4008017CFEC6F988FE14F941" box="[332,439,1645,1671]" country="Ecuador" name="El Chaco" pageId="38" pageNumber="39" stateProvince="Napo">El Chaco</location>
,
<date box="[451,569,1645,1671]" pageId="38" pageNumber="39" value="1983-02-11">
<collectingDate box="[451,569,1645,1671]" pageId="38" pageNumber="39" value="1983-02-11">11.ii.1983</collectingDate>
</date>
[
<date box="[584,793,1644,1671]" pageId="38" pageNumber="39" value="1983-02-11">
<collectingDate box="[584,793,1644,1671]" pageId="38" pageNumber="39" value="1983-02-11">11 February 1983</collectingDate>
</date>
],
<collectorName box="[813,939,1646,1672]" pageId="38" pageNumber="39">L. Huggert</collectorName>
(
<collectionCode box="[954,1039,1645,1671]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="38" pageNumber="39" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="39.[140,1107,1333,1387]" pageId="39" pageNumber="40">
<emphasis bold="true" box="[140,300,1333,1355]" pageId="39" pageNumber="40">Figures 5557.</emphasis>
Female of
<taxonomicName authority="Engel, 2010" authorityName="Engel" authorityYear="2010" box="[402,621,1333,1354]" class="Insecta" family="Halictidae" genus="Chlerogella" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="materdonnae" status="sp. n.">
<emphasis box="[402,621,1333,1354]" italics="true" pageId="39" pageNumber="40">Chlerogella materdonnae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[627,678,1334,1355]" pageId="39" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
<emphasis bold="true" box="[683,709,1333,1355]" pageId="39" pageNumber="40">55</emphasis>
Lateral habitus
<emphasis bold="true" box="[858,884,1333,1355]" pageId="39" pageNumber="40">56</emphasis>
Facial aspect
<emphasis bold="true" box="[1012,1038,1333,1355]" pageId="39" pageNumber="40">57</emphasis>
Lateral aspect of head.
</paragraph>
</subSubSection>
<subSubSection lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="diagnosis">
<paragraph blockId="39.[140,1108,1433,1671]" pageId="39" pageNumber="40">
<emphasis bold="true" box="[185,312,1433,1459]" pageId="39" pageNumber="40">
<emphasis box="[185,307,1433,1459]" inLineHeading="true" pageId="39" pageNumber="40" reason="1">Diagnosis</emphasis>
.
</emphasis>
<taxonomicName authorityName="Engel" authorityYear="2010" box="[320,590,1434,1460]" class="Insecta" family="Halictidae" genus="Chlerogella" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="materdonnae">
<emphasis box="[320,590,1434,1460]" italics="true" pageId="39" pageNumber="40">Chlerogella materdonnae</emphasis>
</taxonomicName>
belongs to a complex of very similar metallic blue species with elongate malar spaces. The female is most similar to
<taxonomicName authorityName="Engel" authorityYear="2010" box="[944,1060,1470,1496]" class="Insecta" family="Halictidae" genus="Chlerogella" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="rostrata">
<emphasis box="[944,963,1470,1496]" italics="true" pageId="39" pageNumber="40">C</emphasis>
.
<emphasis box="[979,1060,1470,1495]" italics="true" pageId="39" pageNumber="40">rostrata</emphasis>
</taxonomicName>
but differs by the brilliant and shiny caerulean blue color of the integument (Figs 5556;
<emphasis box="[140,158,1540,1566]" italics="true" pageId="39" pageNumber="40">cf</emphasis>
. Figs 3031) and the uniformly punctured mesepisternum (not more densely spaced ventrally as in
<taxonomicName authorityName="Engel" authorityYear="2010" box="[308,424,1575,1601]" class="Insecta" family="Halictidae" genus="Chlerogella" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="rostrata">
<emphasis box="[308,327,1575,1601]" italics="true" pageId="39" pageNumber="40">C</emphasis>
.
<emphasis box="[343,424,1575,1600]" italics="true" pageId="39" pageNumber="40">rostrata</emphasis>
</taxonomicName>
,
<emphasis box="[440,551,1575,1601]" italics="true" pageId="39" pageNumber="40">vide supra</emphasis>
). The male can be distinguished from that of
<taxonomicName class="Insecta" family="Halictidae" genus="Chlerogella" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="agalyei">
<emphasis box="[1081,1100,1575,1601]" italics="true" pageId="39" pageNumber="40">C</emphasis>
.
<emphasis box="[140,213,1610,1636]" italics="true" pageId="39" pageNumber="40">agalyei</emphasis>
</taxonomicName>
by the caerulean coloration and the structure of SIV (Fig. 61) and the terminalia (Figs 7071).
</paragraph>
<paragraph blockId="40.[140,1107,1156,1210]" pageId="40" pageNumber="41">
<emphasis bold="true" box="[140,301,1156,1178]" pageId="40" pageNumber="41">Figures 586Ι.</emphasis>
Male of
<taxonomicName authority="Engel, 2010" authorityName="Engel" authorityYear="2010" box="[388,609,1156,1177]" class="Insecta" family="Halictidae" genus="Chlerogella" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="materdonnae" status="sp. n.">
<emphasis box="[388,609,1156,1177]" italics="true" pageId="40" pageNumber="41">Chlerogella materdonnae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[616,668,1157,1178]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
<emphasis bold="true" box="[674,700,1156,1178]" pageId="40" pageNumber="41">58</emphasis>
Lateral habitus
<emphasis bold="true" box="[853,879,1156,1178]" pageId="40" pageNumber="41">59</emphasis>
Facial aspect
<emphasis bold="true" box="[1010,1036,1156,1178]" pageId="40" pageNumber="41">60</emphasis>
Lateral aspect of head
<emphasis bold="true" box="[276,303,1188,1210]" pageId="40" pageNumber="41">6Ι</emphasis>
Apical metasomal sterna.
</paragraph>
</subSubSection>
<subSubSection lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="description">
<paragraph blockId="40.[140,1108,1257,1672]" lastBlockId="41.[140,1108,167,1672]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis bold="true" box="[185,333,1257,1283]" pageId="40" pageNumber="41">
<emphasis box="[185,328,1257,1283]" inLineHeading="true" pageId="40" pageNumber="41" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[340,415,1258,1284]" italics="true" pageId="40" pageNumber="41">Female</emphasis>
: Total body length
<quantity box="[631,737,1258,1284]" metricMagnitude="-3" metricUnit="m" metricValue="9.11" pageId="40" pageNumber="41" unit="mm" value="9.11">9.11 mm</quantity>
; forewing length
<quantity box="[930,1036,1258,1284]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="40" pageNumber="41" unit="mm" value="6.2">6.20 mm</quantity>
. Head length
<quantity box="[220,329,1293,1320]" metricMagnitude="-3" metricUnit="m" metricValue="2.71" pageId="40" pageNumber="41" unit="mm" value="2.71">2.71 mm</quantity>
, width
<quantity box="[416,525,1293,1320]" metricMagnitude="-3" metricUnit="m" metricValue="1.74" pageId="40" pageNumber="41" unit="mm" value="1.74">1.74 mm</quantity>
. Clypeus beginning at lower tangent of compound eyes. Malar space 43.9% compound eye length (malar length
<quantity box="[825,932,1328,1355]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="40" pageNumber="41" unit="mm" value="0.65">0.65 mm</quantity>
; compound eye length
<quantity box="[217,321,1364,1390]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="40" pageNumber="41" unit="mm" value="1.48">1.48 mm</quantity>
) (Figs 5657). Upper interorbital distance
<quantity box="[798,904,1364,1390]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="40" pageNumber="41" unit="mm" value="0.83">0.83 mm</quantity>
; lower interorbital distance
<quantity box="[238,346,1399,1425]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="40" pageNumber="41" unit="mm" value="0.63">0.63 mm</quantity>
. Upper portion of pronotum medially depressed, not elongate, medially less than 0.25 times ocellar diameter in length; ventral portion of preëpisternal sulcus not broad, similar to scrobal sulcus and upper portion of preëpisternal sulcus; intertegular distance
<quantity box="[371,477,1504,1531]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="40" pageNumber="41" unit="mm" value="1.43">1.43 mm</quantity>
; mesoscutellum weakly convex, not bigibbous. Basal vein distad cu-a by three times vein width; 1rs-m distad 1m-cu by two times vein width; 2rsm distad 2m-cu by nine times vein width, 2rs-m weakly arched; first submarginal cell longer than combined lengths of second and third submarginal cells; second submarginal cell slightly narrowed anteriorly, anterior border of second submarginal cell along Rs slightly longer than that of third submarginal cell; posterior border of third submarginal cell slightly more than two times longer than anterior border. Distal hamuli arranged 2-1-2. Inner metatibial spur with six branches (not including apical portion of rachis).
</paragraph>
<paragraph blockId="41.[140,1108,167,1672]" pageId="41" pageNumber="42">Clypeus and supraclypeal area imbricate with weak punctures separated by 0.51.5 times a puncture width; face with small, contiguous punctures, more widely spaced in malar space; punctures of face blending to imbricate integument in ocellocular area and vertex, with punctures separated by 12 times a puncture width; gena finely imbricate with small punctures separated by 0.52 times a puncture width; postgena imbricate and impunctate. Pronotum imbricate with minute punctures separated by 13 times a puncture width; mesoscutum imbricate with punctures separated by a puncture width or less on lateral thirds and posteriorly, anteromedially punctures becoming exceedingly faint to absent; mesoscutellum finely imbricate with small punctures separated by a puncture width or less; metanotum imbricate with minute punctures separated by a puncture width or less. Preëpisternum finely imbricate with small punctures separated by 13 times a puncture width; mesepisternum imbricate with punctures separated by 24 times a puncture width; metepisternum faintly imbricate. Propodeum strongly imbricate. Metasoma finely imbricate.</paragraph>
<paragraph blockId="41.[140,1108,167,1672]" pageId="41" pageNumber="42">Mandible dark brown except reddish at apex; labrum dark brown; clypeal apex dark brown, remainder of clypeus and head light metallic caerulean blue. Antenna dark brown. Mesosoma light metallic caerulean blue (Fig. 55); tegula dark brown. Wing membranes lightly infumate; veins dark brown. Legs dark brown. Metasoma dark brown.</paragraph>
<paragraph blockId="41.[140,1108,167,1672]" pageId="41" pageNumber="42">Pubescence white except intermingled with dark fuscous setae on metatibia and with dark fuscous setae predominant on tarsi, apicalmost metasomal segments, and pro- and mesotibiae.</paragraph>
<paragraph blockId="41.[140,1108,167,1672]" pageId="41" pageNumber="42">
<emphasis box="[185,241,1047,1073]" italics="true" pageId="41" pageNumber="42">Male</emphasis>
: As described for the female except as follows: Total body length 9.0
<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.18" pageId="41" pageNumber="42" unit="mm" value="9.18">9.18 mm</quantity>
; forewing length
<quantity box="[390,566,1082,1109]" metricMagnitude="-3" metricUnit="m" metricValue="6.14" metricValueMax="6.15" metricValueMin="6.13" pageId="41" pageNumber="42" unit="mm" value="6.14" valueMax="6.15" valueMin="6.13">6.136.15 mm</quantity>
. Head length
<quantity box="[728,903,1082,1109]" metricMagnitude="-3" metricUnit="m" metricValue="2.59" metricValueMax="2.6" metricValueMin="2.58" pageId="41" pageNumber="42" unit="mm" value="2.59" valueMax="2.6" valueMin="2.58">2.582.60 mm</quantity>
, width
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.595" metricValueMax="1.6" metricValueMin="1.59" pageId="41" pageNumber="42" unit="mm" value="1.595" valueMax="1.6" valueMin="1.59">1.591.60 mm</quantity>
. Clypeus beginning below lower tangent of compound eyes. Malar space 50.7 51% compound eye length (malar length
<quantity box="[616,724,1152,1179]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="41" pageNumber="42" unit="mm" value="0.73">0.73 mm</quantity>
; compound eye length
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.435" metricValueMax="1.44" metricValueMin="1.43" pageId="41" pageNumber="42" unit="mm" value="1.435" valueMax="1.44" valueMin="1.43">1.431.44 mm</quantity>
) (Figs 5960). Upper interorbital distance
<quantity box="[676,850,1187,1214]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" metricValueMax="7.8" metricValueMin="7.6" pageId="41" pageNumber="42" unit="mm" value="0.77" valueMax="0.78" valueMin="0.76">0.760.78 mm</quantity>
; lower interorbital distance
<quantity box="[206,379,1223,1249]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="41" pageNumber="42" unit="mm" value="0.425" valueMax="0.43" valueMin="0.42">0.420.43 mm</quantity>
. First flagellomere about as long as pedicel, about as long as wide; second flagellomere 2.4 times length of first flagellomere; ventral surfaces of second through eleventh flagellomeres densely covered in placoid sensilla, placoid fields not disrupted. Intertegular distance
<quantity box="[502,676,1328,1355]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" metricValueMax="1.22" metricValueMin="1.2" pageId="41" pageNumber="42" unit="mm" value="1.21" valueMax="1.22" valueMin="1.2">1.201.22 mm</quantity>
; mesoscutellum weakly bigibbous. Inner metatibial spur serrate. Apical margin of SIII entire; apical margin of SIV broadly concave medially and produced into short broad lobes lateral to concavity, without furrow in apical half of disc (Fig. 61); apical margin of SV entire; apical margin of SVI emarginate; terminalia as depicted in figures 70 and 71.
</paragraph>
<paragraph blockId="41.[140,1108,167,1672]" pageId="41" pageNumber="42">Mandible, labrum, and clypeal apex brown. Head and mesosoma light metallic caerulean blue except more deeply azurite colored on face and mesoscutum.</paragraph>
<paragraph blockId="41.[140,1108,167,1672]" lastBlockId="42.[140,1108,167,404]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">Typical gender pilosity except postgena with numerous elongate, sinuate setae, such setae with short apical branches; inner surfaces of trochanters, mesofemur, and metacoxa with elongate, apically-plumose setae, similar setae on inner surfaces of me- tatrochanter, metafemur, and metatibia except largely simple and somewhat sinuate, those of metatibia not particularly elongate, length of such setae not or scarcely longer than metatibial width. Apical margin of SIII with diffuse fringe of moderate-length white setae; SIV with small medioapical pads of short fuscous setae bordering medial emargination; SV with diffuse areas of moderate-length to long white setae.</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="etymology">
<paragraph blockId="42.[140,1108,167,404]" pageId="42" pageNumber="43">
<emphasis bold="true" box="[185,320,343,369]" pageId="42" pageNumber="43">
<emphasis box="[185,315,343,369]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Etymology</emphasis>
.
</emphasis>
The specific epithet is a matronym combining the Latin term
<emphasis box="[1037,1100,343,368]" italics="true" pageId="42" pageNumber="43">mater</emphasis>
, meaning “mother”, and Donna, in loving honor of my mother, Mrs. Donna G. Engel.
</paragraph>
</subSubSection>
</treatment>
</document>